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<document ID-DOI="http://doi.org/10.5281/zenodo.5131745" ID-GBIF-Dataset="09f66541-18e3-4044-b553-7aab04763396" ID-GBIF-Taxon="183733957" ID-ISSN="1982-0224" ID-Zenodo-Dep="5131745" LSID="urn:lsid:zoobank.org:act:10925146-F8F9-4441-AF1F-399F9AD36197" checkinTime="1616760176976" checkinUser="carolina" docAuthor="Menezes, Naércio A. &amp; de Lucena, Carlos Alberto S." docDate="2014" docId="03C4D93834733F48FED8F893FBBF618A" docLanguage="en" docName="NeotropIchthyol.12.2.193-228.pdf" docOrigin="Neotropical Ichthyology 12 (2)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252014000200193&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Charax delimai Menezes &amp; de Lucena 2014, new species" docType="treatment" docUuid="10925146-F8F9-4441-AF1F-399F9AD36197" docUuidSource="ZooBank" docVersion="6" lastPageId="10" lastPageNumber="203" masterDocId="FFFDA140347A3F42FFFBFE69FFCB6623" masterDocTitle="A taxonomic review of the species of Charax Scopoli, 1777 (Teleostei: Characidae: Characinae) with description of a new species from the rio Negro bearing superficial neuromasts on body scales, Amazon basin, Brazil" masterLastPageNumber="228" masterPageNumber="193" pageId="9" pageNumber="202" updateTime="1634698513480" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A taxonomic review of the species of Charax Scopoli, 1777 (Teleostei: Characidae: Characinae) with description of a new species from the rio Negro bearing superficial neuromasts on body scales, Amazon basin, Brazil</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Menezes, Naércio A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>de Lucena, Carlos Alberto S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>12</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>193</mods:start>
<mods:end>228</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252014000200193&amp;lng=en&amp;tlng=en</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/1982-0224-20130175</mods:identifier>
<mods:identifier type="GBIF-Dataset">09f66541-18e3-4044-b553-7aab04763396</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">4638995</mods:identifier>
<mods:identifier type="ZooBank">2CFAB1CE-D283-483B-8C2B-87D95AEADE70</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5131745" ID-GBIF-Taxon="183733957" ID-Zenodo-Dep="5131745" LSID="urn:lsid:zoobank.org:act:10925146-F8F9-4441-AF1F-399F9AD36197" httpUri="http://treatment.plazi.org/id/03C4D93834733F48FED8F893FBBF618A" lastPageId="10" lastPageNumber="203" pageId="9" pageNumber="202">
<subSubSection box="[291,609,1786,1810]" pageId="9" pageNumber="202" type="nomenclature">
<paragraph blockId="9.[291,609,1786,1842]" box="[291,609,1786,1810]" pageId="9" pageNumber="202">
<heading box="[291,609,1786,1810]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="202" reason="2">
<emphasis bold="true" box="[291,609,1786,1810]" pageId="9" pageNumber="202">
<taxonomicName authority="Menezes &amp; de Lucena, 2014" authorityName="Menezes &amp; de Lucena" authorityYear="2014" box="[291,463,1786,1810]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="delimai" status="new species">
<emphasis bold="true" box="[291,463,1786,1810]" italics="true" pageId="9" pageNumber="202">Charax delimai</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[477,609,1786,1810]" pageId="9" pageNumber="202" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[415,485,1818,1842]" pageId="9" pageNumber="202" type="description">
<paragraph blockId="9.[291,609,1786,1842]" box="[415,485,1818,1842]" pageId="9" pageNumber="202">
<figureCitation box="[415,485,1818,1842]" captionStart="Fig" captionStartId="9.[129,168,720,744]" captionTargetBox="[129,1452,176,703]" captionTargetId="figure-577@9.[129,1452,176,703]" captionTargetPageId="9" captionText="Fig. 10. Charax delimai, MZUSP 81505, holotype, mature male, 100 mm SL, Brazil, Amazonas, community of São Pedro, rio Tiquié, rio Negro drainage." figureDoi="http://doi.org/10.5281/zenodo.4639017" httpUri="https://zenodo.org/record/4639017/files/figure.png" pageId="9" pageNumber="202">Fig. 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="202" type="materials_examined">
<paragraph blockId="9.[129,772,1882,1961]" pageId="9" pageNumber="202">
<materialsCitation ID-GBIF-Occurrence="3326898303" collectionCode="MZUSP" collectorName="Adao A. Barbosa." country="Brazil" county="rio Negro" latitude="0.2677778" location="Amazonas" longLatPrecision="21" longitude="-69.9725" municipality="rio Tiquie" pageId="9" pageNumber="202" specimenCode="MZUSP 81505" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[129,229,1882,1903]" pageId="9" pageNumber="202">
<typeStatus box="[129,226,1882,1903]" pageId="9" pageNumber="202">Holotype</typeStatus>
.
</emphasis>
<specimenCode box="[237,387,1882,1904]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="202">MZUSP 81505</specimenCode>
, male,
<quantity box="[460,541,1882,1903]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="202" unit="mm" value="100.0">100 mm</quantity>
SL,
<collectingCountry box="[590,652,1882,1904]" name="Brazil" pageId="9" pageNumber="202">Brazil</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:03C4D93834733F48FED8F893FBBF618A:8EB23EF534733F4BFD6DF933FD34614C" box="[662,767,1882,1903]" country="Brazil" county="rio Negro" latitude="0.2677778" longLatPrecision="21" longitude="-69.9725" municipality="rio Tiquie" name="Amazonas" pageId="9" pageNumber="202">Amazonas</location>
, community of
<location LSID="urn:lsid:plazi:treatment:03C4D93834733F48FED8F893FBBF618A:8EB23EF534733F4BFEF0F91EFEA661AE" box="[267,365,1911,1933]" country="Brazil" county="rio Negro" latitude="0.2677778" longLatPrecision="21" longitude="-69.9725" municipality="rio Tiquie" name="Sao Pedro" pageId="9" pageNumber="202">São Pedro</location>
,
<collectingMunicipality box="[374,470,1911,1933]" pageId="9" pageNumber="202">rio Tiquié</collectingMunicipality>
,
<collectingCounty box="[478,569,1911,1933]" pageId="9" pageNumber="202">rio Negro</collectingCounty>
drainage,
<geoCoordinate box="[667,771,1910,1932]" degrees="0" direction="north" minutes="16" orientation="latitude" pageId="9" pageNumber="202" precision="15" seconds="04" value="0.2677778">0°1604&quot;N</geoCoordinate>
<geoCoordinate box="[129,252,1939,1961]" degrees="69" direction="west" minutes="58" orientation="longitude" pageId="9" pageNumber="202" precision="15" seconds="21" value="-69.9725">69°5821&quot;W</geoCoordinate>
, 2004,
<collectorName box="[322,495,1939,1961]" pageId="9" pageNumber="202">Adão A. Barbosa.</collectorName>
</materialsCitation>
</paragraph>
<paragraph blockId="9.[809,1452,829,1053]" pageId="9" pageNumber="202">
<materialsCitation ID-GBIF-Occurrence="3326898359" collectionCode="MZUSP" collectorName="Sao Pedro" country="Brazil" latitude="0.2678889" location="Amazonas" longLatPrecision="1" longitude="-69.97264" pageId="9" pageNumber="202" specimenCode="MZUSP 84988" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<emphasis bold="true" box="[809,1124,829,851]" pageId="9" pageNumber="202">
<typeStatus box="[809,916,829,850]" pageId="9" pageNumber="202">Paratypes</typeStatus>
.
<collectingCountry box="[930,995,829,851]" name="Brazil" pageId="9" pageNumber="202">Brazil</collectingCountry>
,
<collectingRegion box="[1008,1120,829,850]" pageId="9" pageNumber="202">Amazonas</collectingRegion>
:
</emphasis>
<specimenCode box="[1134,1288,829,850]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="202">MZUSP 84988</specimenCode>
, mature female,
<quantity box="[809,894,858,879]" metricMagnitude="-1" metricUnit="m" metricValue="1.02" pageId="9" pageNumber="202" unit="mm" value="102.0">102 mm</quantity>
SL, near old community of
<collectorName box="[1200,1309,858,880]" pageId="9" pageNumber="202">São Pedro</collectorName>
,
<geoCoordinate box="[1323,1451,858,880]" degrees="0" direction="north" minutes="16" orientation="latitude" pageId="9" pageNumber="202" precision="1" seconds="04.4" value="0.2678889">0°1604.4&quot;N</geoCoordinate>
<geoCoordinate box="[809,952,886,908]" degrees="69" direction="west" minutes="58" orientation="longitude" pageId="9" pageNumber="202" precision="1" seconds="21.5" value="-69.97264">69°5821.5&quot;W</geoCoordinate>
</materialsCitation>
, 2004, F. Lima;
<materialsCitation ID-GBIF-Occurrence="3326898399" collectingDate="2000-10-25" collectingDateMax="2000-10-27" collectingDateMin="2000-10-25" collectionCode="MZUSP" collectorName="Sao Pedro" country="Brazil" latitude="0.2613889" location="rio Tiquie" longLatPrecision="21" longitude="-69.95638" municipality="Umari" pageId="9" pageNumber="202" specimenCode="MZUSP 64359" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<specimenCode box="[1107,1255,886,908]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="202">MZUSP 64359</specimenCode>
, mature female,
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.21" pageId="9" pageNumber="202" unit="mm" value="121.0">121 mm</quantity>
SL, igarapé
<collectingMunicipality box="[966,1030,915,937]" pageId="9" pageNumber="202">Umari</collectingMunicipality>
, tributary of
<location LSID="urn:lsid:plazi:treatment:03C4D93834733F48FED8F893FBBF618A:8EB23EF534733F4BFB7BFDFAFB14658A" box="[1152,1247,915,937]" country="Brazil" latitude="0.2613889" longLatPrecision="21" longitude="-69.95638" municipality="Umari" name="rio Tiquie" pageId="9" pageNumber="202" stateProvince="Amazonas">rio Tiquié</location>
in community of
<collectorName pageId="9" pageNumber="202">São Pedro</collectorName>
,
<geoCoordinate box="[877,984,944,966]" degrees="0" direction="north" minutes="15" orientation="latitude" pageId="9" pageNumber="202" precision="15" seconds="41" value="0.2613889">0°1541&quot;N</geoCoordinate>
<geoCoordinate box="[988,1112,944,966]" degrees="69" direction="west" minutes="57" orientation="longitude" pageId="9" pageNumber="202" precision="15" seconds="23" value="-69.95638">69°5723&quot;W</geoCoordinate>
,
<date box="[1121,1317,944,966]" pageId="9" pageNumber="202" value="2000-10-25" valueMax="2000-10-27" valueMin="2000-10-25">
<collectingDate box="[1121,1317,944,966]" pageId="9" pageNumber="202" value="2000-10-25" valueMax="2000-10-27" valueMin="2000-10-25">25-27 October 2000</collectingDate>
</date>
</materialsCitation>
, F. C.
<materialsCitation ID-GBIF-Occurrence="3326898398" collectionCode="T" country="Brazil" location="Lima" pageId="9" pageNumber="202" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<collectionCode box="[1376,1392,945,966]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="202">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:03C4D93834733F48FED8F893FBBF618A:8EB23EF534733F4BFA83FDD9FA6065E5" box="[1400,1451,944,966]" country="Brazil" name="Lima" pageId="9" pageNumber="202" stateProvince="Amazonas">Lima</location>
and party
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3326898346" collectionCode="MZUSP" collectorName="Sao Pedro" country="Brazil" latitude="0.26666668" location="rio Tiquie" longLatPrecision="21" longitude="-69.96667" municipality="Umari" pageId="9" pageNumber="202" specimenCode="MZUSP 81325" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<specimenCode box="[923,1080,973,995]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="9" pageNumber="202">MZUSP 81325</specimenCode>
, male,
<quantity box="[1158,1243,973,994]" metricMagnitude="-1" metricUnit="m" metricValue="1.02" pageId="9" pageNumber="202" unit="mm" value="102.0">102 mm</quantity>
SL, igarapé
<collectingMunicipality box="[1382,1448,973,995]" pageId="9" pageNumber="202">Umari</collectingMunicipality>
, tributary of
<location LSID="urn:lsid:plazi:treatment:03C4D93834733F48FED8F893FBBF618A:8EB23EF534733F4BFC5DFD83FBC06223" box="[934,1035,1002,1024]" country="Brazil" latitude="0.26666668" longLatPrecision="21" longitude="-69.96667" municipality="Umari" name="rio Tiquie" pageId="9" pageNumber="202" stateProvince="Amazonas">rio Tiquié</location>
in community of
<collectorName box="[1225,1331,1002,1024]" pageId="9" pageNumber="202">São Pedro</collectorName>
,
<geoCoordinate box="[1344,1451,1002,1024]" degrees="0" direction="north" minutes="16" orientation="latitude" pageId="9" pageNumber="202" precision="15" seconds="00" value="0.26666668">0°1600&quot;N</geoCoordinate>
<geoCoordinate box="[809,933,1030,1052]" degrees="69" direction="west" minutes="58" orientation="longitude" pageId="9" pageNumber="202" precision="15" seconds="00" value="-69.96667">69°5800&quot;W</geoCoordinate>
</materialsCitation>
, 2002, F. C.
<materialsCitation ID-GBIF-Occurrence="3326898404" box="[1058,1236,1031,1053]" collectionCode="T" country="Brazil" location="Lima" pageId="9" pageNumber="202" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<collectionCode box="[1058,1074,1031,1052]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="202">T</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:03C4D93834733F48FED8F893FBBF618A:8EB23EF534733F4BFBC7FA6EFBA4623E" box="[1084,1135,1031,1053]" country="Brazil" name="Lima" pageId="9" pageNumber="202" stateProvince="Amazonas">Lima</location>
and party
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="202" type="diagnosis">
<paragraph blockId="9.[809,1452,1104,1641]" pageId="9" pageNumber="202">
<emphasis bold="true" box="[809,927,1104,1128]" pageId="9" pageNumber="202">Diagnosis.</emphasis>
<taxonomicName authorityName="Menezes &amp; de Lucena" authorityYear="2014" box="[934,1103,1105,1129]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="delimai">
<emphasis box="[934,1103,1105,1129]" italics="true" pageId="9" pageNumber="202">Charax delimai</emphasis>
</taxonomicName>
along with
<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1870" box="[1233,1342,1105,1129]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="tectifer">
<emphasis box="[1233,1342,1105,1129]" italics="true" pageId="9" pageNumber="202">C. tectifer</emphasis>
</taxonomicName>
,
<taxonomicName box="[1355,1451,1105,1128]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="metae">
<emphasis box="[1355,1451,1105,1128]" italics="true" pageId="9" pageNumber="202">C. metae</emphasis>
</taxonomicName>
and some specimens of
<taxonomicName box="[1083,1215,1137,1161]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="gibbosus">
<emphasis box="[1083,1215,1137,1161]" italics="true" pageId="9" pageNumber="202">C. gibbosus</emphasis>
</taxonomicName>
,
<taxonomicName box="[1232,1367,1137,1160]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="rupununi">
<emphasis box="[1232,1367,1137,1160]" italics="true" pageId="9" pageNumber="202">C. rupununi</emphasis>
</taxonomicName>
and
<taxonomicName class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="condei">
<emphasis box="[1427,1445,1137,1160]" italics="true" pageId="9" pageNumber="202">C</emphasis>
.
<emphasis box="[809,881,1169,1193]" italics="true" pageId="9" pageNumber="202">condei</emphasis>
</taxonomicName>
are the only members of the genus bearing teeth on the ectopterygoid.
<taxonomicName authorityName="Menezes &amp; de Lucena" authorityYear="2014" box="[1011,1179,1201,1225]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="delimai">
<emphasis box="[1011,1179,1201,1225]" italics="true" pageId="9" pageNumber="202">Charax delimai</emphasis>
</taxonomicName>
differs from
<taxonomicName box="[1321,1424,1201,1225]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="condei">
<emphasis box="[1321,1339,1201,1224]" italics="true" pageId="9" pageNumber="202">C</emphasis>
.
<emphasis box="[1352,1424,1201,1225]" italics="true" pageId="9" pageNumber="202">condei</emphasis>
</taxonomicName>
in having the lateral line complete (
<emphasis box="[1144,1172,1233,1256]" italics="true" pageId="9" pageNumber="202">vs.</emphasis>
incomplete). From
<taxonomicName class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="gibbosus">
<emphasis italics="true" pageId="9" pageNumber="202">Charax gibbosus</emphasis>
</taxonomicName>
and
<taxonomicName box="[965,1099,1265,1288]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="rupununi">
<emphasis box="[965,1099,1265,1288]" italics="true" pageId="9" pageNumber="202">C. rupununi</emphasis>
</taxonomicName>
which have the anal-fin origin anterior to the vertical through the dorsal-fin origin in having the anal-fin origin on vertical or slightly posterior to, dorsalfin origin.
<taxonomicName authorityName="Menezes &amp; de Lucena" authorityYear="2014" box="[922,1089,1361,1385]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="delimai">
<emphasis box="[922,1089,1361,1385]" italics="true" pageId="9" pageNumber="202">Charax delimai</emphasis>
</taxonomicName>
can be readily distinguished from
<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1870" box="[809,927,1393,1417]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="tectifer">
<emphasis box="[809,827,1393,1416]" italics="true" pageId="9" pageNumber="202">C</emphasis>
.
<emphasis box="[846,927,1393,1417]" italics="true" pageId="9" pageNumber="202">tectifer</emphasis>
</taxonomicName>
by the presence (
<emphasis box="[1144,1174,1393,1416]" italics="true" pageId="9" pageNumber="202">vs.</emphasis>
absence) of superficial neuromasts on body, with neuromasts dorso-ventrally arranged on trunk scales except those on the lateral line where neuromasts are absent (
<figureCitation box="[1061,1123,1489,1513]" captionStart="Fig" captionStartId="3.[129,168,636,660]" captionTargetBox="[129,1452,174,615]" captionTargetId="figure-6@3.[129,1452,174,615]" captionTargetPageId="3" captionText="Fig. 1. Superficial neuromasts (black arrows) on lateral body scales of Charax delimai, male (a), MZUSP 81325, paratype, 102 mm SL, Brazil, Amazonas, igarapé Umari, tributary of rio Tiquié in community of São Pedro and female, (b), MZUSP 84988, paratype, 102 mm SL, Brazil, Amazonas, near old community of São Pedro." figureDoi="http://doi.org/10.5281/zenodo.4638997" httpUri="https://zenodo.org/record/4638997/files/figure.png" pageId="9" pageNumber="202">Fig. 1</figureCitation>
) and from
<taxonomicName box="[1243,1338,1489,1512]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="202" phylum="Chordata" rank="species" species="metae">
<emphasis box="[1243,1338,1489,1512]" italics="true" pageId="9" pageNumber="202">C. metae</emphasis>
</taxonomicName>
by having fewer ectopterygoid teeth (3-15
<emphasis box="[1158,1187,1521,1544]" italics="true" pageId="9" pageNumber="202">vs.</emphasis>
20-50), more transverse scale rows between the humeral spot and the supracleithrum (10-12
<emphasis box="[883,911,1585,1608]" italics="true" pageId="9" pageNumber="202">vs.</emphasis>
7-9) and the humeral spot distance 48-51% of SL (
<emphasis box="[1423,1452,1585,1608]" italics="true" pageId="9" pageNumber="202">vs.</emphasis>
41.6-47% of SL,
<figureCitation box="[980,1039,1617,1641]" captionStart="Fig" captionStartId="3.[807,846,1194,1218]" captionTargetBox="[129,1443,738,1174]" captionTargetPageId="3" captionText="Fig. 3. Humeral spot distance as a function of standard length for species of Charax." figureDoi="http://doi.org/10.5281/zenodo.4639001" httpUri="https://zenodo.org/record/4639001/files/figure.png" pageId="9" pageNumber="202">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="203" pageId="9" pageNumber="202" type="description">
<paragraph blockId="9.[809,1452,1680,1961]" lastBlockId="10.[136,779,177,1225]" lastPageId="10" lastPageNumber="203" pageId="9" pageNumber="202">
<emphasis bold="true" box="[809,949,1680,1704]" pageId="9" pageNumber="202">Description.</emphasis>
Morphometrics of
<typeStatus box="[1162,1256,1681,1705]" pageId="9" pageNumber="202">holotype</typeStatus>
and all examined specimens presented in
<tableCitation box="[1066,1148,1713,1737]" captionStart="Table 4" captionStartId="10.[136,199,1281,1305]" captionTargetPageId="10" captionText="Table 4. Morphometrics of Charax delimai. Specimens are from MZUSP 81505 (holotype) and 84988 (paratypes)." pageId="9" pageNumber="202">Table 4</tableCitation>
. Body elongate, moderately large (
<quantity box="[879,1020,1745,1769]" metricMagnitude="-1" metricUnit="m" metricValue="1.105" metricValueMax="1.21" metricValueMin="1.0" pageId="9" pageNumber="202" unit="mm" value="110.5" valueMax="121.0" valueMin="100.0">100-121 mm</quantity>
SL), compressed and moderately deep. Greatest body depth slightly in advance of dorsal-fin origin. Dorsal profile of head and body convex from tip of snout to anterior part of fontanel, concave from that point to base of supraoccipital spine, convex from that point to dorsal-fin origin, nearly straight along dorsal-fin base and from end of dorsal-fin base to caudal peduncle and slightly concave along caudal peduncle. Ventral profile of head and body convex from tip of lower jaw to anal-fin origin, nearly straight along anal-fin base and slightly concave from end of anal-fin base to beginning of procurrent rays. Snout pointed. Lower jaw included in upper jaw when mouth closed. Maxilla extending slightly beyond vertical through middle of orbit.
</paragraph>
<paragraph blockId="10.[136,779,177,1225]" pageId="10" pageNumber="203">
Dorsal-fin rays ii,
<quantity box="[375,417,369,393]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="10" pageNumber="203" unit="in" value="9.0">9 in</quantity>
all specimens posterior most ray unbranched. Adipose fin present. Unbranched anal-fin rays iv or v, usually iv; branched rays 41-45 (iv, 45), 41.9.Anterior anal-fin rays not forming differentiated distinct lobe and not bearing bilateral hooks in two adult males (100 and
<quantity box="[691,779,497,521]" metricMagnitude="-1" metricUnit="m" metricValue="1.02" pageId="10" pageNumber="203" unit="mm" value="102.0">102 mm</quantity>
SL). Pectoral-fin rays i, 13-15 (i, 14), 14.5. Tips of longest pectoral-fin rays reaching about to vertical through middle of pelvic fin. Pelvic-fin rays i, 7. No hooks on pelvic-fin rays of adult males. Tips of longest pelvic-fin rays reaching slightly beyond anal-fin origin. Principal caudal-fin ray count 10/
<quantity box="[740,779,657,681]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="10" pageNumber="203" unit="in" value="9.0">9 in</quantity>
all specimens.
</paragraph>
<paragraph blockId="10.[136,779,177,1225]" pageId="10" pageNumber="203">
Lateral line complete, perforated scales 50-53 (52), 51. Horizontal scale rows from dorsal-fin origin to lateral line 11- 12 (12), 11.7. Horizontal scale rows from pelvic-fin origin to lateral line 9-10 (9), 9.2. Horizontal scale rows from anal-fin origin to lateral line 11-12 (12), 11.7. Predorsal scales 33-35 (34). Scale rows around caudal peduncle 19-20 (20), 19.5. One scale row along anal-fin base, except on last one-fifth of its length. Rostral portions of body scales of both males and females, except for lateral-line scales with vertically oriented superficial neuromasts (
<figureCitation box="[389,450,1009,1033]" captionStart="Fig" captionStartId="3.[129,168,636,660]" captionTargetBox="[129,1452,174,615]" captionTargetId="figure-6@3.[129,1452,174,615]" captionTargetPageId="3" captionText="Fig. 1. Superficial neuromasts (black arrows) on lateral body scales of Charax delimai, male (a), MZUSP 81325, paratype, 102 mm SL, Brazil, Amazonas, igarapé Umari, tributary of rio Tiquié in community of São Pedro and female, (b), MZUSP 84988, paratype, 102 mm SL, Brazil, Amazonas, near old community of São Pedro." figureDoi="http://doi.org/10.5281/zenodo.4638997" httpUri="https://zenodo.org/record/4638997/files/figure.png" pageId="10" pageNumber="203">Fig. 1</figureCitation>
).
</paragraph>
<paragraph blockId="10.[136,779,177,1225]" lastBlockId="10.[816,1456,177,361]" pageId="10" pageNumber="203">Premaxillary with one anterior canine-like tooth followed by set of smaller conical teeth and another canine-like tooth followed by one or two small conical teeth. Total number of premaxillary teeth 13-17 (14) 14.7. Maxillary teeth conical, 55-56 (55), 55.2. Dentary with one canine-like tooth followed by 3-5 (4) 4 conical teeth, another canine-like tooth and posterior row of 21-24 (24) 23 conical teeth. Left ectopterygoid without teeth or with 3 or 9 teeths; right ectopterygoid with 6-9 conical teeth. Total number of ectopterygoid teeth 3-15 (9), 9. Vertebrae 34- 35 (34), 34.2. Eight gill-rakers on lower limb of first gill-arch in all specimens. Branchiostegal rays 4; 3 rays originating from anterior ceratohyal and 1 from posterior ceratohyal</paragraph>
<paragraph blockId="10.[816,1456,401,1513]" pageId="10" pageNumber="203">
<emphasis bold="true" box="[816,1006,401,425]" pageId="10" pageNumber="203">Color in alcohol.</emphasis>
Body pale to light yellow, slightly darker dorsally than ventrally due to concentration of dark chromatophores on predorsal and postdorsal scales. Dark chromatophores mostly concentrated on basal portions of scales leaving light area on remaining portion of each scale and forming alternate pattern of dark and light spots along longitudinal scale rows on body. Whitish color of vertically arranged superficial neuromasts on skin of basal portion of each scale and contrasting strongly with background dark coloration of basal portion of scale (
<figureCitation box="[1198,1259,689,713]" captionStart="Fig" captionStartId="3.[129,168,636,660]" captionTargetBox="[129,1452,174,615]" captionTargetId="figure-6@3.[129,1452,174,615]" captionTargetPageId="3" captionText="Fig. 1. Superficial neuromasts (black arrows) on lateral body scales of Charax delimai, male (a), MZUSP 81325, paratype, 102 mm SL, Brazil, Amazonas, igarapé Umari, tributary of rio Tiquié in community of São Pedro and female, (b), MZUSP 84988, paratype, 102 mm SL, Brazil, Amazonas, near old community of São Pedro." figureDoi="http://doi.org/10.5281/zenodo.4638997" httpUri="https://zenodo.org/record/4638997/files/figure.png" pageId="10" pageNumber="203">Fig. 1</figureCitation>
). Scales on lateral and ventral body portions with fewer chromatophores. Irregularly shaped dark humeral blotch, much closer to dorsalfin origin than to posterior border of opercle, encompassing about 5 scales vertically and horizontally. Dark blotch on caudal peduncle approximately triangular, higher posteriorly and extending over basal portions of central caudal-fin rays. Dorsal portions of head from tip of snout to supraoccipital region darker than remainder of head, dark color extending vertically over first, second, and anterior portion of third infraorbitals below orbit, dorsal portion of prepercle and inner border of opercle. Most of third, fourth and fifth infraorbitals, ventral portion of preopercle, posterior border of opercle and subopercle largely unpigmented with scattered dark chromatophores. Tip of lower jaw dark; lighter posteriorly with scattered chromatophores. Tip of dorsal fin dark. Anal fin with faint basal dark stripe extending from about basal portion of first branched anal-fin ray to end of fin, this stripe separated by light anteriorly wide stripe involving basal and median portions of unbranched rays, narrower from this point backward. Conspicuous dark wider marginal dark stripe extending to end of fins, initially separated by small light stripe extending only to about tenth branched anal-fin ray. Tip of pelvic fin dark, pectorals light with few scattered dark chromatophores. Marginal portion of caudal fin dark, all fin with scattered dark chromatophores.
</paragraph>
<caption pageId="10" pageNumber="203" startId="10.[136,199,1281,1305]" targetBox="[135,776,1371,1956]" targetIsTable="true" targetPageId="10">
<paragraph blockId="10.[136,778,1280,1337]" pageId="10" pageNumber="203">
<emphasis bold="true" box="[136,226,1280,1305]" pageId="10" pageNumber="203">Table 4.</emphasis>
Morphometrics of
<taxonomicName authorityName="Menezes &amp; de Lucena" authorityYear="2014" box="[438,607,1281,1305]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="10" pageNumber="203" phylum="Chordata" rank="species" species="delimai">
<emphasis box="[438,607,1281,1305]" italics="true" pageId="10" pageNumber="203">Charax delimai</emphasis>
</taxonomicName>
. Specimens are from MZUSP 81505 (holotype) and 84988 (paratypes).
</paragraph>
</caption>
<paragraph pageId="10" pageNumber="203">
<table box="[135,776,1371,1956]" gridcols="6" gridrows="24" pageId="10" pageNumber="203" rowsContinueFrom="8.[833,1447,1370,1955]" rowsContinueIn="12.[832,1447,1374,1960]">
<tr box="[135,776,1371,1390]" gridrow="0" pageId="10" pageNumber="203">
<th box="[135,436,1371,1390]" gridcol="0" gridrow="0" pageId="10" pageNumber="203">Characters</th>
<th box="[473,569,1371,1390]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="10" pageNumber="203">Holotype n</th>
<th box="[586,689,1371,1390]" gridcol="3" gridrow="0" pageId="10" pageNumber="203">range</th>
<th box="[698,776,1371,1390]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="10" pageNumber="203">mean SD</th>
</tr>
<tr box="[135,776,1396,1416]" gridrow="1" pageId="10" pageNumber="203">
<th box="[135,436,1396,1416]" gridcol="0" gridrow="1" pageId="10" pageNumber="203">Standard length</th>
<td box="[473,537,1396,1416]" gridcol="1" gridrow="1" pageId="10" pageNumber="203">100.0</td>
<td box="[557,689,1396,1416]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="10" pageNumber="203">4 100.0 - 121.0</td>
<td box="[698,776,1396,1416]" colspan="2" colspanRight="1" gridcol="4" gridrow="1" pageId="10" pageNumber="203">106.2</td>
</tr>
<tr box="[135,776,1421,1440]" gridrow="2" pageId="10" pageNumber="203">
<th box="[135,776,1421,1440]" colspan="6" colspanRight="5" gridcol="0" gridrow="2" pageId="10" pageNumber="203">Percents of standard length</th>
</tr>
<tr box="[135,776,1445,1465]" gridrow="3" pageId="10" pageNumber="203">
<th box="[135,436,1445,1465]" gridcol="0" gridrow="3" pageId="10" pageNumber="203">Depth at dorsal-fin origin</th>
<td box="[473,537,1445,1465]" gridcol="1" gridrow="3" pageId="10" pageNumber="203">39.5</td>
<td box="[557,569,1445,1465]" gridcol="2" gridrow="3" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1445,1465]" gridcol="3" gridrow="3" pageId="10" pageNumber="203">39.5 - 41.1</td>
<td box="[698,739,1445,1465]" gridcol="4" gridrow="3" pageId="10" pageNumber="203">40.2</td>
<td box="[748,776,1445,1465]" gridcol="5" gridrow="3" pageId="10" pageNumber="203">0.7</td>
</tr>
<tr box="[135,776,1470,1490]" gridrow="4" pageId="10" pageNumber="203">
<th box="[135,436,1470,1490]" gridcol="0" gridrow="4" pageId="10" pageNumber="203">Snout to dorsal-fin origin</th>
<td box="[473,537,1470,1490]" gridcol="1" gridrow="4" pageId="10" pageNumber="203">55.0</td>
<td box="[557,569,1470,1490]" gridcol="2" gridrow="4" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1470,1490]" gridcol="3" gridrow="4" pageId="10" pageNumber="203">52.5 - 55.0</td>
<td box="[698,739,1470,1490]" gridcol="4" gridrow="4" pageId="10" pageNumber="203">54.3</td>
<td box="[748,776,1470,1490]" gridcol="5" gridrow="4" pageId="10" pageNumber="203">1.2</td>
</tr>
<tr box="[135,776,1494,1514]" gridrow="5" pageId="10" pageNumber="203">
<th box="[135,436,1494,1514]" gridcol="0" gridrow="5" pageId="10" pageNumber="203">Snout to pectoral-fin origin</th>
<td box="[473,537,1494,1514]" gridcol="1" gridrow="5" pageId="10" pageNumber="203">28.5</td>
<td box="[557,569,1494,1514]" gridcol="2" gridrow="5" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1494,1514]" gridcol="3" gridrow="5" pageId="10" pageNumber="203">28.5 - 30.5</td>
<td box="[698,739,1494,1514]" gridcol="4" gridrow="5" pageId="10" pageNumber="203">29.7</td>
<td box="[748,776,1494,1514]" gridcol="5" gridrow="5" pageId="10" pageNumber="203">1.0</td>
</tr>
<tr box="[135,776,1519,1538]" gridrow="6" pageId="10" pageNumber="203">
<th box="[135,436,1519,1538]" gridcol="0" gridrow="6" pageId="10" pageNumber="203">Snout to pelvic-fin origin</th>
<td box="[473,537,1519,1538]" gridcol="1" gridrow="6" pageId="10" pageNumber="203">40.0</td>
<td box="[557,569,1519,1538]" gridcol="2" gridrow="6" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1519,1538]" gridcol="3" gridrow="6" pageId="10" pageNumber="203">40.0 - 42.1</td>
<td box="[698,739,1519,1538]" gridcol="4" gridrow="6" pageId="10" pageNumber="203">41.0</td>
<td box="[748,776,1519,1538]" gridcol="5" gridrow="6" pageId="10" pageNumber="203">0.9</td>
</tr>
<tr box="[135,776,1544,1563]" gridrow="7" pageId="10" pageNumber="203">
<th box="[135,436,1544,1563]" gridcol="0" gridrow="7" pageId="10" pageNumber="203">Snout to anal-fin origin</th>
<td box="[473,537,1544,1563]" gridcol="1" gridrow="7" pageId="10" pageNumber="203">56.5</td>
<td box="[557,569,1544,1563]" gridcol="2" gridrow="7" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1544,1563]" gridcol="3" gridrow="7" pageId="10" pageNumber="203">55.8 - 57.0</td>
<td box="[698,739,1544,1563]" gridcol="4" gridrow="7" pageId="10" pageNumber="203">56.5</td>
<td box="[748,776,1544,1563]" gridcol="5" gridrow="7" pageId="10" pageNumber="203">0.5</td>
</tr>
<tr box="[135,776,1568,1587]" gridrow="8" pageId="10" pageNumber="203">
<th box="[135,436,1568,1587]" gridcol="0" gridrow="8" pageId="10" pageNumber="203">Caudal peduncle depth</th>
<td box="[473,537,1568,1587]" gridcol="1" gridrow="8" pageId="10" pageNumber="203">9.0</td>
<td box="[557,569,1568,1587]" gridcol="2" gridrow="8" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1568,1587]" gridcol="3" gridrow="8" pageId="10" pageNumber="203">8.8 - 10.0</td>
<td box="[698,739,1568,1587]" gridcol="4" gridrow="8" pageId="10" pageNumber="203">8.8</td>
<td box="[748,776,1568,1587]" gridcol="5" gridrow="8" pageId="10" pageNumber="203">0.4</td>
</tr>
<tr box="[135,776,1593,1612]" gridrow="9" pageId="10" pageNumber="203">
<th box="[135,436,1593,1612]" gridcol="0" gridrow="9" pageId="10" pageNumber="203">Caudal peduncle length</th>
<td box="[473,537,1593,1612]" gridcol="1" gridrow="9" pageId="10" pageNumber="203">7.5</td>
<td box="[557,569,1593,1612]" gridcol="2" gridrow="9" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1593,1612]" gridcol="3" gridrow="9" pageId="10" pageNumber="203">6.6 - 7.5</td>
<td box="[698,739,1593,1612]" gridcol="4" gridrow="9" pageId="10" pageNumber="203">7.8</td>
<td box="[748,776,1593,1612]" gridcol="5" gridrow="9" pageId="10" pageNumber="203">0.4</td>
</tr>
<tr box="[135,776,1617,1637]" gridrow="10" pageId="10" pageNumber="203">
<th box="[135,436,1617,1637]" gridcol="0" gridrow="10" pageId="10" pageNumber="203">Pectoral-fin length</th>
<td box="[473,537,1617,1637]" gridcol="1" gridrow="10" pageId="10" pageNumber="203">19.0</td>
<td box="[557,569,1617,1637]" gridcol="2" gridrow="10" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1617,1637]" gridcol="3" gridrow="10" pageId="10" pageNumber="203">18.0 - 22.0</td>
<td box="[698,739,1617,1637]" gridcol="4" gridrow="10" pageId="10" pageNumber="203">20.0</td>
<td box="[748,776,1617,1637]" gridcol="5" gridrow="10" pageId="10" pageNumber="203">1.8</td>
</tr>
<tr box="[135,776,1642,1661]" gridrow="11" pageId="10" pageNumber="203">
<th box="[135,436,1642,1661]" gridcol="0" gridrow="11" pageId="10" pageNumber="203">Pelvic-fin length</th>
<td box="[473,537,1642,1661]" gridcol="1" gridrow="11" pageId="10" pageNumber="203">18.0</td>
<td box="[557,569,1642,1661]" gridcol="2" gridrow="11" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1642,1661]" gridcol="3" gridrow="11" pageId="10" pageNumber="203">27.2 - 29.0</td>
<td box="[698,739,1642,1661]" gridcol="4" gridrow="11" pageId="10" pageNumber="203">28.1</td>
<td box="[748,776,1642,1661]" gridcol="5" gridrow="11" pageId="10" pageNumber="203">0.9</td>
</tr>
<tr box="[135,776,1666,1686]" gridrow="12" pageId="10" pageNumber="203">
<th box="[135,436,1666,1686]" gridcol="0" gridrow="12" pageId="10" pageNumber="203">Dorsal-fin base length</th>
<td box="[473,537,1666,1686]" gridcol="1" gridrow="12" pageId="10" pageNumber="203">11.5</td>
<td box="[557,569,1666,1686]" gridcol="2" gridrow="12" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1666,1686]" gridcol="3" gridrow="12" pageId="10" pageNumber="203">11.2 - 12.4</td>
<td box="[698,739,1666,1686]" gridcol="4" gridrow="12" pageId="10" pageNumber="203">11.7</td>
<td box="[748,776,1666,1686]" gridcol="5" gridrow="12" pageId="10" pageNumber="203">0.5</td>
</tr>
<tr box="[135,776,1691,1710]" gridrow="13" pageId="10" pageNumber="203">
<th box="[135,436,1691,1710]" gridcol="0" gridrow="13" pageId="10" pageNumber="203">Dorsal-fin height</th>
<td box="[473,537,1691,1710]" gridcol="1" gridrow="13" pageId="10" pageNumber="203">29.0</td>
<td box="[557,569,1691,1710]" gridcol="2" gridrow="13" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1691,1710]" gridcol="3" gridrow="13" pageId="10" pageNumber="203">27.2 - 29.0</td>
<td box="[698,739,1691,1710]" gridcol="4" gridrow="13" pageId="10" pageNumber="203">28.1</td>
<td box="[748,776,1691,1710]" gridcol="5" gridrow="13" pageId="10" pageNumber="203">0.9</td>
</tr>
<tr box="[135,776,1715,1735]" gridrow="14" pageId="10" pageNumber="203">
<th box="[135,436,1715,1735]" gridcol="0" gridrow="14" pageId="10" pageNumber="203">Anal-fin base length</th>
<td box="[473,537,1715,1735]" gridcol="1" gridrow="14" pageId="10" pageNumber="203">47.0</td>
<td box="[557,569,1715,1735]" gridcol="2" gridrow="14" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1715,1735]" gridcol="3" gridrow="14" pageId="10" pageNumber="203">44.6 - 47.0</td>
<td box="[698,739,1715,1735]" gridcol="4" gridrow="14" pageId="10" pageNumber="203">46.0</td>
<td box="[748,776,1715,1735]" gridcol="5" gridrow="14" pageId="10" pageNumber="203">1.2</td>
</tr>
<tr box="[135,776,1740,1759]" gridrow="15" pageId="10" pageNumber="203">
<th box="[135,436,1740,1759]" gridcol="0" gridrow="15" pageId="10" pageNumber="203">Eye to dorsal-fin origin</th>
<td box="[473,537,1740,1759]" gridcol="1" gridrow="15" pageId="10" pageNumber="203">41.0</td>
<td box="[557,569,1740,1759]" gridcol="2" gridrow="15" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1740,1759]" gridcol="3" gridrow="15" pageId="10" pageNumber="203">41.0 - 42.1</td>
<td box="[698,739,1740,1759]" gridcol="4" gridrow="15" pageId="10" pageNumber="203">41.4</td>
<td box="[748,776,1740,1759]" gridcol="5" gridrow="15" pageId="10" pageNumber="203">0.5</td>
</tr>
<tr box="[135,776,1764,1784]" gridrow="16" pageId="10" pageNumber="203">
<th box="[135,436,1764,1784]" gridcol="0" gridrow="16" pageId="10" pageNumber="203">Dorsal-fin origin to caudal-fin base</th>
<td box="[473,537,1764,1784]" gridcol="1" gridrow="16" pageId="10" pageNumber="203">52.0</td>
<td box="[557,569,1764,1784]" gridcol="2" gridrow="16" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1764,1784]" gridcol="3" gridrow="16" pageId="10" pageNumber="203">51.2 - 53.0</td>
<td box="[698,739,1764,1784]" gridcol="4" gridrow="16" pageId="10" pageNumber="203">52.0</td>
<td box="[748,776,1764,1784]" gridcol="5" gridrow="16" pageId="10" pageNumber="203">0.7</td>
</tr>
<tr box="[135,776,1789,1809]" gridrow="17" pageId="10" pageNumber="203">
<th box="[135,436,1789,1809]" gridcol="0" gridrow="17" pageId="10" pageNumber="203">Humeral spot distance</th>
<td box="[473,537,1789,1809]" gridcol="1" gridrow="17" pageId="10" pageNumber="203">48.5</td>
<td box="[557,569,1789,1809]" gridcol="2" gridrow="17" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1789,1809]" gridcol="3" gridrow="17" pageId="10" pageNumber="203">48.0 - 51.0</td>
<td box="[698,776,1789,1809]" colspan="2" colspanRight="1" gridcol="4" gridrow="17" pageId="10" pageNumber="203">49.5 1.4</td>
</tr>
<tr box="[135,776,1813,1833]" gridrow="18" pageId="10" pageNumber="203">
<th box="[135,436,1813,1833]" gridcol="0" gridrow="18" pageId="10" pageNumber="203">Bony head length</th>
<td box="[473,537,1813,1833]" gridcol="1" gridrow="18" pageId="10" pageNumber="203">29.5</td>
<td box="[557,569,1813,1833]" gridcol="2" gridrow="18" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1813,1833]" gridcol="3" gridrow="18" pageId="10" pageNumber="203">29.4 - 30.1</td>
<td box="[698,739,1813,1833]" gridcol="4" gridrow="18" pageId="10" pageNumber="203">29.6</td>
<td box="[748,776,1813,1833]" gridcol="5" gridrow="18" pageId="10" pageNumber="203">0.4</td>
</tr>
<tr box="[135,776,1838,1857]" gridrow="19" pageId="10" pageNumber="203">
<th box="[135,776,1838,1857]" colspan="6" colspanRight="5" gridcol="0" gridrow="19" pageId="10" pageNumber="203">Percents of head length</th>
</tr>
<tr box="[135,776,1863,1882]" gridrow="20" pageId="10" pageNumber="203">
<th box="[135,436,1863,1882]" gridcol="0" gridrow="20" pageId="10" pageNumber="203">Horizontal orbital diameter</th>
<td box="[473,537,1863,1882]" gridcol="1" gridrow="20" pageId="10" pageNumber="203">30.5</td>
<td box="[557,569,1863,1882]" gridcol="2" gridrow="20" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1863,1882]" gridcol="3" gridrow="20" pageId="10" pageNumber="203">30.1 - 33.3</td>
<td box="[698,739,1863,1882]" gridcol="4" gridrow="20" pageId="10" pageNumber="203">31.4</td>
<td box="[748,776,1863,1882]" gridcol="5" gridrow="20" pageId="10" pageNumber="203">1.4</td>
</tr>
<tr box="[135,776,1887,1906]" gridrow="21" pageId="10" pageNumber="203">
<th box="[135,436,1887,1906]" gridcol="0" gridrow="21" pageId="10" pageNumber="203">Snout length</th>
<td box="[473,537,1887,1906]" gridcol="1" gridrow="21" pageId="10" pageNumber="203">27.1</td>
<td box="[557,569,1887,1906]" gridcol="2" gridrow="21" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1887,1906]" gridcol="3" gridrow="21" pageId="10" pageNumber="203">26.0 - 27.1</td>
<td box="[698,739,1887,1906]" gridcol="4" gridrow="21" pageId="10" pageNumber="203">26.6</td>
<td box="[748,776,1887,1906]" gridcol="5" gridrow="21" pageId="10" pageNumber="203">0.4</td>
</tr>
<tr box="[135,776,1912,1931]" gridrow="22" pageId="10" pageNumber="203">
<th box="[135,436,1912,1931]" gridcol="0" gridrow="22" pageId="10" pageNumber="203">Least interorbital width</th>
<td box="[473,537,1912,1931]" gridcol="1" gridrow="22" pageId="10" pageNumber="203">22.0</td>
<td box="[557,569,1912,1931]" gridcol="2" gridrow="22" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1912,1931]" gridcol="3" gridrow="22" pageId="10" pageNumber="203">22.0 - 23.3</td>
<td box="[698,776,1912,1931]" colspan="2" colspanRight="1" gridcol="4" gridrow="22" pageId="10" pageNumber="203">22.6 0.8</td>
</tr>
<tr box="[135,776,1936,1956]" gridrow="23" pageId="10" pageNumber="203">
<th box="[135,436,1936,1956]" gridcol="0" gridrow="23" pageId="10" pageNumber="203">Upper jaw length</th>
<td box="[473,537,1936,1956]" gridcol="1" gridrow="23" pageId="10" pageNumber="203">67.8</td>
<td box="[557,569,1936,1956]" gridcol="2" gridrow="23" pageId="10" pageNumber="203">4</td>
<td box="[586,689,1936,1956]" gridcol="3" gridrow="23" pageId="10" pageNumber="203">65.7 - 67.8</td>
<td box="[698,739,1936,1956]" gridcol="4" gridrow="23" pageId="10" pageNumber="203">66.7</td>
<td box="[748,776,1936,1956]" gridcol="5" gridrow="23" pageId="10" pageNumber="203">0.8</td>
</tr>
</table>
</paragraph>
<paragraph blockId="10.[816,1455,1553,1673]" pageId="10" pageNumber="203">
<emphasis bold="true" box="[816,1044,1553,1577]" pageId="10" pageNumber="203">Sexual dimorphism.</emphasis>
Males of this species much darker than females and both sexes possess superficial neuromasts vertically arranged along basal portions of each scale albeit less numerous on females (
<figureCitation box="[1102,1164,1649,1673]" captionStart="Fig" captionStartId="3.[129,168,636,660]" captionTargetBox="[129,1452,174,615]" captionTargetId="figure-6@3.[129,1452,174,615]" captionTargetPageId="3" captionText="Fig. 1. Superficial neuromasts (black arrows) on lateral body scales of Charax delimai, male (a), MZUSP 81325, paratype, 102 mm SL, Brazil, Amazonas, igarapé Umari, tributary of rio Tiquié in community of São Pedro and female, (b), MZUSP 84988, paratype, 102 mm SL, Brazil, Amazonas, near old community of São Pedro." figureDoi="http://doi.org/10.5281/zenodo.4638997" httpUri="https://zenodo.org/record/4638997/files/figure.png" pageId="10" pageNumber="203">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="203" type="etymology">
<paragraph blockId="10.[816,1455,1713,1865]" pageId="10" pageNumber="203">
<emphasis bold="true" box="[816,947,1713,1737]" pageId="10" pageNumber="203">Etymology.</emphasis>
<taxonomicName authorityName="Menezes &amp; de Lucena" authorityYear="2014" box="[957,1131,1713,1737]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="10" pageNumber="203" phylum="Chordata" rank="species" species="delimai">
<emphasis box="[957,1131,1713,1737]" italics="true" pageId="10" pageNumber="203">Charax delimai</emphasis>
</taxonomicName>
is named after Flávio
<collectingRegion box="[1393,1455,1713,1737]" country="Colombia" name="Cesar" pageId="10" pageNumber="203">César</collectingRegion>
Thadeu de
<collectingRegion box="[932,989,1745,1769]" country="Peru" name="Lima" pageId="10" pageNumber="203">Lima</collectingRegion>
for his great contributions to the knowledge of neotropical freshwater fishes and for collecting most of the specimens that served as the basis for the species description.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="203" type="distribution">
<paragraph blockId="10.[816,1455,1905,1962]" pageId="10" pageNumber="203">
<emphasis bold="true" box="[816,965,1905,1929]" pageId="10" pageNumber="203">Distribution.</emphasis>
<taxonomicName authorityName="Menezes &amp; de Lucena" authorityYear="2014" box="[970,1137,1905,1929]" class="Actinopterygii" family="Characidae" genus="Charax" kingdom="Animalia" order="Characiformes" pageId="10" pageNumber="203" phylum="Chordata" rank="species" species="delimai">
<emphasis box="[970,1137,1905,1929]" italics="true" pageId="10" pageNumber="203">Charax delimai</emphasis>
</taxonomicName>
is known from the rio Tiquié, rio Negro basin,
<collectingCountry box="[988,1052,1937,1961]" name="Brazil" pageId="10" pageNumber="203">Brazil</collectingCountry>
(
<figureCitation box="[1064,1127,1937,1961]" captionStart="Fig" captionStartId="5.[809,847,1648,1672]" captionTargetBox="[809,1452,819,1618]" captionTargetId="figure-305@5.[809,1452,815,1618]" captionTargetPageId="5" captionText="Fig.7. Map of SouthAmerica showing the collecting sites of the Charax species, and the type locality of Charax delimai (purple circle). Charax apurensis - yellow square; C. caudimaculatus - white circle; C. condei - white square; C. gibbosus - purple triangle; C. hemigrammus - red star; C. leticiae - black circle;C. macrolepis-black triangle;C.metae-yellow triangle;C.michaeli - yellow circle;C.niger -blue triangle; C.notulatus - blue circle; C. pauciradiatus - red triangle; C. rupununi - red circle; C. stenopterus - green triangle; C. tectifer - green circle. Some symbols may represent more than one locality." figureDoi="http://doi.org/10.5281/zenodo.4639009" httpUri="https://zenodo.org/record/4639009/files/figure.png" pageId="10" pageNumber="203">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>