766 lines
95 KiB
XML
766 lines
95 KiB
XML
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<mods:title id="1FD1FCC7A028FA5C17A6275A264F229A">Three new scorpion species from the Dominican Republic, Greater Antilles (Scorpiones: Buthidae, Scorpionidae)</mods:title>
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<mods:namePart id="552DF608B8961E03FA7EED60E417BA24">Kovařík, František</mods:namePart>
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<mods:namePart id="1ADE2758461E8760675BE4CD6E011A0D">Teruel, Rolando</mods:namePart>
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<mods:title id="DBC2B9CCBCF44B70ADAC5675C4125909">Euscorpius</mods:title>
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<mods:part id="82552CD7C13E85D1B27FF2D96EDA14A2">
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<mods:date id="554D428BA4532159B7F6C435DB007BB0">2014</mods:date>
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<taxonomicName id="4C6DCCBCFFF66579FCD4F97EFA36F978" ID-CoL="438TC" authority="Kovarik et Teruel" authorityName="Kovarik et Teruel" authorityYear="2014" box="[836,1428,1734,1758]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov." subGenus="Parvabsonus">
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<emphasis id="B9196B2DFFF66579FCD4F97EFC60F978" bold="true" box="[836,962,1734,1758]" italics="true" pageId="2" pageNumber="1">Microtityus</emphasis>
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(
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<emphasis id="B9196B2DFFF66579FC41F97EFBFDF978" bold="true" box="[977,1119,1734,1758]" italics="true" pageId="2" pageNumber="1">Parvabsonus</emphasis>
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)
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<emphasis id="B9196B2DFFF66579FBFFF97EFB6DF978" bold="true" box="[1135,1231,1734,1758]" italics="true" pageId="2" pageNumber="1">minimus</emphasis>
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</emphasis>
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Kovařík et Teruel
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</taxonomicName>
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,
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</paragraph>
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<emphasis id="B9196B2DFFF66579FBC1F95DFB2FF95B" bold="true" box="[1105,1165,1765,1789]" pageId="2" pageNumber="1">sp. n.</emphasis>
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</heading>
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(
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<figureCitation id="1356ABBAFFF66579FC68F8BBFB2BF8BD" box="[1016,1161,1795,1819]" captionStart-0="Figures 1–19" captionStart-1="Figures 20–23" captionStart-2="Figures 24–25" captionStart-3="Figure 26" captionStartId-0="3.[192,278,1736,1760]" captionStartId-1="5.[192,278,1840,1864]" captionStartId-2="6.[192,278,1874,1898]" captionStartId-3="8.[192,267,790,814]" captionTargetBox-0="[280,1357,170,1704]" captionTargetBox-1="[191,1441,215,1806]" captionTargetBox-2="[199,1405,191,1842]" captionTargetBox-3="[191,1440,191,756]" captionTargetId-0="figure-140@3.[276,1357,167,1704]" captionTargetId-1="figure-41@5.[192,1440,216,1805]" captionTargetId-2="figure-31@6.[200,1404,192,1841]" captionTargetId-3="figure-631@8.[192,1439,192,755]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="Figures 1–19: Microtityus minimus sp. n. Figures 1–2, 5, 10, 16–17. Dorsal (1) and ventral (2) views, telson (5), metasoma and telson lateral (10), carapace with chelicerae and tergites I–IV (16) and pectinal area with sternites III–V (17), ♂ paratype (FKCP). Figures 3–4, 6–9, 11–15, 18–19. Dorsal (3) and ventral (4) views, right movable finger internal (6), metasoma and telson ventral (7), dorsal (8), and lateral (9), right pedipalp chela dorsal (11) and external (12), pedipalp patella dorsal (13) and external (14) and pedipalp femur dorsal (15), carapace with chelicerae and tergites I–IV (18) and pectinal area with sternites III– V (19), ♀ paratype (FKCP)." captionText-1="Figures 20–23: Microtityus minimus sp. n. Figure 20. ♀ paratype (FKCP) with newborn. Figures 21–22. ♀ paratype (FKCP). Figure 23. ♂ holotype at the type locality." captionText-2="Figures 24–25: Microtityus minimus sp. n., type locality (subcoastal desert scrub on gypsum-sandy soil)." captionText-3="Figure 26: Geographical distribution of Microtityus minimus sp. n. (black square) and its closest relatives M. paucidentatus (grey square) and M. reini (white squares)." pageId="2" pageNumber="1">Figures 1–26</figureCitation>
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;
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<tableCitation id="C6EF8284FFF66579FB04F8BBFB46F8BD" box="[1172,1252,1795,1819]" captionStart="Table 1" captionStartId="7.[192,257,853,877]" captionText="Table 1: Measurements of four types of Microtityus minimus sp. n. (RTOC). Abbreviations: length (L), width (W), posterior width (Wp), depth (H), left (L), right (R)." pageId="2" pageNumber="1">Table 1</tableCitation>
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)
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<emphasis id="B9196B2DFFF66579FCAFF89AFA02F89C" bold="true" box="[831,1440,1826,1850]" pageId="2" pageNumber="1">http://zoobank.org/urn:lsid:zoobank.org:act:90ABA6</emphasis>
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</paragraph>
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</paragraph>
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<emphasis id="B9196B2DFFF76578FF50F970FCD8F979" bold="true" box="[192,890,1736,1760]" pageId="3" pageNumber="2">
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Figures 1–19:
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<taxonomicName id="4C6DCCBCFFF76578FEF4F972FD91F979" authority="Kovařík & Teruel, 2014" authorityName="Kovařík & Teruel" authorityYear="2014" box="[356,563,1738,1759]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
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<emphasis id="B9196B2DFFF76578FEF4F972FD91F979" bold="true" box="[356,563,1738,1759]" italics="true" pageId="3" pageNumber="2">Microtityus minimus</emphasis>
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Figures 1–2, 5, 10, 16–17
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</emphasis>
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. Dorsal (1) and ventral (2) views, telson (5), metasoma and telson lateral (10), carapace with chelicerae and tergites I–IV (16) and pectinal area with sternites III–V (17), ♂ paratype (FKCP).
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<emphasis id="B9196B2DFFF76578FE8CF8B9FDFDF8B0" bold="true" box="[284,607,1793,1814]" pageId="3" pageNumber="2">Figures 3–4, 6–9, 11–15, 18–19</emphasis>
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. Dorsal (3) and ventral (4) views, right movable finger internal (6), metasoma and telson ventral (7), dorsal (8), and lateral (9), right pedipalp chela dorsal (11) and external (12), pedipalp patella dorsal (13) and external (14) and pedipalp femur dorsal (15), carapace with chelicerae and tergites I–IV (18) and pectinal area with sternites III– V (19), ♀ paratype (FKCP).
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</paragraph>
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</caption>
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et
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<figureCitation id="1356ABBAFFF0657FFDDFFE03FD3CFE75" box="[591,670,443,467]" captionStart="Figures 20–23" captionStartId="5.[192,278,1840,1864]" captionTargetBox="[191,1441,215,1806]" captionTargetId="figure-41@5.[192,1440,216,1805]" captionTargetPageId="5" captionText="Figures 20–23: Microtityus minimus sp. n. Figure 20. ♀ paratype (FKCP) with newborn. Figures 21–22. ♀ paratype (FKCP). Figure 23. ♂ holotype at the type locality." pageId="4" pageNumber="3">Fig. 23</figureCitation>
|
||
,
|
||
<collectionCode id="ED7C2FFAFFF0657FFD39FE03FD50FE75" box="[681,754,443,467]" pageId="4" pageNumber="3">RTOC</collectionCode>
|
||
)
|
||
<specimenCount id="9D6B7CB6FFF0657FFC90FE03FC83FE75" box="[768,801,443,467]" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
|
||
<typeStatus id="54D6099DFFF0657FFF50FE62FEB9FE54" box="[192,283,474,498]" pageId="4" pageNumber="3" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="1356ABBAFFF0657FFEBEFE61FD94FE57" box="[302,566,473,497]" captionStart="Figures 1–19" captionStartId="3.[192,278,1736,1760]" captionTargetBox="[280,1357,170,1704]" captionTargetId="figure-140@3.[276,1357,167,1704]" captionTargetPageId="3" captionText="Figures 1–19: Microtityus minimus sp. n. Figures 1–2, 5, 10, 16–17. Dorsal (1) and ventral (2) views, telson (5), metasoma and telson lateral (10), carapace with chelicerae and tergites I–IV (16) and pectinal area with sternites III–V (17), ♂ paratype (FKCP). Figures 3–4, 6–9, 11–15, 18–19. Dorsal (3) and ventral (4) views, right movable finger internal (6), metasoma and telson ventral (7), dorsal (8), and lateral (9), right pedipalp chela dorsal (11) and external (12), pedipalp patella dorsal (13) and external (14) and pedipalp femur dorsal (15), carapace with chelicerae and tergites I–IV (18) and pectinal area with sternites III– V (19), ♀ paratype (FKCP)." pageId="4" pageNumber="3">Figs. 1–2, 5, 10, 16–17</figureCitation>
|
||
,
|
||
<collectionCode id="ED7C2FFAFFF0657FFDD5FE61FD2BFE57" box="[581,649,473,497]" pageId="4" pageNumber="3">FKCP</collectionCode>
|
||
)
|
||
<specimenCount id="9D6B7CB6FFF0657FFD0BFE61FD1EFE57" box="[667,700,473,497]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
|
||
<typeStatus id="54D6099DFFF0657FFD56FE62FC83FE54" box="[710,801,474,498]" pageId="4" pageNumber="3" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="ED7C2FFAFFF0657FFF58FE40FEB7FDB6" box="[200,277,504,528]" pageId="4" pageNumber="3">RTOC</collectionCode>
|
||
); same data except
|
||
<date id="FFD391FFFFF0657FFE75FE40FDA0FDB6" box="[485,514,504,528]" pageId="4" pageNumber="3" value="2014-03-15">15.</date>
|
||
</materialsCitation>
|
||
<collectionCode id="ED7C2FFAFFF0657FFD91FE40FDBCFDB6" box="[513,542,504,528]" pageId="4" pageNumber="3">III</collectionCode>
|
||
.2014,
|
||
<materialsCitation id="3B05BD62FFF0657FFDFBFE40FEADFDE8" collectionCode="RTOC, FKCP" country="Dominican Republic" pageId="4" pageNumber="3" specimenCount="15" specimenCount-female="12" specimenCount-male="3" stateProvince="Azua" typeStatus="paratype">
|
||
<specimenCount id="9D6B7CB6FFF0657FFDFBFE40FD2FFDB6" box="[619,653,504,528]" pageId="4" pageNumber="3" type="male">3♂</specimenCount>
|
||
<specimenCount id="9D6B7CB6FFF0657FFD03FE41FD17FDB6" box="[659,693,505,528]" pageId="4" pageNumber="3" type="female">7♀</specimenCount>
|
||
<typeStatus id="54D6099DFFF0657FFD2CFE41FC80FDB7" box="[700,802,505,529]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<collectionCode id="ED7C2FFAFFF0657FFF57FDAFFEB3FD89" box="[199,273,535,559]" pageId="4" pageNumber="3">RTOC</collectionCode>
|
||
)
|
||
<specimenCount id="9D6B7CB6FFF0657FFEB5FDAFFEE4FD89" box="[293,326,535,559]" pageId="4" pageNumber="3" type="female">5♀</specimenCount>
|
||
<typeStatus id="54D6099DFFF0657FFEC2FDA0FE1AFD96" box="[338,440,536,560]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<figureCitation id="1356ABBAFFF0657FFE5CFDAFFCBCFD89" box="[460,798,535,559]" captionStart-0="Figures 1–19" captionStart-1="Figures 20–23" captionStartId-0="3.[192,278,1736,1760]" captionStartId-1="5.[192,278,1840,1864]" captionTargetBox-0="[280,1357,170,1704]" captionTargetBox-1="[191,1441,215,1806]" captionTargetId-0="figure-140@3.[276,1357,167,1704]" captionTargetId-1="figure-41@5.[192,1440,216,1805]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Figures 1–19: Microtityus minimus sp. n. Figures 1–2, 5, 10, 16–17. Dorsal (1) and ventral (2) views, telson (5), metasoma and telson lateral (10), carapace with chelicerae and tergites I–IV (16) and pectinal area with sternites III–V (17), ♂ paratype (FKCP). Figures 3–4, 6–9, 11–15, 18–19. Dorsal (3) and ventral (4) views, right movable finger internal (6), metasoma and telson ventral (7), dorsal (8), and lateral (9), right pedipalp chela dorsal (11) and external (12), pedipalp patella dorsal (13) and external (14) and pedipalp femur dorsal (15), carapace with chelicerae and tergites I–IV (18) and pectinal area with sternites III– V (19), ♀ paratype (FKCP)." captionText-1="Figures 20–23: Microtityus minimus sp. n. Figure 20. ♀ paratype (FKCP) with newborn. Figures 21–22. ♀ paratype (FKCP). Figure 23. ♂ holotype at the type locality." pageId="4" pageNumber="3">Figs. 3–4, 6–9, 11–15, 18–22</figureCitation>
|
||
,
|
||
<collectionCode id="ED7C2FFAFFF0657FFF50FD8EFEA5FDE8" box="[192,263,566,590]" pageId="4" pageNumber="3">FKCP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377E4B4FFF0657FFF50FDCCFE2FFD6E" pageId="4" pageNumber="3" type="etymology">
|
||
<paragraph id="8BD2B73FFFF0657FFF50FDCCFE2FFD6E" blockId="4.[192,797,627,712]" pageId="4" pageNumber="3">ETYMOLOGY. The selected name is a Latin adjective that alludes to the extremely small adult size of this species, see remarks below.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377E4B4FFF0657FFF50FD56FECBF9D9" pageId="4" pageNumber="3" type="diagnosis">
|
||
<paragraph id="8BD2B73FFFF0657FFF50FD56FECBF9D9" blockId="4.[192,802,750,1663]" pageId="4" pageNumber="3">
|
||
DIAGNOSIS. Adult size very small (males
|
||
<quantity id="4C951ADAFFF0657FFD3DFD56FCBCFCA3" box="[685,798,750,774]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="4" pageNumber="3" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
|
||
, females
|
||
<quantity id="4C951ADAFFF0657FFEB4FCB4FE01FC82" box="[292,419,780,804]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="4" pageNumber="3" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
|
||
) for the genus. Coloration pale yellow to yellowish brown, moderately spotted with blackish brown all over the body and appendages, tergites without clearly defined longitudinal dark bands, pedipalp hand immaculate (males) to vestigially spotted (females), fingers blackish with yellowish tips, metasomal segment V and telson only slightly more reddish; in general, coloration in males is paler and less densely spotted. Pedipalps neobothriotaxic (femoral trichobothrium
|
||
<emphasis id="B9196B2DFFF0657FFF6FFB98FEB4FB9E" box="[255,278,1056,1080]" italics="true" pageId="4" pageNumber="3">
|
||
d
|
||
<subScript id="17E9B57AFFF0657FFE9DFB92FEB4FB9E" attach="left" box="[269,278,1066,1080]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
|
||
</emphasis>
|
||
absent), fixed and movable fingers with 8–9 principal rows of denticles. Sternites IV–V with two carinae, VI–VII with four; posterior margin of III widely convex, with the smooth patch large, subtriangular, flat, and not protruding beyond margin; V with the smooth patch moderately large, cordiform and bulky (males) or very small, fusiform, and flat (females). Metasoma moderately short, with 10-10-8-8-5 complete carinae, most of which are sharply serrate to denticulate, segments II– IV dorsolateral carinae with 2–3 terminal denticles unevenly enlarged, all intercarinal spaces finely and densely granulose; segment V prismatic in cross-section, with ordinary carination and tegumentary sculpture. Telson short oval, vesicle minutely (males) to coarsely (females) granulose, subaculear tubercle moderate and irregular, aculeus as long as vesicle (males) or slightly shorter (females). Pectines with 10–11 teeth (mode 10) in males, 8–9 (mode 9) in females; basal plate in females slightly longer than wide, spatulate, and with posterior margin straight.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377E4B4FFF06573FF50F91DFD42F884" lastPageId="8" lastPageNumber="7" pageId="4" pageNumber="3" type="description">
|
||
<paragraph id="8BD2B73FFFF0657FFF50F91DFC28FBD0" blockId="4.[192,802,1700,1908]" lastBlockId="4.[831,1441,198,1908]" pageId="4" pageNumber="3">
|
||
DESCRIPTION (adult male
|
||
<typeStatus id="54D6099DFFF0657FFE66F91CFDF1F91A" box="[502,595,1700,1724]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
unless otherwise noted). Coloration (
|
||
<figureCitation id="1356ABBAFFF0657FFE36F97BFE5CF97D" box="[422,510,1731,1755]" captionStart="Figures 20–23" captionStartId="5.[192,278,1840,1864]" captionTargetBox="[191,1441,215,1806]" captionTargetId="figure-41@5.[192,1440,216,1805]" captionTargetPageId="5" captionText="Figures 20–23: Microtityus minimus sp. n. Figure 20. ♀ paratype (FKCP) with newborn. Figures 21–22. ♀ paratype (FKCP). Figure 23. ♂ holotype at the type locality." pageId="4" pageNumber="3">Fig. 23</figureCitation>
|
||
) base pale yellowish to yellow, densely spotted with blackish brown all over the body and appendages except on the ventral region, which is essentially immaculate. Chelicerae pale yellowish; manus reticulate with blackish brown all over, but with pattern becoming denser and darker distally: fingers with basal half deeply infuscate. Pedipalp femur yellow, sparsely spotted with blackish brown on all surfaces except ventral, which is essentially immaculate; patella yellowish, very densely spotted with blackish brown on all surfaces except ventral, which possesses spots only in the distal half; chela pale yellow, with hand immaculate, fingers blackish with distal one-third yellowish. Carapace symmetrically and very densely spotted with blackish brown (most spots are interconnected), with frontal lobes essentially immaculate but separated by a large blackish spot; tergites densely spotted with blackish brown, but without clearly defined pattern of longitudinal bands; venter essentially immaculate, sternum with a wide, discontinuous transverse blackish stripe. Legs yellowish, becoming progressively paler distally, and with internal surface immaculate except on basitarsi; coxa and trochanter essentially immaculate yellow; femur and patella yellowish, with a single large, irregular blackish spot than surrounds dorsal, external, and ventral surfaces; basitarsi each conspicuously annulated: basal half blackish brown, distal half pale yellowish; telotarsi immaculate whitish. Metasoma pale yellowish, sparsely spotted with blackish brown on distal half of every segment, giving an irregularly annulated appearance; spots become progressively larger and denser towards segment V, which has distal half densely spotted; dorsal surface on I–IV with a large, arrowhead-shaped blackish spot medially; telson vesicle essentially immaculate yellowish, aculeus immaculate, with the distal two-thirds dark reddish brown.
|
||
</paragraph>
|
||
<paragraph id="8BD2B73FFFF0657FFCAFFBC5FB66FB77" blockId="4.[831,1441,198,1908]" pageId="4" pageNumber="3">
|
||
CHELICERAE (
|
||
<figureCitation id="1356ABBAFFF0657FFC4CFBC5FB90FB33" box="[988,1074,1149,1173]" captionStart="Figures 1–19" captionStartId="3.[192,278,1736,1760]" captionTargetBox="[280,1357,170,1704]" captionTargetId="figure-140@3.[276,1357,167,1704]" captionTargetPageId="3" captionText="Figures 1–19: Microtityus minimus sp. n. Figures 1–2, 5, 10, 16–17. Dorsal (1) and ventral (2) views, telson (5), metasoma and telson lateral (10), carapace with chelicerae and tergites I–IV (16) and pectinal area with sternites III–V (17), ♂ paratype (FKCP). Figures 3–4, 6–9, 11–15, 18–19. Dorsal (3) and ventral (4) views, right movable finger internal (6), metasoma and telson ventral (7), dorsal (8), and lateral (9), right pedipalp chela dorsal (11) and external (12), pedipalp patella dorsal (13) and external (14) and pedipalp femur dorsal (15), carapace with chelicerae and tergites I–IV (18) and pectinal area with sternites III– V (19), ♀ paratype (FKCP)." pageId="4" pageNumber="3">Fig. 16</figureCitation>
|
||
, male
|
||
<typeStatus id="54D6099DFFF0657FFB15FBC5FB40FB33" box="[1157,1250,1149,1173]" pageId="4" pageNumber="3" type="paratype">paratype</typeStatus>
|
||
). With dentition typical for the genus. Tegument smooth and shiny, weakly granulose distally on manus.
|
||
</paragraph>
|
||
<paragraph id="8BD2B73FFFF0657FFCAFFB60FCD7F9A3" blockId="4.[831,1441,198,1908]" pageId="4" pageNumber="3">
|
||
CARAPACE (
|
||
<figureCitation id="1356ABBAFFF0657FFC54FB60FBB7FB56" box="[964,1045,1240,1264]" captionStart="Figures 1–19" captionStartId="3.[192,278,1736,1760]" captionTargetBox="[280,1357,170,1704]" captionTargetId="figure-140@3.[276,1357,167,1704]" captionTargetPageId="3" captionText="Figures 1–19: Microtityus minimus sp. n. Figures 1–2, 5, 10, 16–17. Dorsal (1) and ventral (2) views, telson (5), metasoma and telson lateral (10), carapace with chelicerae and tergites I–IV (16) and pectinal area with sternites III–V (17), ♂ paratype (FKCP). Figures 3–4, 6–9, 11–15, 18–19. Dorsal (3) and ventral (4) views, right movable finger internal (6), metasoma and telson ventral (7), dorsal (8), and lateral (9), right pedipalp chela dorsal (11) and external (12), pedipalp patella dorsal (13) and external (14) and pedipalp femur dorsal (15), carapace with chelicerae and tergites I–IV (18) and pectinal area with sternites III– V (19), ♀ paratype (FKCP)." pageId="4" pageNumber="3">Fig. 16</figureCitation>
|
||
, male
|
||
<typeStatus id="54D6099DFFF0657FFBCDFB60FB18FB56" box="[1117,1210,1240,1264]" pageId="4" pageNumber="3" type="paratype">paratype</typeStatus>
|
||
). Slightly wider than long, markedly triangular. Anterior margin very narrow, deeply bilobed. Carinae: anterior median, lateral ocular, central median and posterior median granulose, superciliary subgranulose, other carinae obsolete to absent. Tegument very finely and densely granulose, with many coarse granules scattered. Median eyes separated by slightly more than one ocular diameter; three pairs of lateral eyes, which are all much smaller than median eyes.
|
||
</paragraph>
|
||
<paragraph id="8BD2B73FFFF0657CFCAFF9B4FE62FAA2" blockId="4.[831,1441,198,1908]" lastBlockId="7.[192,802,956,1894]" lastPageId="7" lastPageNumber="6" pageId="4" pageNumber="3">
|
||
MESOSOMA (
|
||
<figureCitation id="1356ABBAFFF0657FFC5DF9B3FBF2F985" box="[973,1104,1547,1571]" captionStart="Figures 1–19" captionStartId="3.[192,278,1736,1760]" captionTargetBox="[280,1357,170,1704]" captionTargetId="figure-140@3.[276,1357,167,1704]" captionTargetPageId="3" captionText="Figures 1–19: Microtityus minimus sp. n. Figures 1–2, 5, 10, 16–17. Dorsal (1) and ventral (2) views, telson (5), metasoma and telson lateral (10), carapace with chelicerae and tergites I–IV (16) and pectinal area with sternites III–V (17), ♂ paratype (FKCP). Figures 3–4, 6–9, 11–15, 18–19. Dorsal (3) and ventral (4) views, right movable finger internal (6), metasoma and telson ventral (7), dorsal (8), and lateral (9), right pedipalp chela dorsal (11) and external (12), pedipalp patella dorsal (13) and external (14) and pedipalp femur dorsal (15), carapace with chelicerae and tergites I–IV (18) and pectinal area with sternites III– V (19), ♀ paratype (FKCP)." pageId="4" pageNumber="3">Figs. 16–17</figureCitation>
|
||
, male
|
||
<typeStatus id="54D6099DFFF0657FFB0CF9B3FB5FF985" box="[1180,1277,1547,1571]" pageId="4" pageNumber="3" type="paratype">paratype</typeStatus>
|
||
). Tergites with the same sculpture as on carapace; I–VI with three longitudinal carinae, which are granulose sharply projected beyond the posterior margin, VII with five serrate longitudinal carinae. Sternum type 1, pentagonal, wider than long. Pectines reaching coxa-trochanter joint of leg IV; tooth count 10/10, fulcra very well developed; basal middle lamella slightly enlarged, angulose; basal plate unmodified, about as long as wide, and with posterior margin essentially straight. Sternites III–VI with round to very short-oval spiracles and a narrow, smooth to subgranulose longitudinal carina along midline of IV– VII; III with posterior margin strongly convex, lateral areas very shallowly depressed and essentially smooth, and with the smooth patch large, subtriangular, flat, and not protruding beyond posterior margin; IV–VI coriaceous to finely granulose, with coarser granules scattered; VII finely and densely granulose; V with posterior margin essentially straight and the smooth patch moderately large, cordiform, slightly longer than wide, translucent, and bulky, essentially not protruding from margin of sternite; IV–V without carinae, VI with two, VII with four, all granulose.
|
||
</paragraph>
|
||
<caption id="DF12E7B7FFF1657EFF50F888FDFBF8C5" pageId="5" pageNumber="4" startId="5.[192,278,1840,1864]" targetBox="[191,1441,215,1806]" targetPageId="5">
|
||
<paragraph id="8BD2B73FFFF1657EFF50F888FDFBF8C5" blockId="5.[192,1441,1840,1891]" pageId="5" pageNumber="4">
|
||
<emphasis id="B9196B2DFFF1657EFF50F888FD45F8E1" bold="true" box="[192,743,1840,1864]" pageId="5" pageNumber="4">
|
||
Figures 20–23:
|
||
<taxonomicName id="4C6DCCBCFFF1657EFEFFF88AFD9FF8E1" authority="Kovařík & Teruel, 2014" authorityName="Kovařík & Teruel" authorityYear="2014" box="[367,573,1842,1863]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
|
||
<emphasis id="B9196B2DFFF1657EFEFFF88AFD9FF8E1" bold="true" box="[367,573,1842,1863]" italics="true" pageId="5" pageNumber="4">Microtityus minimus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22AD656FFF1657EFDD3F88AFDDBF8E1" box="[579,633,1842,1863]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
Figure 20.
|
||
</emphasis>
|
||
♀ paratype (FKCP) with newborn.
|
||
<emphasis id="B9196B2DFFF1657EFBD2F88AFB7AF8E1" bold="true" box="[1090,1240,1842,1863]" pageId="5" pageNumber="4">Figures 21–22.</emphasis>
|
||
♀ paratype (FKCP).
|
||
<emphasis id="B9196B2DFFF1657EFF50F8F5FE8AF8C4" bold="true" box="[192,296,1869,1890]" pageId="5" pageNumber="4">Figure 23.</emphasis>
|
||
♂ holotype at the type locality.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF12E7B7FFF2657DFF50F8EAFB6CF8CF" box="[192,1230,1874,1898]" pageId="6" pageNumber="5" startId="6.[192,278,1874,1898]" targetBox="[199,1405,191,1842]" targetPageId="6">
|
||
<paragraph id="8BD2B73FFFF2657DFF50F8EAFB6CF8CF" blockId="6.[192,1230,1874,1898]" box="[192,1230,1874,1898]" pageId="6" pageNumber="5">
|
||
<emphasis id="B9196B2DFFF2657DFF50F8EAFDDBF8CF" bold="true" box="[192,633,1874,1898]" pageId="6" pageNumber="5">
|
||
Figures 24–25:
|
||
<taxonomicName id="4C6DCCBCFFF2657DFEFEF8ECFD9EF8CF" authority="Kovařík & Teruel, 2014" authorityName="Kovařík & Teruel" authorityYear="2014" box="[366,572,1876,1897]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
|
||
<emphasis id="B9196B2DFFF2657DFEFEF8ECFD9EF8CF" bold="true" box="[366,572,1876,1897]" italics="true" pageId="6" pageNumber="5">Microtityus minimus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22AD656FFF2657DFDD3F8ECFDDBF8CF" box="[579,633,1876,1897]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, type locality (subcoastal desert scrub on gypsum-sandy soil).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD2B73FFFF3657CFE84FF73FAEBFC97" pageId="7" pageNumber="6">
|
||
<table id="F96D459FFFF39A84FF49FF72FA24FC97" box="[217,1414,202,817]" gridcols="6" gridrows="20" pageId="7" pageNumber="6">
|
||
<tr id="355DB57DFFF39A84FF49FF72FA24FF45" box="[217,1414,202,227]" gridrow="0" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FF72FDBDFF45" box="[217,543,202,227]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="7" pageNumber="6">
|
||
<emphasis id="B9196B2DFFF3657CFE84FF73FE7FFF45" bold="true" box="[276,477,203,227]" pageId="7" pageNumber="6">Dimensions (mm)</emphasis>
|
||
</th>
|
||
<th id="768CDC01FFF39A84FDA5FF72FD49FF45" box="[565,747,202,227]" gridcol="2" gridrow="0" pageId="7" pageNumber="6">
|
||
♂
|
||
<emphasis id="B9196B2DFFF3657CFDFAFF72FD72FF44" bold="true" box="[618,720,202,226]" pageId="7" pageNumber="6">paratype</emphasis>
|
||
</th>
|
||
<th id="768CDC01FFF39A84FC97FF72FC1FFF45" box="[775,957,202,227]" gridcol="3" gridrow="0" pageId="7" pageNumber="6">
|
||
♂
|
||
<emphasis id="B9196B2DFFF3657CFCAEFF72FC02FF44" bold="true" box="[830,928,202,226]" pageId="7" pageNumber="6">holotype</emphasis>
|
||
</th>
|
||
<th id="768CDC01FFF39A84FC7BFF72FB02FF45" box="[1003,1184,202,227]" gridcol="4" gridrow="0" pageId="7" pageNumber="6">
|
||
♀
|
||
<emphasis id="B9196B2DFFF3657CFB8FFF72FB27FF44" bold="true" box="[1055,1157,202,226]" pageId="7" pageNumber="6">paratype</emphasis>
|
||
</th>
|
||
<th id="768CDC01FFF39A84FB40FF72FA24FF45" box="[1232,1414,202,227]" gridcol="5" gridrow="0" pageId="7" pageNumber="6">
|
||
♀
|
||
<emphasis id="B9196B2DFFF3657CFA95FF72FAC9FF44" bold="true" box="[1285,1387,202,226]" pageId="7" pageNumber="6">paratype</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FF54FA24FEA2" box="[217,1414,236,260]" gridrow="1" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FF54FE01FEA2" box="[217,419,236,260]" gridcol="0" gridrow="1" pageId="7" pageNumber="6">Carapace</th>
|
||
<td id="768CDC01FFF39A84FE2AFF54FDBDFEA2" box="[442,543,236,260]" gridcol="1" gridrow="1" pageId="7" pageNumber="6">L / Wp</td>
|
||
<td id="768CDC01FFF39A84FDA5FF54FD49FEA2" box="[565,747,236,260]" gridcol="2" gridrow="1" pageId="7" pageNumber="6">1.32 / 1.40</td>
|
||
<td id="768CDC01FFF39A84FC97FF54FC1FFEA2" box="[775,957,236,260]" gridcol="3" gridrow="1" pageId="7" pageNumber="6">1.45 / 1.47</td>
|
||
<td id="768CDC01FFF39A84FC7BFF54FB02FEA2" box="[1003,1184,236,260]" gridcol="4" gridrow="1" pageId="7" pageNumber="6">1.55 / 1.65</td>
|
||
<td id="768CDC01FFF39A84FB40FF54FA24FEA2" box="[1232,1414,236,260]" gridcol="5" gridrow="1" pageId="7" pageNumber="6">1.60 / 1.94</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FEB3FA24FE85" box="[217,1414,267,291]" gridrow="2" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FEB3FE01FE85" box="[217,419,267,291]" gridcol="0" gridrow="2" pageId="7" pageNumber="6">Mesosoma</th>
|
||
<td id="768CDC01FFF39A84FE2AFEB3FDBDFE85" box="[442,543,267,291]" gridcol="1" gridrow="2" pageId="7" pageNumber="6">L</td>
|
||
<td id="768CDC01FFF39A84FDA5FEB3FD49FE85" box="[565,747,267,291]" gridcol="2" gridrow="2" pageId="7" pageNumber="6">2.00</td>
|
||
<td id="768CDC01FFF39A84FC97FEB3FC1FFE85" box="[775,957,267,291]" gridcol="3" gridrow="2" pageId="7" pageNumber="6">2.55</td>
|
||
<td id="768CDC01FFF39A84FC7BFEB3FB02FE85" box="[1003,1184,267,291]" gridcol="4" gridrow="2" pageId="7" pageNumber="6">3.25</td>
|
||
<td id="768CDC01FFF39A84FB40FEB3FA24FE85" box="[1232,1414,267,291]" gridcol="5" gridrow="2" pageId="7" pageNumber="6">3.50</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FE91FA24FEE7" box="[217,1414,297,321]" gridrow="3" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FE91FE01FEE7" box="[217,419,297,321]" gridcol="0" gridrow="3" pageId="7" pageNumber="6">Tergite VII</th>
|
||
<td id="768CDC01FFF39A84FE2AFE91FDBDFEE7" box="[442,543,297,321]" gridcol="1" gridrow="3" pageId="7" pageNumber="6">L / W</td>
|
||
<td id="768CDC01FFF39A84FDA5FE91FD49FEE7" box="[565,747,297,321]" gridcol="2" gridrow="3" pageId="7" pageNumber="6">0.65 / 1.25</td>
|
||
<td id="768CDC01FFF39A84FC97FE91FC1FFEE7" box="[775,957,297,321]" gridcol="3" gridrow="3" pageId="7" pageNumber="6">0.75 / 1.30</td>
|
||
<td id="768CDC01FFF39A84FC7BFE91FB02FEE7" box="[1003,1184,297,321]" gridcol="4" gridrow="3" pageId="7" pageNumber="6">0.80 / 1.50</td>
|
||
<td id="768CDC01FFF39A84FB40FE91FA24FEE7" box="[1232,1414,297,321]" gridcol="5" gridrow="3" pageId="7" pageNumber="6">0.84 / 1.52</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FEF0FA24FEC6" box="[217,1414,328,352]" gridrow="4" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FEF0FE01FEC6" box="[217,419,328,352]" gridcol="0" gridrow="4" pageId="7" pageNumber="6">Metasoma</th>
|
||
<td id="768CDC01FFF39A84FE2AFEF0FDBDFEC6" box="[442,543,328,352]" gridcol="1" gridrow="4" pageId="7" pageNumber="6">L</td>
|
||
<td id="768CDC01FFF39A84FDA5FEF0FD49FEC6" box="[565,747,328,352]" gridcol="2" gridrow="4" pageId="7" pageNumber="6">5.91</td>
|
||
<td id="768CDC01FFF39A84FC97FEF0FC1FFEC6" box="[775,957,328,352]" gridcol="3" gridrow="4" pageId="7" pageNumber="6">6.31</td>
|
||
<td id="768CDC01FFF39A84FC7BFEF0FB02FEC6" box="[1003,1184,328,352]" gridcol="4" gridrow="4" pageId="7" pageNumber="6">6.64</td>
|
||
<td id="768CDC01FFF39A84FB40FEF0FA24FEC6" box="[1232,1414,328,352]" gridcol="5" gridrow="4" pageId="7" pageNumber="6">6.75</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FEDFFA24FED9" box="[217,1414,359,383]" gridrow="5" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FEDFFE01FED9" box="[217,419,359,383]" gridcol="0" gridrow="5" pageId="7" pageNumber="6">Segment I</th>
|
||
<td id="768CDC01FFF39A84FE2AFEDFFDBDFED9" box="[442,543,359,383]" gridcol="1" gridrow="5" pageId="7" pageNumber="6">L / W</td>
|
||
<td id="768CDC01FFF39A84FDA5FEDFFD49FED9" box="[565,747,359,383]" gridcol="2" gridrow="5" pageId="7" pageNumber="6">0.65 / 0.73</td>
|
||
<td id="768CDC01FFF39A84FC97FEDFFC1FFED9" box="[775,957,359,383]" gridcol="3" gridrow="5" pageId="7" pageNumber="6">0.67 / 0.80</td>
|
||
<td id="768CDC01FFF39A84FC7BFEDFFB02FED9" box="[1003,1184,359,383]" gridcol="4" gridrow="5" pageId="7" pageNumber="6">0.74 / 0.84</td>
|
||
<td id="768CDC01FFF39A84FB40FEDFFA24FED9" box="[1232,1414,359,383]" gridcol="5" gridrow="5" pageId="7" pageNumber="6">0.69 / 0.84</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FE3EFA24FE38" box="[217,1414,390,414]" gridrow="6" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FE3EFE01FE38" box="[217,419,390,414]" gridcol="0" gridrow="6" pageId="7" pageNumber="6">Segment II</th>
|
||
<td id="768CDC01FFF39A84FE2AFE3EFDBDFE38" box="[442,543,390,414]" gridcol="1" gridrow="6" pageId="7" pageNumber="6">L / W</td>
|
||
<td id="768CDC01FFF39A84FDA5FE3EFD49FE38" box="[565,747,390,414]" gridcol="2" gridrow="6" pageId="7" pageNumber="6">0.75 / 0.68</td>
|
||
<td id="768CDC01FFF39A84FC97FE3EFC1FFE38" box="[775,957,390,414]" gridcol="3" gridrow="6" pageId="7" pageNumber="6">0.82 / 0.73</td>
|
||
<td id="768CDC01FFF39A84FC7BFE3EFB02FE38" box="[1003,1184,390,414]" gridcol="4" gridrow="6" pageId="7" pageNumber="6">0.84 / 0.74</td>
|
||
<td id="768CDC01FFF39A84FB40FE3EFA24FE38" box="[1232,1414,390,414]" gridcol="5" gridrow="6" pageId="7" pageNumber="6">0.86 / 0.75</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FE1CFA24FE1A" box="[217,1414,420,444]" gridrow="7" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FE1CFE01FE1A" box="[217,419,420,444]" gridcol="0" gridrow="7" pageId="7" pageNumber="6">Segment III</th>
|
||
<td id="768CDC01FFF39A84FE2AFE1CFDBDFE1A" box="[442,543,420,444]" gridcol="1" gridrow="7" pageId="7" pageNumber="6">L / W</td>
|
||
<td id="768CDC01FFF39A84FDA5FE1CFD49FE1A" box="[565,747,420,444]" gridcol="2" gridrow="7" pageId="7" pageNumber="6">0.87 / 0.63</td>
|
||
<td id="768CDC01FFF39A84FC97FE1CFC1FFE1A" box="[775,957,420,444]" gridcol="3" gridrow="7" pageId="7" pageNumber="6">0.95 / 0.68</td>
|
||
<td id="768CDC01FFF39A84FC7BFE1CFB02FE1A" box="[1003,1184,420,444]" gridcol="4" gridrow="7" pageId="7" pageNumber="6">0.97 / 0.68</td>
|
||
<td id="768CDC01FFF39A84FB40FE1CFA24FE1A" box="[1232,1414,420,444]" gridcol="5" gridrow="7" pageId="7" pageNumber="6">0.99 / 0.69</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FE7BFA24FE7D" box="[217,1414,451,475]" gridrow="8" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FE7BFE01FE7D" box="[217,419,451,475]" gridcol="0" gridrow="8" pageId="7" pageNumber="6">Segment IV</th>
|
||
<td id="768CDC01FFF39A84FE2AFE7BFDBDFE7D" box="[442,543,451,475]" gridcol="1" gridrow="8" pageId="7" pageNumber="6">L / W</td>
|
||
<td id="768CDC01FFF39A84FDA5FE7BFD49FE7D" box="[565,747,451,475]" gridcol="2" gridrow="8" pageId="7" pageNumber="6">1.03 / 0.61</td>
|
||
<td id="768CDC01FFF39A84FC97FE7BFC1FFE7D" box="[775,957,451,475]" gridcol="3" gridrow="8" pageId="7" pageNumber="6">1.12 / 0.65</td>
|
||
<td id="768CDC01FFF39A84FC7BFE7BFB02FE7D" box="[1003,1184,451,475]" gridcol="4" gridrow="8" pageId="7" pageNumber="6">1.20 / 0.67</td>
|
||
<td id="768CDC01FFF39A84FB40FE7BFA24FE7D" box="[1232,1414,451,475]" gridcol="5" gridrow="8" pageId="7" pageNumber="6">1.22 / 0.67</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FE59FA24FE5F" box="[217,1414,481,505]" gridrow="9" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FE59FE01FE5F" box="[217,419,481,505]" gridcol="0" gridrow="9" pageId="7" pageNumber="6">Segment V</th>
|
||
<td id="768CDC01FFF39A84FE2AFE59FDBDFE5F" box="[442,543,481,505]" gridcol="1" gridrow="9" pageId="7" pageNumber="6">L / W</td>
|
||
<td id="768CDC01FFF39A84FDA5FE59FD49FE5F" box="[565,747,481,505]" gridcol="2" gridrow="9" pageId="7" pageNumber="6">1.37 / 0.58</td>
|
||
<td id="768CDC01FFF39A84FC97FE59FC1FFE5F" box="[775,957,481,505]" gridcol="3" gridrow="9" pageId="7" pageNumber="6">1.45 / 0.63</td>
|
||
<td id="768CDC01FFF39A84FC7BFE59FB02FE5F" box="[1003,1184,481,505]" gridcol="4" gridrow="9" pageId="7" pageNumber="6">1.50 / 0.65</td>
|
||
<td id="768CDC01FFF39A84FB40FE59FA24FE5F" box="[1232,1414,481,505]" gridcol="5" gridrow="9" pageId="7" pageNumber="6">1.54 / 0.65</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FDB8FA24FDBE" box="[217,1414,512,536]" gridrow="10" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FDB8FE01FDBE" box="[217,419,512,536]" gridcol="0" gridrow="10" pageId="7" pageNumber="6">Telson</th>
|
||
<td id="768CDC01FFF39A84FE2AFDB8FDBDFDBE" box="[442,543,512,536]" gridcol="1" gridrow="10" pageId="7" pageNumber="6">L</td>
|
||
<td id="768CDC01FFF39A84FDA5FDB8FD49FDBE" box="[565,747,512,536]" gridcol="2" gridrow="10" pageId="7" pageNumber="6">1.24</td>
|
||
<td id="768CDC01FFF39A84FC97FDB8FC1FFDBE" box="[775,957,512,536]" gridcol="3" gridrow="10" pageId="7" pageNumber="6">1.30</td>
|
||
<td id="768CDC01FFF39A84FC7BFDB8FB02FDBE" box="[1003,1184,512,536]" gridcol="4" gridrow="10" pageId="7" pageNumber="6">1.39</td>
|
||
<td id="768CDC01FFF39A84FB40FDB8FA24FDBE" box="[1232,1414,512,536]" gridcol="5" gridrow="10" pageId="7" pageNumber="6">1.45</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FDA7FA24FD91" box="[217,1414,543,567]" gridrow="11" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FDA7FE01FD91" box="[217,419,543,567]" gridcol="0" gridrow="11" pageId="7" pageNumber="6">Vesicle</th>
|
||
<td id="768CDC01FFF39A84FE2AFDA7FDBDFD91" box="[442,543,543,567]" gridcol="1" gridrow="11" pageId="7" pageNumber="6">L / W / H</td>
|
||
<td id="768CDC01FFF39A84FDA5FDA7FD49FD91" box="[565,747,543,567]" gridcol="2" gridrow="11" pageId="7" pageNumber="6">0.62 / 0.41 / 0.41</td>
|
||
<td id="768CDC01FFF39A84FC97FDA7FC1FFD91" box="[775,957,543,567]" gridcol="3" gridrow="11" pageId="7" pageNumber="6">0.65 / 0.42 / 0.42</td>
|
||
<td id="768CDC01FFF39A84FC7BFDA7FB02FD91" box="[1003,1184,543,567]" gridcol="4" gridrow="11" pageId="7" pageNumber="6">0.70 / 0.47 / 0.46</td>
|
||
<td id="768CDC01FFF39A84FB40FDA7FA24FD91" box="[1232,1414,543,567]" gridcol="5" gridrow="11" pageId="7" pageNumber="6">0.75 / 0.47 / 0.46</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FD86FA24FDF0" box="[217,1414,574,598]" gridrow="12" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FD86FE01FDF0" box="[217,419,574,598]" gridcol="0" gridrow="12" pageId="7" pageNumber="6">Aculeus</th>
|
||
<td id="768CDC01FFF39A84FE2AFD86FDBDFDF0" box="[442,543,574,598]" gridcol="1" gridrow="12" pageId="7" pageNumber="6">L</td>
|
||
<td id="768CDC01FFF39A84FDA5FD86FD49FDF0" box="[565,747,574,598]" gridcol="2" gridrow="12" pageId="7" pageNumber="6">0.62</td>
|
||
<td id="768CDC01FFF39A84FC97FD86FC1FFDF0" box="[775,957,574,598]" gridcol="3" gridrow="12" pageId="7" pageNumber="6">0.65</td>
|
||
<td id="768CDC01FFF39A84FC7BFD86FB02FDF0" box="[1003,1184,574,598]" gridcol="4" gridrow="12" pageId="7" pageNumber="6">0.69</td>
|
||
<td id="768CDC01FFF39A84FB40FD86FA24FDF0" box="[1232,1414,574,598]" gridcol="5" gridrow="12" pageId="7" pageNumber="6">0.70</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FDE4FA24FDD2" box="[217,1414,604,628]" gridrow="13" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FDE4FE01FDD2" box="[217,419,604,628]" gridcol="0" gridrow="13" pageId="7" pageNumber="6">Pedipalp</th>
|
||
<td id="768CDC01FFF39A84FE2AFDE4FDBDFDD2" box="[442,543,604,628]" gridcol="1" gridrow="13" pageId="7" pageNumber="6">L</td>
|
||
<td id="768CDC01FFF39A84FDA5FDE4FD49FDD2" box="[565,747,604,628]" gridcol="2" gridrow="13" pageId="7" pageNumber="6">3.86</td>
|
||
<td id="768CDC01FFF39A84FC97FDE4FC1FFDD2" box="[775,957,604,628]" gridcol="3" gridrow="13" pageId="7" pageNumber="6">4.07</td>
|
||
<td id="768CDC01FFF39A84FC7BFDE4FB02FDD2" box="[1003,1184,604,628]" gridcol="4" gridrow="13" pageId="7" pageNumber="6">4.59</td>
|
||
<td id="768CDC01FFF39A84FB40FDE4FA24FDD2" box="[1232,1414,604,628]" gridcol="5" gridrow="13" pageId="7" pageNumber="6">4.61</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FDC3FA24FD35" box="[217,1414,635,659]" gridrow="14" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FDC3FE01FD35" box="[217,419,635,659]" gridcol="0" gridrow="14" pageId="7" pageNumber="6">Femur</th>
|
||
<td id="768CDC01FFF39A84FE2AFDC3FDBDFD35" box="[442,543,635,659]" gridcol="1" gridrow="14" pageId="7" pageNumber="6">L / W</td>
|
||
<td id="768CDC01FFF39A84FDA5FDC3FD49FD35" box="[565,747,635,659]" gridcol="2" gridrow="14" pageId="7" pageNumber="6">0.97 / 0.34</td>
|
||
<td id="768CDC01FFF39A84FC97FDC3FC1FFD35" box="[775,957,635,659]" gridcol="3" gridrow="14" pageId="7" pageNumber="6">1.00 / 0.38</td>
|
||
<td id="768CDC01FFF39A84FC7BFDC3FB02FD35" box="[1003,1184,635,659]" gridcol="4" gridrow="14" pageId="7" pageNumber="6">1.15 / 0.40</td>
|
||
<td id="768CDC01FFF39A84FB40FDC3FA24FD35" box="[1232,1414,635,659]" gridcol="5" gridrow="14" pageId="7" pageNumber="6">1.15 / 0.41</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FD22FA24FD14" box="[217,1414,666,690]" gridrow="15" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FD22FE01FD14" box="[217,419,666,690]" gridcol="0" gridrow="15" pageId="7" pageNumber="6">Patella</th>
|
||
<td id="768CDC01FFF39A84FE2AFD22FDBDFD14" box="[442,543,666,690]" gridcol="1" gridrow="15" pageId="7" pageNumber="6">L / W</td>
|
||
<td id="768CDC01FFF39A84FDA5FD22FD49FD14" box="[565,747,666,690]" gridcol="2" gridrow="15" pageId="7" pageNumber="6">1.13 / 0.44</td>
|
||
<td id="768CDC01FFF39A84FC97FD22FC1FFD14" box="[775,957,666,690]" gridcol="3" gridrow="15" pageId="7" pageNumber="6">1.22 / 0.47</td>
|
||
<td id="768CDC01FFF39A84FC7BFD22FB02FD14" box="[1003,1184,666,690]" gridcol="4" gridrow="15" pageId="7" pageNumber="6">1.36 / 0.54</td>
|
||
<td id="768CDC01FFF39A84FB40FD22FA24FD14" box="[1232,1414,666,690]" gridcol="5" gridrow="15" pageId="7" pageNumber="6">1.36 / 0.54</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FD00FA24FD76" box="[217,1414,696,720]" gridrow="16" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FD00FE01FD76" box="[217,419,696,720]" gridcol="0" gridrow="16" pageId="7" pageNumber="6">Chela</th>
|
||
<td id="768CDC01FFF39A84FE2AFD00FDBDFD76" box="[442,543,696,720]" gridcol="1" gridrow="16" pageId="7" pageNumber="6">L</td>
|
||
<td id="768CDC01FFF39A84FDA5FD00FD49FD76" box="[565,747,696,720]" gridcol="2" gridrow="16" pageId="7" pageNumber="6">1.76</td>
|
||
<td id="768CDC01FFF39A84FC97FD00FC1FFD76" box="[775,957,696,720]" gridcol="3" gridrow="16" pageId="7" pageNumber="6">1.85</td>
|
||
<td id="768CDC01FFF39A84FC7BFD00FB02FD76" box="[1003,1184,696,720]" gridcol="4" gridrow="16" pageId="7" pageNumber="6">2.08</td>
|
||
<td id="768CDC01FFF39A84FB40FD00FA24FD76" box="[1232,1414,696,720]" gridcol="5" gridrow="16" pageId="7" pageNumber="6">2.10</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FD6FFA24FD49" box="[217,1414,727,751]" gridrow="17" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FD6FFE01FD49" box="[217,419,727,751]" gridcol="0" gridrow="17" pageId="7" pageNumber="6">Hand</th>
|
||
<td id="768CDC01FFF39A84FE2AFD6FFDBDFD49" box="[442,543,727,751]" gridcol="1" gridrow="17" pageId="7" pageNumber="6">L / W / H</td>
|
||
<td id="768CDC01FFF39A84FDA5FD6FFD49FD49" box="[565,747,727,751]" gridcol="2" gridrow="17" pageId="7" pageNumber="6">0.59 / 0.34 / 0.34</td>
|
||
<td id="768CDC01FFF39A84FC97FD6FFC1FFD49" box="[775,957,727,751]" gridcol="3" gridrow="17" pageId="7" pageNumber="6">0.60 / 0.35 / 0.35</td>
|
||
<td id="768CDC01FFF39A84FC7BFD6FFB02FD49" box="[1003,1184,727,751]" gridcol="4" gridrow="17" pageId="7" pageNumber="6">0.69 / 0.44 / 0.44</td>
|
||
<td id="768CDC01FFF39A84FB40FD6FFA24FD49" box="[1232,1414,727,751]" gridcol="5" gridrow="17" pageId="7" pageNumber="6">0.70 / 0.45 / 0.45</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FD4EFA24FCA8" box="[217,1414,758,782]" gridrow="18" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FD4EFE01FCA8" box="[217,419,758,782]" gridcol="0" gridrow="18" pageId="7" pageNumber="6">Movable finger</th>
|
||
<td id="768CDC01FFF39A84FE2AFD4EFDBDFCA8" box="[442,543,758,782]" gridcol="1" gridrow="18" pageId="7" pageNumber="6">L</td>
|
||
<td id="768CDC01FFF39A84FDA5FD4EFD49FCA8" box="[565,747,758,782]" gridcol="2" gridrow="18" pageId="7" pageNumber="6">1.17</td>
|
||
<td id="768CDC01FFF39A84FC97FD4EFC1FFCA8" box="[775,957,758,782]" gridcol="3" gridrow="18" pageId="7" pageNumber="6">1.25</td>
|
||
<td id="768CDC01FFF39A84FC7BFD4EFB02FCA8" box="[1003,1184,758,782]" gridcol="4" gridrow="18" pageId="7" pageNumber="6">1.39</td>
|
||
<td id="768CDC01FFF39A84FB40FD4EFA24FCA8" box="[1232,1414,758,782]" gridcol="5" gridrow="18" pageId="7" pageNumber="6">1.40</td>
|
||
</tr>
|
||
<tr id="355DB57DFFF39A84FF49FCA1FA24FC97" box="[217,1414,793,817]" gridrow="19" pageId="7" pageNumber="6">
|
||
<th id="768CDC01FFF39A84FF49FCA1FE01FC97" box="[217,419,793,817]" gridcol="0" gridrow="19" pageId="7" pageNumber="6">Total</th>
|
||
<td id="768CDC01FFF39A84FE2AFCA1FDBDFC97" box="[442,543,793,817]" gridcol="1" gridrow="19" pageId="7" pageNumber="6">L</td>
|
||
<td id="768CDC01FFF39A84FDA5FCA1FD49FC97" box="[565,747,793,817]" gridcol="2" gridrow="19" pageId="7" pageNumber="6">9.23</td>
|
||
<td id="768CDC01FFF39A84FC97FCA1FC1FFC97" box="[775,957,793,817]" gridcol="3" gridrow="19" pageId="7" pageNumber="6">10.31</td>
|
||
<td id="768CDC01FFF39A84FC7BFCA1FB02FC97" box="[1003,1184,793,817]" gridcol="4" gridrow="19" pageId="7" pageNumber="6">11.44</td>
|
||
<td id="768CDC01FFF39A84FB40FCA1FA24FC97" box="[1232,1414,793,817]" gridcol="5" gridrow="19" pageId="7" pageNumber="6">11.85</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF12E7B7FFF3657CFF50FCEDFDECFC2E" pageId="7" pageNumber="6" startId="7.[192,257,853,877]">
|
||
<paragraph id="8BD2B73FFFF3657CFF50FCEDFDECFC2E" blockId="7.[192,1440,853,904]" pageId="7" pageNumber="6">
|
||
<emphasis id="B9196B2DFFF3657CFF50FCEDFE82FCCB" bold="true" box="[192,288,853,877]" pageId="7" pageNumber="6">Table 1:</emphasis>
|
||
Measurements of four types of
|
||
<taxonomicName id="4C6DCCBCFFF3657CFDF0FCEFFC8AFCCA" authority="Kovařík & Teruel, 2014" authorityName="Kovařík & Teruel" authorityYear="2014" box="[608,808,855,876]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
|
||
<emphasis id="B9196B2DFFF3657CFDF0FCEFFC8AFCCA" box="[608,808,855,876]" italics="true" pageId="7" pageNumber="6">Microtityus minimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9196B2DFFF3657CFCA1FCEFFCCBFCCA" bold="true" box="[817,873,855,876]" pageId="7" pageNumber="6">
|
||
<taxonomicNameLabel id="A22AD656FFF3657CFCA1FCEFFCCBFCCA" box="[817,873,855,876]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(RTOC). Abbreviations: length (L), width (W), posterior width (Wp), depth (H), left (L), right (R).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD2B73FFFF3657CFF50FAB3FD4DF96B" blockId="7.[192,802,956,1894]" pageId="7" pageNumber="6">
|
||
METASOMA AND TELSON (
|
||
<figureCitation id="1356ABBAFFF3657CFE7BFAB3FD3AFA85" box="[491,664,1291,1315]" captionStart="Figures 1–19" captionStartId="3.[192,278,1736,1760]" captionTargetBox="[280,1357,170,1704]" captionTargetId="figure-140@3.[276,1357,167,1704]" captionTargetPageId="3" captionText="Figures 1–19: Microtityus minimus sp. n. Figures 1–2, 5, 10, 16–17. Dorsal (1) and ventral (2) views, telson (5), metasoma and telson lateral (10), carapace with chelicerae and tergites I–IV (16) and pectinal area with sternites III–V (17), ♂ paratype (FKCP). Figures 3–4, 6–9, 11–15, 18–19. Dorsal (3) and ventral (4) views, right movable finger internal (6), metasoma and telson ventral (7), dorsal (8), and lateral (9), right pedipalp chela dorsal (11) and external (12), pedipalp patella dorsal (13) and external (14) and pedipalp femur dorsal (15), carapace with chelicerae and tergites I–IV (18) and pectinal area with sternites III– V (19), ♀ paratype (FKCP)." pageId="7" pageNumber="6">Figs. 5 and 10</figureCitation>
|
||
, male
|
||
<typeStatus id="54D6099DFFF3657CFD7AFAB3FF4DFAE7" pageId="7" pageNumber="6" type="paratype">paratype</typeStatus>
|
||
). Metasoma with intercarinal tegument finely and densely granulose, with many coarser granules scattered; segments I–II with ten complete carinae, II–IV with eight (even though lateral supramedian carinae are indicated by irregular granulation), V with five, all sharply serrate to denticulate, dorsolateral carinae on I–IV with 2–3 terminal denticles unevenly enlarged and very sharp. Telson vesicle short-oval, coriaceous with small granules scattered, and with a granulose ventromedian carina that abruptly rises into the subaculear tubercle, which points towards the basal third of the aculeus and is moderately-sized, irregularly conical, essentially smooth, and possesses two vestigial dorsal granules; aculeus as long as vesicle, sharp, and evenly curved.
|
||
</paragraph>
|
||
<paragraph id="8BD2B73FFFF3657CFF50F96CFD3BF8AF" blockId="7.[192,802,956,1894]" pageId="7" pageNumber="6">LEGS. Legs with all carinae finely serrate; intercarinal tegument very finely and densely granulose.</paragraph>
|
||
<paragraph id="8BD2B73FFFF3657CFF50F8A9FBE9FA83" blockId="7.[192,802,956,1894]" lastBlockId="7.[831,1440,956,1317]" pageId="7" pageNumber="6">
|
||
PEDIPALPS (
|
||
<figureCitation id="1356ABBAFFF3657CFED5F8A8FE12F88E" box="[325,432,1808,1832]" captionStart="Figures 1–19" captionStartId="3.[192,278,1736,1760]" captionTargetBox="[280,1357,170,1704]" captionTargetId="figure-140@3.[276,1357,167,1704]" captionTargetPageId="3" captionText="Figures 1–19: Microtityus minimus sp. n. Figures 1–2, 5, 10, 16–17. Dorsal (1) and ventral (2) views, telson (5), metasoma and telson lateral (10), carapace with chelicerae and tergites I–IV (16) and pectinal area with sternites III–V (17), ♂ paratype (FKCP). Figures 3–4, 6–9, 11–15, 18–19. Dorsal (3) and ventral (4) views, right movable finger internal (6), metasoma and telson ventral (7), dorsal (8), and lateral (9), right pedipalp chela dorsal (11) and external (12), pedipalp patella dorsal (13) and external (14) and pedipalp femur dorsal (15), carapace with chelicerae and tergites I–IV (18) and pectinal area with sternites III– V (19), ♀ paratype (FKCP)." pageId="7" pageNumber="6">Figs. 1–2</figureCitation>
|
||
, male
|
||
<typeStatus id="54D6099DFFF3657CFE6FF8A8FDFEF88E" box="[511,604,1808,1832]" pageId="7" pageNumber="6" type="paratype">paratype</typeStatus>
|
||
). Neobothriotaxic A-α (femoral
|
||
<emphasis id="B9196B2DFFF3657CFEC4F897FEC8F8E1" box="[340,362,1839,1863]" italics="true" pageId="7" pageNumber="6">
|
||
d
|
||
<subScript id="17E9B57AFFF3657CFEF1F881FEC8F8E1" attach="left" box="[353,362,1849,1863]" fontSize="6" pageId="7" pageNumber="6">2</subScript>
|
||
</emphasis>
|
||
absent). Femur with five serrate carinae; intercarinal tegument coriaceous to finely granulose, with coarser granules scattered; internal surface with the four
|
||
<emphasis id="B9196B2DFFF3657CFCE6FC64FCDFFC55" box="[886,893,988,1011]" italics="true" pageId="7" pageNumber="6">i</emphasis>
|
||
trichobothria surrounding a large, truncate spur. Patella with seven serrate carinae; intercarinal tegument with the same granular sculpture as on femur, but finer; internal surface with granules becoming progressively larger distally. Chela oval and much narrower than patella; hand with nine finely serrate carinae, intercarinal tegument very finely and densely granulose; fingers without basal lobe/notch combination, fixed finger with 9/9 principal rows of denticles, movable finger with 8/8 plus apical subrow composed by four granules aligned similar to principal rows.
|
||
</paragraph>
|
||
<paragraph id="8BD2B73FFFF3657CFCAFFAEAFB66F924" blockId="7.[831,1441,1361,1667]" pageId="7" pageNumber="6">
|
||
FEMALE (
|
||
<figureCitation id="1356ABBAFFF3657CFC36FAE9FB4BFACF" box="[934,1257,1361,1385]" captionStart-0="Figures 1–19" captionStart-1="Figures 20–23" captionStartId-0="3.[192,278,1736,1760]" captionStartId-1="5.[192,278,1840,1864]" captionTargetBox-0="[280,1357,170,1704]" captionTargetBox-1="[191,1441,215,1806]" captionTargetId-0="figure-140@3.[276,1357,167,1704]" captionTargetId-1="figure-41@5.[192,1440,216,1805]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Figures 1–19: Microtityus minimus sp. n. Figures 1–2, 5, 10, 16–17. Dorsal (1) and ventral (2) views, telson (5), metasoma and telson lateral (10), carapace with chelicerae and tergites I–IV (16) and pectinal area with sternites III–V (17), ♂ paratype (FKCP). Figures 3–4, 6–9, 11–15, 18–19. Dorsal (3) and ventral (4) views, right movable finger internal (6), metasoma and telson ventral (7), dorsal (8), and lateral (9), right pedipalp chela dorsal (11) and external (12), pedipalp patella dorsal (13) and external (14) and pedipalp femur dorsal (15), carapace with chelicerae and tergites I–IV (18) and pectinal area with sternites III– V (19), ♀ paratype (FKCP)." captionText-1="Figures 20–23: Microtityus minimus sp. n. Figure 20. ♀ paratype (FKCP) with newborn. Figures 21–22. ♀ paratype (FKCP). Figure 23. ♂ holotype at the type locality." pageId="7" pageNumber="6">Figs. 3–4, 6–9, 11–15, 18–22</figureCitation>
|
||
;
|
||
<tableCitation id="C6EF8284FFF3657CFB65FAE9FA98FACF" box="[1269,1338,1361,1385]" captionStart="Table 1" captionStartId="7.[192,257,853,877]" captionText="Table 1: Measurements of four types of Microtityus minimus sp. n. (RTOC). Abbreviations: length (L), width (W), posterior width (Wp), depth (H), left (L), right (R)." pageId="7" pageNumber="6">Tab. 1</tableCitation>
|
||
). Similar to the male, but sexual dimorphism is evident by:
|
||
<emphasis id="B9196B2DFFF3657CFAC9FAC9FACDFA2F" bold="true" box="[1369,1391,1393,1417]" pageId="7" pageNumber="6">1)</emphasis>
|
||
size larger;
|
||
<emphasis id="B9196B2DFFF3657CFC05FA28FC09FA0E" bold="true" box="[917,939,1424,1448]" pageId="7" pageNumber="6">2)</emphasis>
|
||
color pattern somewhat darker and more densely spotted;
|
||
<emphasis id="B9196B2DFFF3657CFC64FA17FBA8FA61" bold="true" box="[1012,1034,1455,1479]" pageId="7" pageNumber="6">3)</emphasis>
|
||
genital papillae absent;
|
||
<emphasis id="B9196B2DFFF3657CFA82FA17FA8AFA61" bold="true" box="[1298,1320,1455,1479]" pageId="7" pageNumber="6">4)</emphasis>
|
||
mesosoma remarkably wider, with sides convex;
|
||
<emphasis id="B9196B2DFFF3657CFB74FA77FB58FA41" bold="true" box="[1252,1274,1487,1511]" pageId="7" pageNumber="6">5)</emphasis>
|
||
pectines smaller, narrower, with lower tooth counts, and with basal plate longer than wide, spatulate;
|
||
<emphasis id="B9196B2DFFF3657CFB4BF9B5FB53F983" bold="true" box="[1243,1265,1549,1573]" pageId="7" pageNumber="6">6)</emphasis>
|
||
pedipalps and metasoma with segments slightly shorter and wider;
|
||
<emphasis id="B9196B2DFFF3657CFA1AF995FA02F9E3" bold="true" box="[1418,1440,1581,1605]" pageId="7" pageNumber="6">7)</emphasis>
|
||
sternite V with the smooth patch much smaller, fusiform, flat, and with a median suture.
|
||
</paragraph>
|
||
<paragraph id="8BD2B73FFFF3657CFCAFF908FBDDF8C5" blockId="7.[831,1441,1711,1891]" pageId="7" pageNumber="6">
|
||
VARIATION. The size differences among adults are so small (directly correlated to the minuteness of this scorpion), that a clear-cut recognition of size classes is not reliable. Nevertheless, there are probably two size classes in each sex; inside each class, males are consistently smaller than females (
|
||
<tableCitation id="C6EF8284FFF3657CFBB9F8F3FBCCF8C5" box="[1065,1134,1867,1891]" captionStart="Table 1" captionStartId="7.[192,257,853,877]" captionText="Table 1: Measurements of four types of Microtityus minimus sp. n. (RTOC). Abbreviations: length (L), width (W), posterior width (Wp), depth (H), left (L), right (R)." pageId="7" pageNumber="6">Tab. 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF12E7B7FFFC6573FF50FCAEFDFCFCEE" pageId="8" pageNumber="7" startId="8.[192,267,790,814]" targetBox="[191,1440,191,756]" targetPageId="8">
|
||
<paragraph id="8BD2B73FFFFC6573FF50FCAEFDFCFCEE" blockId="8.[192,1441,790,841]" pageId="8" pageNumber="7">
|
||
<emphasis id="B9196B2DFFFC6573FF50FCAEFE95FC88" bold="true" box="[192,311,790,814]" pageId="8" pageNumber="7">Figure 26:</emphasis>
|
||
Geographical distribution of
|
||
<emphasis id="B9196B2DFFFC6573FDCCFCAFFCCEFC8A" bold="true" box="[604,876,791,812]" pageId="8" pageNumber="7">
|
||
<taxonomicName id="4C6DCCBCFFFC6573FDCCFCAFFC8EFC8A" authority="Kovařík & Teruel, 2014" authorityName="Kovařík & Teruel" authorityYear="2014" box="[604,812,791,812]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
|
||
<emphasis id="B9196B2DFFFC6573FDCCFCAFFC8EFC8A" bold="true" box="[604,812,791,812]" italics="true" pageId="8" pageNumber="7">Microtityus minimus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22AD656FFFC6573FCA4FCAFFCCEFC8A" box="[820,876,791,812]" pageId="8" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(black square) and its closest relatives
|
||
<taxonomicName id="4C6DCCBCFFFC6573FB62FCAFFA03FC8A" authorityName="Armas et Marcano" authorityYear="1992" box="[1266,1441,791,812]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="paucidentatus">
|
||
<emphasis id="B9196B2DFFFC6573FB62FCAFFA03FC8A" bold="true" box="[1266,1441,791,812]" italics="true" pageId="8" pageNumber="7">M. paucidentatus</emphasis>
|
||
</taxonomicName>
|
||
(grey square) and
|
||
<taxonomicName id="4C6DCCBCFFFC6573FEFEFC8BFE1DFCEE" authorityName="Armas et Teruel" authorityYear="2012" box="[366,447,819,840]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="reini">
|
||
<emphasis id="B9196B2DFFFC6573FEFEFC8BFE1DFCEE" bold="true" box="[366,447,819,840]" italics="true" pageId="8" pageNumber="7">M. reini</emphasis>
|
||
</taxonomicName>
|
||
(white squares).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD2B73FFFFC6573FF60FCC5FD39FB88" blockId="8.[192,802,893,1070]" pageId="8" pageNumber="7">Count of teeth per pecten was also little variable: 10 (n=8) and 11 (n=2) in males, 8 (n=4) and 9 (n=22) in females. The number of principal rows of denticles was almost constant too: nine and eight in fixed and movable fingers, respectively; a few specimens had eight and seven due to poorly defined basalmost rows.</paragraph>
|
||
<paragraph id="8BD2B73FFFFC6573FF50FBE2FD5DFA6C" blockId="8.[192,803,1114,1826]" pageId="8" pageNumber="7">
|
||
AFFINITIES. Only two other Hispaniolan species lack femoral trichobothrium
|
||
<emphasis id="B9196B2DFFFC6573FE43FBC1FE4BFB37" box="[467,489,1145,1169]" italics="true" pageId="8" pageNumber="7">
|
||
d
|
||
<subScript id="17E9B57AFFFC6573FE70FB3BFE4BFB37" attach="left" box="[480,489,1155,1169]" fontSize="6" pageId="8" pageNumber="7">2</subScript>
|
||
</emphasis>
|
||
:
|
||
<taxonomicName id="4C6DCCBCFFFC6573FD91FBC2FE72FB16" authority="Armas et Marcano, 1992" authorityName="Armas et Marcano" authorityYear="1992" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="paucidentatus">
|
||
<emphasis id="B9196B2DFFFC6573FD91FBC2FC80FB37" box="[513,802,1145,1169]" italics="true" pageId="8" pageNumber="7">Microtityus paucidentatus</emphasis>
|
||
Armas et Marcano, 1992
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6DCCBCFFFC6573FD9BFB21FF5BFB76" authority="Armas et Teruel, 2012" authorityName="Armas et Teruel" authorityYear="2012" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="reini">
|
||
<emphasis id="B9196B2DFFFC6573FD9BFB21FDC0FB16" box="[523,610,1177,1200]" italics="true" pageId="8" pageNumber="7">M. reini</emphasis>
|
||
Armas et Teruel, 2012
|
||
</taxonomicName>
|
||
. The former is known only from a single locality in the southern slope of the Sierra de Neiba (
|
||
<collectingRegion id="49A979DDFFFC6573FD14FB6FFE83FAA8" country="Dominican Republic" name="Independencia" pageId="8" pageNumber="7">Independencia Province</collectingRegion>
|
||
, roughly
|
||
<quantity id="4C951ADAFFFC6573FE1EFB4EFE46FAA8" box="[398,484,1270,1294]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="7" unit="km" value="100.0">100 km</quantity>
|
||
to the west of El Número), and is easy to distinguish from
|
||
<taxonomicName id="4C6DCCBCFFFC6573FD85FAAEFD36FA8B" authority="Kovařík & Teruel, 2014" authorityName="Kovařík & Teruel" authorityYear="2014" box="[533,660,1302,1325]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
|
||
<emphasis id="B9196B2DFFFC6573FD85FAAEFD36FA8B" box="[533,660,1302,1325]" italics="true" pageId="8" pageNumber="7">M. minimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9196B2DFFFC6573FD0CFAAEFD7BFA88" bold="true" box="[668,729,1302,1326]" pageId="8" pageNumber="7">
|
||
<taxonomicNameLabel id="A22AD656FFFC6573FD0CFAAEFD7BFA88" box="[668,729,1302,1326]" pageId="8" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by:
|
||
<emphasis id="B9196B2DFFFC6573FC9BFAAEFC80FA88" bold="true" box="[779,802,1302,1326]" pageId="8" pageNumber="7">1)</emphasis>
|
||
size larger (male
|
||
<quantity id="4C951ADAFFFC6573FE11FA8DFE72FAEB" box="[385,464,1333,1357]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="7" unit="mm" value="13.0">13 mm</quantity>
|
||
, female
|
||
<quantity id="4C951ADAFFFC6573FDA0FA8DFD20FAEB" box="[560,642,1333,1357]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="7" unit="mm" value="16.0">16 mm</quantity>
|
||
);
|
||
<emphasis id="B9196B2DFFFC6573FD08FA8DFD0CFAEB" bold="true" box="[664,686,1333,1357]" pageId="8" pageNumber="7">2)</emphasis>
|
||
color pattern darker and much more densely spotted;
|
||
<emphasis id="B9196B2DFFFC6573FD3EFAEDFD66FACB" bold="true" box="[686,708,1365,1389]" pageId="8" pageNumber="7">3)</emphasis>
|
||
pectinal tooth count higher (male 11/11, female 9/10);
|
||
<emphasis id="B9196B2DFFFC6573FC9CFACCFC80FA2A" bold="true" box="[780,802,1396,1420]" pageId="8" pageNumber="7">4)</emphasis>
|
||
metasoma slightly longer and less robust;
|
||
<emphasis id="B9196B2DFFFC6573FD04FA2BFD08FA0D" bold="true" box="[660,682,1427,1451]" pageId="8" pageNumber="7">5)</emphasis>
|
||
male sternite V with the smooth patch much wider and shorter.
|
||
</paragraph>
|
||
<paragraph id="8BD2B73FFFFC6573FF60FA69FD42F884" blockId="8.[192,803,1114,1826]" pageId="8" pageNumber="7">
|
||
The most similar morphologically is actually
|
||
<taxonomicName id="4C6DCCBCFFFC6573FC95FA6AFF50F9AF" authorityName="Armas et Teruel" authorityYear="2012" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="reini">
|
||
<emphasis id="B9196B2DFFFC6573FC95FA6AFF50F9AF" italics="true" pageId="8" pageNumber="7">M. reini</emphasis>
|
||
</taxonomicName>
|
||
, which occurs in the southern foothills of Cordillera Central (
|
||
<collectingRegion id="49A979DDFFFC6573FEB6F9A8FE4BF98E" box="[294,489,1552,1576]" country="Dominican Republic" name="Peravia" pageId="8" pageNumber="7">Peravia Province</collectingRegion>
|
||
, about
|
||
<quantity id="4C951ADAFFFC6573FDDAF9A8FD3AF98E" box="[586,664,1552,1576]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="7" unit="km" value="25.0">25 km</quantity>
|
||
east of El Número), and can be unequivocally separated on the basis of the following characters (females only, as males are unknown):
|
||
<emphasis id="B9196B2DFFFC6573FEFBF9D6FE23F920" bold="true" box="[363,385,1646,1670]" pageId="8" pageNumber="7">1)</emphasis>
|
||
size 13–14% larger (
|
||
<quantity id="4C951ADAFFFC6573FDE8F9D6FD56F920" box="[632,756,1646,1670]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="8" pageNumber="7" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
|
||
);
|
||
<emphasis id="B9196B2DFFFC6573FC9BF9D6FC80F920" bold="true" box="[779,802,1646,1670]" pageId="8" pageNumber="7">2)</emphasis>
|
||
coloration basically darker, but with the spots lighter (body and appendages brownish orange, patterned with medium brown);
|
||
<emphasis id="B9196B2DFFFC6573FE13F974FE3BF942" bold="true" box="[387,409,1740,1764]" pageId="8" pageNumber="7">3)</emphasis>
|
||
basal pectinal plate shorter, trapezoidal, and posteriorly narrower;
|
||
<emphasis id="B9196B2DFFFC6573FDB7F953FD9CF8A5" bold="true" box="[551,574,1771,1795]" pageId="8" pageNumber="7">4)</emphasis>
|
||
sternite III with posterior margin widely bilobed, with a median notch.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377E4B4FFFC6573FF50F8F1FAC1F8BD" pageId="8" pageNumber="7" type="distribution">
|
||
<paragraph id="8BD2B73FFFFC6573FF50F8F1FC83F8D9" blockId="8.[192,802,1864,1919]" pageId="8" pageNumber="7">
|
||
DISTRIBUTION (
|
||
<figureCitation id="1356ABBAFFFC6573FEFBF8F0FE1CF8C6" box="[363,446,1864,1888]" captionStart="Figure 26" captionStartId="8.[192,267,790,814]" captionTargetBox="[191,1440,191,756]" captionTargetId="figure-631@8.[192,1439,192,755]" captionTargetPageId="8" captionText="Figure 26: Geographical distribution of Microtityus minimus sp. n. (black square) and its closest relatives M. paucidentatus (grey square) and M. reini (white squares)." pageId="8" pageNumber="7">Fig. 26</figureCitation>
|
||
). Known only from the
|
||
<typeStatus id="54D6099DFFFC6573FD5CF8F1FD58F8C7" box="[716,762,1865,1889]" pageId="8" pageNumber="7">type</typeStatus>
|
||
locality, in the southernmost foothills of the Cordillera
|
||
</paragraph>
|
||
<paragraph id="8BD2B73FFFFC6573FCAFFCC5FCC2FC15" blockId="8.[831,1440,893,947]" pageId="8" pageNumber="7">
|
||
Central mountain range, in southern
|
||
<collectingCountry id="F37AF7AFFFFC6573FB41FCC5FCF8FC15" name="Dominican Republic" pageId="8" pageNumber="7">Dominican Republic</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD2B73FFFFC6573FCAFFC61FB9CFB58" blockId="8.[831,1441,984,1819]" pageId="8" pageNumber="7">
|
||
ECOLOGICAL NOTES. This species lives in subcoastal desert scrub on gypsum-sandy soil, at an elevation of
|
||
<quantity id="4C951ADAFFFC6573FCAFFBACFC19FB8A" box="[831,955,1044,1068]" metricMagnitude="2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="8" pageNumber="7" unit="m" value="165.0" valueMax="170.0" valueMin="160.0">160–170 m</quantity>
|
||
a. s. l. (
|
||
<figureCitation id="1356ABBAFFFC6573FBB3FBACFBDAFB8A" box="[1059,1144,1044,1068]" captionStart="Figures 24–25" captionStartId="6.[192,278,1874,1898]" captionTargetBox="[199,1405,191,1842]" captionTargetId="figure-31@6.[200,1404,192,1841]" captionTargetPageId="6" captionText="Figures 24–25: Microtityus minimus sp. n., type locality (subcoastal desert scrub on gypsum-sandy soil)." pageId="8" pageNumber="7">Fig. 24</figureCitation>
|
||
). All specimens were collected during diurnal searches, hanging to the underside of small limestone rocks half-buried in the dry leaf litter of thorny shrubs and cacti, at the base of hills (
|
||
<figureCitation id="1356ABBAFFFC6573FAD1FBD6FA36FB20" box="[1345,1428,1134,1158]" captionStart="Figures 24–25" captionStartId="6.[192,278,1874,1898]" captionTargetBox="[199,1405,191,1842]" captionTargetId="figure-31@6.[200,1404,192,1841]" captionTargetPageId="6" captionText="Figures 24–25: Microtityus minimus sp. n., type locality (subcoastal desert scrub on gypsum-sandy soil)." pageId="8" pageNumber="7">Fig. 25</figureCitation>
|
||
). As observed previously in a few members of the genus, when the rocks were turned the individuals remained still but started to run very quickly after 10–20 seconds or at the slightest touch.
|
||
</paragraph>
|
||
<paragraph id="8BD2B73FFFFC6573FCFFFABDFABEFA77" blockId="8.[831,1441,984,1819]" pageId="8" pageNumber="7">
|
||
Other scorpions that occur at this locality are the scorpionid
|
||
<taxonomicName id="4C6DCCBCFFFC6573FC2EFA9BFA9AFA9D" authority="(Pocock, 1898)" baseAuthorityName="Pocock" baseAuthorityYear="1898" box="[958,1336,1315,1339]" class="Arachnida" family="Diplocentridae" genus="Cazierius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="politus">
|
||
<emphasis id="B9196B2DFFFC6573FC2EFA9BFBD9FA9D" box="[958,1147,1315,1339]" italics="true" pageId="8" pageNumber="7">Cazierius politus</emphasis>
|
||
(Pocock, 1898)
|
||
</taxonomicName>
|
||
, and the buthids
|
||
<taxonomicName id="4C6DCCBCFFFC6573FC05FAF9FACCFAFF" authority="Armas et Marcano, 1987" authorityName="Armas et Marcano" authorityYear="1987" box="[917,1390,1345,1369]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="bani">
|
||
<emphasis id="B9196B2DFFFC6573FC05FAF9FBFBFAFF" box="[917,1113,1345,1369]" italics="true" pageId="8" pageNumber="7">Centruroides bani</emphasis>
|
||
Armas et Marcano, 1987
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6DCCBCFFFC6573FCAFFAE6FB6DFAD0" authority="(Karsch, 1879)" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[831,1231,1374,1398]" class="Arachnida" family="Buthidae" genus="Rhopalurus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="princeps">
|
||
<emphasis id="B9196B2DFFFC6573FCAFFAE6FB82FAD0" box="[831,1056,1374,1398]" italics="true" pageId="8" pageNumber="7">Rhopalurus princeps</emphasis>
|
||
(Karsch, 1879)
|
||
</taxonomicName>
|
||
. All of them occur syntopically under rocks with
|
||
<taxonomicName id="4C6DCCBCFFFC6573FB04FAC5FAD6FA32" authority="Kovařík & Teruel, 2014" authorityName="Kovařík & Teruel" authorityYear="2014" box="[1172,1396,1405,1428]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
|
||
<emphasis id="B9196B2DFFFC6573FB04FAC5FAD6FA32" box="[1172,1396,1405,1428]" italics="true" pageId="8" pageNumber="7">Microtityus minimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9196B2DFFFC6573FA10FAC5FCF6FA15" bold="true" pageId="8" pageNumber="7">
|
||
<taxonomicNameLabel id="A22AD656FFFC6573FA10FAC5FCF6FA15" pageId="8" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but
|
||
<taxonomicName id="4C6DCCBCFFFC6573FC1CFA23FBF2FA15" authorityName="Armas et Marcano" authorityYear="1987" box="[908,1104,1435,1459]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="bani">
|
||
<emphasis id="B9196B2DFFFC6573FC1CFA23FBF2FA15" box="[908,1104,1435,1459]" italics="true" pageId="8" pageNumber="7">Centruroides bani</emphasis>
|
||
</taxonomicName>
|
||
is also found under dead barks of shrubs and trees, both standing and fallen.
|
||
</paragraph>
|
||
<paragraph id="8BD2B73FFFFC6573FCFFFA6FFAC1F8BD" blockId="8.[831,1441,984,1819]" pageId="8" pageNumber="7">
|
||
All females were late pregnant when collected, with embryos clearly visible through sternites and pleural membranes. Seven were kept alive and gave birth during the first month in captivity (
|
||
<figureCitation id="1356ABBAFFFC6573FBFCF989FB1EF9EF" box="[1132,1212,1585,1609]" captionStart="Figures 20–23" captionStartId="5.[192,278,1840,1864]" captionTargetBox="[191,1441,215,1806]" captionTargetId="figure-41@5.[192,1440,216,1805]" captionTargetPageId="5" captionText="Figures 20–23: Microtityus minimus sp. n. Figure 20. ♀ paratype (FKCP) with newborn. Figures 21–22. ♀ paratype (FKCP). Figure 23. ♂ holotype at the type locality." pageId="8" pageNumber="7">Fig. 20</figureCitation>
|
||
); the litters consisted of 1, 2, 2, 3, 3, 4, and 5 newborn, and the first instar lasted for 4–6 days. Also, five of the 15 individuals collected on
|
||
<date id="FFD391FFFFFC6573FC5FF933FB23F905" box="[975,1153,1675,1699]" pageId="8" pageNumber="7" value="2014-03-15">March 15, 2014</date>
|
||
(
|
||
<specimenCount id="9D6B7CB6FFFC6573FB03F933FAA4F905" box="[1171,1286,1675,1699]" pageId="8" pageNumber="7" type="male">two males</specimenCount>
|
||
and
|
||
<specimenCount id="9D6B7CB6FFFC6573FAD2F933FCDCF967" pageId="8" pageNumber="7" type="female">three females</specimenCount>
|
||
) were subadults and showed obvious signs of being about to carry out their final ecdysis; all of them were kept alive and molted successfully into perfect adults within 5–12 days, without need of being fed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377E4B4FFFC6570FCAFF8FFFDEDFE33" lastPageId="11" lastPageNumber="10" pageId="8" pageNumber="7" type="discussion">
|
||
<paragraph id="8BD2B73FFFFC6570FCAFF8FFFE20FEFE" blockId="8.[831,1440,1862,1917]" lastBlockId="11.[192,801,198,344]" lastPageId="11" lastPageNumber="10" pageId="8" pageNumber="7">
|
||
REMARKS. Despite the tiny size, maturity of the specimens herein declared as adults was clearly demonstrated: captive males mated and produced spermatophores, and females gave birth. Thus,
|
||
<taxonomicName id="4C6DCCBCFFFD6572FDCDF8F9FD7EF8FE" authority="Kovařík & Teruel, 2014" authorityName="Kovařík & Teruel" authorityYear="2014" box="[605,732,1857,1880]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
|
||
<emphasis id="B9196B2DFFFD6572FDCDF8F9FD7EF8FE" box="[605,732,1857,1880]" italics="true" pageId="9" pageNumber="8">M. minimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9196B2DFFFD6572FD74F8F9FC83F8FF" bold="true" box="[740,801,1857,1881]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="A22AD656FFFD6572FD74F8F9FC83F8FF" box="[740,801,1857,1881]" pageId="9" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the smallest confirmed scorpion; a few other similarly sized (
|
||
<quantity id="4C951ADAFFFD6572FC18F899FC75F89F" box="[904,983,1825,1849]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="8" unit="mm" value="10.0">10 mm</quantity>
|
||
or less) specimens of the genera
|
||
<taxonomicName id="4C6DCCBCFFFD6572FAC6F89AFBF5F8FE" authority="Lourenco, 1995" authorityName="Lourenco" authorityYear="1995" class="Arachnida" family="Microcharmidae" genus="Microcharmus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9196B2DFFFD6572FAC6F89AFC39F8FE" italics="true" pageId="9" pageNumber="8">Microcharmus</emphasis>
|
||
Lourenço, 1995
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6DCCBCFFFD6572FB07F8F8FCD7F8D1" authority="Mitchell, 1968" authorityName="Mitchell" authorityYear="1968" class="Arachnida" family="Typhlochactidae" genus="Typhlochactas" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9196B2DFFFD6572FB07F8F8FA91F8FE" box="[1175,1331,1856,1880]" italics="true" pageId="9" pageNumber="8">Typhlochactas</emphasis>
|
||
Mitchell, 1968
|
||
</taxonomicName>
|
||
have been alleged to be adults (e. g.,
|
||
<bibRefCitation id="EFFCCACEFFFD6572FA94F8E7FA3AF8D1" author="SISSOM" box="[1284,1432,1887,1911]" pageId="9" pageNumber="8" pagination="365 - 371" refId="ref13411" refString="SISSOM, W. D. 1988. Typhlochactas mitchelli, a new species of eyeless, montane forest litter scorpion from northeastern Oaxaca, Mexico (Chactidae, Superstitioninae, Typhlochactini). The Journal of Arachnology, 16: 365 - 371." type="journal article" year="1988">Sissom, 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFCCACEFFFF6570FF50FF7EFE1AFF78" author="LOURENCO" box="[192,440,198,222]" pageId="11" pageNumber="10" pagination="751 - 783" refId="ref13247" refString="LOURENCO, W. R., S. M. GOODMAN & B. L. FISHER. 2006. A reappraisal of the geographical distribution of the endemic family Microcharmidae Lourenco (Scorpiones) in Madagascar and description of eight new species and subspecies. Proceedings of the California Academy of Sciences, 4 th series, 57 (26): 751 - 783." type="journal article" year="2006">Lourenço et al., 2006</bibRefCitation>
|
||
; Vignoli & Prendini, 2009) but their maturity, although apparent and probably real, has been never demonstrated.
|
||
<figureCitation id="1356ABBAFFFF6570FE70FEBBFDE8FEBD" box="[480,586,259,283]" captionStart="Figures 20–23" captionStartId="5.[192,278,1840,1864]" captionTargetBox="[191,1441,215,1806]" captionTargetId="figure-41@5.[192,1440,216,1805]" captionTargetPageId="5" captionText="Figures 20–23: Microtityus minimus sp. n. Figure 20. ♀ paratype (FKCP) with newborn. Figures 21–22. ♀ paratype (FKCP). Figure 23. ♂ holotype at the type locality." pageId="11" pageNumber="10">Figure 20</figureCitation>
|
||
shows the smallest scorpion female (total length
|
||
<quantity id="4C951ADAFFFF6570FD84FE99FDD8FE9F" box="[532,634,289,313]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="11" pageNumber="10" unit="mm" value="11.4">11.4 mm</quantity>
|
||
) photographed with the newborn.
|
||
</paragraph>
|
||
<caption id="DF12E7B7FFFD6572FF50F92AFBA5F95A" pageId="9" pageNumber="8" startId="9.[192,278,1682,1706]" targetBox="[196,1431,172,1635]" targetPageId="9">
|
||
<paragraph id="8BD2B73FFFFD6572FF50F92AFBA5F95A" blockId="9.[192,1440,1682,1789]" pageId="9" pageNumber="8">
|
||
<emphasis id="B9196B2DFFFD6572FF50F92AFE43F90F" bold="true" box="[192,481,1682,1706]" pageId="9" pageNumber="8">Figures 27–39: Figure 27</emphasis>
|
||
.
|
||
<taxonomicName id="4C6DCCBCFFFD6572FE63F92CFD61F90F" authorityName=": Teruel" authorityYear="2005" box="[499,707,1684,1705]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="quisqueyanus">
|
||
<emphasis id="B9196B2DFFFD6572FE63F92CFD61F90F" bold="true" box="[499,707,1684,1705]" italics="true" pageId="9" pageNumber="8">Tityus quisqueyanus</emphasis>
|
||
</taxonomicName>
|
||
, dorsal view, ♂ topotype (FKCP).
|
||
<emphasis id="B9196B2DFFFD6572FBA6F92CFADFF90F" bold="true" box="[1078,1405,1684,1705]" pageId="9" pageNumber="8">
|
||
Figures 28–39.
|
||
<taxonomicName id="4C6DCCBCFFFD6572FB4CF92CFA94F90F" authority="Kovařík & Teruel, 2014" authorityName="Kovařík & Teruel" authorityYear="2014" box="[1244,1334,1684,1705]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="kindli" status="sp. nov.">
|
||
<emphasis id="B9196B2DFFFD6572FB4CF92CFA94F90F" bold="true" box="[1244,1334,1684,1705]" italics="true" pageId="9" pageNumber="8">T. kindli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22AD656FFFD6572FAD2F92CFADFF90F" box="[1346,1405,1684,1705]" pageId="9" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ♂ holotype, dorsal (28) and ventral (29) views, pectinal area and sternites (30), carapace with chelicerae and tergites I–IV (31), right movable finger internal (32), pedipalp chela dorsal (33) and external (34), pedipalp patella dorsal (35) and external (36), pedipalp femur dorsal (37), tibia and tarsomeres of third (38) and fourth (39) right legs.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF12E7B7FFFE6571FF50F8A8FF5DF8F9" pageId="10" pageNumber="9" startId="10.[192,278,1808,1832]" targetBox="[355,1326,206,1768]" targetPageId="10">
|
||
<paragraph id="8BD2B73FFFFE6571FF50F8A8FF5DF8F9" blockId="10.[192,1440,1808,1887]" pageId="10" pageNumber="9">
|
||
<emphasis id="B9196B2DFFFE6571FF50F8A8FD65F881" bold="true" box="[192,711,1808,1832]" pageId="10" pageNumber="9">
|
||
Figures 40–45: Figures 40–42.
|
||
<taxonomicName id="4C6DCCBCFFFE6571FD9CF8AAFD2BF881" authority="Kovařík & Teruel, 2014" authorityName="Kovařík & Teruel" authorityYear="2014" box="[524,649,1810,1831]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="kindli" status="sp. nov.">
|
||
<emphasis id="B9196B2DFFFE6571FD9CF8AAFD2BF881" bold="true" box="[524,649,1810,1831]" italics="true" pageId="10" pageNumber="9">Tityus kindli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22AD656FFFE6571FD00F8AAFD65F881" box="[656,711,1810,1831]" pageId="10" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ♂ holotype, metasoma and telson dorsal (40), ventral (41) and lateral (42) views.
|
||
<emphasis id="B9196B2DFFFE6571FE97F895FDF2F8E4" bold="true" box="[263,592,1837,1858]" pageId="10" pageNumber="9">
|
||
Figures 43–45.
|
||
<taxonomicName id="4C6DCCBCFFFE6571FE3AF895FDF2F8E4" authorityName=": Teruel" authorityYear="2005" box="[426,592,1837,1858]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="quisqueyanus">
|
||
<emphasis id="B9196B2DFFFE6571FE3AF895FDF2F8E4" bold="true" box="[426,592,1837,1858]" italics="true" pageId="10" pageNumber="9">T. quisqueyanus</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
, ♂ topotype (FKCP), metasoma and telson dorsal (43), ventral (44) and lateral (45) views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD2B73FFFFF6570FF50FEC6FDEDFE33" blockId="11.[192,591,382,406]" box="[192,591,382,406]" pageId="11" pageNumber="10">COMPARATIVE MATERIAL EXAMINED.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C377E4B4FFFF6570FF50FE04FED7FDCA" pageId="11" pageNumber="10" type="description">
|
||
<paragraph id="8BD2B73FFFFF6570FF50FE04FED7FDCA" blockId="11.[192,802,443,621]" pageId="11" pageNumber="10">
|
||
<taxonomicName id="4C6DCCBCFFFF6570FF50FE04FD4DFE75" authority="Armas et Marcano, 1992" authorityName="Armas et Marcano" authorityYear="1992" box="[192,751,443,468]" class="Arachnida" family="Buthidae" genus="Microtityus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="paucidentatus">
|
||
<emphasis id="B9196B2DFFFF6570FF50FE04FE7CFE72" bold="true" box="[192,478,444,468]" italics="true" pageId="11" pageNumber="10">Microtityus paucidentatus</emphasis>
|
||
<materialsCitation id="3B05BD62FFFF6570FE76FE03FD4DFE75" box="[486,751,443,467]" collectingDate="1987-10-03" collectorName="L. F. de Armas & de Armas, A & P. Rivera" country="Dominican Republic" county="Neiba Municipality" elevation="150" location="Apolinar Perdomo" municipality="Sierra de Neiba" pageId="11" pageNumber="10" specimenCount="1" specimenCount-female="1" stateProvince="Baoruco" typeStatus="holotype">Armas et Marcano, 1992</materialsCitation>
|
||
</taxonomicName>
|
||
<collectingCountry id="F37AF7AFFFFF6570FF50FE61FE06FE57" box="[192,420,473,497]" name="Dominican Republic" pageId="11" pageNumber="10">Dominican Republic</collectingCountry>
|
||
,
|
||
<collectingRegion id="49A979DDFFFF6570FE22FE61FD24FE57" box="[434,646,473,497]" country="Dominican Republic" name="Baoruco" pageId="11" pageNumber="10">Bahoruco Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="62B3CFB3FFFF6570FD04FE61FEB0FDB6" pageId="11" pageNumber="10">Neiba Municipality</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BB62D45FFFF6570FEB3FE40FE74FDB6" box="[291,470,504,528]" pageId="11" pageNumber="10">Sierra de Neiba</collectingMunicipality>
|
||
,
|
||
<location id="8EB2E1E4FFFF6570FE76FE40FD16FDB6" LSID="urn:lsid:plazi:treatment:03C40629FFF66570FCD4F97EFED7FDCA:8EB2E1E4FFFF6570FE76FE40FD16FDB6" box="[486,692,504,528]" country="Dominican Republic" county="Neiba Municipality" municipality="Sierra de Neiba" name="Apolinar Perdomo" pageId="11" pageNumber="10" stateProvince="Baoruco">Apolinar Perdomo</location>
|
||
,
|
||
<location id="8EB2E1E4FFFF6570FD54FE40FF57FD89" LSID="urn:lsid:plazi:treatment:03C40629FFF66570FCD4F97EFED7FDCA:8EB2E1E4FFFF6570FD54FE40FF57FD89" country="Dominican Republic" county="Neiba Municipality" municipality="Sierra de Neiba" name="Segundo Paso" pageId="11" pageNumber="10" stateProvince="Baoruco">Segundo Paso</location>
|
||
,
|
||
<elevation id="0040500CFFFF6570FE93FDAFFE3BFD89" box="[259,409,535,559]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="10" unit="m" value="150.0">
|
||
<quantity id="4C951ADAFFFF6570FE93FDAFFEEBFD89" box="[259,329,535,559]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="10" unit="m" value="150.0">150 m</quantity>
|
||
a. s. l.
|
||
</elevation>
|
||
,
|
||
<date id="FFD391FFFFFF6570FE38FDAFFDAFFD89" box="[424,525,535,559]" pageId="11" pageNumber="10" value="1987-10-03">
|
||
<collectingDate id="EF976817FFFF6570FE38FDAFFDAFFD89" box="[424,525,535,559]" pageId="11" pageNumber="10" value="1987-10-03">3.X.1987</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="2698D2E9FFFF6570FDDDFDA0FD58FD89" box="[589,762,535,559]" pageId="11" pageNumber="10">L. F. de Armas</collectorName>
|
||
, A. Abud et
|
||
<collectorName id="2698D2E9FFFF6570FEBFFD8EFE3EFDE8" box="[303,412,566,590]" pageId="11" pageNumber="10">P. Rivera</collectorName>
|
||
,
|
||
<specimenCount id="9D6B7CB6FFFF6570FE23FD8EFE76FDE8" box="[435,468,566,590]" pageId="11" pageNumber="10" type="female">1♀</specimenCount>
|
||
<typeStatus id="54D6099DFFFF6570FE74FD8EFDE3FDE8" box="[484,577,566,590]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
(IESC-3.2815)
|
||
<materialsCitation id="3B05BD62FFFF6570FC91FD8EFED3FDCA" collectionCode="IESC" country="Dominican Republic" pageId="11" pageNumber="10" specimenCount="1" specimenCount-male="1" stateProvince="Baoruco" typeStatus="paratype">
|
||
<specimenCount id="9D6B7CB6FFFF6570FC91FD8EFC80FDE8" box="[769,802,566,590]" pageId="11" pageNumber="10" type="male">1♂</specimenCount>
|
||
<typeStatus id="54D6099DFFFF6570FF50FDEDFEB9FDCB" box="[192,283,597,621]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="ED7C2FFAFFFF6570FEBAFDECFECBFDCA" box="[298,361,596,620]" pageId="11" pageNumber="10">IESC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |