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<document id="0AB8DE4A4D595984258D9A4E346C1035" ID-CLB-Dataset="53812" ID-DOI="10.11646/zootaxa.5250.1.1" ID-GBIF-Dataset="da8acfed-df61-403e-9a49-7f856ce56196" ID-ISSN="1175­5334" ID-Zenodo-Dep="7705657" ID-ZooBank="4729EAF3-8B78-4AAD-B373-9179D008F239" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1678178160677" checkinUser="plazi" docAuthor="WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI &amp; HUANG, JIA" docDate="2023" docId="0393914C2A0D8757C282FD7269B0FEA6" docLanguage="en" docName="zootaxa.5250.1.1.pdf" docOrigin="Zootaxa 5250 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5250.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Stegana (Steganina) nigripalpula Cui &amp; Chen, sp. nov." docType="treatment" docVersion="8" lastPageNumber="48" masterDocId="FFAAE9342A228778C215FF896958FB43" masterDocTitle="Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)" masterLastPageNumber="109" masterPageNumber="1" pageNumber="48" updateTime="1699476226253" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3053D1E972D643BE0A3349D9CE1CEFA8">Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)</mods:title>
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<treatment id="0393914C2A0D8757C282FD7269B0FEA6" ID-DOI="http://doi.org/10.5281/zenodo.7705537" ID-GBIF-Taxon="209579477" ID-Zenodo-Dep="7705537" LSID="urn:lsid:plazi:treatment:0393914C2A0D8757C282FD7269B0FEA6" httpUri="http://treatment.plazi.org/id/0393914C2A0D8757C282FD7269B0FEA6" lastPageNumber="48" pageId="47" pageNumber="48">
<subSubSection id="C32073D12A0D8757C282FD726A6FF856" box="[151,823,762,789]" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="8B85205A2A0D8757C282FD726A6FF856" blockId="47.[151,823,762,825]" box="[151,823,762,789]" pageId="47" pageNumber="48">
<heading id="D0CD97362A0D8757C282FD726A6FF856" bold="true" box="[151,823,762,789]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
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<taxonomicName id="4C3A5BD92A0D8757C282FD726B93F857" ID-CoL="BDMSD" authority="Cui &amp; Chen" authorityName="Cui &amp; Chen" box="[151,715,762,789]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="nigripalpula" status="sp. nov." subGenus="Steganina">
<emphasis id="B94EFC482A0D8757C282FD7269A1F856" bold="true" box="[151,249,763,789]" italics="true" pageId="47" pageNumber="48">Stegana</emphasis>
(
<emphasis id="B94EFC482A0D8757C31FFD7268DCF856" bold="true" box="[266,388,763,789]" italics="true" pageId="47" pageNumber="48">Steganina</emphasis>
)
<emphasis id="B94EFC482A0D8757C380FD736B73F856" bold="true" box="[405,555,762,789]" italics="true" pageId="47" pageNumber="48">nigripalpula</emphasis>
Cui &amp; Chen
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,
<taxonomicNameLabel id="A27D41332A0D8757C0CCFD726A6FF856" box="[729,823,763,789]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C32073D12A0D8757C282FC9668D7F87A" box="[151,399,798,825]" pageId="47" pageNumber="48" type="description">
<paragraph id="8B85205A2A0D8757C282FC9668D7F87A" blockId="47.[151,823,762,825]" box="[151,399,798,825]" pageId="47" pageNumber="48">
(
<figureCitation id="13013CDF2A0D8757C28AFC96686AF87A" box="[159,306,798,825]" captionStart="FIGURE 5" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@28.[173,1415,181,1919]" captionTargetPageId="28" captionText="FIGURE 5. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov.; C, D. S. (S.) S. (S.) nigripalpula Cui &amp; Chen, sp. nov.; E, F. S. (S.) glabra Chen &amp; Chen, 2012; G, H. S. (S.) masanoritodai Okada &amp; Sidorenko, 1992; I, J. S. (S.) multidentata Chen, Gao, &amp; Chen, 2009." figureDoi="http://doi.org/10.5281/zenodo.7705675" httpUri="https://zenodo.org/record/7705675/files/figure.png" pageId="47" pageNumber="48">Figs. 5C, 5D</figureCitation>
,
<figureCitation id="13013CDF2A0D8757C32BFC9668DFF87A" box="[318,391,799,825]" captionStart="FIGURE 20" captionStartId="45.[151,250,1851,1875]" captionTargetBox="[157,1431,931,1818]" captionTargetId="figure-260@45.[151,1436,920,1828]" captionTargetPageId="45" captionText="FIGURE 20. Female terminalia in Stegana (Steganina) nigripes Zhang &amp; Chen, 2015 (AC) from Botanic Garden, Ruili, Yunnan, S. (S.) cardua Cui &amp; Chen, sp. nov. (DF), and S. (S.) nigripalpula Cui &amp; Chen, sp. nov. (GI). A, D. G. Epiproct and cercus in dorsal view; B, E. H. Eighth sternite in ventral view; C, F, I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705713" httpUri="https://zenodo.org/record/7705713/files/figure.png" pageId="47" pageNumber="48">20GI</figureCitation>
)
</paragraph>
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<subSubSection id="C32073D12A0D8757C282FCEF6BA9F8E6" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="8B85205A2A0D8757C282FCEF6BA9F8E6" blockId="47.[151,1437,870,1509]" pageId="47" pageNumber="48">
<emphasis id="B94EFC482A0D8757C282FCEF684AF8C3" bold="true" box="[151,274,870,896]" pageId="47" pageNumber="48">Diagnosis.</emphasis>
This species differs from the other species of the
<taxonomicName id="4C3A5BD92A0D8757C151FCEE6AF6F8C3" authorityName="Wheeler &amp; Takada" authorityYear="1964" box="[836,942,871,896]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="ornatipes">
<emphasis id="B94EFC482A0D8757C151FCEE6AF6F8C3" box="[836,942,871,896]" italics="true" pageId="47" pageNumber="48">ornatipes</emphasis>
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species group in having the large and black palpus (
<figureCitation id="13013CDF2A0D8757C2E5FC026811F8E6" box="[240,329,906,933]" captionStart="FIGURE 5" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@28.[173,1415,181,1919]" captionTargetPageId="28" captionText="FIGURE 5. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov.; C, D. S. (S.) S. (S.) nigripalpula Cui &amp; Chen, sp. nov.; E, F. S. (S.) glabra Chen &amp; Chen, 2012; G, H. S. (S.) masanoritodai Okada &amp; Sidorenko, 1992; I, J. S. (S.) multidentata Chen, Gao, &amp; Chen, 2009." figureDoi="http://doi.org/10.5281/zenodo.7705675" httpUri="https://zenodo.org/record/7705675/files/figure.png" pageId="47" pageNumber="48">Fig. 5D</figureCitation>
); the first flagellomere entirely black.
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A0D8757C2D2FC276B98FE72" pageId="47" pageNumber="48" type="description">
<paragraph id="8B85205A2A0D8757C2D2FC276A2AFFE2" blockId="47.[151,1437,870,1509]" pageId="47" pageNumber="48">
<emphasis id="B94EFC482A0D8757C2D2FC276801F88B" bold="true" box="[199,345,942,968]" pageId="47" pageNumber="48">Description.</emphasis>
Female: Frons entirely black. Face black, with 1 yellow band medially; facial carina absent. Clypeus black. Mesonotum and scutellum dark brown (
<figureCitation id="13013CDF2A0D8757C117FC5A6A25F8AE" box="[770,893,978,1005]" captionStart="FIGURE 5" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@28.[173,1415,181,1919]" captionTargetPageId="28" captionText="FIGURE 5. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov.; C, D. S. (S.) S. (S.) nigripalpula Cui &amp; Chen, sp. nov.; E, F. S. (S.) glabra Chen &amp; Chen, 2012; G, H. S. (S.) masanoritodai Okada &amp; Sidorenko, 1992; I, J. S. (S.) multidentata Chen, Gao, &amp; Chen, 2009." figureDoi="http://doi.org/10.5281/zenodo.7705675" httpUri="https://zenodo.org/record/7705675/files/figure.png" pageId="47" pageNumber="48">Fig. 5C, D</figureCitation>
). Katepisternum dark brown above, yellowish to yellow below (
<figureCitation id="13013CDF2A0D8757C32AFC7E68C1FF52" box="[319,409,1014,1041]" captionStart="FIGURE 5" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@28.[173,1415,181,1919]" captionTargetPageId="28" captionText="FIGURE 5. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov.; C, D. S. (S.) S. (S.) nigripalpula Cui &amp; Chen, sp. nov.; E, F. S. (S.) glabra Chen &amp; Chen, 2012; G, H. S. (S.) masanoritodai Okada &amp; Sidorenko, 1992; I, J. S. (S.) multidentata Chen, Gao, &amp; Chen, 2009." figureDoi="http://doi.org/10.5281/zenodo.7705675" httpUri="https://zenodo.org/record/7705675/files/figure.png" pageId="47" pageNumber="48">Fig. 5D</figureCitation>
). Halter mostly black. Fore femur with 3 long setae on distal half of posteroventral surface. All abdominal tergites black medially, dark brown laterally (
<figureCitation id="13013CDF2A0D8757C12AFB926AE1FF76" box="[831,953,1050,1077]" captionStart="FIGURE 5" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-30@28.[173,1415,181,1919]" captionTargetPageId="28" captionText="FIGURE 5. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) marenubila Cui &amp; Chen, sp. nov.; C, D. S. (S.) S. (S.) nigripalpula Cui &amp; Chen, sp. nov.; E, F. S. (S.) glabra Chen &amp; Chen, 2012; G, H. S. (S.) masanoritodai Okada &amp; Sidorenko, 1992; I, J. S. (S.) multidentata Chen, Gao, &amp; Chen, 2009." figureDoi="http://doi.org/10.5281/zenodo.7705675" httpUri="https://zenodo.org/record/7705675/files/figure.png" pageId="47" pageNumber="48">Fig. 5C, D</figureCitation>
); sternites brown, about 3 times as wide as long. Female terminalia (
<figureCitation id="13013CDF2A0D8757C3A1FBB66B6EFF1A" box="[436,566,1087,1113]" captionStart="FIGURE 20" captionStartId="45.[151,250,1851,1875]" captionTargetBox="[157,1431,931,1818]" captionTargetId="figure-260@45.[151,1436,920,1828]" captionTargetPageId="45" captionText="FIGURE 20. Female terminalia in Stegana (Steganina) nigripes Zhang &amp; Chen, 2015 (AC) from Botanic Garden, Ruili, Yunnan, S. (S.) cardua Cui &amp; Chen, sp. nov. (DF), and S. (S.) nigripalpula Cui &amp; Chen, sp. nov. (GI). A, D. G. Epiproct and cercus in dorsal view; B, E. H. Eighth sternite in ventral view; C, F, I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705713" httpUri="https://zenodo.org/record/7705713/files/figure.png" pageId="47" pageNumber="48">Fig. 20GI</figureCitation>
): Epiproct and cercus lacking pubescence (
<figureCitation id="13013CDF2A0D8757C608FBB66DDFFF1A" box="[1053,1159,1087,1113]" captionStart="FIGURE 20" captionStartId="45.[151,250,1851,1875]" captionTargetBox="[157,1431,931,1818]" captionTargetId="figure-260@45.[151,1436,920,1828]" captionTargetPageId="45" captionText="FIGURE 20. Female terminalia in Stegana (Steganina) nigripes Zhang &amp; Chen, 2015 (AC) from Botanic Garden, Ruili, Yunnan, S. (S.) cardua Cui &amp; Chen, sp. nov. (DF), and S. (S.) nigripalpula Cui &amp; Chen, sp. nov. (GI). A, D. G. Epiproct and cercus in dorsal view; B, E. H. Eighth sternite in ventral view; C, F, I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705713" httpUri="https://zenodo.org/record/7705713/files/figure.png" pageId="47" pageNumber="48">Fig. 20G</figureCitation>
). Eighth sternite about 2 times as wide as long, glabrous on basal 2/3 portion, with numerous prensisetae near posterior margin (
<figureCitation id="13013CDF2A0D8757C732FBEA6CC8FF3E" box="[1319,1424,1123,1149]" captionStart="FIGURE 20" captionStartId="45.[151,250,1851,1875]" captionTargetBox="[157,1431,931,1818]" captionTargetId="figure-260@45.[151,1436,920,1828]" captionTargetPageId="45" captionText="FIGURE 20. Female terminalia in Stegana (Steganina) nigripes Zhang &amp; Chen, 2015 (AC) from Botanic Garden, Ruili, Yunnan, S. (S.) cardua Cui &amp; Chen, sp. nov. (DF), and S. (S.) nigripalpula Cui &amp; Chen, sp. nov. (GI). A, D. G. Epiproct and cercus in dorsal view; B, E. H. Eighth sternite in ventral view; C, F, I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705713" httpUri="https://zenodo.org/record/7705713/files/figure.png" pageId="47" pageNumber="48">Fig. 20H</figureCitation>
). Hypoproct with numerous setae and much pubescence (
<figureCitation id="13013CDF2A0D8757C11DFB0E6A3EFFE2" box="[776,870,1159,1185]" captionStart="FIGURE 20" captionStartId="45.[151,250,1851,1875]" captionTargetBox="[157,1431,931,1818]" captionTargetId="figure-260@45.[151,1436,920,1828]" captionTargetPageId="45" captionText="FIGURE 20. Female terminalia in Stegana (Steganina) nigripes Zhang &amp; Chen, 2015 (AC) from Botanic Garden, Ruili, Yunnan, S. (S.) cardua Cui &amp; Chen, sp. nov. (DF), and S. (S.) nigripalpula Cui &amp; Chen, sp. nov. (GI). A, D. G. Epiproct and cercus in dorsal view; B, E. H. Eighth sternite in ventral view; C, F, I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705713" httpUri="https://zenodo.org/record/7705713/files/figure.png" pageId="47" pageNumber="48">Fig. 20I</figureCitation>
).
</paragraph>
<paragraph id="8B85205A2A0D8757C2D2FB226B98FE72" blockId="47.[151,1437,870,1509]" pageId="47" pageNumber="48">
<emphasis id="B94EFC482A0D8757C2D2FB226B52FF87" bold="true" box="[199,522,1194,1221]" pageId="47" pageNumber="48">Measurements and indices.</emphasis>
BL =
<quantity id="4CC28DBF2A0D8757C045FB236BEBFF86" box="[592,691,1194,1221]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" pageId="47" pageNumber="48" unit="mm" value="2.65">2.65 mm</quantity>
in
<typeStatus id="54819EF82A0D8757C0C0FB226A62FF86" box="[725,826,1195,1221]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
, ThL =
<quantity id="4CC28DBF2A0D8757C185FB226AAEFF86" box="[912,1014,1195,1221]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="47" pageNumber="48" unit="mm" value="1.33">1.33 mm</quantity>
, WL =
<quantity id="4CC28DBF2A0D8757C652FB226DF5FF86" box="[1095,1197,1195,1221]" metricMagnitude="-3" metricUnit="m" metricValue="2.61" pageId="47" pageNumber="48" unit="mm" value="2.61">2.61 mm</quantity>
, WW =
<quantity id="4CC28DBF2A0D8757C71DFB226C36FF86" box="[1288,1390,1195,1221]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" pageId="47" pageNumber="48" unit="mm" value="1.09">1.09 mm</quantity>
, arb = 8/6, avd = 0.73, adf = 1.56, flw = 2.83, FW/HW = 0.35, ch/o = 0.11, prorb = 0.95, rcorb = 0.74, vb = 0.57, dcl = 0.59, presctl = 0.67, sctl = damaged, sterno = 0.62, orbito = 1.62, dcp = 0.27, sctlp = 1.46, C = 1.79, 4c = 1.14, 4v = 1.45, 5x = 1.11, ac = 8.00, M = 0.61, C3F = 0.62.
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A0D8757C2D2FAB26A87FE3A" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8B85205A2A0D8757C2D2FAB26A87FE3A" blockId="47.[151,1437,870,1509]" pageId="47" pageNumber="48">
<emphasis id="B94EFC482A0D8757C2D2FAB2682CFE17" bold="true" box="[199,372,1338,1365]" pageId="47" pageNumber="48">Type material.</emphasis>
<materialsCitation id="3B522A072A0D8757C36FFAB26A87FE3A" ID-GBIF-Occurrence="4097216316" collectingDate="2011-09-29" collectionCode="SCAU" collectorName="H. W. Chen." country="China" elevation="570" latitude="21.6025" location="Mengla" longLatPrecision="21" longitude="101.585" municipality="Wangtianshu" pageId="47" pageNumber="48" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54819EF82A0D8757C36FFAB268BAFE16" box="[378,482,1339,1365]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3CEBD32A0D8757C3FCFAB268A6FE16" box="[489,510,1339,1365]" pageId="47" pageNumber="48" type="female"></specimenCount>
(
<collectionCode id="ED2BB89F2A0D8757C019FAB26B03FE16" box="[524,603,1339,1365]" pageId="47" pageNumber="48">SCAU</collectionCode>
, No. 125416, DNA #1876, 2787),
<collectingCountry id="F32D60CA2A0D8757C1CCFAB26D6EFE16" box="[985,1078,1339,1365]" name="China" pageId="47" pageNumber="48">CHINA</collectingCountry>
:
<collectingMunicipality id="6BE1BA202A0D8757C62AFAB26D8CFE16" box="[1087,1236,1339,1365]" pageId="47" pageNumber="48">Wangtianshu</collectingMunicipality>
,
<location id="8EE576812A0D8757C6CBFAB26C6EFE16" LSID="urn:lsid:plazi:treatment:0393914C2A0D8757C282FD7269B0FEA6:8EE576812A0D8757C6CBFAB26C6EFE16" box="[1246,1334,1339,1365]" country="China" latitude="21.6025" longLatPrecision="21" longitude="101.585" municipality="Wangtianshu" name="Mengla" pageId="47" pageNumber="48" stateProvince="Yunnan">Mengla</location>
,
<collectingRegion id="49FEEEB82A0D8757C755FAB26CC1FE16" box="[1344,1433,1339,1365]" country="China" name="Yunnan" pageId="47" pageNumber="48">Yunnan</collectingRegion>
,
<geoCoordinate id="EE0E469D2A0D8757C282FAD66844FE3A" box="[151,284,1375,1401]" degrees="21" direction="north" minutes="36" orientation="latitude" pageId="47" pageNumber="48" precision="15" seconds="09" value="21.6025">21°36'09''N</geoCoordinate>
,
<geoCoordinate id="EE0E469D2A0D8757C33DFAD768EFFE3A" box="[296,439,1374,1401]" degrees="101" direction="east" minutes="35" orientation="longitude" pageId="47" pageNumber="48" precision="15" seconds="06" value="101.585">101°35'06''E</geoCoordinate>
, alt.
<quantity id="4CC28DBF2A0D8757C3F8FAD76B68FE3A" box="[493,560,1374,1401]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="47" pageNumber="48" unit="m" value="570.0">
<elevation id="0017C7692A0D8757C3F8FAD76B68FE3A" box="[493,560,1374,1401]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="47" pageNumber="48" unit="m" value="570.0">570m</elevation>
</quantity>
,
<date id="FF84069A2A0D8757C02EFAD66BECFE3A" box="[571,692,1375,1401]" pageId="47" pageNumber="48" value="2011-09-29">
<collectingDate id="EFC0FF722A0D8757C02EFAD66BECFE3A" box="[571,692,1375,1401]" pageId="47" pageNumber="48" value="2011-09-29">29.ix.2011</collectingDate>
</date>
, ex tree trunk,
<collectorName id="26CF458C2A0D8757C14EFAD66A87FE3A" box="[859,991,1375,1401]" pageId="47" pageNumber="48">H.W. Chen.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A0D8757C2D2FA0B6B79FEDE" box="[199,545,1410,1437]" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8B85205A2A0D8757C2D2FA0B6B79FEDE" blockId="47.[151,1437,870,1509]" box="[199,545,1410,1437]" pageId="47" pageNumber="48">
<emphasis id="B94EFC482A0D8757C2D2FA0B6839FEDF" bold="true" box="[199,353,1410,1436]" pageId="47" pageNumber="48">Distribution.</emphasis>
<collectingCountry id="F32D60CA2A0D8757C372FA0A68F2FEDE" box="[359,426,1411,1437]" name="China" pageId="47" pageNumber="48">China</collectingCountry>
(
<collectingRegion id="49FEEEB82A0D8757C3ACFA0A6B4DFEDE" box="[441,533,1411,1437]" country="China" name="Yunnan" pageId="47" pageNumber="48">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C32073D12A0D8757C2D2FA2E69B0FEA6" pageId="47" pageNumber="48" type="etymology">
<paragraph id="8B85205A2A0D8757C2D2FA2E69B0FEA6" blockId="47.[151,1437,870,1509]" pageId="47" pageNumber="48">
<emphasis id="B94EFC482A0D8757C2D2FA2E6816FE82" bold="true" box="[199,334,1447,1473]" pageId="47" pageNumber="48">Etymology.</emphasis>
A combination of the Latin words &quot;
<emphasis id="B94EFC482A0D8757C0C2FA2E6A4AFE83" box="[727,786,1447,1472]" italics="true" pageId="47" pageNumber="48">niger</emphasis>
&quot; (= black) and &quot;
<emphasis id="B94EFC482A0D8757C1D2FA2E6D7FFE82" box="[967,1063,1447,1473]" italics="true" pageId="47" pageNumber="48">palpulus</emphasis>
'' (= palpus), referring to the black palpus.
</paragraph>
</subSubSection>
</treatment>
</document>