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<document id="3BE1E177244C217D750EE095D12883EE" ID="10.11646/zootaxa.4136.1.1" ID-DOI="10.11646/zootaxa.4136.1.1" ID-GBIF-Dataset="98dfcf93-7454-4d14-b34e-9b39b6bbfd1c" ID-ISSN="1175-5326" ID-Zenodo-Dep="399782" ID-ZooBank="F8A4AE64-AD35-4FC5-966C-90FAC6588827" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1467724316376" checkinUser="plazi" docAuthor="Zhang, Ruilei, Song, Chao, Qi, Xin &amp; Wang, Xinhua" docDate="2016" docId="03ED87CDA630FFDCFF3DFBABFAB26DAB" docLanguage="en" docName="zootaxa.4136.1.1.pdf" docOrigin="Zootaxa 4136 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polypedilum (Tripodura) bilamella Zhang &amp; Wang, sp. n." docType="treatment" docVersion="15" lastPageNumber="12" masterDocId="FFD4FFB5A639FFD7FFAAFF93FB7A6929" masterDocTitle="Taxonomic review on the subgenus Tripodura Townes (Diptera: Chironomidae: Polypedilum) from China with eleven new species and a supplementary world checklist" masterLastPageNumber="53" masterPageNumber="1" pageNumber="10" updateTime="1698363515033" updateUser="plazi">
<mods:mods id="DA4046F017B617D4E64A3ADCC1C5F169" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0DECEC7D6DFE7DFA80192D25B0B0EE5B">
<mods:title id="377AA8E6BFACEB0B4DE4462168BBA11F">Taxonomic review on the subgenus Tripodura Townes (Diptera: Chironomidae: Polypedilum) from China with eleven new species and a supplementary world checklist</mods:title>
</mods:titleInfo>
<mods:name id="45FB8BCC7E94884C8429B0488231C156" type="personal">
<mods:role id="7BECD3D3BD594EE742722B8C226E8BBA">
<mods:roleTerm id="B09C887D641B5E580677B9C8D8DEA69C">Author</mods:roleTerm>
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<mods:namePart id="EDBD15E7D9A701FBB1BDCD4D8B55339B">Zhang, Ruilei</mods:namePart>
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<mods:namePart id="82A61432BD93638C8F370251946D9BB9">Song, Chao</mods:namePart>
</mods:name>
<mods:name id="D3EAC08A35BF905FDE7E3DC6B2A103BD" type="personal">
<mods:role id="B29DA3EA41B84D67E9244206F6DA9C97">
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<mods:namePart id="C0DE9AD53C5A9862CBD5F54F4BA309CD">Qi, Xin</mods:namePart>
</mods:name>
<mods:name id="7AF45A2D9B992F54B7FD59F1B1744E52" type="personal">
<mods:role id="D3AF0B59676F74CD12D9B377BEA62D05">
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<mods:namePart id="6E83E4F16EEE367788A8F4B70777E25A">Wang, Xinhua</mods:namePart>
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<mods:title id="9524B327FFCE2CD8AFEC33DEC573C3F3">Zootaxa</mods:title>
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<mods:part id="9BC1C07FAAF4B0FA1A821D0A883629CE">
<mods:date id="9F688C1C7387FD93CF420DD561050379">2016</mods:date>
<mods:detail id="BC56BB2FE602143430953B79F812FB7D" type="volume">
<mods:number id="4EBB2E0C0193186D57A0228C9F54261D">4136</mods:number>
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<mods:detail id="D257917A8AE6CFEDA4526EFD0EFED888" type="issue">
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<mods:classification id="887F6F608336E95DAC56814A29456ABC">journal article</mods:classification>
<mods:identifier id="D04587591D5A7101401F5A845A939D07" type="DOI">10.11646/zootaxa.4136.1.1</mods:identifier>
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<treatment id="03ED87CDA630FFDCFF3DFBABFAB26DAB" ID-DOI="http://doi.org/10.5281/zenodo.5634969" ID-GBIF-Taxon="121269058" ID-Zenodo-Dep="5634969" LSID="urn:lsid:plazi:treatment:03ED87CDA630FFDCFF3DFBABFAB26DAB" httpUri="http://treatment.plazi.org/id/03ED87CDA630FFDCFF3DFBABFAB26DAB" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
<subSubSection id="C35E6550A630FFDEFF3DFBABFA7D6D5A" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BFB36DBA630FFDEFF3DFBABF8346D7B" blockId="9.[151,846,1080,1139]" box="[151,846,1080,1106]" pageId="9" pageNumber="10">
<heading id="D0B381B7A630FFDEFF3DFBABF8346D7B" bold="true" box="[151,846,1080,1106]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B930EAC9A630FFDEFF3DFBABF8346D7B" bold="true" box="[151,846,1080,1106]" pageId="9" pageNumber="10">
<taxonomicName id="4C444D58A630FFDEFF3DFBABF87E6D7B" ID-CoL="9HK44" authority="Zhang &amp; Wang" authorityName="Zhang &amp; Wang" box="[151,772,1080,1106]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bilamella" status="sp. nov." subGenus="Tripodura">
<emphasis id="B930EAC9A630FFDEFF3DFBABFA556D7B" bold="true" box="[151,303,1080,1106]" italics="true" pageId="9" pageNumber="10">Polypedilum</emphasis>
(
<emphasis id="B930EAC9A630FFDEFE95FBABFAC06D7B" bold="true" box="[319,442,1080,1106]" italics="true" pageId="9" pageNumber="10">Tripodura</emphasis>
)
<emphasis id="B930EAC9A630FFDEFE61FBABF9466D7B" bold="true" box="[459,572,1080,1106]" italics="true" pageId="9" pageNumber="10">bilamella</emphasis>
Zhang &amp; Wang
</taxonomicName>
<taxonomicNameLabel id="A20357B2A630FFDEFCA6FBABF8346D7B" box="[780,846,1080,1106]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36DBA630FFDEFF3DFBC9FA7D6D5A" blockId="9.[151,846,1080,1139]" box="[151,263,1114,1139]" pageId="9" pageNumber="10">
(
<figureCitation id="137F2A5EA630FFDEFF34FBC9FB856D5A" box="[158,255,1114,1139]" captionStart="FIGURE 4 A – E" captionStartId="10.[151,250,1811,1833]" captionTargetBox="[160,1424,205,1785]" captionTargetId="figure@10.[150,1436,193,1790]" captionTargetPageId="10" captionText="FIGURE 4 A – E. Polypedilum (Tripodura) bilamella Zhang &amp; Wang sp. n. A. Wing. B. Fore tibial scale. C. Superior volsella. D. Hypopygium, dorsal and ventral view. E. Inferior volsella." httpUri="https://zenodo.org/record/399786/files/figure.png" pageId="9" pageNumber="10">Figure 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35E6550A630FFDEFF3DFB30F82A6D90" box="[151,848,1187,1209]" pageId="9" pageNumber="10" type="description">
<paragraph id="8BFB36DBA630FFDEFF3DFB30F82A6D90" blockId="9.[151,848,1187,1209]" box="[151,848,1187,1209]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB5411F5A630FFDEFF3DFB30F82A6D90" box="[151,848,1187,1209]" pageId="9" pageNumber="10">
<taxonomicName id="4C444D58A630FFDEFF3DFB30F92E6D90" ID-CoL="8DGF9" authority="Zettstedt" authorityName="Zettstedt" box="[151,596,1187,1209]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pullum" subGenus="Tripodura">
<emphasis id="B930EAC9A630FFDEFF3DFB30FA6D6D90" box="[151,279,1187,1209]" italics="true" pageId="9" pageNumber="10">Polypedilum</emphasis>
(
<emphasis id="B930EAC9A630FFDEFE8FFB30FAF66D90" box="[293,396,1187,1209]" italics="true" pageId="9" pageNumber="10">Tripodura</emphasis>
)
<emphasis id="B930EAC9A630FFDEFE31FB30FA9B6D90" box="[411,481,1187,1209]" italics="true" pageId="9" pageNumber="10">pullum</emphasis>
(Zettstedt)
</taxonomicName>
, sensu
<treatmentCitation id="0AE510CAA630FFDEFD36FB37F8366D90" author="Wang" box="[668,844,1187,1209]" page="646" pageId="9" pageNumber="10" year="2000">
<bibRefCitation id="EFD54B2AA630FFDEFD36FB37F8606D90" author="Wang" box="[668,794,1187,1209]" pageId="9" pageNumber="10" refString="Wang, X. H. (2000) A revised checklist of Chironomidae from China (Diptera). In: Hoffrichter, O. (Eds.), Late 20 th Century Research on Chironomidae. - An Anthology from 13 th International Symposium on Chironomidae. Shaker Verlag, Aachen, pp. 629 - 652." type="book chapter" year="2000">Wang, 2000</bibRefCitation>
: 646
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35E6550A630FFDEFF3DFB76FA3D6E4B" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BFB36DBA630FFDEFF3DFB76FA3D6E4B" blockId="9.[151,1437,1253,2034]" pageId="9" pageNumber="10">
<emphasis id="B930EAC9A630FFDEFF3DFB76FA3C6DD7" bold="true" box="[151,326,1253,1278]" pageId="9" pageNumber="10">
<typeStatus id="54FF8879A630FFDEFF3DFB76FBA86DD7" box="[151,210,1253,1278]" pageId="9" pageNumber="10">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54FF8879A630FFDEFEE5FB76FACD6DD7" box="[335,439,1253,1278]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
(BDN.10066), male,
<collectingCountry id="F353764BA630FFDEFD1AFB75F8766DD7" box="[688,780,1254,1278]" name="China" pageId="9" pageNumber="10">CHINA</collectingCountry>
, Xizang Autonomous Region, Ya’dong County, Yacheng,
<date id="FFFA101BA630FFDEFF3DFA9AFA336C0B" box="[151,329,1289,1314]" pageId="9" pageNumber="10" value="2003-08-26">26. VIII. 2003</date>
, light trap, H. Xue.
<typeStatus id="54FF8879A630FFDEFDE6FA99F9B86C0B" box="[588,706,1290,1314]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D42FD52A630FFDEFD6CFA99F86C6C0B" box="[710,790,1289,1314]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
, Hubei Province, Shengnongjia Natural Conversation, Hongping,
<quantity id="4CBC9B3EA630FFDEFEBCFABDFA1D6C6F" box="[278,359,1326,1350]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="10" unit="m" value="1800.0">1800m</quantity>
,
<date id="FFFA101BA630FFDEFEDEFABDF96B6C6F" box="[372,529,1326,1350]" pageId="9" pageNumber="10" value="1997-07-18">18. VII. 1997</date>
, light trap, Y. Du.
<specimenCount id="9D42FD52A630FFDEFD42FABDF8396C6F" box="[744,835,1325,1350]" pageId="9" pageNumber="10" type="male">3 males</specimenCount>
, Sichuan Province, Guanxian County,
<date id="FFFA101BA630FFDEFB56FABDFEE26C6F" box="[1276,1432,1325,1350]" pageId="9" pageNumber="10" value="1985-07-11">11. VII. 1985</date>
, light trap, X. Wang.
<specimenCount id="9D42FD52A630FFDEFED6FAC1FAAF6C43" box="[380,469,1361,1386]" pageId="9" pageNumber="10" type="male">3 males</specimenCount>
, Sichuan Province, Luding County,
<date id="FFFA101BA630FFDEFCC0FAC1F8976C43" box="[874,1005,1361,1386]" pageId="9" pageNumber="10" value="1996-06-07">7. VI. 1996</date>
, light trap, X. Wang.
<specimenCount id="9D42FD52A630FFDEFB77FAC2FE4D6C43" box="[1245,1335,1361,1386]" pageId="9" pageNumber="10" type="male">6 males</specimenCount>
, Sichuan Province, Kangding County, Zheduo River,
<quantity id="4CBC9B3EA630FFDEFD26FAE5F9A76CA7" box="[652,733,1398,1422]" metricMagnitude="3" metricUnit="m" metricValue="2.12" pageId="9" pageNumber="10" unit="m" value="2120.0">2120m</quantity>
, light trap,
<date id="FFFA101BA630FFDEFCCBFAE5F89C6CA7" box="[865,998,1397,1422]" pageId="9" pageNumber="10" value="1996-06-08">8. VI. 1996</date>
, X. Wang.
<specimenCount id="9D42FD52A630FFDEFBCDFAE5FFB86CA7" box="[1127,1218,1397,1422]" pageId="9" pageNumber="10" type="male">3 males</specimenCount>
, Sichuan Province, Yajiang County, Sandaoqiao,
<quantity id="4CBC9B3EA630FFDEFE40FA0AF9416C9B" box="[490,571,1433,1458]" metricMagnitude="3" metricUnit="m" metricValue="2.46" pageId="9" pageNumber="10" unit="m" value="2460.0">2460m</quantity>
,
<date id="FFFA101BA630FFDEFDE2FA09F9B56C9B" box="[584,719,1433,1458]" pageId="9" pageNumber="10" value="1996-06-09">9. VI. 1996</date>
, sweep net, X. Wang.
<specimenCount id="9D42FD52A630FFDEFC7EFA09FF4A6C9B" box="[980,1072,1433,1458]" pageId="9" pageNumber="10" type="male">8 males</specimenCount>
, Sichuan Province, Xiangcheng County, Shuoqu River,
<quantity id="4CBC9B3EA630FFDEFE01FA2EFA866CFF" box="[427,508,1469,1494]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="9" pageNumber="10" unit="m" value="2750.0">2750m</quantity>
,
<date id="FFFA101BA630FFDEFDA6FA2DF9DF6CFF" box="[524,677,1469,1494]" pageId="9" pageNumber="10" value="1996-06-12">12. VI. 1996</date>
, light trap, X. Wang.
<specimenCount id="9D42FD52A630FFDEFC06FA2DFF706CFF" box="[940,1034,1469,1494]" pageId="9" pageNumber="10" type="male">3 males</specimenCount>
, Sichuan Province, Garzê County, Yajiang,
<quantity id="4CBC9B3EA630FFDEFF51FA71FA366CD3" box="[251,332,1506,1530]" metricMagnitude="3" metricUnit="m" metricValue="2.83" pageId="9" pageNumber="10" unit="m" value="2830.0">2830m</quantity>
,
<date id="FFFA101BA630FFDEFEFCFA71FA9D6CD3" box="[342,487,1505,1530]" pageId="9" pageNumber="10" value="1996-06-14">14. VI. 1996</date>
, light trap, X. Wang.
<specimenCount id="9D42FD52A630FFDEFD7DFA71F85F6CD3" box="[727,805,1505,1530]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
, Sichuan Province, Kangding County, Wasigou,
<quantity id="4CBC9B3EA630FFDEFAEDFA71FEE26CD3" box="[1351,1432,1506,1530]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="m" value="2000.0">2000m</quantity>
, light trap,
<date id="FFFA101BA630FFDEFEA0F996FAE06F37" box="[266,410,1541,1566]" pageId="9" pageNumber="10" value="1996-06-15">15. VI. 1996</date>
, X. Wang.
<specimenCount id="9D42FD52A630FFDEFDBCF995F9146F37" box="[534,622,1541,1566]" pageId="9" pageNumber="10" type="male">2 males</specimenCount>
, Sichuan Ya’an City, Zhougong River,
<date id="FFFA101BA630FFDEFB8EF995FFCE6F37" box="[1060,1204,1541,1566]" pageId="9" pageNumber="10" value="1996-06-18">18. VI. 1996</date>
, light trap, X. Wang.
<specimenCount id="9D42FD52A630FFDEFF3DF9BAFB886F6B" box="[151,242,1577,1602]" pageId="9" pageNumber="10" type="male">6 males</specimenCount>
, Sichuan Province, Baoxing County, Xilong Town,
<quantity id="4CBC9B3EA630FFDEFCE1F9B9F8E06F6B" box="[843,922,1578,1602]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="10" unit="m" value="1100.0">1100m</quantity>
,
<date id="FFFA101BA630FFDEFC02F9B9FF446F6B" box="[936,1086,1577,1602]" pageId="9" pageNumber="10" value="1996-06-19">19. VI. 1996</date>
, light trap, X. Wang.
<specimenCount id="9D42FD52A630FFDEFA96F9B9FEE26F6B" box="[1340,1432,1577,1602]" pageId="9" pageNumber="10" type="male">4 males</specimenCount>
, Yunnan Province, Dali City, Diancangshan,
<quantity id="4CBC9B3EA630FFDEFD25F9DEF99A6F4F" box="[655,736,1613,1638]" metricMagnitude="3" metricUnit="m" metricValue="2.52" pageId="9" pageNumber="10" unit="m" value="2520.0">2520m</quantity>
, 23,
<date id="FFFA101BA630FFDEFCB2F9DDF80E6F4F" box="[792,884,1613,1638]" pageId="9" pageNumber="10" value="1996-05">V. 1996</date>
, light trap, B. Wang.
<specimenCount id="9D42FD52A630FFDEFBC1F9DDFFBC6F4F" box="[1131,1222,1613,1638]" pageId="9" pageNumber="10" type="male">3 males</specimenCount>
, Yunnan Province, Eryuan County, Niujie Township, Futianlin, Meici River,
<quantity id="4CBC9B3EA630FFDEFCB4F9E2F8156FA3" box="[798,879,1649,1674]" metricMagnitude="3" metricUnit="m" metricValue="2.2619999999999996" pageId="9" pageNumber="10" unit="m" value="2262.0">2262m</quantity>
, light trap,
<date id="FFFA101BA630FFDEFC44F9E1FF086FA3" box="[1006,1138,1649,1674]" pageId="9" pageNumber="10" value="1996-05-24">24. V. 1996</date>
. C. Zhou.
<specimenCount id="9D42FD52A630FFDEFB4DF9E1FE4F6FA3" box="[1255,1333,1649,1674]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
, Shannxi Province, Zhouzhi County, Banfangzi Town,
<date id="FFFA101BA630FFDEFD30F905F8436F87" box="[666,825,1686,1710]" pageId="9" pageNumber="10" value="1994-08-10">10. VIII 1994</date>
, W. Bu.
<specimenCount id="9D42FD52A630FFDEFC37F905F8976F87" box="[925,1005,1685,1710]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
, Shannxi Province, Ningshan County, Xunyangba,
<date id="FFFA101BA630FFDEFE82F92AFAAB6FFB" box="[296,465,1721,1746]" pageId="9" pageNumber="10" value="1994-08-16">16. VIII. 1994</date>
, sweep net, B. Wen.
<specimenCount id="9D42FD52A630FFDEFD6DF92AF8586FFB" box="[711,802,1721,1746]" pageId="9" pageNumber="10" type="male">5 males</specimenCount>
, Gansu Province, Yuzhong County, Xinlong Mountain,
<quantity id="4CBC9B3EA630FFDEFF3DF94DFB926FDF" box="[151,232,1758,1782]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="10" unit="m" value="2200.0">2200m</quantity>
,
<date id="FFFA101BA630FFDEFF58F94DFACC6FDF" box="[242,438,1758,1782]" pageId="9" pageNumber="10" value="1993-07-30" valueMax="1993-07-31" valueMin="1993-07-30">30–31. VII. 1993</date>
, sweep net, W. Bu.
<specimenCount id="9D42FD52A630FFDEFD3BF94DF9A56FDF" box="[657,735,1757,1782]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
, Gansu Province, Yuzhong County, Maxian Mountain,
<date id="FFFA101BA630FFDEFAE3F94EFBA86E33" pageId="9" pageNumber="10" value="1993-08-05">5. VIII. 1993</date>
, sweep net, W. Bu.
<specimenCount id="9D42FD52A630FFDEFE1DF891F97D6E33" box="[439,519,1793,1818]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
, Gansu Province, Yongdeng County, Liancheng Township, Tulugou,
<quantity id="4CBC9B3EA630FFDEFAB6F891FE176E33" box="[1308,1389,1794,1818]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="10" unit="m" value="2300.0">2300m</quantity>
,
<date id="FFFA101BA630FFDEFAD3F892FA6A6E17" pageId="9" pageNumber="10" value="1993-08-16">16. VIII. 1993</date>
, light trap, W. Bu.
<specimenCount id="9D42FD52A630FFDEFE49F8B5F9306E17" box="[483,586,1829,1854]" pageId="9" pageNumber="10" type="male">13 males</specimenCount>
, Ningxia Autonomous Region, Liuban Mountain Natural Converse,
<date id="FFFA101BA630FFDEFAE3F8B6FBA86E4B" pageId="9" pageNumber="10" value="1987-08-06">6. VIII. 1987</date>
, X. Wang.
</paragraph>
</subSubSection>
<subSubSection id="C35E6550A630FFDEFF6DF8FEF90A6EDB" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8BFB36DBA630FFDEFF6DF8FEFA396E83" blockId="9.[151,1437,1253,2034]" pageId="9" pageNumber="10">
<emphasis id="B930EAC9A630FFDEFF6DF8FEFA346EAF" bold="true" box="[199,334,1901,1926]" pageId="9" pageNumber="10">Etymology.</emphasis>
From Latin,
<emphasis id="B930EAC9A630FFDEFE42F8FEFA876EAF" box="[488,509,1901,1926]" italics="true" pageId="9" pageNumber="10">bi</emphasis>
, two and
<emphasis id="B930EAC9A630FFDEFDDEF8FEF9B26EAF" box="[628,712,1901,1926]" italics="true" pageId="9" pageNumber="10">lamella</emphasis>
, lamella, thin plate, referring to the anal point with two dorsal lobed lamellae.
</paragraph>
<paragraph id="8BFB36DBA630FFDEFF6DF826F90A6EDB" blockId="9.[151,1437,1253,2034]" pageId="9" pageNumber="10">
<emphasis id="B930EAC9A630FFDEFF6DF826FAAF6EE7" bold="true" box="[199,469,1973,1998]" pageId="9" pageNumber="10">Diagnostic characters.</emphasis>
Wing is full of gray marking. Anal point bears tow dosal lobed lamellae. Lateral projection of anal point is well developed.
</paragraph>
</subSubSection>
<caption id="DF3B6653A633FFDDFF3DF880F87C6E61" httpUri="https://zenodo.org/record/399786/files/figure.png" pageId="10" pageNumber="11" targetBox="[160,1424,205,1785]" targetPageId="10">
<paragraph id="8BFB36DBA633FFDDFF3DF880F87C6E61" blockId="10.[151,1437,1811,1864]" pageId="10" pageNumber="11">
<emphasis id="B930EAC9A633FFDDFF3DF880FA336E01" bold="true" box="[151,329,1811,1833]" pageId="10" pageNumber="11">FIGURE 4 A–E.</emphasis>
<taxonomicName id="4C444D58A633FFDDFEE5F880F82C6E00" authority="Zhang &amp; Wang" authorityName="Zhang &amp; Wang" box="[335,854,1811,1833]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bilamella" status="sp. nov." subGenus="Tripodura">
<emphasis id="B930EAC9A633FFDDFEE5F880FAB56E00" box="[335,463,1811,1833]" italics="true" pageId="10" pageNumber="11">Polypedilum</emphasis>
(
<emphasis id="B930EAC9A633FFDDFE74F880F93E6E00" box="[478,580,1811,1833]" italics="true" pageId="10" pageNumber="11">Tripodura</emphasis>
)
<emphasis id="B930EAC9A633FFDDFDFEF880F9CE6E00" box="[596,692,1811,1833]" italics="true" pageId="10" pageNumber="11">bilamella</emphasis>
Zhang &amp; Wang
</taxonomicName>
<taxonomicNameLabel id="A20357B2A633FFDDFCF6F880F8EF6E01" box="[860,917,1811,1832]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B930EAC9A633FFDDFCF6F880F8EF6E01" bold="true" box="[860,917,1811,1832]" pageId="10" pageNumber="11">sp. n.</emphasis>
</taxonomicNameLabel>
A. Wing. B. Fore tibial scale. C. Superior volsella. D. Hypopygium, dorsal and ventral view. E. Inferior volsella.
</paragraph>
</caption>
<subSubSection id="C35E6550A633FFDCFF6DF8FCF8B86A17" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
<paragraph id="8BFB36DBA633FFDDFF6DF8FCF8B16E85" blockId="10.[151,1436,1903,2036]" pageId="10" pageNumber="11">
<emphasis id="B930EAC9A633FFDDFF6DF8FCFA2C6EA1" bold="true" box="[199,342,1903,1928]" pageId="10" pageNumber="11">Male imago</emphasis>
(n=20). Total length 2.10–3.60,
<quantity id="4CBC9B3EA633FFDDFD66F8E3F84E6EAE" box="[716,820,1904,1928]" metricMagnitude="-3" metricUnit="m" metricValue="2.89" pageId="10" pageNumber="11" unit="mm" value="2.89">2.89 mm</quantity>
. Wing length 1.43–2.10,
<quantity id="4CBC9B3EA633FFDDFBFCF8E3FFC56EAE" box="[1110,1215,1904,1928]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="10" pageNumber="11" unit="mm" value="1.78">1.78 mm</quantity>
. Total length/ wing length 1.39–1.88, 1.63. Wing length/ length of profemur 2.17–2.82, 2.56.
</paragraph>
<paragraph id="8BFB36DBA633FFDDFF6DF824FF4E6EDD" blockId="10.[151,1436,1903,2036]" pageId="10" pageNumber="11">
<emphasis id="B930EAC9A633FFDDFF6DF824FA386EF9" box="[199,322,1975,2000]" italics="true" pageId="10" pageNumber="11">Coloration</emphasis>
. Head dark brown. Thorax yellowish brown to dark brown. Legs yellowish brown to brown, femora with pale sub-apical ring. Abdominal segments yellowish brown to brown.
</paragraph>
<paragraph id="8BFB36DBA632FFDCFF6DFF04FA0B6835" blockId="11.[151,1437,151,1154]" pageId="11" pageNumber="12">
<emphasis id="B930EAC9A632FFDCFF6DFF04FA7E6999" box="[199,260,151,176]" italics="true" pageId="11" pageNumber="12">Head</emphasis>
. AR 0.40–0.73, 0.54. Ultimate flagellomere 90–410, 250 µm long. Temporal setae 10–14, 12; including 2–4, 3 inner verticals; 4–8, 5 outer verticals and 2–6, 4 postorbitals. Clypeus with 13–27, 19 setae. Tentorium 100– 150, 118 µm long; 20–35, 26 µm wide. Palpomere lengths (in µm): 28–50, 37; 30–48, 38; 75–113, 96; 90–128, 109; 173–228, 192.
</paragraph>
<paragraph id="8BFB36DBA632FFDCFF6DFEBAF802684E" blockId="11.[151,1437,151,1154]" pageId="11" pageNumber="12">
<emphasis id="B930EAC9A632FFDCFF6DFEBAFB856869" box="[199,255,297,320]" italics="true" pageId="11" pageNumber="12">Wing</emphasis>
(
<figureCitation id="137F2A5EA632FFDCFEA5FEB4FAFD6869" box="[271,391,295,320]" captionStart="FIGURE 4 A – E" captionStartId="10.[151,250,1811,1833]" captionTargetBox="[160,1424,205,1785]" captionTargetId="figure@10.[150,1436,193,1790]" captionTargetPageId="10" captionText="FIGURE 4 A – E. Polypedilum (Tripodura) bilamella Zhang &amp; Wang sp. n. A. Wing. B. Fore tibial scale. C. Superior volsella. D. Hypopygium, dorsal and ventral view. E. Inferior volsella." httpUri="https://zenodo.org/record/399786/files/figure.png" pageId="11" pageNumber="12">Figure 4A</figureCitation>
). Wing widely coved with obscure markings. VR 1.19–1.35, 1.29. R with 14–25, 20 setae; R1 with 11–20, 15; R4+5 with 22–38, 29. Squama with 6–16, 10 setae.
</paragraph>
<paragraph id="8BFB36DBA632FFDCFF6DFEE6FA856898" blockId="11.[151,1437,151,1154]" pageId="11" pageNumber="12">
<emphasis id="B930EAC9A632FFDCFF6DFEE6FA6268A7" box="[199,280,373,398]" italics="true" pageId="11" pageNumber="12">Thorax</emphasis>
. Dorsocentrals 17–24, 20, including 2–4, 3 humerals; acrostichals 13–22, 18; prealars 4–7, 5. Scutellum with 12–28, 18 setae.
</paragraph>
<paragraph id="8BFB36DBA632FFDCFF6DFE2DF97C6B4F" blockId="11.[151,1437,151,1154]" pageId="11" pageNumber="12">
<emphasis id="B930EAC9A632FFDCFF6DFE2DFB8668FC" box="[199,252,446,469]" italics="true" pageId="11" pageNumber="12">Legs</emphasis>
. Terminal scale (
<figureCitation id="137F2A5EA632FFDCFE6EFE2EF94068FF" box="[452,570,445,470]" captionStart="FIGURE 4 A – E" captionStartId="10.[151,250,1811,1833]" captionTargetBox="[160,1424,205,1785]" captionTargetId="figure@10.[150,1436,193,1790]" captionTargetPageId="10" captionText="FIGURE 4 A – E. Polypedilum (Tripodura) bilamella Zhang &amp; Wang sp. n. A. Wing. B. Fore tibial scale. C. Superior volsella. D. Hypopygium, dorsal and ventral view. E. Inferior volsella." httpUri="https://zenodo.org/record/399786/files/figure.png" pageId="11" pageNumber="12">Figure 4B</figureCitation>
) of front tibia 33–55, 44 µm long, triangular and apically pointed. Spur of middle tibia 40–65, 49 um long including 20–33, 26 µm long comb, un-spurred comb 18–28, 23 µm long, spur of hind tibia 40–65, 52 µm long including 25–40, 30 µm long comb, un-spurred comb 23–38, 28 µm long. Width at apex of front tibia 40–53, 46 µm, of middle tibia 43–65, 51 µm, of hind tibia 45–70, 59 µm. Lengths (in µm) and proportions of legs as in
<tableCitation id="C6C60360A632FFDCFE06FDDEF97A6B4F" box="[428,512,589,614]" captionStart="TABLE 1" captionStartId="3.[125,201,606,626]" captionTargetBox="[126,1891,656,1541]" captionTargetPageId="3" captionText="TABLE 1. Lengths (in µm) and proportions of legs." httpUri="http://table.plazi.org/id/DF3B6653A63AFFD4FFD7FDCDF9236B5B" pageId="11" pageNumber="12" tableUuid="DF3B6653A63AFFD4FFD7FDCDF9236B5B">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BFB36DBA632FFDCFF6DFDE1F8B86A17" blockId="11.[151,1437,151,1154]" pageId="11" pageNumber="12">
<emphasis id="B930EAC9A632FFDCFF6DFDE1FA2C6BA0" box="[199,342,626,649]" italics="true" pageId="11" pageNumber="12">Hypopygium</emphasis>
(
<figureCitation id="137F2A5EA632FFDCFEC7FDE2F9766BA3" box="[365,524,625,650]" captionStart="FIGURE 4 A – E" captionStartId="10.[151,250,1811,1833]" captionTargetBox="[160,1424,205,1785]" captionTargetId="figure@10.[150,1436,193,1790]" captionTargetPageId="10" captionText="FIGURE 4 A – E. Polypedilum (Tripodura) bilamella Zhang &amp; Wang sp. n. A. Wing. B. Fore tibial scale. C. Superior volsella. D. Hypopygium, dorsal and ventral view. E. Inferior volsella." httpUri="https://zenodo.org/record/399786/files/figure.png" pageId="11" pageNumber="12">Figure 4C–E</figureCitation>
). Anal tergite bands fused basal. Tergite IX with 6–18, 10 median setae. Laterosternite with 2–3, 2 setae. Anal point 53–88, 72 µm long, lateral projection 13–33, 22 µm long. Phallapodeme 65–100, 89 µm long; transverse sternapodeme 25–53, 42 µm long. Gonocoxite 103–163, 136 µm long. Superior volsella (
<figureCitation id="137F2A5EA632FFDCFE07FD4EF95E6BDF" box="[429,548,733,758]" captionStart="FIGURE 4 A – E" captionStartId="10.[151,250,1811,1833]" captionTargetBox="[160,1424,205,1785]" captionTargetId="figure@10.[150,1436,193,1790]" captionTargetPageId="10" captionText="FIGURE 4 A – E. Polypedilum (Tripodura) bilamella Zhang &amp; Wang sp. n. A. Wing. B. Fore tibial scale. C. Superior volsella. D. Hypopygium, dorsal and ventral view. E. Inferior volsella." httpUri="https://zenodo.org/record/399786/files/figure.png" pageId="11" pageNumber="12">Figure 4C</figureCitation>
) pad–like, expanded distally, with 5–6 short inner setae and 2–3 long apical setae. Inferior volsella 95–140, 120 µm long, with 8–13, 11 setae. Gonostylus 120–193, 164 µm long, with 5–7 long setae along inner margin. HR 0.69–0.92, 0.83. HV 1.53–1.94, 1.77.
</paragraph>
</subSubSection>
<subSubSection id="C35E6550A632FFDCFF6DFCDAF8896A4B" box="[199,1011,841,866]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8BFB36DBA632FFDCFF6DFCDAF8896A4B" blockId="11.[151,1437,151,1154]" box="[199,1011,841,866]" pageId="11" pageNumber="12">
<emphasis id="B930EAC9A632FFDCFF6DFCDAFA1A6A4B" bold="true" box="[199,352,841,866]" pageId="11" pageNumber="12">Distribution.</emphasis>
It was collected from both Palaearctic and Oriental
<collectingCountry id="F353764BA632FFDCFC02FCDAF8956A4B" box="[936,1007,841,866]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E6550A632FFDCFF6DFCFEFAB26DAB" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8BFB36DBA632FFDCFF6DFCFEFAB26DAB" blockId="11.[151,1437,151,1154]" pageId="11" pageNumber="12">
<emphasis id="B930EAC9A632FFDCFF6DFCFEFA416AAF" bold="true" box="[199,315,877,902]" pageId="11" pageNumber="12">Remarks.</emphasis>
The present new species is very close to
<taxonomicName id="4C444D58A632FFDCFCBBFCFDFFCD6AAF" authority="Tokunaga" authorityName="Tokunaga" box="[785,1207,877,902]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="decematoguttatum" subGenus="Tripodura">
<emphasis id="B930EAC9A632FFDCFCBBFCFDF8656AAC" box="[785,799,878,901]" italics="true" pageId="11" pageNumber="12">P</emphasis>
. (
<emphasis id="B930EAC9A632FFDCFC9DFCFDF83E6AAC" box="[823,836,878,901]" italics="true" pageId="11" pageNumber="12">T</emphasis>
.)
<emphasis id="B930EAC9A632FFDCFCF6FCFEFF516AAF" box="[860,1067,877,902]" italics="true" pageId="11" pageNumber="12">decematoguttatum</emphasis>
(Tokunaga)
</taxonomicName>
,
<taxonomicName id="4C444D58A632FFDCFB69FCFDFA396A83" authority="Zhang &amp; Wang" authorityName="Zhang &amp; Wang" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="napahaiense" status="sp. nov." subGenus="Tripodura">
<emphasis id="B930EAC9A632FFDCFB69FCFDFFAB6AAC" box="[1219,1233,878,901]" italics="true" pageId="11" pageNumber="12">P</emphasis>
. (
<emphasis id="B930EAC9A632FFDCFB43FCFDFF8C6AAC" box="[1257,1270,878,901]" italics="true" pageId="11" pageNumber="12">T</emphasis>
.)
<emphasis id="B930EAC9A632FFDCFAA4FCFEFEE66AAF" box="[1294,1436,877,902]" italics="true" pageId="11" pageNumber="12">napahaiense</emphasis>
Zhang &amp; Wang
</taxonomicName>
<taxonomicNameLabel id="A20357B2A632FFDCFEE3FC02FAF26A83" box="[329,392,913,938]" pageId="11" pageNumber="12" rank="species">
<emphasis id="B930EAC9A632FFDCFEE3FC02FAF26A83" bold="true" box="[329,392,913,938]" pageId="11" pageNumber="12">sp. n.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C444D58A632FFDCFE14FC01F8236A83" authority="Zhang &amp; Wang" authorityName="Zhang &amp; Wang" box="[446,857,913,938]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mengmanense" status="sp. nov." subGenus="Tripodura">
<emphasis id="B930EAC9A632FFDCFE14FC01FAB66A80" box="[446,460,914,937]" italics="true" pageId="11" pageNumber="12">P</emphasis>
. (
<emphasis id="B930EAC9A632FFDCFE49FC01FA8A6A80" box="[483,496,914,937]" italics="true" pageId="11" pageNumber="12">T</emphasis>
.)
<emphasis id="B930EAC9A632FFDCFDA2FC01F9DD6A80" box="[520,679,914,937]" italics="true" pageId="11" pageNumber="12">mengmanense</emphasis>
Zhang &amp; Wang
</taxonomicName>
<taxonomicNameLabel id="A20357B2A632FFDCFCF5FC02F8E56A83" box="[863,927,913,938]" pageId="11" pageNumber="12" rank="species">
<emphasis id="B930EAC9A632FFDCFCF5FC02F8E56A83" bold="true" box="[863,927,913,938]" pageId="11" pageNumber="12">sp. n.</emphasis>
</taxonomicNameLabel>
in male hypopygial morphology that the anal point bears two dorsal lobed lamellae. We must be very careful when we observe above 4 species under microscope, because the dorsal lobed lamellae is transparent and easy to ignore in dorsal view. The above 4 species show a bit difference in wing markings and the length of lateral projection of anal point;
<bibRefCitation id="EFD54B2AA632FFDCFB3BFC6DFE256D3F" box="[1169,1375,1021,1046]" pageId="11" pageNumber="12" refString="Song, C., Wang, Q., Zhang, R. L., Sun, B. J. &amp; Wang, X. H. (2016) Exploring the utility of DNA barcoding in species delimitation of Polypedilum (Tripodura) non-biting midges (Diptera: Chironomidae). Zootaxa, 4079 (5), 534 - 550. http: // dx. doi. org / 10.11646 / zootaxa. 4079.5.2" type="journal article">
Song
<emphasis id="B930EAC9A632FFDCFB7EFC6CFF926D3F" box="[1236,1256,1023,1046]" italics="true" pageId="11" pageNumber="12">et</emphasis>
al. (2016)
</bibRefCitation>
have certified these bit differences can result in inter-specific divergence by molecular data.
<bibRefCitation id="EFD54B2AA632FFDCFBD9FBB1FE7C6D13" author="Wang" box="[1139,1286,1058,1082]" pageId="11" pageNumber="12" refString="Wang, X. H. (2000) A revised checklist of Chironomidae from China (Diptera). In: Hoffrichter, O. (Eds.), Late 20 th Century Research on Chironomidae. - An Anthology from 13 th International Symposium on Chironomidae. Shaker Verlag, Aachen, pp. 629 - 652." type="book chapter" year="2000">Wang (2000)</bibRefCitation>
had wrongly identified some specimens of
<taxonomicName id="4C444D58A632FFDCFE4EFBD4F8336D77" authority="Zhang &amp; Wang" authorityName="Zhang &amp; Wang" box="[484,841,1093,1118]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bilamella" status="sp. nov." subGenus="Tripodura">
<emphasis id="B930EAC9A632FFDCFE4EFBD4FA8C6D77" box="[484,502,1095,1118]" italics="true" pageId="11" pageNumber="12">P.</emphasis>
(
<emphasis id="B930EAC9A632FFDCFDA3FBD4F9666D77" box="[521,540,1095,1118]" italics="true" pageId="11" pageNumber="12">T.</emphasis>
)
<emphasis id="B930EAC9A632FFDCFD87FBD6F9ED6D77" box="[557,663,1093,1118]" italics="true" pageId="11" pageNumber="12">bilamella</emphasis>
Zhang &amp; Wang
</taxonomicName>
<taxonomicNameLabel id="A20357B2A632FFDCFCFAFBD6F8F56D77" box="[848,911,1093,1118]" pageId="11" pageNumber="12" rank="species">
<emphasis id="B930EAC9A632FFDCFCFAFBD6F8F56D77" bold="true" box="[848,911,1093,1118]" pageId="11" pageNumber="12">sp. n.</emphasis>
</taxonomicNameLabel>
collected from Ningxia Autonomous region as
<taxonomicName id="4C444D58A632FFDCFF3DFBF9FABE6DAB" authority="Zetterstedt" authorityName="Zetterstedt" box="[151,452,1129,1154]" class="Insecta" family="Chironomidae" genus="Polypedilum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pullum" subGenus="Tripodura">
<emphasis id="B930EAC9A632FFDCFF3DFBF9FBDF6DA8" box="[151,165,1130,1153]" italics="true" pageId="11" pageNumber="12">P</emphasis>
. (
<emphasis id="B930EAC9A632FFDCFF16FBF9FBB36DA8" box="[188,201,1130,1153]" italics="true" pageId="11" pageNumber="12">T</emphasis>
.)
<emphasis id="B930EAC9A632FFDCFF4BFBFAFA556DAB" box="[225,303,1129,1154]" italics="true" pageId="11" pageNumber="12">pullum</emphasis>
(Zetterstedt)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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