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<document id="82B720E133A9E473A824124F377339A5" ID-CLB-Dataset="5062" ID-DOI="10.5281/zenodo.5170621" ID-GBIF-Dataset="af75a762-387c-4dc9-850c-293eb7c3e15e" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170621" ID-ZooBank="5A3A8AB7-CC4D-476E-9351-7D41830FEEA5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628286353334" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2016" docId="03948B2CFFF6FFF8388D683CFAC465AE" docLanguage="en" docName="InsectaMundi.0484.1-4.pdf" docOrigin="Insecta Mundi 2016 (484)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Dinonigidius bartolozzii Paulsen 2016, new species" docType="treatment" docUuid="37859D6E-BB61-4DCE-A994-380E12998A0C" docUuidSource="ZooBank" docVersion="8" lastPageNumber="3" masterDocId="FFADF354FFF5FFFC384D693DFFBF6714" masterDocTitle="A new species of Dinonigidius de Lisle from Sri Lanka (Coleoptera: Lucanidae)" masterLastPageNumber="4" masterPageNumber="1" pageNumber="2" updateTime="1699057894213" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="B80F21050571E195544B70A49330FF0D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F2977B71862FD53CEE07D29E409879C">
<mods:title id="4A2377126A714D938BF6168D5C957207">A new species of Dinonigidius de Lisle from Sri Lanka (Coleoptera: Lucanidae)</mods:title>
</mods:titleInfo>
<mods:name id="0ED7860FD09B719021614797B737949F" type="personal">
<mods:role id="D845F46C0CACA2FD55CB43003AAEDA9A">
<mods:roleTerm id="2D316F7AE43FF0F62BE9DA49128FB97A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4A89B49D43A2846346E2D682D5E24531">Paulsen, M. J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B75E347570616D86C075D40A634C4E0F">text</mods:typeOfResource>
<mods:relatedItem id="E76FF7FB30A8E2E26C6484C2EF959F8B" type="host">
<mods:titleInfo id="B8993F0D95C609B69029864C40766B78">
<mods:title id="16B8FE36C97558AC490F4C3392EE2940">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="38B797A2D6BC65833627F8239999B529">
<mods:date id="B8E9E992E88876D57EB05C9754BE47C7">2016</mods:date>
<mods:detail id="41E612CBE14E833AF8EDD0984314F21A" type="pubDate">
<mods:number id="A8821BBC80615F65088FED6D8F74A6FE">2016-05-04</mods:number>
</mods:detail>
<mods:detail id="129F096882C1BD0DC715C96B6CF57EEE" type="volume">
<mods:number id="39ECBD4D4E8E44AB2BD5C53AC828CF37">2016</mods:number>
</mods:detail>
<mods:detail id="F0CF6BCFC16F846A34938B32D170E227" type="issue">
<mods:number id="CF1760E9AB8CA6D482D972BFA9B599CD">484</mods:number>
</mods:detail>
<mods:extent id="532D412059F1E4137D6513DCA7E30B59" unit="page">
<mods:start id="D3D3F305547A68CDC872791242F22D36">1</mods:start>
<mods:end id="55CBE3BD1C9CFE00D2B3089EA976065F">4</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B7D367544BA5155E482FEF9674EF699D">journal article</mods:classification>
<mods:identifier id="AEDA8BE7E855FE00776B943EB4482B99" type="CLB-Dataset">5062</mods:identifier>
<mods:identifier id="BB6BD4D22CE1ED8F188956DCBE56EF02" type="DOI">10.5281/zenodo.5170621</mods:identifier>
<mods:identifier id="989BF97E3FBA3A1D7147B2C3DA35162A" type="GBIF-Dataset">af75a762-387c-4dc9-850c-293eb7c3e15e</mods:identifier>
<mods:identifier id="A403F17892BD9956840AA364D0D625D9" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="658F4DE5B59779CCD3540B8EE82AE93D" type="Zenodo-Dep">5170621</mods:identifier>
<mods:identifier id="AD361E8F6986937D4665BF469DA4F331" type="ZooBank">5A3A8AB7-CC4D-476E-9351-7D41830FEEA5</mods:identifier>
</mods:mods>
<treatment id="03948B2CFFF6FFF8388D683CFAC465AE" ID-DOI="http://doi.org/10.5281/zenodo.5185231" ID-GBIF-Taxon="183927156" ID-Zenodo-Dep="5185231" LSID="urn:lsid:zoobank.org:act:37859D6E-BB61-4DCE-A994-380E12998A0C" httpUri="http://treatment.plazi.org/id/03948B2CFFF6FFF8388D683CFAC465AE" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
<subSubSection id="C32769B1FFF6FFFF388D683CFC84660D" box="[192,827,257,282]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="8B823A3AFFF6FFFF388D683CFC84660D" blockId="3.[192,827,257,282]" box="[192,827,257,282]" pageId="3" pageNumber="2">
<heading id="D0CA8D56FFF6FFFF388D683CFC84660D" bold="true" box="[192,827,257,282]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
<emphasis id="B949E628FFF6FFFF388D683CFC84660D" bold="true" box="[192,827,257,282]" pageId="3" pageNumber="2">
<taxonomicName id="4C3D41B9FFF6FFFF388D683CFD37660E" ID-CoL="3695S" authority="Paulsen" authorityName="Paulsen" authorityYear="2016" box="[192,648,257,282]" class="Insecta" family="Lucanidae" genus="Dinonigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="bartolozzii" status="sp. nov.">
<emphasis id="B949E628FFF6FFFF388D683CFDAF660D" bold="true" box="[192,528,257,281]" italics="true" pageId="3" pageNumber="2">Dinonigidius bartolozzii</emphasis>
Paulsen
</taxonomicName>
,
<taxonomicNameLabel id="A27A5B53FFF6FFFF3ADB683FFC84660D" box="[662,827,257,282]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32769B1FFF6FFFF388D687CFAB565EE" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="8B823A3AFFF6FFFF388D687CFAB565EE" blockId="3.[192,1441,321,762]" pageId="3" pageNumber="2">
<emphasis id="B949E628FFF6FFFF388D687CFE62664D" bold="true" box="[192,477,321,345]" pageId="3" pageNumber="2">Materials Examined.</emphasis>
<materialsCitation id="3B553067FFF6FFFF39AE687FFA23666E" ID-GBIF-Occurrence="3332178305" collectingDate="1990-01-01" collectionCode="MZUF" collectorName="Rautenstrauch" country="Sri Lanka" location="Kandy" pageId="3" pageNumber="2" specimenCount="2" typeStatus="holotype">
<typeStatus id="54868498FFF6FFFF39AE687FFDED664E" box="[483,594,322,346]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
male, deposited at
<collectionCode id="ED2CA2FFFFF6FFFF3B77687FFC31664D" box="[826,910,322,345]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="3" pageNumber="2">MZUF</collectionCode>
, labeled: a) “
<collectingCountry id="F32A7AAAFFF6FFFF3C6B687FFB01664E" box="[1062,1214,322,346]" name="Sri Lanka" pageId="3" pageNumber="2">SRI LANKA</collectingCountry>
/
<location id="8EE26CE1FFF6FFFF3C9D687FFA9B664E" LSID="urn:lsid:plazi:treatment:03948B2CFFF6FFF8388D683CFAC465AE:8EE26CE1FFF6FFFF3C9D687FFA9B664E" box="[1232,1316,322,346]" country="Sri Lanka" name="Kandy" pageId="3" pageNumber="2">Kandy</location>
,
<date id="FF831CFAFFF6FFFF3D62687FFA2C664E" box="[1327,1427,322,346]" pageId="3" pageNumber="2" value="1990-01-01">
<collectingDate id="EFC7E512FFF6FFFF3D62687FFA2C664E" box="[1327,1427,322,346]" pageId="3" pageNumber="2" value="1990-01-01">1.I.1990</collectingDate>
</date>
/
<collectorName id="26C85FECFFF6FFFF388D685FFEC8666E" box="[192,375,354,378]" pageId="3" pageNumber="2">Rautenstrauch</collectorName>
leg.”; b) on red paper, “
<taxonomicName id="4C3D41B9FFF6FFFF3AC7685FFC95666E" authorityName="de Lisle" authorityYear="1974" box="[650,810,354,378]" class="Insecta" family="Lucanidae" genus="Dinonigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B949E628FFF6FFFF3AC7685FFC95666E" box="[650,810,354,378]" italics="true" pageId="3" pageNumber="2">Dinonigidius</emphasis>
</taxonomicName>
/
<taxonomicName id="4C3D41B9FFF6FFFF3B70685FFC01666E" authorityName="Paulsen" authorityYear="2016" box="[829,958,354,378]" class="Insecta" family="Lucanidae" genus="Dinonigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="bartolozzii">
<emphasis id="B949E628FFF6FFFF3B70685FFC01666E" box="[829,958,354,378]" italics="true" pageId="3" pageNumber="2">bartolozzii</emphasis>
</taxonomicName>
/ [male symbol] Paulsen /
<typeStatus id="54868498FFF6FFFF3CB9685FFA2C666E" box="[1268,1427,354,378]" pageId="3" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
”
</materialsCitation>
.
<materialsCitation id="3B553067FFF6FFFF388D68BFFC2D668E" ID-GBIF-Occurrence="3332178301" box="[192,914,386,410]" collectionCode="LBC" country="Sri Lanka" pageId="3" pageNumber="2" specimenCount="4" specimenCount-female="1" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="54868498FFF6FFFF388D68BFFE82668E" box="[192,317,386,410]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D3BF1B3FFF6FFFF390268BFFE1A668E" box="[335,421,386,410]" pageId="3" pageNumber="2" type="male">3 male</specimenCount>
,
<specimenCount id="9D3BF1B3FFF6FFFF39FE68BFFD9F668E" box="[435,544,386,410]" pageId="3" pageNumber="2" type="female">1 female</specimenCount>
;
<collectionCode id="ED2CA2FFFFF6FFFF3A6068BFFDD9668E" box="[557,614,386,410]" country="Philippines" lsid="urn:lsid:biocol.org:col:14633" name="University of the Philippines at Los Baños" pageId="3" pageNumber="2" type="Herbarium">LBC</collectionCode>
) labeled: a) as
<typeStatus id="54868498FFF6FFFF3B6868BFFC2D668E" box="[805,914,386,410]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
</materialsCitation>
.
<materialsCitation id="3B553067FFF6FFFF3BD368BFFB3B66AE" ID-GBIF-Occurrence="3332178303" collectingDate="1990-01-01" collectionCode="NHM, LBC, PGC, GBC" country="Sri Lanka" location="Kandy" municipality="Rautenstrauch" pageId="3" pageNumber="2" specimenCount="8" specimenCount-female="3" specimenCount-male="5" typeStatus="paratype">
<typeStatus id="54868498FFF6FFFF3BD368BFFBA4668E" box="[926,1051,386,410]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D3BF1B3FFF6FFFF3C6068BCFB3C668E" box="[1069,1155,385,410]" pageId="3" pageNumber="2" type="male">5 male</specimenCount>
,
<specimenCount id="9D3BF1B3FFF6FFFF3CDD68BFFB42668E" box="[1168,1277,386,410]" pageId="3" pageNumber="2" type="female">3 female</specimenCount>
;
<collectionCode id="ED2CA2FFFFF6FFFF3D4668BFFAEC668D" box="[1291,1363,386,409]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="3" pageNumber="2" type="Herbarium">NHM</collectionCode>
,
<collectionCode id="ED2CA2FFFFF6FFFF3D2D68BFFA24668E" box="[1376,1435,386,410]" country="Philippines" lsid="urn:lsid:biocol.org:col:14633" name="University of the Philippines at Los Baños" pageId="3" pageNumber="2" type="Herbarium">LBC</collectionCode>
,
<collectionCode id="ED2CA2FFFFF6FFFF388D689FFF4266AE" box="[192,253,418,442]" country="Russia" httpUri="http://grbio.org/cool/f49n-4xcd" name="Peterhof Genetic Collection of Microalgae" pageId="3" pageNumber="2">PGC</collectionCode>
,
<collectionCode id="ED2CA2FFFFF6FFFF3947689FFEF966AE" box="[266,326,418,442]" pageId="3" pageNumber="2">GBC</collectionCode>
) labeled: a) “
<collectingCountry id="F32A7AAAFFF6FFFF39A1689FFD3866AE" box="[492,647,418,442]" name="Sri Lanka" pageId="3" pageNumber="2">SRI LANKA</collectingCountry>
/
<location id="8EE26CE1FFF6FFFF3AEC689FFD4D66AE" LSID="urn:lsid:plazi:treatment:03948B2CFFF6FFF8388D683CFAC465AE:8EE26CE1FFF6FFFF3AEC689FFD4D66AE" box="[673,754,418,442]" country="Sri Lanka" municipality="Rautenstrauch" name="Kandy" pageId="3" pageNumber="2">Kandy</location>
/
<date id="FF831CFAFFF6FFFF3B41689FFCCF66AE" box="[780,880,418,442]" pageId="3" pageNumber="2" value="1990-01-01">
<collectingDate id="EFC7E512FFF6FFFF3B41689FFCCF66AE" box="[780,880,418,442]" pageId="3" pageNumber="2" value="1990-01-01">1.I.1990</collectingDate>
</date>
leg. /
<collectingMunicipality id="6BE6A040FFF6FFFF3BF2689FFBC666AE" box="[959,1145,418,442]" pageId="3" pageNumber="2">Rautenstrauch</collectingMunicipality>
”
</materialsCitation>
;
<collectionCode id="ED2CA2FFFFF6FFFF3CDF689FFB6866AD" box="[1170,1239,418,441]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="3" pageNumber="2" type="Herbarium">NHM</collectionCode>
specimens with additional handwritten labels “12954-a”, “12954-b”, “17540a”, and “17540b”.
<typeStatus id="54868498FFF6FFFF3C3268FFFB4F66CE" box="[1151,1264,450,474]" pageId="3" pageNumber="2" type="paratype">Paratype</typeStatus>
female (
<collectionCode id="ED2CA2FFFFF6FFFF3D1168FFFA2766CE" box="[1372,1432,450,474]" country="Russia" httpUri="http://grbio.org/cool/f49n-4xcd" name="Peterhof Genetic Collection of Microalgae" pageId="3" pageNumber="2">PGC</collectionCode>
) labeled: a) “
<collectingCountry id="F32A7AAAFFF6FFFF391D68DFFE5666EE" box="[336,489,482,506]" name="Sri Lanka" pageId="3" pageNumber="2">SRI LANKA</collectingCountry>
/ Kandy /
<date id="FF831CFAFFF6FFFF3A2868DFFD7666EE" box="[613,713,482,506]" pageId="3" pageNumber="2" value="1990-01-01">1.I.1990</date>
”.
<typeStatus id="54868498FFF6FFFF3AAC68DFFCED66EE" box="[737,850,482,506]" pageId="3" pageNumber="2" type="paratype">Paratype</typeStatus>
male (
<collectionCode id="ED2CA2FFFFF6FFFF3BE968DFFC4D66EE" box="[932,1010,482,506]" pageId="3" pageNumber="2">MJPC</collectionCode>
) labeled: a) “
<collectingCountry id="F32A7AAAFFF6FFFF3CDF68DFFA9466EE" box="[1170,1323,482,506]" name="Sri Lanka" pageId="3" pageNumber="2">SRI LANKA</collectingCountry>
/ Kandy /
<date id="FF831CFAFFF6FFFF388D6B3FFE8A650E" box="[192,309,514,538]" pageId="3" pageNumber="2" value="2001-01-12">12.I.2001</date>
”.
<materialsCitation id="3B553067FFF6FFFF39016B3FFB12650E" ID-GBIF-Occurrence="3332178302" box="[332,1197,514,538]" collectingDate="2001-01-12" collectionCode="MJPC" country="Sri Lanka" location="Kandy" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
<typeStatus id="54868498FFF6FFFF39016B3FFE02650E" box="[332,445,514,538]" pageId="3" pageNumber="2" type="paratype">Paratype</typeStatus>
female (
<collectionCode id="ED2CA2FFFFF6FFFF3A6B6B3FFDCB650E" box="[550,628,514,538]" pageId="3" pageNumber="2">MJPC</collectionCode>
) labeled: a) “
<collectingCountry id="F32A7AAAFFF6FFFF3B5B6B3FFC10650E" box="[790,943,514,538]" name="Sri Lanka" pageId="3" pageNumber="2">SRI LANKA</collectingCountry>
/
<location id="8EE26CE1FFF6FFFF3B886B3FFBA9650E" LSID="urn:lsid:plazi:treatment:03948B2CFFF6FFF8388D683CFAC465AE:8EE26CE1FFF6FFFF3B886B3FFBA9650E" box="[965,1046,514,538]" country="Sri Lanka" name="Kandy" pageId="3" pageNumber="2">Kandy</location>
/
<date id="FF831CFAFFF6FFFF3C606B3FFB1E650E" box="[1069,1185,514,538]" pageId="3" pageNumber="2" value="2001-01-12">
<collectingDate id="EFC7E512FFF6FFFF3C606B3FFB1E650E" box="[1069,1185,514,538]" pageId="3" pageNumber="2" value="2001-01-12">12.I.2001</collectingDate>
</date>
”
</materialsCitation>
; b) on orange paper “DNA VOUCHER / P466 2014 / MJ Paulsen –
<collectionCode id="ED2CA2FFFFF6FFFF3B4B6B1FFCE0652E" box="[774,863,546,570]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="3" pageNumber="2">UNSM</collectionCode>
”.
<materialsCitation id="3B553067FFF6FFFF3B3B6B1FFDBF654D" ID-GBIF-Occurrence="3332178306" collectingDate="1995-02" collectionCode="MJPC" country="Sri Lanka" location="SRI LANKA" pageId="3" pageNumber="2" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54868498FFF6FFFF3B3B6B1FFC4C652E" box="[886,1011,546,570]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D3BF1B3FFF6FFFF3C4B6B1FFBE5652E" box="[1030,1114,546,570]" pageId="3" pageNumber="2" type="male">1 male</specimenCount>
,
<specimenCount id="9D3BF1B3FFF6FFFF3C256B1FFB6B652E" box="[1128,1236,546,570]" pageId="3" pageNumber="2" type="female">1 female</specimenCount>
;
<collectionCode id="ED2CA2FFFFF6FFFF3CAF6B1FFA8F652E" box="[1250,1328,546,570]" pageId="3" pageNumber="2">MJPC</collectionCode>
) labeled: a) “
<collectingCountry id="F32A7AAAFFF6FFFF38A76B7FFE3C654E" box="[234,387,578,602]" name="Sri Lanka" pageId="3" pageNumber="2">SRI LANKA</collectingCountry>
/
<date id="FF831CFAFFF6FFFF39D76B7CFE4B654D" box="[410,500,577,601]" pageId="3" pageNumber="2" value="1995-02">
<collectingDate id="EFC7E512FFF6FFFF39D76B7CFE4B654D" box="[410,500,577,601]" pageId="3" pageNumber="2" value="1995-02">II.1995</collectingDate>
</date>
”
</materialsCitation>
.
<materialsCitation id="3B553067FFF6FFFF3A416B7FFE76656E" ID-GBIF-Occurrence="3332178304" collectionCode="NHM, MJPC" collectorName="A. Maier" country="Sri Lanka" location="Sri Lanka" pageId="3" pageNumber="2" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="54868498FFF6FFFF3A416B7FFD36654E" box="[524,649,578,602]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D3BF1B3FFF6FFFF3AD76B7FFD50654E" box="[666,751,578,602]" pageId="3" pageNumber="2" type="male">3 male</specimenCount>
,
<specimenCount id="9D3BF1B3FFF6FFFF3AB66B7FFCD8654E" box="[763,871,578,602]" pageId="3" pageNumber="2" type="female">2 female</specimenCount>
;
<collectionCode id="ED2CA2FFFFF6FFFF3B3E6B7FFC04654D" box="[883,955,578,601]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="3" pageNumber="2" type="Herbarium">NHM</collectionCode>
,
<collectionCode id="ED2CA2FFFFF6FFFF3B856B7FFBA9654E" box="[968,1046,578,602]" pageId="3" pageNumber="2">MJPC</collectionCode>
) labeled: a)
<collectingCountry id="F32A7AAAFFF6FFFF3CE16B7FFA96654E" box="[1196,1321,578,602]" name="Sri Lanka" pageId="3" pageNumber="2">Sri Lanka</collectingCountry>
/ Kandy / 5.1994 / leg.
<collectorName id="26C85FECFFF6FFFF39176B5FFE76656E" box="[346,457,610,634]" pageId="3" pageNumber="2">A. Maier</collectorName>
</materialsCitation>
”;
<collectionCode id="ED2CA2FFFFF6FFFF39926B5FFD9B656D" box="[479,548,610,633]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="3" pageNumber="2" type="Herbarium">NHM</collectionCode>
specimens with additional handwritten labels “18129-a”, “18129-b”, and “18129-c”.
<typeStatus id="54868498FFF6FFFF39086BBFFE09658E" box="[325,438,642,666]" pageId="3" pageNumber="2" type="paratype">Paratype</typeStatus>
male (
<collectionCode id="ED2CA2FFFFF6FFFF3A406BBFFDE7658E" box="[525,600,642,666]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="3" pageNumber="2" type="Museum">FSCA</collectionCode>
) labeled: a) “
<collectingCountry id="F32A7AAAFFF6FFFF3AB26BBFFC24658E" box="[767,923,642,666]" name="Sri Lanka" pageId="3" pageNumber="2">SRI LANKA</collectingCountry>
/ Kandy /
<date id="FF831CFAFFF6FFFF3C6F6BBFFB6E658E" box="[1058,1233,642,666]" pageId="3" pageNumber="2" value="1999-01">January 1999</date>
”.
<typeStatus id="54868498FFF6FFFF3CA46BBFFAE5658E" box="[1257,1370,642,666]" pageId="3" pageNumber="2" type="paratype">Paratype</typeStatus>
male (
<collectionCode id="ED2CA2FFFFF6FFFF38856B9FFEA965AE" box="[200,278,674,698]" pageId="3" pageNumber="2">MJPC</collectionCode>
) labeled: a) “
<collectingCountry id="F32A7AAAFFF6FFFF39F56B9FFDEE65AE" box="[440,593,674,698]" name="Sri Lanka" pageId="3" pageNumber="2">SRI LANKA</collectingCountry>
/
<collectingRegion id="49F9F4D8FFF6FFFF3A2A6B9FFC8765AE" box="[615,824,674,698]" country="Sri Lanka" name="Central" pageId="3" pageNumber="2">Central Province</collectingRegion>
/ Kandy District /
<date id="FF831CFAFFF6FFFF3C516B9FFB6A65AD" box="[1052,1237,673,698]" pageId="3" pageNumber="2" value="1995-02">February 1995</date>
”.
<typeStatus id="54868498FFF6FFFF3CA16B9FFAE265AE" box="[1260,1373,674,698]" pageId="3" pageNumber="2" type="paratype">Paratype</typeStatus>
male (
<collectionCode id="ED2CA2FFFFF6FFFF38856BFFFEBF65CE" box="[200,256,706,730]" country="Spain" lsid="urn:lsid:biocol.org:col:14286" name="Universidad de La Laguna" pageId="3" pageNumber="2" type="Herbarium">TFC</collectionCode>
) with handwritten label: a) “Kandy /
<collectingCountry id="F32A7AAAFFF6FFFF3A856BFFFCFB65CE" box="[712,836,706,730]" name="Sri Lanka" pageId="3" pageNumber="2">Sri Lanka</collectingCountry>
/ 14.05.94”. All
<typeStatus id="54868498FFF6FFFF3C496BFFFBC165CE" box="[1028,1150,706,730]" pageId="3" pageNumber="2" type="paratype">paratypes</typeStatus>
with
<typeStatus id="54868498FFF6FFFF3C896BFFFA8D65CE" box="[1220,1330,706,730]" pageId="3" pageNumber="2" type="paratype">paratype</typeStatus>
label, on yellow paper: “
<taxonomicName id="4C3D41B9FFF6FFFF393B6BDFFDA865EE" authorityName="de Lisle" authorityYear="1974" box="[374,535,738,762]" class="Insecta" family="Lucanidae" genus="Dinonigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B949E628FFF6FFFF393B6BDFFDA865EE" box="[374,535,738,762]" italics="true" pageId="3" pageNumber="2">Dinonigidius</emphasis>
</taxonomicName>
/
<taxonomicName id="4C3D41B9FFF6FFFF3A606BDFFD1165EE" authorityName="Paulsen" authorityYear="2016" box="[557,686,738,762]" class="Insecta" family="Lucanidae" genus="Dinonigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="bartolozzii">
<emphasis id="B949E628FFF6FFFF3A606BDFFD1165EE" box="[557,686,738,762]" italics="true" pageId="3" pageNumber="2">bartolozzii</emphasis>
</taxonomicName>
/ [male or female symbol] Paulsen /
<typeStatus id="54868498FFF6FFFF3C216BDFFABA65EE" box="[1132,1285,738,762]" pageId="3" pageNumber="2" type="paratype">PARATYPE</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C32769B1FFF6FFFF388D6A1CFA92602E" pageId="3" pageNumber="2" type="description">
<paragraph id="8B823A3AFFF6FFFF388D6A1CFD73610D" blockId="3.[192,1441,801,1562]" pageId="3" pageNumber="2">
<emphasis id="B949E628FFF6FFFF388D6A1CFDE6642D" bold="true" box="[192,601,801,826]" pageId="3" pageNumber="2">
Description,
<typeStatus id="54868498FFF6FFFF393F6A1FFE54642E" box="[370,491,802,826]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
(
<figureCitation id="130626BFFFF6FFFF39B66A1CFDF8642D" box="[507,583,801,825]" captionStart="Figures 1–4" captionStartId="5.[192,288,1479,1501]" captionTargetBox="[382,1260,205,1450]" captionTargetId="figure-41@5.[376,1279,192,1464]" captionTargetPageId="5" captionText="Figures 1–4. Dorsal habitus and male genitalia of Dinonigidius species, ventral view. Arrows indicate median lobe. 1, 3) D. bartolozzii Paulsen, n.sp. 2, 4) D. ahenobarbus de Lisle." figureDoi="http://doi.org/10.5281/zenodo.5170625" httpUri="https://zenodo.org/record/5170625/files/figure.png" pageId="3" pageNumber="2">Fig. 1</figureCitation>
).
</emphasis>
<taxonomicName id="4C3D41B9FFF6FFFF3A2D6A1FFD56642E" authorityName="Linnaeus" authorityYear="1758" box="[608,745,802,826]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
: Scarabaeoidea:
<taxonomicName id="4C3D41B9FFF6FFFF3BF86A1FFB85642E" authorityName="Latreille" authorityYear="1804" box="[949,1082,802,826]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="family">Lucanidae</taxonomicName>
.
<emphasis id="B949E628FFF6FFFF3C096A1FFB1D642E" box="[1092,1186,802,826]" italics="true" pageId="3" pageNumber="2">Length:</emphasis>
19.0 mm.
<emphasis id="B949E628FFF6FFFF3D6C6A1FFACB642E" box="[1313,1396,802,826]" italics="true" pageId="3" pageNumber="2">Width:</emphasis>
7.0 mm (elytra).
<emphasis id="B949E628FFF6FFFF39136A7FFE18644E" box="[350,423,834,858]" italics="true" pageId="3" pageNumber="2">Color:</emphasis>
Piceous, with clypeus, mandibles, legs, and venter lighter red.
<emphasis id="B949E628FFF6FFFF3CEF6A7FFB54644E" box="[1186,1259,834,858]" italics="true" pageId="3" pageNumber="2">Head:</emphasis>
Surface shiny, punctate, punctures with short setae; punctures coarse and dense (distance between punctures about 1 diameter) on frons, fine and sparse (punctures separated by distinctly more than 1 diameter) on clypeus. Clypeus declivous, narrowing between mandibles to subtruncate apex; apex subdentate medially. Eyes completely divided by ocular canthus; canthus rounded, narrow and uniform in width, reaching temporal process at distinct suture. Temporal process large, projecting posteriorly as a triangular lobe; lobe divided into anterolateral and posterior faces by an impunctate ridge; both faces coarsely, contiguously punctate. Antennal club small, short (shorter than scape), antennomeres 8–10 (club) entirely tomentose. Mandibles (to true apex) shorter than head, apex acute, internally tridentate (right) or quadridentate (left), teeth abraded; dorsal surface produced into long, vertical curved ramus (false apex); ramus with apex obtuse. Mentum broad, three times wider than long, bilobed, surface coarsely but irregularly densely punctate.
<emphasis id="B949E628FFF6FFFF3A116D9FFD6163AD" box="[604,734,1186,1209]" italics="true" pageId="3" pageNumber="2">Pronotum:</emphasis>
Form short, as wide as elytra, with anterior tumosity but lacking anteriorly bifurcate projection. Anterior angles emarginate. Surface variable, disc sparsely punctate, area laterad of disc with areas of coarse, dense to contiguous punctures.
<emphasis id="B949E628FFF6FFFF3CDA6DDFFB5363EE" box="[1175,1260,1250,1274]" italics="true" pageId="3" pageNumber="2">Elytra:</emphasis>
Form parallelsided. Surface striate; striae impressed with coarse, oval, almost contiguous punctures; striae distinct on disc, becoming obsolete laterally; interstriae 1–3 with fine (smaller than strial punctures) punctures in 2–3 irregular rows; interstriae 4 and greater with punctures subequal in size to strial punctures, obscuring striae.
<emphasis id="B949E628FFF6FFFF39D86C5FFE6A626D" box="[405,469,1378,1401]" italics="true" pageId="3" pageNumber="2">Legs:</emphasis>
Protibia with acute apex and with 5 acute teeth decreasing in size proximally. Mesotibiae with 1 large external tooth below middle with 1–2 smaller accessory teeth proximally. Metatibiae with 1 small tooth medially.
<emphasis id="B949E628FFF6FFFF3AE36C9FFCB162AE" box="[686,782,1442,1466]" italics="true" pageId="3" pageNumber="2">Thorax:</emphasis>
Prosternal process behind procoxae strongly pointed. Metasternum entirely punctate, lacking glabrous area anteriorly just behind metacoxae.
<emphasis id="B949E628FFF6FFFF3D6B6CFFFA1F62CE" box="[1318,1440,1474,1498]" italics="true" pageId="3" pageNumber="2">Abdomen:</emphasis>
Male genitalia as in (
<figureCitation id="130626BFFFF6FFFF39896CDFFDB462EE" box="[452,523,1506,1530]" captionStart="Figures 1–4" captionStartId="5.[192,288,1479,1501]" captionTargetBox="[382,1260,205,1450]" captionTargetId="figure-41@5.[376,1279,192,1464]" captionTargetPageId="5" captionText="Figures 1–4. Dorsal habitus and male genitalia of Dinonigidius species, ventral view. Arrows indicate median lobe. 1, 3) D. bartolozzii Paulsen, n.sp. 2, 4) D. ahenobarbus de Lisle." figureDoi="http://doi.org/10.5281/zenodo.5170625" httpUri="https://zenodo.org/record/5170625/files/figure.png" pageId="3" pageNumber="2">Fig. 2</figureCitation>
), flagellum asymmetrical and curved at apex, shorter than basal piece and parameres combined; median lobe narrow.
</paragraph>
<paragraph id="8B823A3AFFF6FFFF388D6F7FFDD9616E" blockId="3.[192,1440,1601,1658]" pageId="3" pageNumber="2">
<emphasis id="B949E628FFF6FFFF388D6F7FFD8C614E" bold="true" box="[192,563,1601,1626]" pageId="3" pageNumber="2">
<typeStatus id="54868498FFF6FFFF388D6F7FFE80614E" box="[192,319,1602,1626]" pageId="3" pageNumber="2" type="paratype">Paratype</typeStatus>
variation, males.
</emphasis>
Length:
<quantity id="4CC597DFFFF6FFFF3AEB6F7CFCE6614D" box="[678,857,1601,1625]" metricMagnitude="-2" metricUnit="m" metricValue="1.845" metricValueMax="2.1399999999999997" metricValueMin="1.55" pageId="3" pageNumber="2" unit="mm" value="18.45" valueMax="21.4" valueMin="15.5">15.5–21.4 mm</quantity>
. Width: 6.0–
<quantity id="4CC597DFFFF6FFFF3BB56F7FFBE5614D" box="[1016,1114,1602,1626]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="3" pageNumber="2" unit="mm" value="7.2">7.2 mm</quantity>
. Males with color varying from red (teneral) to almost black.
</paragraph>
<paragraph id="8B823A3AFFF6FFFF388D6F9FFA92602E" blockId="3.[192,1440,1697,1850]" pageId="3" pageNumber="2">
<emphasis id="B949E628FFF6FFFF388D6F9FFDFB61AD" bold="true" box="[192,580,1697,1722]" pageId="3" pageNumber="2">
<typeStatus id="54868498FFF6FFFF388D6F9FFE8061AE" box="[192,319,1698,1722]" pageId="3" pageNumber="2" type="paratype">Paratype</typeStatus>
variation, females.
</emphasis>
Length:
<quantity id="4CC597DFFFF6FFFF3AFD6F9FFCE061AD" box="[688,863,1698,1722]" metricMagnitude="-2" metricUnit="m" metricValue="1.6649999999999998" metricValueMax="1.86" metricValueMin="1.47" pageId="3" pageNumber="2" unit="mm" value="16.65" valueMax="18.6" valueMin="14.7">14.7–18.6 mm</quantity>
. Width:
<quantity id="4CC597DFFFF6FFFF3B8F6F9CFBEB61AD" box="[962,1108,1697,1721]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="7.2" metricValueMin="5.9" pageId="3" pageNumber="2" unit="mm" value="6.55" valueMax="7.2" valueMin="5.9">5.9–7.2 mm</quantity>
. Females with simple mandibles lacking vertical ramus of male or any vertical development (small dorsal projection at base present in
<taxonomicName id="4C3D41B9FFF6FFFF39416FDFFE7261EE" box="[268,461,1762,1786]" class="Insecta" family="Lucanidae" genus="Dinonigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="ahenobarbus">
<emphasis id="B949E628FFF6FFFF39416FDFFE7261EE" box="[268,461,1762,1786]" italics="true" pageId="3" pageNumber="2">D. ahenobarbus</emphasis>
</taxonomicName>
); apex acute, internally with 1–2 teeth. Females with anterior pronotal tumosity weakly indicated, lateral contiguously punctate areas subfoveate. Females with temporal process not strongly produced posteriorly, instead narrow, rounded and similar in shape to the canthus.
</paragraph>
</subSubSection>
<subSubSection id="C32769B1FFF1FFF8388D69FFFE60664E" pageId="4" pageNumber="3" type="etymology">
<paragraph id="8B823A3AFFF1FFF8388D69FFFE60664E" blockId="4.[192,1440,194,346]" pageId="4" pageNumber="3">
<emphasis id="B949E628FFF1FFF8388D69FFFEE367CE" bold="true" box="[192,348,194,218]" pageId="4" pageNumber="3">Etymology.</emphasis>
The species is named in honor of my friend and colleague Luca Bartolozzi, Natural History Museum “La Specola”, University of Florence,
<collectingCountry id="F32A7AAAFFF1FFF83B7369DFFCC467EE" box="[830,891,226,250]" name="Italy" pageId="4" pageNumber="3">Italy</collectingCountry>
, who provided an excellent treatment of the Sri Lankan stag beetle fauna with H. E. Bomans just before specimens of this species began to be discovered. His knowledge, insights, and generous loans of material have contributed greatly to my research on
<taxonomicName id="4C3D41B9FFF1FFF8391B687FFE64664E" authorityName="Latreille" authorityYear="1804" box="[342,475,322,346]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="family">Lucanidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32769B1FFF1FFF8388D68BCFD1066EE" pageId="4" pageNumber="3" type="distribution">
<paragraph id="8B823A3AFFF1FFF8388D68BCFD1066EE" blockId="4.[192,1441,385,506]" pageId="4" pageNumber="3">
<emphasis id="B949E628FFF1FFF8388D68BCFECB668D" bold="true" box="[192,372,385,409]" pageId="4" pageNumber="3">Distribution.</emphasis>
Almost all specimens known are from Kandy,
<collectingCountry id="F32A7AAAFFF1FFF83BE468BFFB97668E" box="[937,1064,386,410]" name="Sri Lanka" pageId="4" pageNumber="3">Sri Lanka</collectingCountry>
. A photograph of
<specimenCount id="9D3BF1B3FFF1FFF83CB168BFFA1E668E" box="[1276,1441,386,410]" pageId="4" pageNumber="3" type="generic">one specimen</specimenCount>
from Elpitiya in southern
<collectingCountry id="F32A7AAAFFF1FFF83A4A689FFD3A66AE" box="[519,645,418,442]" name="Sri Lanka" pageId="4" pageNumber="3">Sri Lanka</collectingCountry>
is considered to be referable to
<emphasis id="B949E628FFF1FFF83C59689FFB7D66AE" box="[1044,1218,418,442]" italics="true" pageId="4" pageNumber="3">
<taxonomicName id="4C3D41B9FFF1FFF83C59689FFB0266AE" authorityName="Paulsen" authorityYear="2016" box="[1044,1213,418,442]" class="Insecta" family="Lucanidae" genus="Dinonigidius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="bartolozzii">D. bartolozzii</taxonomicName>
,
</emphasis>
but the specimen was not physically examined and because it is from a locality distinct from the remainder of the type series it was not included as a
<typeStatus id="54868498FFF1FFF83A7468DFFD1466EE" box="[569,683,482,506]" pageId="4" pageNumber="3" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C32769B1FFF1FFF8388D6B1FFAC465AE" pageId="4" pageNumber="3" type="discussion">
<paragraph id="8B823A3AFFF1FFF8388D6B1FFAC465AE" blockId="4.[192,1440,546,698]" pageId="4" pageNumber="3">
<emphasis id="B949E628FFF1FFF8388D6B1FFEFC652E" bold="true" box="[192,323,546,570]" pageId="4" pageNumber="3">Remarks.</emphasis>
The new species differs from
<taxonomicName id="4C3D41B9FFF1FFF83AE96B1FFCDC652E" box="[676,867,546,570]" class="Insecta" family="Lucanidae" genus="Dinonigidius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="ahenobarbus">
<emphasis id="B949E628FFF1FFF83AE96B1FFCDC652E" box="[676,867,546,570]" italics="true" pageId="4" pageNumber="3">D. ahenobarbus</emphasis>
</taxonomicName>
(
<figureCitation id="130626BFFFF1FFF83B3F6B1FFC0A652E" box="[882,949,546,570]" captionStart="Figures 1–4" captionStartId="5.[192,288,1479,1501]" captionTargetBox="[382,1260,205,1450]" captionTargetId="figure-41@5.[376,1279,192,1464]" captionTargetPageId="5" captionText="Figures 1–4. Dorsal habitus and male genitalia of Dinonigidius species, ventral view. Arrows indicate median lobe. 1, 3) D. bartolozzii Paulsen, n.sp. 2, 4) D. ahenobarbus de Lisle." figureDoi="http://doi.org/10.5281/zenodo.5170625" httpUri="https://zenodo.org/record/5170625/files/figure.png" pageId="4" pageNumber="3">Fig. 3</figureCitation>
) in the following ways: anterior pronotal process lacking in males; male genitalia with a shorter, asymmetrical flagellum and much narrower median lobe (flagellum longer than parameres + basal piece, median lobe broad in
<taxonomicName id="4C3D41B9FFF1FFF83CEF6B5FFADD656E" box="[1186,1378,610,634]" class="Insecta" family="Lucanidae" genus="Dinonigidius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="ahenobarbus">
<emphasis id="B949E628FFF1FFF83CEF6B5FFADD656E" box="[1186,1378,610,634]" italics="true" pageId="4" pageNumber="3">D. ahenobarbus</emphasis>
</taxonomicName>
(
<figureCitation id="130626BFFFF1FFF83D3D6B5FFF70658E" captionStart="Figures 1–4" captionStartId="5.[192,288,1479,1501]" captionTargetBox="[382,1260,205,1450]" captionTargetId="figure-41@5.[376,1279,192,1464]" captionTargetPageId="5" captionText="Figures 1–4. Dorsal habitus and male genitalia of Dinonigidius species, ventral view. Arrows indicate median lobe. 1, 3) D. bartolozzii Paulsen, n.sp. 2, 4) D. ahenobarbus de Lisle." figureDoi="http://doi.org/10.5281/zenodo.5170625" httpUri="https://zenodo.org/record/5170625/files/figure.png" pageId="4" pageNumber="3">Fig. 4</figureCitation>
)); prosternal process acute in both sexes, not bluntly rounded; metasternum lacking anterior glabrous area in both sexes; temporal process in males with posterior face distinctly punctate, not glabrous.
</paragraph>
</subSubSection>
</treatment>
</document>
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