447 lines
78 KiB
XML
447 lines
78 KiB
XML
<document id="BEE95D0534FCB0273006B5835CF4A2A8" ID-DOI="10.5281/zenodo.278414" ID-GBIF-Dataset="1a59c750-bd1e-46c7-bbb4-05c0138d3e0b" ID-ISSN="1175-5326" ID-Zenodo-Dep="278414" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460337975097" checkinUser="plazi" docAuthor="Nogueira, João Miguel De Matos, Harris, Leslie, Hutchings, Pat & Fukuda, Marcelo Veronesi" docDate="2011" docId="03C4D91AD514FFB617FFFB9AFDC696A9" docLanguage="en" docName="zt02995p026.pdf" docOrigin="Zootaxa 2995" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Eupolymnia turgidula Ehlers 1887" docType="treatment" docVersion="8" lastPageNumber="24" masterDocId="FFFDA162D507FFA11768FF80FFCD972E" masterDocTitle="Four terebellines (Polychaeta, Terebellidae) with problematic taxonomic histories" masterLastPageNumber="26" masterPageNumber="1" pageNumber="20" updateTime="1698260250247" updateUser="plazi">
|
||
<mods:mods id="972D3E2BFAECD6366F9EFCC90B017517" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="ECB763491D9C5ADC5BB9846EBE823847">
|
||
<mods:title id="EEBE13C45B3CD4BD631C1E304F428339">Four terebellines (Polychaeta, Terebellidae) with problematic taxonomic histories</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="5DC3064105677CB464E05C781599ED95" type="personal">
|
||
<mods:role id="80406122DF5D30B9071256D54376E7CB">
|
||
<mods:roleTerm id="52BDCACEE5D277ADA11BDF2A5A78676C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A552BAC810F85A3E339A17E3F048E03C">Nogueira, João Miguel De Matos</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="41FF8CE8CCB44FB8A4B96F8111CFD0BF" type="personal">
|
||
<mods:role id="8865F050B61DEF759D2C62445D067F4B">
|
||
<mods:roleTerm id="24CC2BEA28C69236EF5A06DD587062D3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A9835A7D495E90AB67695764A68818FB">Harris, Leslie</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="DAEE30B286A656D940215BED72ABB927" type="personal">
|
||
<mods:role id="6FF71797FCE616433DF2D88DDC48CAA1">
|
||
<mods:roleTerm id="E1996675C33D7AE99E7BE4A5346421F4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A5BE4F04ED856990A065DDB798B37023">Hutchings, Pat</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="DAAF9F5DF8AA77EC647D7C33D6DB5C5D" type="personal">
|
||
<mods:role id="F9CED52A6AAA818D4E566CE389A6974D">
|
||
<mods:roleTerm id="A1501DF1AA9776B7D7D4B5FD11297536">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E636C8937858447E00755AF75E2E4259">Fukuda, Marcelo Veronesi</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="E8C25C38F7037FD91E4526A18870B37E">text</mods:typeOfResource>
|
||
<mods:relatedItem id="97100159F0064F72DBB33F24CE688A8A" type="host">
|
||
<mods:titleInfo id="A2AC29A3D081EA759ACA2BF4ECBFC570">
|
||
<mods:title id="AF3F6BCDE57DE4A94B8741C6B2A673FC">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="A9A558BB86EF6A7A003AFD1157AA9F3D">
|
||
<mods:date id="0812A76C34A581E23044AFCF5EBF838B">2011</mods:date>
|
||
<mods:detail id="6D16759923985F3EA68B0B5DC197BC6E" type="volume">
|
||
<mods:number id="C8A683105D41DF888E847BBAB693E7E7">2995</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="9B713E165DD64381EFA5FD9BA2BFB4E0" unit="page">
|
||
<mods:start id="72302F6C11AD59C52CA214F4B0CB46C2">1</mods:start>
|
||
<mods:end id="07BBD4C82D195FD3C8C8CDFF14E05670">26</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="BE0B40C71D7F047CCB503ED83CC27F31">journal article</mods:classification>
|
||
<mods:identifier id="476EE7C23D732059958CD4430AA8B443" type="DOI">10.5281/zenodo.278414</mods:identifier>
|
||
<mods:identifier id="6AA5D8FC8929EB1EE63AD026C0ABF21F" type="GBIF-Dataset">1a59c750-bd1e-46c7-bbb4-05c0138d3e0b</mods:identifier>
|
||
<mods:identifier id="C5A74414CB87AEFE116DC4C971CEDCB0" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="543B92BE1F8B3D6F29C28400A4E421F4" type="Zenodo-Dep">278414</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C4D91AD514FFB617FFFB9AFDC696A9" ID-DOI="http://doi.org/10.5281/zenodo.5633384" ID-GBIF-Taxon="119405526" ID-Zenodo-Dep="5633384" LSID="urn:lsid:plazi:treatment:03C4D91AD514FFB617FFFB9AFDC696A9" httpUri="http://treatment.plazi.org/id/03C4D91AD514FFB617FFFB9AFDC696A9" lastPageId="23" lastPageNumber="24" pageId="19" pageNumber="20">
|
||
<subSubSection id="C3773B87D514FFB217FFFB9AFEE7937B" pageId="19" pageNumber="20" type="nomenclature">
|
||
<paragraph id="8BD2680CD514FFB217FFFB9AFD95931A" blockId="19.[151,600,1050,1109]" box="[151,600,1050,1076]" pageId="19" pageNumber="20">
|
||
<heading id="D09ADF60D514FFB217FFFB9AFD95931A" bold="true" box="[151,600,1050,1076]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
|
||
<taxonomicName id="4C6D138FD514FFB217FFFB9AFD95931A" ID-CoL="7BQ9H" authority="Ehlers, 1887" authorityName="Ehlers" authorityYear="1887" box="[151,600,1050,1076]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED514FFB217FFFB9AFD95931A" bold="true" box="[151,600,1050,1076]" pageId="19" pageNumber="20">
|
||
<emphasis id="B919B41ED514FFB217FFFB9AFE6F931A" bold="true" box="[151,418,1050,1076]" italics="true" pageId="19" pageNumber="20">Eupolymnia turgidula</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFC15FDD514FFB216DAFB9AFD9D931A" author="Ehlers" box="[434,592,1050,1076]" pageId="19" pageNumber="20" refString="Ehlers, E. (1887) Florida-Anneliden. (Report on the annelids of the dredging expedition of the U. S. coast survey steamer Blake). Memoirs of the Museum of Comparative Zoology of Harvard, 15, VI + 335 p, pl. 1 - 60." type="book chapter" year="1887">Ehlers, 1887</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD2680CD514FFB217FFFBBDFEE7937B" blockId="19.[151,600,1050,1109]" box="[151,298,1085,1109]" pageId="19" pageNumber="20">
|
||
<figureCitation id="13567489D514FFB217FFFBBDFEE7937B" box="[151,298,1085,1109]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="20.[151,250,1825,1848]" captionStartId-1="22.[151,250,1951,1974]" captionTargetBox-0="[230,1353,198,1791]" captionTargetBox-1="[157,1431,307,1923]" captionTargetId-0="figure@20.[225,1359,193,1797]" captionTargetId-1="figure@22.[151,1436,301,1930]" captionTargetPageId-0="20" captionTargetPageId-1="22" captionText-0="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." captionText-1="FIGURE 10. Eupolymnia turgidula, holotype (MCZ 846). A: notochaetae, segment 18; B: tips of some notochaetae from posterior tier, segment 18; C: uncini, segment 7; D – E: uncini, segment 12; F: uncini, segment 23. Scale bars: A, E = 80 µm; B = 40 µm; C = 15 µm; D = 10 µm; F = 30 µm." httpUri-0="https://zenodo.org/record/278423/files/figure.png" httpUri-1="https://zenodo.org/record/278424/files/figure.png" pageId="19" pageNumber="20">Figures 9–10</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3773B87D514FFB217FFFB00FCFB93DF" pageId="19" pageNumber="20" type="reference_group">
|
||
<paragraph id="8BD2680CD514FFB217FFFB00FCDA93BB" blockId="19.[151,960,1152,1266]" box="[151,791,1152,1174]" pageId="19" pageNumber="20">
|
||
<treatmentCitationGroup id="AB7D4F22D514FFB217FFFB00FCDA93BB" box="[151,791,1152,1174]" pageId="19" pageNumber="20">
|
||
<taxonomicName id="4C6D138FD514FFB217FFFB00FDDF93BB" ID-CoL="7BQ9H" authority="Ehlers, 1887: 241" authorityName="Ehlers" authorityPageNumber="241" authorityYear="1887" box="[151,530,1152,1174]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED514FFB217FFFB00FE9593B8" box="[151,344,1152,1174]" italics="true" pageId="19" pageNumber="20">Terebella turgidula</emphasis>
|
||
<treatmentCitation id="0ACC4E1DD514FFB21636FB00FDDF93BB" author="Ehlers" box="[350,530,1152,1174]" page="241" pageId="19" pageNumber="20" year="1887">
|
||
<bibRefCitation id="EFFC15FDD514FFB21636FB00FE2F93BB" author="Ehlers" box="[350,482,1152,1174]" pageId="19" pageNumber="20" refString="Ehlers, E. (1887) Florida-Anneliden. (Report on the annelids of the dredging expedition of the U. S. coast survey steamer Blake). Memoirs of the Museum of Comparative Zoology of Harvard, 15, VI + 335 p, pl. 1 - 60." type="book chapter" year="1887">Ehlers, 1887</bibRefCitation>
|
||
: 241
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
– 245, Taf. 52,
|
||
<figureCitation id="13567489D514FFB215DAFB00FCDE93BB" box="[690,787,1152,1174]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1828,1851]" captionStartId-1="4.[151,250,1959,1982]" captionStartId-2="6.[151,250,1853,1876]" captionStartId-3="8.[151,250,1789,1812]" captionTargetBox-0="[197,1392,183,1791]" captionTargetBox-1="[154,1429,1493,1932]" captionTargetBox-2="[187,1400,199,1825]" captionTargetBox-3="[156,1431,408,1762]" captionTargetId-0="figure@3.[193,1398,177,1796]" captionTargetId-1="figure@4.[151,1436,1489,1938]" captionTargetId-2="figure@6.[181,1405,193,1832]" captionTargetId-3="figure@8.[151,1436,403,1768]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="FIGURE 1. Pista corrientis, holotype (BMNH 1885.12.1.348). A: anterior end, ventral view; B: anterior end, right lateral view; C: anterior end, left lateral view; D: anterior parapodia of region with neuropodia only; E: posterior end, left lateral view; F: detail of anterior end, dorsal view; G: detail of anterior end, left lateral view; H: detail of anterior end, ventral view; I: detail of anterior end, right lateral view; J: close up view of anterior end, dorsal view; K: close up view of anterior end, left lateral view; L: close up view of anterior end, ventral view. Numbers refer to segments; unspecified arrows show nephridial or genital papillae; ll = lower lip; P = prostomium; po = pharyngeal organ; ul = upper lip. Scale bars: A = 0.8 mm; B – C = 0.6 mm; D = 0.3 mm; E = 0.2 mm; F – L = 0.5 mm." captionText-1="FIGURE 2. Pista corrientis, holotype (BMNH 1885.12.1.348). A: uncini, segment 9; B: uncini, segment 23; C: uncinus, segment 19; D: uncinus, segment 83. Scale bars: A = 20 µm; B – C = 10 µm; D = 5 µm." captionText-2="FIGURE 3. Pista nonatoi sp. nov., holotype (MZUSP 00950). A: worm inside the tube, anterior end, right lateral view; B: anterior end, right ventrolateral view; C: anterior end, right lateral view; D: anterior end, right dorsolateral view; E: detail of the anterior end, right lateral view; F: detail of the anterior end, right dorsolateral view; arrows point to genital papillae; G: neuropodia from anterior region with neuropodia only; H: neuropodia from posterior region with neuropodia only; I – J: close up views of anterior end, ventral view; K: close up view of anterior end, dorsal view. Numbers refer to segments; ll = lower lip; P = prostomium; ul = upper lip. Scale bars: A = 2 mm; B = 1.5 mm; C – F, I = 1 mm; G = 0.4 mm; H, J = 0.5 mm; K = 0.7 mm." captionText-3="FIGURE 4. Pista nonatoi sp. nov., (spec. AM W 34750). A: anterior end, ventral view; B: anterior end, right dorsolateral view; C: anterior end, dorsal view; D: anterior end, right lateral view; E: anterior end, oral area and left lateral, superior view; F: oral area, superior view; G: close up view of oral area, ventral view; H: close up view of anterior end, left lateral view; I: close up view of anterior end, dorsal view; J: close up view of anterior end, ventral view; K: close up view of anterior end, left ventrolateral view; L: posterior segments, left lateral view. Numbers refer to segments; unspecified arrows point to genital papillae; ll = lower lip; P = prostomium; ul = upper lip; arrows point to nephridial and genital papillae. Scale bars: A – B, D, G – J = 0.4 mm; C, E – F, K, L = 0.3 mm." httpUri-0="https://zenodo.org/record/278415/files/figure.png" httpUri-1="https://zenodo.org/record/278416/files/figure.png" httpUri-2="https://zenodo.org/record/278417/files/figure.png" httpUri-3="https://zenodo.org/record/278418/files/figure.png" pageId="19" pageNumber="20">Figs. 1–4</figureCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD2680CD514FFB217FFFB1FFCDC93FD" blockId="19.[151,960,1152,1266]" pageId="19" pageNumber="20">
|
||
<treatmentCitationGroup id="AB7D4F22D514FFB217FFFB1FFCDC93FD" pageId="19" pageNumber="20">
|
||
?
|
||
<taxonomicName id="4C6D138FD514FFB217C0FB1FFD91939A" ID-CoL="7BQ5Z" authority="Schmarda, 1861: 43" authorityName="Schmarda" authorityPageNumber="43" authorityYear="1861" box="[168,604,1182,1205]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="crassicornis">
|
||
<emphasis id="B919B41ED514FFB217C0FB1FFE44939B" box="[168,393,1183,1205]" italics="true" pageId="19" pageNumber="20">Terebella crassicornis</emphasis>
|
||
<treatmentCitation id="0ACC4E1DD514FFB216E6FB1EFD91939A" author="Schmarda" box="[398,604,1182,1204]" page="43" pageId="19" pageNumber="20" year="1861">
|
||
<bibRefCitation id="EFFC15FDD514FFB216E6FB1EFDFB939A" author="Schmarda" box="[398,566,1182,1204]" pageId="19" pageNumber="20" refString="Schmarda, L. K. (1861) Neue wirbellose Thiere beobachtet und gesammelt auf einer Reise um die Erdr 1853 bis 1857. 1. Turbellarien, Rotatorien und Anneliden, pt. 2. Wilhelm Engelmann, Leipzig, 164 pp." type="book" year="1861">Schmarda, 1861</bibRefCitation>
|
||
: 43
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, Figs. A–C;
|
||
<treatmentCitation id="0ACC4E1DD514FFB215B4FB1FFC5E939A" author="Augener" box="[732,915,1182,1204]" page="36" pageId="19" pageNumber="20" year="1925">
|
||
<bibRefCitation id="EFFC15FDD514FFB215B4FB1FFCA2939A" author="Augener" box="[732,879,1182,1204]" pageId="19" pageNumber="20" refString="Augener, H. (1925) Uber westindische und einige andere Polychaeten-Typen von Grube (Oersted), Kroyer, Morch und Schmarda. Publikationer fra Universitetets Zoologiske Museum Kobenhavn, 39, 1 - 47." type="journal article" year="1925">Augener 1925</bibRefCitation>
|
||
: 36
|
||
</treatmentCitation>
|
||
–37.?
|
||
<taxonomicName id="4C6D138FD514FFB217C0FB3DFDF393FD" ID-CoL="7BQ97" authority="Webster, 1884: 324" authorityName="Webster" authorityPageNumber="324" authorityYear="1884" box="[168,574,1213,1235]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="magnifica">
|
||
<emphasis id="B919B41ED514FFB217C0FB3DFEBC93FD" box="[168,369,1213,1235]" italics="true" pageId="19" pageNumber="20">Terebella magnifica</emphasis>
|
||
<treatmentCitation id="0ACC4E1DD514FFB21610FB3DFDF393FD" author="Webster" box="[376,574,1213,1235]" page="324" pageId="19" pageNumber="20" year="1884">
|
||
<bibRefCitation id="EFFC15FDD514FFB21610FB3DFDC193FD" author="Webster" box="[376,524,1213,1235]" pageId="19" pageNumber="20" refString="Webster, H. E. (1884) Annelida from Bermuda collected by G. Brown Goode. Bulletin of the United States National Museum, 25, 307 - 327." type="journal article" year="1884">Webster, 1884</bibRefCitation>
|
||
: 324
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, Pl. XI, Figs. 58–60.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BD2680CD514FFB217FFFB5DFCFB93DF" blockId="19.[151,960,1152,1266]" box="[151,822,1243,1266]" pageId="19" pageNumber="20">
|
||
<treatmentCitationGroup id="AB7D4F22D514FFB217FFFB5DFCFB93DF" box="[151,822,1243,1266]" pageId="19" pageNumber="20">
|
||
Not
|
||
<taxonomicName id="4C6D138FD514FFB217ABFB5CFE4993DC" ID-CoL="7BQ9H" box="[195,388,1244,1266]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="19" pageNumber="20" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED514FFB217ABFB5CFE4993DC" box="[195,388,1244,1266]" italics="true" pageId="19" pageNumber="20">Terebella turgidula</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<treatmentCitation id="0ACC4E1DD514FFB216F9FB5BFD4693DF" ID-DOI="http://doi.org/10.5281/zenodo.5324818" ID-Zenodo-Dep="5324818" author="Londono-Mesa" box="[401,651,1243,1265]" httpUri="http://treatment.plazi.org/id/03F75303AE60FF81FF7EFA633D8587B5" page="63" pageId="19" pageNumber="20" year="2009">
|
||
<bibRefCitation id="EFFC15FDD514FFB216F9FB5BFDA493DF" author="Londono-Mesa" box="[401,617,1243,1265]" pageId="19" pageNumber="20" refString="Londono-Mesa, M. H. (2009) Terebellidae (Polychaeta: Terebellida) from the Grand Caribbean Region. Zootaxa, 2320, 1 - 93." type="journal article" year="2009">Londoño-Mesa 2009</bibRefCitation>
|
||
: 63
|
||
</treatmentCitation>
|
||
–64, Fig. 18D–J.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3773B87D514FFB217FFFA9EFC1D9250" pageId="19" pageNumber="20" type="materials_examined">
|
||
<paragraph id="8BD2680CD514FFB217FFFA9EFC1D9250" blockId="19.[151,1436,1310,2018]" pageId="19" pageNumber="20">
|
||
<emphasis id="B919B41ED514FFB217FFFA9EFE4E9219" bold="true" box="[151,387,1310,1335]" pageId="19" pageNumber="20">Material examined.</emphasis>
|
||
<typeStatus id="54D6D6AED514FFB216E4FA9EFE399218" box="[396,500,1310,1334]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED7CF0C9D514FFB2156EFA9EFD889218" LSID="urn:lsid:biocol.org:col:33791" box="[518,581,1310,1334]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="19" pageNumber="20">MCZ</collectionCode>
|
||
846): incomplete specimen in poor state of preservation (see below), ~
|
||
<quantity id="4C95C5E9D514FFB212E8FA9EFF0E9274" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="20" unit="mm" value="15.0">15 mm</quantity>
|
||
long,
|
||
<quantity id="4C95C5E9D514FFB21662FAC2FE869274" box="[266,331,1346,1370]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="mm" value="2.0">2 mm</quantity>
|
||
wide; coll.
|
||
<collectingCountry id="F37A289CD514FFB216B8FAC2FDC69274" box="[464,523,1346,1370]" name="United States of America" pageId="19" pageNumber="20">USA</collectingCountry>
|
||
, Florida, Key West,
|
||
<quantity id="4C95C5E9D514FFB21599FAC2FCF79274" box="[753,826,1346,1370]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="19" pageNumber="20" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">2–4 m</quantity>
|
||
,
|
||
<emphasis id="B919B41ED514FFB2142EFAC3FC4B9275" box="[838,902,1347,1371]" italics="true" pageId="19" pageNumber="20">Blake</emphasis>
|
||
Expedition, by A. Agassiz,
|
||
<date id="FFD34ECCD514FFB213ABFAC2FA889274" box="[1219,1349,1346,1370]" pageId="19" pageNumber="20" value="1877" valueMax="1878">1877–1878</date>
|
||
. Slides: notochaetae from segment 18; neurochaetae from segments 7, 12, and 23.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3773B87D514FFB417AFFA0AFECA95AA" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="description">
|
||
<paragraph id="8BD2680CD514FFB417AFFA0AFECA95AA" blockId="19.[151,1436,1310,2018]" lastBlockId="21.[151,1437,151,2013]" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
|
||
<emphasis id="B919B41ED514FFB217AFFA0AFE94928D" bold="true" box="[199,345,1418,1443]" pageId="19" pageNumber="20">Description.</emphasis>
|
||
<typeStatus id="54D6D6AED514FFB2160BFA0AFE06928C" box="[355,459,1418,1442]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
|
||
incomplete specimen, with region with biramous parapodia and ~26 segments with neuropodia only, in poor state of preservation, nearly torn midway along region with biramous parapodia and also at transition between regions with biramous parapodia and with neuropodia only. Pieces of tube together with specimen; mucous, transparent, with some embedded coarse shells (
|
||
<figureCitation id="13567489D514FFB2140DFA76FC649120" box="[869,937,1526,1550]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
|
||
K). Body slightly inflated dorsally after segment 4 (
|
||
<figureCitation id="13567489D514FFB2179CF99AFEF5911C" box="[244,312,1562,1586]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
|
||
B, G–H). Ventral shields on segments 3–18, all corrugated, more evident on anterior segments (
|
||
<figureCitation id="13567489D514FFB21207F99AFF699178" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
|
||
C–G, I); shields progressively wider on segments 3–5, then progressively narrower until last shield; anterior shields short and compact, progressively longer from segment 10, last shield distinctly shorter (
|
||
<figureCitation id="13567489D514FFB213ADF9E2FACB9154" box="[1221,1286,1634,1658]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
|
||
C–G, I); after segment 18, shields continue as mid-ventral stripe extending posteriorly. Prostomium at base of upper lip; distal part forming shelf-like process from which buccal tentacles originate (
|
||
<figureCitation id="13567489D514FFB214AAF92AFBCA91EC" box="[962,1031,1706,1730]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
|
||
B–H); eyespots still conspicuous at least at left lateral of basal part of prostomium, apparently forming continuous row, laterally broader, distinctly thinner mid-dorsally. Peristomium restricted to lips; upper lip short, broader than long; lower lip short, almost completely covered by ventral lobe on segment 1 (
|
||
<figureCitation id="13567489D514FFB215F3F896FD109000" box="[667,733,1814,1838]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
|
||
B–H). Segment 1 short, more developed ventrally, with ventral lobe around and below lower lip (
|
||
<figureCitation id="13567489D514FFB21571F8BAFD90907C" box="[537,605,1850,1874]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
|
||
C–G). Segment 2 with one pair of relatively large, rounded ventro-lateral lobes, connected to each other by low flap across ventrum (
|
||
<figureCitation id="13567489D514FFB2145CF8DEFCBA9058" box="[820,887,1886,1910]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
|
||
C–G). Segments 3–4 with short, rounded ventrolateral lobes, progressively more laterally placed, those on segment 3 larger and connected to each other by crest across ventrum (
|
||
<figureCitation id="13567489D514FFB21638F826FE5F9090" box="[336,402,1958,1982]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
|
||
C, E–G). Three pairs of branchiae on segments 2–4, with short, stout and annulated cylindrical basal stems branching to short distal filaments; branchiae progressively shorter and vertically aligned, last pair distinctly shorter than other pairs (
|
||
<figureCitation id="13567489D512FFB41545FF17FDBE9781" box="[557,627,151,175]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="21" pageNumber="22">Fig. 9</figureCitation>
|
||
B–C, E–H). Seventeen pairs of notopodia, starting on segment 4 and extending to segment 20; notopodia relatively long. Narrowly winged, distally smooth notochaetae on both tiers, those from posterior tier about twice as long as those on anterior tier, with limbation on distal half of chaetae (
|
||
<figureCitation id="13567489D512FFB41207FF5FFF7E9632" captionStart="FIGURE 10" captionStartId="22.[151,250,1951,1974]" captionTargetBox="[157,1431,307,1923]" captionTargetId="figure@22.[151,1436,301,1930]" captionTargetPageId="22" captionText="FIGURE 10. Eupolymnia turgidula, holotype (MCZ 846). A: notochaetae, segment 18; B: tips of some notochaetae from posterior tier, segment 18; C: uncini, segment 7; D – E: uncini, segment 12; F: uncini, segment 23. Scale bars: A, E = 80 µm; B = 40 µm; C = 15 µm; D = 10 µm; F = 30 µm." httpUri="https://zenodo.org/record/278424/files/figure.png" pageId="21" pageNumber="22">Fig. 10</figureCitation>
|
||
A–B). Neuropodia starting on segment 5, as low rectangular ridges slightly raised from surface of body until segment on which notopodia terminate (
|
||
<figureCitation id="13567489D512FFB41508FEA7FD6F9611" box="[608,674,295,319]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="21" pageNumber="22">Fig. 9</figureCitation>
|
||
C–G, I), as long, foliaceous pinnules thereafter, with internal shafts and small, spherical dorsal lobe, first two pinnules much shorter than following ones (
|
||
<figureCitation id="13567489D512FFB413EAFECCFB0C964A" box="[1154,1217,332,356]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="21" pageNumber="22">
|
||
Fig
|
||
<date id="FFD34ECCD512FFB413DBFECCFB0C964A" box="[1203,1217,332,356]" pageId="21" pageNumber="22">9</date>
|
||
</figureCitation>
|
||
I–J). Short-handled uncini throughout, about as long as high on region with biramous parapodia, with pronounced dorsal button situated closer to tip of prow than base of main fang, distally rounded prow and crest with 2 rows of secondary teeth, first row with two large teeth, second row with single minute tooth placed between teeth of first row, often not visible in lateral view (
|
||
<figureCitation id="13567489D512FFB41610FE5CFE0496DA" box="[376,457,476,500]" captionStart="FIGURE 10" captionStartId="22.[151,250,1951,1974]" captionTargetBox="[157,1431,307,1923]" captionTargetId="figure@22.[151,1436,301,1930]" captionTargetPageId="22" captionText="FIGURE 10. Eupolymnia turgidula, holotype (MCZ 846). A: notochaetae, segment 18; B: tips of some notochaetae from posterior tier, segment 18; C: uncini, segment 7; D – E: uncini, segment 12; F: uncini, segment 23. Scale bars: A, E = 80 µm; B = 40 µm; C = 15 µm; D = 10 µm; F = 30 µm." httpUri="https://zenodo.org/record/278424/files/figure.png" pageId="21" pageNumber="22">Fig. 10</figureCitation>
|
||
C–E); uncini in completely intercalated double rows from segment 11 until segment on which notopodia terminate (
|
||
<figureCitation id="13567489D512FFB416BAFE7FFDEE9539" box="[466,547,511,535]" captionStart="FIGURE 10" captionStartId="22.[151,250,1951,1974]" captionTargetBox="[157,1431,307,1923]" captionTargetId="figure@22.[151,1436,301,1930]" captionTargetPageId="22" captionText="FIGURE 10. Eupolymnia turgidula, holotype (MCZ 846). A: notochaetae, segment 18; B: tips of some notochaetae from posterior tier, segment 18; C: uncini, segment 7; D – E: uncini, segment 12; F: uncini, segment 23. Scale bars: A, E = 80 µm; B = 40 µm; C = 15 µm; D = 10 µm; F = 30 µm." httpUri="https://zenodo.org/record/278424/files/figure.png" pageId="21" pageNumber="22">Fig. 10</figureCitation>
|
||
E); from segment 21, uncini longer than high, with distally pointed prow, otherwise as those on region with biramous parapodia (
|
||
<figureCitation id="13567489D512FFB415ADFDA4FCDA9512" box="[709,791,548,572]" captionStart="FIGURE 10" captionStartId="22.[151,250,1951,1974]" captionTargetBox="[157,1431,307,1923]" captionTargetId="figure@22.[151,1436,301,1930]" captionTargetPageId="22" captionText="FIGURE 10. Eupolymnia turgidula, holotype (MCZ 846). A: notochaetae, segment 18; B: tips of some notochaetae from posterior tier, segment 18; C: uncini, segment 7; D – E: uncini, segment 12; F: uncini, segment 23. Scale bars: A, E = 80 µm; B = 40 µm; C = 15 µm; D = 10 µm; F = 30 µm." httpUri="https://zenodo.org/record/278424/files/figure.png" pageId="21" pageNumber="22">Fig. 10</figureCitation>
|
||
F). Nephridial papillae on segments 3–5, between parapodial lobes on segment 5, in equivalent position on segments 3–4 (
|
||
<figureCitation id="13567489D512FFB41406FDC7FC7D9571" box="[878,944,583,607]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="21" pageNumber="22">Fig. 9</figureCitation>
|
||
E–G), genital papillae not visible. Pygidium unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF123884D513FFB517FFF8A1FCE590DE" httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="20" pageNumber="21" targetBox="[230,1353,198,1791]" targetPageId="20">
|
||
<paragraph id="8BD2680CD513FFB517FFF8A1FCE590DE" blockId="20.[151,1436,1825,2032]" pageId="20" pageNumber="21">
|
||
<emphasis id="B919B41ED513FFB517FFF8A1FEDF9019" bold="true" box="[151,274,1825,1848]" pageId="20" pageNumber="21">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C6D138FD513FFB51670F8A2FE389016" box="[280,501,1826,1848]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="20" pageNumber="21" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED513FFB51670F8A2FE389016" box="[280,501,1826,1848]" italics="true" pageId="20" pageNumber="21">Eupolymnia turgidula</emphasis>
|
||
</taxonomicName>
|
||
, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E–F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P(dp) = prostomium, distal part; ul = upper lip. Scale bars: B–C = 1.5 mm; D–I = 1 mm; J = 0.6 mm; K = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3773B87D512FFB617AFFD10FDC696A9" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="discussion">
|
||
<paragraph id="8BD2680CD512FFB417AFFD10FE1795C1" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
|
||
<emphasis id="B919B41ED512FFB417AFFD10FEF69586" bold="true" box="[199,315,656,680]" pageId="21" pageNumber="22">Remarks.</emphasis>
|
||
The state of preservation prevents the proper evaluation of some characters, such as the alignment of the first pairs of notopodia and the position of neuropodial papillae in relation to the mid-ventral stripe in the region with neuropodia only.
|
||
</paragraph>
|
||
<paragraph id="8BD2680CD512FFB417AFFD7CFDBF9451" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
|
||
<collectingCountry id="F37A289CD512FFB417AFFD7CFF2B943A" box="[199,230,764,788]" name="American Samoa" pageId="21" pageNumber="22">As</collectingCountry>
|
||
discussed by
|
||
<bibRefCitation id="EFFC15FDD512FFB416EEFD7CFD4B943A" author="Londono-Mesa" box="[390,646,764,788]" pageId="21" pageNumber="22" refString="Londono-Mesa, M. H. (2009) Terebellidae (Polychaeta: Terebellida) from the Grand Caribbean Region. Zootaxa, 2320, 1 - 93." type="journal article" year="2009">Londoño-Mesa (2009)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C6D138FD512FFB415FBFD7DFCD7943A" box="[659,794,764,788]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED512FFB415FBFD7DFCD7943A" box="[659,794,764,788]" italics="true" pageId="21" pageNumber="22">E. turgidula</emphasis>
|
||
</taxonomicName>
|
||
was described as a species of
|
||
<taxonomicName id="4C6D138FD512FFB4131FFD7CFA55943A" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" box="[1143,1432,764,788]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="genus">
|
||
<emphasis id="B919B41ED512FFB4131FFD7CFB13943A" box="[1143,1246,764,788]" italics="true" pageId="21" pageNumber="22">Terebella</emphasis>
|
||
<bibRefCitation id="EFFC15FDD512FFB4138FFD7CFA55943A" author="Linnaeus" box="[1255,1432,764,788]" pageId="21" pageNumber="22" refString="Linnaeus, C. (1767) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis, 12 ed. Vol. 1. L. Salvius, Holmiae, 533 - 1327." type="journal article" year="1767">Linnaeus, 1767</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFFC15FDD512FFB417FFFC9FFEF99419" author="Hessle" box="[151,308,799,823]" pageId="21" pageNumber="22" refString="Hessle, C. (1917) Zur Kenntnis der terebellomorphen Polychaeten. Zoologiska Bidrag fran Uppsala, 5, 39 - 258." type="journal article" year="1917">Hessle (1917)</bibRefCitation>
|
||
transferred it to the genus
|
||
<taxonomicName id="4C6D138FD512FFB4150CFCA0FC599419" authority="Malmgren, 1867" authorityName="Malmgren" authorityYear="1867" box="[612,916,799,824]" class="Magnoliopsida" family="Asteraceae" genus="Polymnia" kingdom="Animalia" order="Asterales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B919B41ED512FFB4150CFCA0FD029416" box="[612,719,800,824]" italics="true" pageId="21" pageNumber="22">Polymnia</emphasis>
|
||
<bibRefCitation id="EFFC15FDD512FFB415BEFC9FFC599419" author="Malmgren" box="[726,916,799,823]" pageId="21" pageNumber="22" refString="Malmgren, A. J. (1867) Annulata Polychaeta Spetsbergiae, Groenlandiae, Islandiae et Scandinaviae hactenus cognita. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, Stockholm, 24, 127 - 235." type="journal article" year="1867">Malmgren, 1867</bibRefCitation>
|
||
</taxonomicName>
|
||
, which was changed to
|
||
<taxonomicName id="4C6D138FD512FFB413CAFCA0FAE49416" box="[1186,1321,800,824]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="genus">
|
||
<emphasis id="B919B41ED512FFB413CAFCA0FAE49416" box="[1186,1321,800,824]" italics="true" pageId="21" pageNumber="22">Eupolymnia</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFFC15FDD512FFB4123DFC9FFF299472" author="Verrill" pageId="21" pageNumber="22" refString="Verrill, A. E. (1900) Additions to the Turbellaria, Nemertina, and Annelida of the Bermudas, with a revision of the New England genera and species. Transactions of the Connecticut Academy of Arts and Sciences, 10, 595 - 671." type="journal article" year="1900">Verrill (1900)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="EFFC15FDD512FFB41799FCC4FE699472" author="Augener" box="[241,420,836,860]" pageId="21" pageNumber="22" refString="Augener, H. (1925) Uber westindische und einige andere Polychaeten-Typen von Grube (Oersted), Kroyer, Morch und Schmarda. Publikationer fra Universitetets Zoologiske Museum Kobenhavn, 39, 1 - 47." type="journal article" year="1925">Augener (1925)</bibRefCitation>
|
||
then synonymized it with
|
||
<taxonomicName id="4C6D138FD512FFB415BDFCC5FCB29472" box="[725,895,837,860]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="crassicornis">
|
||
<emphasis id="B919B41ED512FFB415BDFCC5FCB29472" box="[725,895,837,860]" italics="true" pageId="21" pageNumber="22">E. crassicornis</emphasis>
|
||
</taxonomicName>
|
||
, which was subsequently followed by
|
||
<bibRefCitation id="EFFC15FDD512FFB41252FCC5FF2E9451" author="Hartman" pageId="21" pageNumber="22" refString="Hartman, O. (1938) Annotated list of the types of polychaetous annelids in the Museum of Comparative Zoology. Bulletin of the Museum of Comparative Zoology of Harvard, 85, 1 - 31." type="journal article" year="1938">Hartman (1938)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFFC15FDD512FFB41787FCE7FE5C9451" author="Rullier" box="[239,401,871,895]" pageId="21" pageNumber="22" refString="Rullier, F. (1974) Quelques annelides polychetes de Cuba recueillies dans des eponges. Travaux du Museum d'Histoire Naturelle Grigore Antipa, 14, 9 - 77." type="journal article" year="1974">Rullier (1974)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="EFFC15FDD512FFB416A4FCE7FDA09451" author="Holthe" box="[460,621,871,895]" pageId="21" pageNumber="22" refString="Holthe, T. (1986) Evolution, systematics and distribution of the Polychaeta Terebellomorpha, with a catalogue of the taxa and a bibliography. Gunneria, 55, 1 - 236." type="journal article" year="1986">Holthe (1986)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD2680CD512FFB417AFFC0CFBAC9352" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
|
||
Recently,
|
||
<bibRefCitation id="EFFC15FDD512FFB4165FFC0CFDF9948A" author="Londono-Mesa" box="[311,564,908,932]" pageId="21" pageNumber="22" refString="Londono-Mesa, M. H. (2009) Terebellidae (Polychaeta: Terebellida) from the Grand Caribbean Region. Zootaxa, 2320, 1 - 93." type="journal article" year="2009">Londoño-Mesa (2009)</bibRefCitation>
|
||
resurrected this taxon as a species of
|
||
<taxonomicName id="4C6D138FD512FFB41488FC0CFB8A948A" box="[992,1095,908,932]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="genus">
|
||
<emphasis id="B919B41ED512FFB41488FC0CFB8A948A" box="[992,1095,908,932]" italics="true" pageId="21" pageNumber="22">Terebella</emphasis>
|
||
</taxonomicName>
|
||
. However, the specimen Londoño-Mesa described and illustrated (his Figs. 18D–J) as the
|
||
<typeStatus id="54D6D6AED512FFB4142AFC2FFC6994E9" box="[834,932,943,967]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
is not the same specimen that was sent to us, although the catalog number quoted by
|
||
<bibRefCitation id="EFFC15FDD512FFB4150AFC54FCAC94C2" author="Londono-Mesa" box="[610,865,980,1004]" pageId="21" pageNumber="22" refString="Londono-Mesa, M. H. (2009) Terebellidae (Polychaeta: Terebellida) from the Grand Caribbean Region. Zootaxa, 2320, 1 - 93." type="journal article" year="2009">Londoño-Mesa (2009)</bibRefCitation>
|
||
is the same as the specimen we examined (MCZ 846). We are at a loss to explain this discrepancy. Londoño-Mesa’s specimen is
|
||
<quantity id="4C95C5E9D512FFB4134CFC77FB4C933E" box="[1060,1153,1015,1040]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" pageId="21" pageNumber="22" unit="mm" value="210.0">210 mm</quantity>
|
||
long with 140 segments, it does not have lobes on anterior segments, it has 27 pairs of notopodia and distally serrated notochaetae. Ehlers stated that his specimen was
|
||
<quantity id="4C95C5E9D512FFB4168CFBC0FDF99376" box="[484,564,1088,1112]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="21" pageNumber="22" unit="mm" value="31.0">31 mm</quantity>
|
||
long for 53 segments, had 18 pairs of notopodia, and had lobes on anterior segments (clearly illustrated in
|
||
<bibRefCitation id="EFFC15FDD512FFB4169FFBE4FD4A9352" author="Ehlers" box="[503,647,1124,1148]" pageId="21" pageNumber="22" refString="Ehlers, E. (1887) Florida-Anneliden. (Report on the annelids of the dredging expedition of the U. S. coast survey steamer Blake). Memoirs of the Museum of Comparative Zoology of Harvard, 15, VI + 335 p, pl. 1 - 60." type="book chapter" year="1887">Ehlers, 1887</bibRefCitation>
|
||
, Taf. 52,
|
||
<figureCitation id="13567489D512FFB41587FBE4FCFD9352" box="[751,816,1124,1148]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1851]" captionTargetBox="[197,1392,183,1791]" captionTargetId="figure@3.[193,1398,177,1796]" captionTargetPageId="3" captionText="FIGURE 1. Pista corrientis, holotype (BMNH 1885.12.1.348). A: anterior end, ventral view; B: anterior end, right lateral view; C: anterior end, left lateral view; D: anterior parapodia of region with neuropodia only; E: posterior end, left lateral view; F: detail of anterior end, dorsal view; G: detail of anterior end, left lateral view; H: detail of anterior end, ventral view; I: detail of anterior end, right lateral view; J: close up view of anterior end, dorsal view; K: close up view of anterior end, left lateral view; L: close up view of anterior end, ventral view. Numbers refer to segments; unspecified arrows show nephridial or genital papillae; ll = lower lip; P = prostomium; po = pharyngeal organ; ul = upper lip. Scale bars: A = 0.8 mm; B – C = 0.6 mm; D = 0.3 mm; E = 0.2 mm; F – L = 0.5 mm." httpUri="https://zenodo.org/record/278415/files/figure.png" pageId="21" pageNumber="22">Fig. 1</figureCitation>
|
||
, included here as
|
||
<figureCitation id="13567489D512FFB41493FBE4FBF09352" box="[1019,1085,1124,1148]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="21" pageNumber="22">Fig. 9</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="8BD2680CD512FFB417AFFB08FD2793C6" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
|
||
Although the specimen sent to us as the
|
||
<typeStatus id="54D6D6AED512FFB415E4FB08FD23938E" box="[652,750,1160,1184]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6D138FD512FFB4147CFB09FC57938E" box="[788,922,1160,1184]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED512FFB4147CFB09FC57938E" box="[788,922,1160,1184]" italics="true" pageId="21" pageNumber="22">E. turgidula</emphasis>
|
||
</taxonomicName>
|
||
is in poor condition compared to the drawing provided by Ehlers, and is no longer complete and with the body severely damaged in places, we have no doubt we are dealing with the same specimen (see
|
||
<figureCitation id="13567489D512FFB41509FB50FD6F93C6" box="[609,674,1232,1256]" captionStart="FIGURE 9" captionStartId="20.[151,250,1825,1848]" captionTargetBox="[230,1353,198,1791]" captionTargetId="figure@20.[225,1359,193,1797]" captionTargetPageId="20" captionText="FIGURE 9. Eupolymnia turgidula, holotype (MCZ 846). A: Ehlers’ drawing of the holotype, entire worm, ventral view (Ehlers 1887: Taf. 52, Fig. 1); B: entire worm, dorsal view; C: entire worm, left ventrolateral view; D: anterior end, ventral view; E – F: close up views of anterior end, left ventrolateral view; G: anterior end, left lateral view; H: anterior end, dorsal view; I: transition between regions with biramous parapodia and neuropodia only, right lateral view; J: close up view of anterior segments of region with neuropodia only, right lateral view; K: piece of tube. Numbers refer to segments, uspecified arrows point to nephridial papillae; * = lower lip; ** = prostomium, basal part; P = prostomium; P (dp) = prostomium, distal part; ul = upper lip. Scale bars: B – C = 1.5 mm; D – I = 1 mm; J = 0.6 mm; K = 2 mm." httpUri="https://zenodo.org/record/278423/files/figure.png" pageId="21" pageNumber="22">Fig. 9</figureCitation>
|
||
A–G).
|
||
</paragraph>
|
||
<paragraph id="8BD2680CD512FFB417AFFB74FC56927A" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
|
||
The specimen
|
||
<bibRefCitation id="EFFC15FDD512FFB41606FB74FDA09222" author="Londono-Mesa" box="[366,621,1268,1292]" pageId="21" pageNumber="22" refString="Londono-Mesa, M. H. (2009) Terebellidae (Polychaeta: Terebellida) from the Grand Caribbean Region. Zootaxa, 2320, 1 - 93." type="journal article" year="2009">Londoño-Mesa (2009)</bibRefCitation>
|
||
examined and illustrated definitely belongs to
|
||
<taxonomicName id="4C6D138FD512FFB413E3FB74FB3D9222" box="[1163,1264,1268,1292]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="genus">
|
||
<emphasis id="B919B41ED512FFB413E3FB74FB3D9222" box="[1163,1264,1268,1292]" italics="true" pageId="21" pageNumber="22">Terebella</emphasis>
|
||
</taxonomicName>
|
||
, as it does not present lobes on anterior segments and has a large number of pairs of notopodia (27 pairs, against 17 as in most, if not all, species of
|
||
<taxonomicName id="4C6D138FD512FFB4160AFABCFE24927A" box="[354,489,1340,1364]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="genus">
|
||
<emphasis id="B919B41ED512FFB4160AFABCFE24927A" box="[354,489,1340,1364]" italics="true" pageId="21" pageNumber="22">Eupolymnia</emphasis>
|
||
</taxonomicName>
|
||
) bearing distally serrated notochaetae.
|
||
</paragraph>
|
||
<paragraph id="8BD2680CD512FFB417AFFAE0FAEB9192" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
|
||
The question of the validity of the taxon
|
||
<taxonomicName id="4C6D138FD512FFB415CCFAE1FCE19256" box="[676,812,1376,1400]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED512FFB415CCFAE1FCE19256" box="[676,812,1376,1400]" italics="true" pageId="21" pageNumber="22">E. turgidula</emphasis>
|
||
</taxonomicName>
|
||
is complicated.
|
||
<bibRefCitation id="EFFC15FDD512FFB41484FAE0FB629256" author="Schmarda" box="[1004,1199,1376,1400]" pageId="21" pageNumber="22" refString="Schmarda, L. K. (1861) Neue wirbellose Thiere beobachtet und gesammelt auf einer Reise um die Erdr 1853 bis 1857. 1. Turbellarien, Rotatorien und Anneliden, pt. 2. Wilhelm Engelmann, Leipzig, 164 pp." type="book" year="1861">Schmarda (1861)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C6D138FD512FFB41246FAE1FF1092B2" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="crassicornis">
|
||
<emphasis id="B919B41ED512FFB41246FAE1FF1092B2" italics="true" pageId="21" pageNumber="22">E. crassicornis</emphasis>
|
||
</taxonomicName>
|
||
as having 25 pairs of notopodia and uncini with a single row of three secondary teeth.
|
||
<bibRefCitation id="EFFC15FDD512FFB413CCFA04FA9892B2" author="Augener" box="[1188,1365,1412,1436]" pageId="21" pageNumber="22" refString="Augener, H. (1925) Uber westindische und einige andere Polychaeten-Typen von Grube (Oersted), Kroyer, Morch und Schmarda. Publikationer fra Universitetets Zoologiske Museum Kobenhavn, 39, 1 - 47." type="journal article" year="1925">Augener (1925)</bibRefCitation>
|
||
based the synonymization of
|
||
<taxonomicName id="4C6D138FD512FFB416FEFA29FDD192EE" box="[406,540,1448,1472]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED512FFB416FEFA29FDD192EE" box="[406,540,1448,1472]" italics="true" pageId="21" pageNumber="22">E. turgidula</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C6D138FD512FFB41533FA29FCCE92EE" box="[603,771,1449,1472]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="crassicornis">
|
||
<emphasis id="B919B41ED512FFB41533FA29FCCE92EE" box="[603,771,1449,1472]" italics="true" pageId="21" pageNumber="22">E. crassicornis</emphasis>
|
||
</taxonomicName>
|
||
on the similarity between the uncini of both species and his belief that
|
||
<taxonomicName id="4C6D138FD512FFB41678FA4DFE7592CA" box="[272,440,1485,1508]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="crassicornis">
|
||
<emphasis id="B919B41ED512FFB41678FA4DFE7592CA" box="[272,440,1485,1508]" italics="true" pageId="21" pageNumber="22">E. crassicornis</emphasis>
|
||
</taxonomicName>
|
||
should have only 17 pairs of notopodia despite Schmarda’s statement to the contrary. In addition to the problem concerning the numbers of pairs of notopodia present in both species, the
|
||
<typeStatus id="54D6D6AED512FFB41391FA70FA969126" box="[1273,1371,1520,1544]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6D138FD512FFB412ECFA71FF339102" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED512FFB412ECFA71FF339102" italics="true" pageId="21" pageNumber="22">E. turgidula</emphasis>
|
||
</taxonomicName>
|
||
has uncini with two rows of secondary teeth and, although the second row is difficult to discern, depending on the position of the uncini on the slide, the first row clearly has only two teeth (see
|
||
<figureCitation id="13567489D512FFB413E7F9B8FB12917E" box="[1167,1247,1592,1616]" captionStart="FIGURE 10" captionStartId="22.[151,250,1951,1974]" captionTargetBox="[157,1431,307,1923]" captionTargetId="figure@22.[151,1436,301,1930]" captionTargetPageId="22" captionText="FIGURE 10. Eupolymnia turgidula, holotype (MCZ 846). A: notochaetae, segment 18; B: tips of some notochaetae from posterior tier, segment 18; C: uncini, segment 7; D – E: uncini, segment 12; F: uncini, segment 23. Scale bars: A, E = 80 µm; B = 40 µm; C = 15 µm; D = 10 µm; F = 30 µm." httpUri="https://zenodo.org/record/278424/files/figure.png" pageId="21" pageNumber="22">Fig. 10</figureCitation>
|
||
C–F). Moreover, the general morphology of the uncini of these two taxa, which Augener considered very similar to each other, is considerably different in our opinion, especially considering the shape of the dorsal button, prow, base, and heel (compare Schmarda’s illustration provided by
|
||
<bibRefCitation id="EFFC15FDD512FFB415F7F924FBE59192" author="Londono-Mesa" box="[671,1064,1700,1724]" pageId="21" pageNumber="22" refString="Londono-Mesa, M. H. (2009) Terebellidae (Polychaeta: Terebellida) from the Grand Caribbean Region. Zootaxa, 2320, 1 - 93." type="journal article" year="2009">
|
||
Londoño-Mesa [2009: 30,
|
||
<figureCitation id="13567489D512FFB414A3F924FBC19192" box="[971,1036,1700,1724]" captionStart="FIGURE 7" captionStartId="16.[151,250,1953,1976]" captionTargetBox="[157,1430,379,1926]" captionTargetId="figure@16.[151,1436,373,1932]" captionTargetPageId="16" captionText="FIGURE 7. Pista sombreriana, holotype (BMNH 1885.12.1.344). A: entire worm, dorsal view; B; entire worm, ventral view; C: notochaetae from anterior tier, segment 14; D: uncini, segment 6; E: uncini from posterior region with biramous parapodia; F: uncini from anterior region with neuropodia only. Scale bars: A – B = 1.5 mm; C = 50 µm; D – E = 20 µm; F = 15 µm." httpUri="https://zenodo.org/record/278421/files/figure.png" pageId="21" pageNumber="22">Fig. 7</figureCitation>
|
||
C]
|
||
</bibRefCitation>
|
||
with our
|
||
<figureCitation id="13567489D512FFB413FCF924FB299192" box="[1172,1252,1700,1724]" captionStart="FIGURE 10" captionStartId="22.[151,250,1951,1974]" captionTargetBox="[157,1431,307,1923]" captionTargetId="figure@22.[151,1436,301,1930]" captionTargetPageId="22" captionText="FIGURE 10. Eupolymnia turgidula, holotype (MCZ 846). A: notochaetae, segment 18; B: tips of some notochaetae from posterior tier, segment 18; C: uncini, segment 7; D – E: uncini, segment 12; F: uncini, segment 23. Scale bars: A, E = 80 µm; B = 40 µm; C = 15 µm; D = 10 µm; F = 30 µm." httpUri="https://zenodo.org/record/278424/files/figure.png" pageId="21" pageNumber="22">Fig. 10</figureCitation>
|
||
C–E).
|
||
</paragraph>
|
||
<paragraph id="8BD2680CD512FFB417AFF948FB019006" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
|
||
<bibRefCitation id="EFFC15FDD512FFB417AFF948FE0A91CE" author="Londono-Mesa" box="[199,455,1736,1760]" pageId="21" pageNumber="22" refString="Londono-Mesa, M. H. (2009) Terebellidae (Polychaeta: Terebellida) from the Grand Caribbean Region. Zootaxa, 2320, 1 - 93." type="journal article" year="2009">Londoño-Mesa (2009)</bibRefCitation>
|
||
listed three species of
|
||
<taxonomicName id="4C6D138FD512FFB415B3F948FCAF91CE" box="[731,866,1736,1760]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="genus">
|
||
<emphasis id="B919B41ED512FFB415B3F948FCAF91CE" box="[731,866,1736,1760]" italics="true" pageId="21" pageNumber="22">Eupolymnia</emphasis>
|
||
</taxonomicName>
|
||
for the Caribbean:
|
||
<taxonomicName id="4C6D138FD512FFB41323F949FB3A91CE" box="[1099,1271,1737,1760]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="crassicornis">
|
||
<emphasis id="B919B41ED512FFB41323F949FB3A91CE" box="[1099,1271,1737,1760]" italics="true" pageId="21" pageNumber="22">E. crassicornis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6D138FD512FFB41261F949FE80902A" authority="Webster, 1884" authorityName="Webster" authorityYear="1884" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="magnifica">
|
||
<emphasis id="B919B41ED512FFB41261F949FA5191CE" box="[1289,1436,1736,1760]" italics="true" pageId="21" pageNumber="22">E. magnifica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFC15FDD512FFB417F6F96CFE88902A" author="Webster" box="[158,325,1772,1796]" pageId="21" pageNumber="22" refString="Webster, H. E. (1884) Annelida from Bermuda collected by G. Brown Goode. Bulletin of the United States National Museum, 25, 307 - 327." type="journal article" year="1884">Webster, 1884</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6D138FD512FFB416E1F96DFD38902A" authority="Londono-Mesa, 2009" authorityName="Londono-Mesa" authorityYear="2009" box="[393,757,1772,1796]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="rullieri">
|
||
<emphasis id="B919B41ED512FFB416E1F96DFE35902A" box="[393,504,1772,1796]" italics="true" pageId="21" pageNumber="22">E. rullieri</emphasis>
|
||
<bibRefCitation id="EFFC15FDD512FFB41568F96CFD38902A" author="Londono-Mesa" box="[512,757,1772,1796]" pageId="21" pageNumber="22" refString="Londono-Mesa, M. H. (2009) Terebellidae (Polychaeta: Terebellida) from the Grand Caribbean Region. Zootaxa, 2320, 1 - 93." type="journal article" year="2009">Londoño-Mesa, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C6D138FD512FFB41469F96CFC12902A" box="[769,991,1772,1796]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="rullieri">
|
||
<emphasis id="B919B41ED512FFB41469F96CFC12902A" box="[769,991,1772,1796]" italics="true" pageId="21" pageNumber="22">Eupolymnia rullieri</emphasis>
|
||
</taxonomicName>
|
||
is clearly a separate taxon from
|
||
<taxonomicName id="4C6D138FD512FFB4123BF96DFF129006" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED512FFB4123BF96DFF129006" italics="true" pageId="21" pageNumber="22">E. turgidula</emphasis>
|
||
</taxonomicName>
|
||
, differing mostly in the position of the genital papillae and in the uncinial dental formula.
|
||
</paragraph>
|
||
<paragraph id="8BD2680CD512FFB717AFF8B5FE2397FA" blockId="21.[151,1437,151,2013]" lastBlockId="22.[151,1436,151,212]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
However, we could not detect any consistent difference between the redescription of
|
||
<taxonomicName id="4C6D138FD512FFB41317F8B5FAC39062" box="[1151,1294,1844,1868]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="magnifica">
|
||
<emphasis id="B919B41ED512FFB41317F8B5FAC39062" box="[1151,1294,1844,1868]" italics="true" pageId="21" pageNumber="22">E. magnifica</emphasis>
|
||
</taxonomicName>
|
||
provided by
|
||
<bibRefCitation id="EFFC15FDD512FFB417FFF8D8FE59905E" author="Londono-Mesa" box="[151,404,1880,1904]" pageId="21" pageNumber="22" refString="Londono-Mesa, M. H. (2009) Terebellidae (Polychaeta: Terebellida) from the Grand Caribbean Region. Zootaxa, 2320, 1 - 93." type="journal article" year="2009">Londoño-Mesa (2009)</bibRefCitation>
|
||
and our redescription above of the
|
||
<typeStatus id="54D6D6AED512FFB4144DF8D8FC4A905E" box="[805,903,1880,1904]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6D138FD512FFB414C4F8D9FBFE905E" box="[940,1075,1880,1904]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED512FFB414C4F8D9FBFE905E" box="[940,1075,1880,1904]" italics="true" pageId="21" pageNumber="22">E. turgidula</emphasis>
|
||
</taxonomicName>
|
||
, except for a great difference in body size.
|
||
<bibRefCitation id="EFFC15FDD512FFB41678F8FCFDC090BA" author="Londono-Mesa" box="[272,525,1916,1940]" pageId="21" pageNumber="22" refString="Londono-Mesa, M. H. (2009) Terebellidae (Polychaeta: Terebellida) from the Grand Caribbean Region. Zootaxa, 2320, 1 - 93." type="journal article" year="2009">Londoño-Mesa (2009)</bibRefCitation>
|
||
said the best
|
||
<typeStatus id="54D6D6AED512FFB415CDF8FDFD3190BB" box="[677,764,1917,1941]" pageId="21" pageNumber="22" type="syntype">syntype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6D138FD512FFB4144AF8FDFC7F90BA" box="[802,946,1916,1940]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="magnifica">
|
||
<emphasis id="B919B41ED512FFB4144AF8FDFC7F90BA" box="[802,946,1916,1940]" italics="true" pageId="21" pageNumber="22">E. magnifica</emphasis>
|
||
</taxonomicName>
|
||
had nephridial papillae on segments 3–5 as also occurs in the
|
||
<typeStatus id="54D6D6AED512FFB4160BF820FE089096" box="[355,453,1952,1976]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6D138FD512FFB41683F821FDBC9096" box="[491,625,1952,1976]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="21" pageNumber="22" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED512FFB41683F821FDBC9096" box="[491,625,1952,1976]" italics="true" pageId="21" pageNumber="22">E. turgidula</emphasis>
|
||
</taxonomicName>
|
||
. Other specimens examined by
|
||
<bibRefCitation id="EFFC15FDD512FFB414B3F820FB159096" author="Londono-Mesa" box="[987,1240,1952,1976]" pageId="21" pageNumber="22" refString="Londono-Mesa, M. H. (2009) Terebellidae (Polychaeta: Terebellida) from the Grand Caribbean Region. Zootaxa, 2320, 1 - 93." type="journal article" year="2009">Londoño-Mesa (2009)</bibRefCitation>
|
||
also had papillae on segments 6–10 but it was not specified as to whether these were in addition to the nephridial papillae on segments 3–5 or if they were on
|
||
<typeStatus id="54D6D6AED511FFB71681FF18FDD4979E" box="[489,537,152,176]" pageId="22" pageNumber="23">type</typeStatus>
|
||
or non-type material. These specimens need to be re-examined to confirm that they belong to the same taxon.
|
||
</paragraph>
|
||
<caption id="DF123884D511FFB717FFF81FFDFD90DC" httpUri="https://zenodo.org/record/278424/files/figure.png" pageId="22" pageNumber="23" targetBox="[157,1431,307,1923]" targetPageId="22">
|
||
<paragraph id="8BD2680CD511FFB717FFF81FFDFD90DC" blockId="22.[151,1437,1951,2034]" pageId="22" pageNumber="23">
|
||
<emphasis id="B919B41ED511FFB717FFF81FFDDF909B" bold="true" box="[151,530,1951,1974]" pageId="22" pageNumber="23">
|
||
FIGURE 10.
|
||
<taxonomicName id="4C6D138FD511FFB7164DF81FFDC1909B" box="[293,524,1951,1973]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="22" pageNumber="23" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED511FFB7164DF81FFDC1909B" bold="true" box="[293,524,1951,1973]" italics="true" pageId="22" pageNumber="23">Eupolymnia turgidula</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holotype (MCZ 846). A: notochaetae, segment 18; B: tips of some notochaetae from posterior tier, segment 18; C: uncini, segment 7; D–E: uncini, segment 12; F: uncini, segment 23. Scale bars: A, E = 80 µm; B = 40 µm; C = 15 µm; D = 10 µm; F = 30 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD2680CD510FFB617AFFF18FDC696A9" blockId="23.[151,1436,151,391]" pageId="23" pageNumber="24">
|
||
We consider that
|
||
<taxonomicName id="4C6D138FD510FFB616E0FF19FDC3979E" box="[392,526,152,176]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED510FFB616E0FF19FDC3979E" box="[392,526,152,176]" italics="true" pageId="23" pageNumber="24">E. turgidula</emphasis>
|
||
</taxonomicName>
|
||
is very likely a junior synonym of
|
||
<taxonomicName id="4C6D138FD510FFB614FEFF19FBE9979E" box="[918,1060,152,176]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="magnifica">
|
||
<emphasis id="B919B41ED510FFB614FEFF19FBE9979E" box="[918,1060,152,176]" italics="true" pageId="23" pageNumber="24">E. magnifica</emphasis>
|
||
</taxonomicName>
|
||
, instead of
|
||
<taxonomicName id="4C6D138FD510FFB613CFFF19FA82979E" box="[1191,1359,153,176]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="crassicornis">
|
||
<emphasis id="B919B41ED510FFB613CFFF19FA82979E" box="[1191,1359,153,176]" italics="true" pageId="23" pageNumber="24">E. crassicornis</emphasis>
|
||
</taxonomicName>
|
||
. However, we prefer to keep
|
||
<taxonomicName id="4C6D138FD510FFB616CCFF3DFDE697FA" box="[420,555,188,212]" class="Polychaeta" family="Terebellidae" genus="Eupolymnia" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="turgidula">
|
||
<emphasis id="B919B41ED510FFB616CCFF3DFDE697FA" box="[420,555,188,212]" italics="true" pageId="23" pageNumber="24">E. turgidula</emphasis>
|
||
</taxonomicName>
|
||
as a valid taxon until more detailed studies on these taxa are carried out. The
|
||
<typeStatus id="54D6D6AED510FFB617FFFF5FFF3497D9" box="[151,249,223,247]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6D138FD510FFB6164EFF61FE1F97D6" box="[294,466,225,248]" class="Polychaeta" family="Terebellidae" genus="Terebella" kingdom="Animalia" order="Terebellida" pageId="23" pageNumber="24" phylum="Annelida" rank="species" species="crassicornis">
|
||
<emphasis id="B919B41ED510FFB6164EFF61FE1F97D6" box="[294,466,225,248]" italics="true" pageId="23" pageNumber="24">E. crassicornis</emphasis>
|
||
</taxonomicName>
|
||
, described as dried by
|
||
<bibRefCitation id="EFFC15FDD510FFB6158FFF60FC5297D9" author="Augener" box="[743,927,223,248]" pageId="23" pageNumber="24" refString="Augener, H. (1925) Uber westindische und einige andere Polychaeten-Typen von Grube (Oersted), Kroyer, Morch und Schmarda. Publikationer fra Universitetets Zoologiske Museum Kobenhavn, 39, 1 - 47." type="journal article" year="1925">Augener (1925)</bibRefCitation>
|
||
, now appears to be missing (pers. com., S. Szeiler, Naturhistorisches Museum, Wien) and a
|
||
<typeStatus id="54D6D6AED510FFB615B8FE85FCE59633" box="[720,808,261,285]" pageId="23" pageNumber="24" type="neotype">neotype</typeStatus>
|
||
would have to be erected to stabilize the species and resolve the question of synonymy. However, considering all the problems to characterize this taxon discussed above, it is not possible to erect a
|
||
<typeStatus id="54D6D6AED510FFB61566FECDFDAA964B" box="[526,615,333,357]" pageId="23" pageNumber="24" type="neotype">neotype</typeStatus>
|
||
at this stage and therefore this taxon should be considered as undeterminable, if the
|
||
<typeStatus id="54D6D6AED510FFB6167DFEEFFEBA96A9" box="[277,375,367,391]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
is not found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |