227 lines
20 KiB
XML
227 lines
20 KiB
XML
<document ID-DOI="10.25221/fee.374.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7165031" approvalRequired="81" approvalRequired_for_textStreams="81" checkinTime="1665246964924" checkinUser="felipe" docAuthor="Mikhaljova, E. V." docDate="2019" docId="03EDCF74FF82793397ECFDF8FC7093BB" docLanguage="en" docName="FarEastEntomol.374.1-9.pdf" docOrigin="Far Eastern Entomologist 374" docSource="http://dx.doi.org/10.25221/fee.374.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Altajosoma kultukensis Mikhaljova 2019, sp. n." docType="treatment" docVersion="2" lastPageNumber="5" masterDocId="FFD4B70CFF807937970FFFC0FFAC973E" masterDocTitle="Millipedes of the order Chordeumatida (Diplopoda) from the Russian Far East and Siberia in the collection of the Natural History Museum of Denmark" masterLastPageNumber="9" masterPageNumber="1" pageNumber="3" updateTime="1665260623001" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Millipedes of the order Chordeumatida (Diplopoda) from the Russian Far East and Siberia in the collection of the Natural History Museum of Denmark</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mikhaljova, E. V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Far Eastern Entomologist</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2019-01-09</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>374</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>9</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.25221/fee.374.1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.25221/fee.374.1</mods:identifier>
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<mods:identifier type="ISSN">2713-2196</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7165031</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03EDCF74FF82793397ECFDF8FC7093BB" httpUri="http://treatment.plazi.org/id/03EDCF74FF82793397ECFDF8FC7093BB" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
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<subSubSection pageId="2" pageNumber="3" type="nomenclature">
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<paragraph blockId="2.[227,1048,568,652]" box="[227,696,568,592]" pageId="2" pageNumber="3">
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<heading box="[227,696,568,592]" fontSize="9" level="2" pageId="2" pageNumber="3" reason="2">
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<emphasis bold="true" box="[227,696,568,592]" pageId="2" pageNumber="3">
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<taxonomicName authority="Mikhaljova" authorityName="Mikhaljova" authorityYear="2019" box="[227,625,568,592]" class="Diplopoda" family="Diplomaragnidae" genus="Altajosoma" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="kultukensis" status="sp. n.">
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<emphasis bold="true" box="[227,484,568,592]" italics="true" pageId="2" pageNumber="3">Altajosoma kultukensis</emphasis>
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Mikhaljova
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</taxonomicName>
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,
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<taxonomicNameLabel box="[636,696,568,592]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="2.[227,1048,568,652]" box="[227,1048,598,622]" pageId="2" pageNumber="3">
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http/
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<uri box="[280,1048,598,622]" pageId="2" pageNumber="3">
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urn:lsid:zoobank.org:act:
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<uuid box="[538,1048,598,622]" pageId="2" pageNumber="3">A17B6265-B2BD-40CC-97D6-0B5328307009</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection box="[227,320,628,652]" pageId="2" pageNumber="3" type="description">
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<paragraph blockId="2.[227,1048,568,652]" box="[227,320,628,652]" pageId="2" pageNumber="3">
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<figureCitation box="[227,320,628,652]" captionStart="Figs 1–5" captionStartId="3.[265,306,1410,1432]" captionTargetBox="[227,1099,227,1367]" captionTargetId="figure-0@3.[227,1114,227,1378]" captionTargetPageId="3" captionText="Figs 1–5. Altajosoma kultukensis sp. n., male holotype. 1 – coxa and trochanter 10; 2 –" figureDoi="http://doi.org/10.5281/zenodo.7165037" httpUri="https://zenodo.org/record/7165037/files/figure.png" pageId="2" pageNumber="3">Figs 1–5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="materials_examined">
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<paragraph blockId="2.[227,1128,688,1552]" box="[265,1126,688,712]" pageId="2" pageNumber="3">
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<materialsCitation box="[265,1126,688,712]" collectionCode="FSCB" country="Russia" location="Rio Bistraya" municipality="Kurtuk" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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MATERIAL.
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<typeStatus box="[417,516,688,712]" pageId="2" pageNumber="3">Holotype</typeStatus>
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–
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<specimenCount box="[546,566,688,711]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
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(
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<collectionCode box="[582,652,688,712]" pageId="2" pageNumber="3">FSCB</collectionCode>
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),
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<location LSID="urn:lsid:plazi:treatment:03EDCF74FF82793397ECFDF8FC7093BB:8E9B28B9FF82793595AFFD70FC8695F6" box="[672,810,688,712]" country="Russia" municipality="Kurtuk" name="Rio Bistraya" pageId="2" pageNumber="3">Rio Bistraya</location>
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, 10.IX. 95,
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<collectingMunicipality box="[939,1017,688,712]" pageId="2" pageNumber="3">Kurtuk</collectingMunicipality>
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,
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<collectingCountry box="[1030,1092,688,712]" name="Russia" pageId="2" pageNumber="3">Rusia</collectingCountry>
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, F.
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</materialsCitation>
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</paragraph>
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<paragraph blockId="2.[227,1128,688,1552]" pageId="2" pageNumber="3">
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Ruano leg. (=
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<emphasis bold="true" box="[378,463,718,742]" pageId="2" pageNumber="3">
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<collectingCountry box="[378,458,718,742]" name="Russia" pageId="2" pageNumber="3">Russia</collectingCountry>
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:
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</emphasis>
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Irkutskaya oblast: Slyudyanskii District, environs of Kultuk village and Bistraya River,
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<date box="[520,644,748,772]" pageId="2" pageNumber="3" value="1995-09-10">10.IX 1995</date>
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, leg. F. Ruano).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
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<paragraph blockId="2.[227,1128,688,1552]" box="[265,1128,778,802]" pageId="2" pageNumber="3">DESCRIPTION. MALE. The specimen is broken into four separate body portions.</paragraph>
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<paragraph blockId="2.[227,1128,688,1552]" pageId="2" pageNumber="3">Length about 19, width 2.0 mm with paraterga. Coloration in alcohol including legs and antennae uniformly dark brown (probably the color was distorted by prolonged storage). Eyes black.</paragraph>
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<paragraph blockId="2.[227,1128,688,1552]" box="[265,1127,898,923]" pageId="2" pageNumber="3">Body with 32 segments. Head covered both with relatively long and short setae.</paragraph>
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<paragraph blockId="2.[227,1128,688,1552]" pageId="2" pageNumber="3">Eye patches subtriangular, with 28–29 ocelli. Collum semicircular. Both collum and somite 2 narrower than head with genae. Somite 2 wider than collum. Body width gradually increasing until somite 6–7, body parallel-sided on somites 21–22,</paragraph>
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<paragraph blockId="2.[227,1128,688,1552]" pageId="2" pageNumber="3">thereafter gradually tapering. Paraterga beginning on collum, well developed on somites 6–26, reduced on somite 27, onward missing. Metazonital macrochaetae in a transverse row on somites 28–31, like an extended triangle on preceding somites.</paragraph>
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<paragraph blockId="2.[227,1128,688,1552]" pageId="2" pageNumber="3">Anterolateral and medial macrochaetae subequal in length, caudolateral ones longest, distal part of all macrochaetae filiform, apex pointed. Axial suture well developed.</paragraph>
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<paragraph blockId="2.[227,1128,688,1552]" pageId="2" pageNumber="3">Legs long and slender. Leg pairs 1–2 typically reduced in size, with tarsal brushes as usual. Legs 3–7 increasingly enlarged toward gonopods. Femora 5–7 swollen and curved.</paragraph>
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<paragraph blockId="2.[227,1128,688,1552]" pageId="2" pageNumber="3">Leg pairs 3–7 with a small group of funnel–shaped tarsal papillae apically near claw. Claw of leg pairs 3–7 at base with a long setoid outgrowth ventrally but without additional claws dorsally. Postgonopodal legs (including leg pairs 10 and 11) with small group of funnel-shaped tarsal papillae apically near claw, however tarsal papillae gradually reduced toward posterior part of body. Claws of postgonopodal legs (including leg pairs 10 and 11) at base with two small additional claws dorsally and a relatively long setoid outgrowth ventrally, but small additional claws gradually missing toward posterior part of body; at least claws of hindmost legs at base devoid of such additional claw.</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7165037" ID-Zenodo-Dep="7165037" box="[265,1127,1410,1432]" httpUri="https://zenodo.org/record/7165037/files/figure.png" pageId="3" pageNumber="4" startId="3.[265,306,1410,1432]" targetBox="[227,1099,227,1367]" targetPageId="3">
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<paragraph blockId="3.[227,1127,1410,1598]" box="[265,1127,1410,1432]" pageId="3" pageNumber="4">
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Figs 1–5.
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<taxonomicName authority="Mikhaljova, 2019" authorityName="Mikhaljova" authorityYear="2019" box="[364,587,1410,1431]" class="Diplopoda" family="Diplomaragnidae" genus="Altajosoma" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kultukensis" status="sp. n.">
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<emphasis box="[364,587,1410,1431]" italics="true" pageId="3" pageNumber="4">Altajosoma kultukensis</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[595,651,1411,1432]" pageId="3" pageNumber="4">
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<taxonomicNameLabel box="[595,651,1411,1432]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, male holotype. 1 – coxa and trochanter 10; 2 –
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</paragraph>
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</caption>
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<paragraph blockId="3.[227,1127,1410,1598]" box="[227,1127,1438,1460]" pageId="3" pageNumber="4">gonopods (caudal view, telopodites of posterior gonopod omitted); 3 – gonopods (front view,</paragraph>
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<paragraph blockId="3.[227,1127,1410,1598]" pageId="3" pageNumber="4">
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telopodites of posterior gonopod omitted); 4 – gonopods (lateral view, telopodites of posterior gonopod omitted); 5 – telopodite of posterior gonopod;
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<emphasis bold="true" box="[873,900,1493,1514]" pageId="3" pageNumber="4">pp</emphasis>
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– posterior angiocoxal process of posterior gonopod;
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<emphasis bold="true" box="[527,552,1521,1542]" pageId="3" pageNumber="4">ap</emphasis>
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– anterior angiocoxal process of posterior gonopod;
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<emphasis bold="true" box="[1077,1106,1521,1542]" pageId="3" pageNumber="4">ms</emphasis>
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–
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</paragraph>
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<paragraph blockId="3.[227,1127,1410,1598]" pageId="3" pageNumber="4">
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mesal sheath process of colpocoxite;
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<emphasis bold="true" box="[590,610,1549,1570]" pageId="3" pageNumber="4">lp</emphasis>
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– lateral sheath process of colpocoxite;
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<emphasis bold="true" box="[1011,1022,1549,1570]" pageId="3" pageNumber="4">r</emphasis>
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– remnant of posterior gonopod telopodite. Scale in mm.
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</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" pageId="4" pageNumber="5">
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Legs 10 and 11 with coxal glands. Coxa 10 with a caudoventral conical process setose apically; trochanter 10 with a ventral low setose outgrowth (
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<figureCitation box="[949,1013,263,287]" captionStart="Figs 1–5" captionStartId="3.[265,306,1410,1432]" captionTargetBox="[227,1099,227,1367]" captionTargetId="figure-0@3.[227,1114,227,1378]" captionTargetPageId="3" captionText="Figs 1–5. Altajosoma kultukensis sp. n., male holotype. 1 – coxa and trochanter 10; 2 –" figureDoi="http://doi.org/10.5281/zenodo.7165037" httpUri="https://zenodo.org/record/7165037/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
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). Coxa 11
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</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" box="[227,1040,293,317]" pageId="4" pageNumber="5">with a caudoventral low setose outgrowth. Trochanter 11 without processes.</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" pageId="4" pageNumber="5">
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Gonopods large, curved posteriad distally (
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<figureCitation box="[749,851,323,347]" captionStart="Figs 1–5" captionStartId="3.[265,306,1410,1432]" captionTargetBox="[227,1099,227,1367]" captionTargetId="figure-0@3.[227,1114,227,1378]" captionTargetPageId="3" captionText="Figs 1–5. Altajosoma kultukensis sp. n., male holotype. 1 – coxa and trochanter 10; 2 –" figureDoi="http://doi.org/10.5281/zenodo.7165037" httpUri="https://zenodo.org/record/7165037/files/figure.png" pageId="4" pageNumber="5">Figs 2–4</figureCitation>
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; posterior gonopod telopodites broken). Anterior gonopods with subflagelliform 2-segmented telopodites,
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</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" box="[227,1127,383,407]" pageId="4" pageNumber="5">latter not hidden inside a narrow sheath groove; pointed tips of second telopodito-</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" box="[227,1127,413,437]" pageId="4" pageNumber="5">meres beset with cuticular spinules. Posterior gonopod colpocoxites fused to 2/3</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" box="[227,1127,443,467]" pageId="4" pageNumber="5">extent. Colpocoxites entire. Each angiocoxite with a globule in posterior view.</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" pageId="4" pageNumber="5">
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Posterior angiocoxal processes (
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<emphasis bold="true" box="[575,605,473,497]" pageId="4" pageNumber="5">pp</emphasis>
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) large, curved posteriad, rounded at tip, lacking teeth. Mesal (
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<emphasis bold="true" box="[374,407,503,527]" pageId="4" pageNumber="5">ms</emphasis>
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) and lateral
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<emphasis bold="true" box="[543,574,502,526]" pageId="4" pageNumber="5">(lp</emphasis>
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) sheath processes of colpocoxite small, originating near apex (see Remarks). Angiocoxites with depressions and ridges in anterior view,
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</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" pageId="4" pageNumber="5">
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each supplied with digitiform process (
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<emphasis bold="true" box="[665,693,563,587]" pageId="4" pageNumber="5">ap</emphasis>
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) blunted apically. Each colpocoxite in anterior view with longitudinal crest with broad apical portion. Posterior gonopod telopodite 2-segmented, setose, with a long femur and claw vestige (
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<figureCitation box="[958,1023,623,647]" captionStart="Figs 1–5" captionStartId="3.[265,306,1410,1432]" captionTargetBox="[227,1099,227,1367]" captionTargetId="figure-0@3.[227,1114,227,1378]" captionTargetPageId="3" captionText="Figs 1–5. Altajosoma kultukensis sp. n., male holotype. 1 – coxa and trochanter 10; 2 –" figureDoi="http://doi.org/10.5281/zenodo.7165037" httpUri="https://zenodo.org/record/7165037/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
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).
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</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" box="[265,485,653,677]" pageId="4" pageNumber="5">FEMALE unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="diagnosis">
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<paragraph blockId="4.[227,1127,233,1158]" box="[265,1127,683,707]" pageId="4" pageNumber="5">
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DIAGNOSIS. New species closely related to
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<taxonomicName authority="(Shear, 1990)" baseAuthorityName="Shear" baseAuthorityYear="1990" box="[744,1123,683,707]" class="Diplopoda" family="Diplomaragnidae" genus="Altajosoma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="shilenkovi">
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<emphasis box="[744,971,683,707]" italics="true" pageId="4" pageNumber="5">Altajosoma shilenkovi</emphasis>
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(Shear, 1990)
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</taxonomicName>
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,
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</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" pageId="4" pageNumber="5">but distinguished by the presence of anterior angiocoxal processes, configuration of posterior gonopod colpocoxites, telopodites without subapical triangular lamella,</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" box="[227,1072,773,797]" pageId="4" pageNumber="5">colpocoxites fused to 2/3 extent, small lateral sheath processes of colpocoxites.</paragraph>
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</subSubSection>
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<subSubSection box="[265,740,803,827]" pageId="4" pageNumber="5" type="distribution">
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<paragraph blockId="4.[227,1127,233,1158]" box="[265,740,803,827]" pageId="4" pageNumber="5">
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DISTRIBUTION.
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<collectingCountry box="[465,540,803,827]" name="Russia" pageId="4" pageNumber="5">Russia</collectingCountry>
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: Irkutskaya oblast.
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</paragraph>
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</subSubSection>
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<subSubSection box="[265,938,833,858]" pageId="4" pageNumber="5" type="etymology">
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<paragraph blockId="4.[227,1127,233,1158]" box="[265,938,833,858]" pageId="4" pageNumber="5">
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ETYMOLOGY. The specific epithet refers to the
|
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<typeStatus box="[798,844,834,858]" pageId="4" pageNumber="5">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="discussion">
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<paragraph blockId="4.[227,1127,233,1158]" pageId="4" pageNumber="5">REMARKS. It should be noted that distal parts of colpocoxites of a single examined male-holotype are dark brown and look worn like those of old specimens. As result the shape of mesal and lateral sheath processes of colpocoxite of young individuals may be some differ from that described above. However, to estimate the possible variations of the gonopod processes, additional material needs to be studied.</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" box="[265,1127,1013,1037]" pageId="4" pageNumber="5">
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<taxonomicName authority="Mikhaljova, 2019" authorityName="Mikhaljova" authorityYear="2019" box="[265,507,1013,1037]" class="Diplopoda" family="Diplomaragnidae" genus="Altajosoma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kultukensis" status="sp. n.">
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<emphasis box="[265,507,1013,1037]" italics="true" pageId="4" pageNumber="5">Altajosoma kultukensis</emphasis>
|
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</taxonomicName>
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<emphasis bold="true" box="[514,574,1013,1037]" pageId="4" pageNumber="5">
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<taxonomicNameLabel box="[514,574,1013,1037]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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can be conspecific with the
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<taxonomicName baseAuthorityName="Gerstfeldt" baseAuthorityYear="1859" box="[876,1127,1013,1037]" class="Diplopoda" family="Craspedosomatidae" genus="Craspedosoma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="armatus">
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<emphasis box="[876,1127,1013,1037]" italics="true" pageId="4" pageNumber="5">Craspedosoma armatus</emphasis>
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</taxonomicName>
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</paragraph>
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<paragraph blockId="4.[227,1127,233,1158]" box="[227,1127,1043,1068]" pageId="4" pageNumber="5">
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(Gerstfeldt, 1859). Latter was very poorly described from
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<collectingRegion box="[852,928,1043,1067]" country="Russia" name="Irkutsk" pageId="4" pageNumber="5">Irkutsk</collectingRegion>
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(Gerstfeldt, 1859)
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</paragraph>
|
||
<paragraph blockId="4.[227,1127,233,1158]" box="[227,1126,1073,1098]" pageId="4" pageNumber="5">
|
||
and recorded from Kultuk (Haase, 1880). But in the absence of
|
||
<typeStatus box="[950,996,1074,1098]" pageId="4" pageNumber="5">type</typeStatus>
|
||
material of
|
||
</paragraph>
|
||
<paragraph blockId="4.[227,1127,233,1158]" box="[227,1126,1103,1127]" pageId="4" pageNumber="5">
|
||
<taxonomicName baseAuthorityName="Gerstfeldt" baseAuthorityYear="1859" box="[227,492,1103,1127]" class="Diplopoda" family="Craspedosomatidae" genus="Craspedosoma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="armatum">
|
||
<emphasis box="[227,492,1103,1127]" italics="true" pageId="4" pageNumber="5">Craspedosoma armatum</emphasis>
|
||
</taxonomicName>
|
||
, ultimate solution can be achieved only when strict topo-
|
||
</paragraph>
|
||
<paragraph blockId="4.[227,1127,233,1158]" box="[227,988,1133,1158]" pageId="4" pageNumber="5">
|
||
<typeStatus box="[227,283,1134,1158]" pageId="4" pageNumber="5">types</typeStatus>
|
||
and material studied by Haase become available for examination.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |