treatments-xml/data/03/93/80/03938025FFADA933A7D8B7BEFDC7FC58.xml
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<document id="74029587BF8E37B016F633E34893DEE3" ID-CLB-Dataset="8556" ID-DOI="10.11646/zootaxa.4843.1.1" ID-GBIF-Dataset="a2aef684-44f1-429d-9dd5-dd394539fa36" ID-ISSN="1175-5326" ID-Zenodo-Dep="4405729" ID-ZooBank="57044928-1950-4525-850E-E381A52599A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609419131752" checkinUser="plazi" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="03938025FFADA933A7D8B7BEFDC7FC58" docLanguage="en" docName="zootaxa.4843.1.1.pdf" docOrigin="Zootaxa 4843 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole madecassa Forel 1892" docType="treatment" docVersion="10" lastPageNumber="38" masterDocId="FFAAF85DFF8EA916A74FB608FFC6FFFD" masterDocTitle="The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group" masterLastPageNumber="64" masterPageNumber="1" pageNumber="36" updateTime="1698850390174" updateUser="ExternalLinkService">
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<mods:title id="B6B570FC2333D183BABB694F1D828DF4">The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group</mods:title>
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<mods:namePart id="5B298D23D397D5A2C843DCB0395D3FA0">Salata, Sebastian</mods:namePart>
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<mods:namePart id="7F119DCF43D671375A2A7F2B841104C8">Fisher, Brian L.</mods:namePart>
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<mods:title id="2784FFE7E6A90E03F9A298B3D6E91B4D">Zootaxa</mods:title>
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<mods:part id="FC78CDFF00C5EA96AF373936FC7512F7">
<mods:date id="7FA7134EF736C4F261CF2A40A1B84690">2020</mods:date>
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<treatment id="03938025FFADA933A7D8B7BEFDC7FC58" ID-DOI="http://doi.org/10.5281/zenodo.4488393" ID-GBIF-Taxon="177202118" ID-Zenodo-Dep="4488393" LSID="urn:lsid:plazi:treatment:03938025FFADA933A7D8B7BEFDC7FC58" httpUri="http://treatment.plazi.org/id/03938025FFADA933A7D8B7BEFDC7FC58" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36">
<subSubSection id="C32062B8FFADA935A7D8B7BEFDD8FE2D" box="[151,542,438,465]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8B853133FFADA935A7D8B7BEFDD8FE2D" blockId="35.[151,542,438,501]" box="[151,542,438,465]" pageId="35" pageNumber="36">
<heading id="D0CD865FFFADA935A7D8B7BEFDD8FE2D" bold="true" box="[151,542,438,465]" fontSize="11" level="2" pageId="35" pageNumber="36" reason="4">
<taxonomicName id="4C3A4AB0FFADA935A7D8B7BEFDD8FE2D" ID-CoL="4FYFF" authority="Forel, 1892" authorityName="Forel" authorityYear="1892" box="[151,542,438,465]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FFADA935A7D8B7BEFDD8FE2D" bold="true" box="[151,542,438,465]" pageId="35" pageNumber="36">
<emphasis id="B94EED21FFADA935A7D8B7BEFE4CFE2C" bold="true" box="[151,394,438,465]" italics="true" pageId="35" pageNumber="36">Pheidole madecassa</emphasis>
<bibRefCitation id="EFAB4CC2FFADA935A6DDB7BFFDD8FE2D" author="Forel, A." box="[402,542,438,465]" pageId="35" pageNumber="36" pagination="516 - 535" refId="ref52583" refString="Forel, A. (1892) Nouvelles especes de Formicides de Madagascar (recoltees par M. Sikora). Premiere serie. Annales de la Societe Entomologique de Belgique, 36, 516 - 535." type="journal article" year="1892">Forel, 1892</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFADA935A7D8B7D3FE46FE08" box="[151,384,475,501]" pageId="35" pageNumber="36" type="description">
<paragraph id="8B853133FFADA935A7D8B7D3FE46FE08" blockId="35.[151,542,438,501]" box="[151,384,475,501]" pageId="35" pageNumber="36">
<figureCitation id="13012DB6FFADA935A7D8B7D3FF21FE08" box="[151,231,475,501]" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="35" pageNumber="36">Figs 17</figureCitation>
AF, 27I, 28I
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFADA935A7D8B417FEC0FD11" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="8B853133FFADA935A7D8B417FD95FDCA" blockId="35.[151,595,543,567]" box="[151,595,543,567]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="AB2A161DFFADA935A7D8B417FD95FDCA" box="[151,595,543,567]" pageId="35" pageNumber="36">
<treatmentCitation id="0A9B1722FFADA935A7D8B417FDD3FDCA" ID-DOI="http://doi.org/10.5281/zenodo.6293267" ID-Zenodo-Dep="6293267" author="Forel, A." box="[151,533,543,567]" httpUri="http://treatment.plazi.org/id/D711212AFFE50CD3E7A886B8604AA8B9" page="525" pageId="35" pageNumber="36" year="1892">
<taxonomicName id="4C3A4AB0FFADA935A7D8B417FDD3FDCA" ID-CoL="4FYFF" authority="Forel, 1892: 525" authorityName="Forel" authorityPageNumber="525" authorityYear="1892" box="[151,533,543,567]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FFADA935A7D8B417FEA1FDCA" box="[151,359,543,567]" italics="true" pageId="35" pageNumber="36">Pheidole madecassa</emphasis>
<bibRefCitation id="EFAB4CC2FFADA935A622B417FDD3FDCA" author="Forel, A." box="[365,533,543,567]" pageId="35" pageNumber="36" pagination="516 - 535" refId="ref52583" refString="Forel, A. (1892) Nouvelles especes de Formicides de Madagascar (recoltees par M. Sikora). Premiere serie. Annales de la Societe Entomologique de Belgique, 36, 516 - 535." type="journal article" year="1892">Forel, 1892: 525</bibRefCitation>
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(s.w.)
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</paragraph>
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<materialsCitation id="3B523B6EFFADA935A7D8B46DFD00FD59" ID-GBIF-Occurrence="3027954468" collectionCode="MHNG" collectorName="Sikora" country="Madagascar" location="Mangoro river" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Toamasina" typeStatus="lectotype">
<typeStatus id="54818F91FFADA935A7D8B46DFEFDFD82" box="[151,315,613,639]" pageId="35" pageNumber="36" type="lectotype">LECTOTYPE</typeStatus>
: [designated here]:1s., bottom specimen on the pin,
<collectingCountry id="F32D71A3FFADA935A4D8B46DFBE6FD82" box="[919,1056,613,639]" name="Madagascar" pageId="35" pageNumber="36">Madagascar</collectingCountry>
,
<collectingRegion id="49FEFFD1FFADA935A362B46DFB6FFD82" box="[1069,1193,613,639]" country="Madagascar" name="Toamasina" pageId="35" pageNumber="36">Toamasina</collectingRegion>
,
<location id="8EE567E8FFADA935A3F8B46EFA98FD82" LSID="urn:lsid:plazi:treatment:03938025FFADA933A7D8B7BEFDC7FC58:8EE567E8FFADA935A3F8B46EFA98FD82" box="[1207,1374,613,640]" country="Madagascar" name="Mangoro river" pageId="35" pageNumber="36" stateProvince="Toamasina">Mangoro river</location>
, coll.
<collectorName id="26CF54E5FFADA935A7D8B481FF25FD5E" box="[151,227,649,675]" pageId="35" pageNumber="36">Sikora</collectorName>
, CASENT0101707, ANTC3400 (
<collectionCode id="ED2BA9F6FFADA935A52BB481FD7DFD5E" box="[612,699,649,675]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="35" pageNumber="36" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B523B6EFFADA935A59AB481FD31FD3A" ID-GBIF-Occurrence="3027954397" collectionCode="MHNG" country="Madagascar" pageId="35" pageNumber="36" specimenCount="2" stateProvince="Toamasina" typeStatus="paralectotype">
<typeStatus id="54818F91FFADA935A59AB481FC17FD5E" box="[725,977,649,675]" pageId="35" pageNumber="36" type="paralectotype">PARALECTOTYPES</typeStatus>
: 1s., top specimen, the same pin as
<typeStatus id="54818F91FFADA935A23DB481FF26FD35" pageId="35" pageNumber="36" type="lectotype">lectotype</typeStatus>
, CASENT0876545, ANTC3400 (
<collectionCode id="ED2BA9F6FFADA935A512B4A5FD72FD3A" box="[605,692,685,711]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="35" pageNumber="36" type="Museum">MHNG</collectionCode>
);
<specimenCount id="9D3CFABAFFADA935A582B4A5FD31FD3A" box="[717,759,685,711]" pageId="35" pageNumber="36" type="generic">2w.</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B523B6EFFADA935A44BB4A5FEC7FD11" ID-GBIF-Occurrence="3027954449" collectionCode="MHNG" collectorName="Sikora" country="Madagascar" location="Mangoro river" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Toamasina" typeStatus="paralectotype">
the same data as lectotype, CASENT0101897, ANTC3401 (
<collectionCode id="ED2BA9F6FFADA935A7D0B4D9FF30FD16" box="[159,246,721,747]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="35" pageNumber="36" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFADA935A788B4FDFAD6FBAF" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8B853133FFADA935A788B4FDFD04FBD2" blockId="35.[151,1437,613,2037]" pageId="35" pageNumber="36">
<emphasis id="B94EED21FFADA935A788B4FDFE44FCF2" bold="true" box="[199,386,757,783]" pageId="35" pageNumber="36">Other material.</emphasis>
<materialsCitation id="3B523B6EFFADA935A6C5B4FDFD17FCCE" ID-GBIF-Occurrence="3027954435" collectionCode="CASC" country="Madagascar" elevation="1180" location="Fisher" pageId="35" pageNumber="36" specimenCount="2" stateProvince="Antsiranana">
<collectingCountry id="F32D71A3FFADA935A6C5B4FDFDD5FCF2" box="[394,531,757,783]" name="Madagascar" pageId="35" pageNumber="36">Madagascar</collectingCountry>
.
<collectingRegion id="49FEFFD1FFADA935A552B4FDFD6FFCF2" box="[541,681,757,783]" country="Madagascar" name="Antsiranana" pageId="35" pageNumber="36">Antsiranana</collectingRegion>
:
<specimenCount id="9D3CFABAFFADA935A5FBB4FDFD18FCF2" box="[692,734,757,783]" pageId="35" pageNumber="36" type="generic">2w.</specimenCount>
, 1s.,
<locationDeviation id="9A372811FFADA935A456B4FDFBF0FCF2" box="[793,1078,757,783]" location="Befingotra" pageId="35" pageNumber="36">
<quantity id="4CC29CD6FFADA935A456B4FDFCAEFCF2" box="[793,872,757,783]" metricMagnitude="3" metricUnit="m" metricValue="9.2" pageId="35" pageNumber="36" unit="km" value="9.2">9.2 km</quantity>
WSW
<locationDeviation id="9A372811FFADA935A4F4B4FDFBF0FCF2" box="[955,1078,757,783]" pageId="35" pageNumber="36">Befingotra</locationDeviation>
</locationDeviation>
,
<locationDeviation id="9A372811FFADA935A30DB4FDFA8EFCF2" box="[1090,1352,757,783]" pageId="35" pageNumber="36">Rés. Anjanaharibe-Sud</locationDeviation>
, -14.75 49.46667,
<quantity id="4CC29CD6FFADA935A640B511FEA0FCC9" box="[271,358,793,820]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="35" pageNumber="36" unit="m" value="1180.0">
<elevation id="0017D600FFADA935A640B511FEA0FCC9" box="[271,358,793,820]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="35" pageNumber="36" unit="m" value="1180.0">1180 m</elevation>
</quantity>
, B. L.
<location id="8EE567E8FFADA935A6FCB511FE3FFCCE" LSID="urn:lsid:plazi:treatment:03938025FFADA933A7D8B7BEFDC7FC58:8EE567E8FFADA935A6FCB511FE3FFCCE" box="[435,505,793,819]" country="Madagascar" name="Fisher" pageId="35" pageNumber="36" stateProvince="Antsiranana">Fisher</location>
<emphasis id="B94EED21FFADA935A54EB512FDF5FCCE" box="[513,563,793,819]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. leg. (
<collectionCode id="ED2BA9F6FFADA935A535B511FD0FFCCE" box="[634,713,793,819]" pageId="35" pageNumber="36">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523B6EFFADA935A592B511FD1FFCAA" ID-GBIF-Occurrence="3027954444" collectionCode="CASC" country="Madagascar" elevation="1200" location="Fisher" pageId="35" pageNumber="36" specimenCount="1">
<specimenCount id="9D3CFABAFFADA935A592B511FCC0FCCE" box="[733,774,793,819]" pageId="35" pageNumber="36" type="generic">1w.</specimenCount>
,
<locationDeviation id="9A372811FFADA935A45BB511FBF6FCCE" box="[788,1072,793,819]" location="Befingotra" pageId="35" pageNumber="36">
<quantity id="4CC29CD6FFADA935A45BB511FCA4FCCE" box="[788,866,793,819]" metricMagnitude="3" metricUnit="m" metricValue="9.2" pageId="35" pageNumber="36" unit="km" value="9.2">9.2 km</quantity>
WSW
<locationDeviation id="9A372811FFADA935A4FAB511FBF6FCCE" box="[949,1072,793,819]" pageId="35" pageNumber="36">Befingotra</locationDeviation>
</locationDeviation>
,
<locationDeviation id="9A372811FFADA935A374B511FA87FCCE" box="[1083,1345,793,819]" pageId="35" pageNumber="36">Rés. Anjanaharibe-Sud</locationDeviation>
, -14.75, 49.46667
<quantity id="4CC29CD6FFADA935A646B535FEA5FCA5" box="[265,355,829,856]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="35" pageNumber="36" unit="m" value="1200.0">
<elevation id="0017D600FFADA935A646B535FEA5FCA5" box="[265,355,829,856]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="35" pageNumber="36" unit="m" value="1200.0">1200 m</elevation>
</quantity>
, B. L.
<location id="8EE567E8FFADA935A6FAB535FE3DFCAA" LSID="urn:lsid:plazi:treatment:03938025FFADA933A7D8B7BEFDC7FC58:8EE567E8FFADA935A6FAB535FE3DFCAA" box="[437,507,829,855]" country="Madagascar" name="Fisher" pageId="35" pageNumber="36">Fisher</location>
<emphasis id="B94EED21FFADA935A54BB536FDF1FCAA" box="[516,567,829,855]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. leg. (
<collectionCode id="ED2BA9F6FFADA935A5CDB535FD17FCAA" box="[642,721,829,855]" pageId="35" pageNumber="36">CASC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B523B6EFFADA935A5AAB535FC41FC86" ID-GBIF-Occurrence="3027954325" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="825" latitude="-15.58506" location="Ambohitsitondroina" longLatPrecision="1" longitude="50.00952" pageId="35" pageNumber="36" specimenCount="3" stateProvince="Toamasina">
<collectingRegion id="49FEFFD1FFADA935A5AAB535FCA4FCAA" box="[741,866,829,855]" country="Madagascar" name="Toamasina" pageId="35" pageNumber="36">Toamasina</collectingRegion>
:
<specimenCount id="9D3CFABAFFADA935A420B535FC5EFCAA" box="[879,920,829,855]" pageId="35" pageNumber="36" type="generic">3w.</specimenCount>
, 1s.,
<locationDeviation id="9A372811FFADA935A498B535FB2BFCAA" box="[983,1261,829,856]" location="Ambanizana" pageId="35" pageNumber="36">
<quantity id="4CC29CD6FFADA935A498B535FBE1FCAA" box="[983,1063,829,855]" metricMagnitude="3" metricUnit="m" metricValue="6.9" pageId="35" pageNumber="36" unit="km" value="6.9">6.9 km</quantity>
NE Ambanizana
</locationDeviation>
,
<location id="8EE567E8FFADA935A3B6B535FF27FC86" LSID="urn:lsid:plazi:treatment:03938025FFADA933A7D8B7BEFDC7FC58:8EE567E8FFADA935A3B6B535FF27FC86" country="Madagascar" latitude="-15.58506" longLatPrecision="1" longitude="50.00952" name="Ambohitsitondroina" pageId="35" pageNumber="36" stateProvince="Toamasina">Ambohitsitondroina</location>
,
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,
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<emphasis id="B94EED21FFADA935A5F5B56AFD2DFC86" box="[698,747,865,891]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
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, F.C.
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<quantity id="4CC29CD6FFADA935A262B569FAB0FC81" box="[1325,1398,865,892]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="35" pageNumber="36" unit="m" value="960.0">
<elevation id="0017D600FFADA935A262B569FAB0FC81" box="[1325,1398,865,892]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="35" pageNumber="36" unit="m" value="960.0">960 m</elevation>
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,
<collectorName id="26CF54E5FFADA935A2CEB56AFEE9FC62" pageId="35" pageNumber="36">H. J. Ratsirarson</collectorName>
leg. (
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<materialsCitation id="3B523B6EFFADA935A683B58DFE8DFC3E" ID-GBIF-Occurrence="3027954686" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="1075" location="Moramanga" municipality="Foret Ambatovy" pageId="35" pageNumber="36" specimenCount="1">
1s.,
<collectingMunicipality id="6BE1AB49FFADA935A6B6B58EFD69FC62" box="[505,687,901,928]" pageId="35" pageNumber="36">Forêt Ambatovy</collectingMunicipality>
,
<quantity id="4CC29CD6FFADA935A5F5B58DFCD2FC62" box="[698,788,901,927]" metricMagnitude="4" metricUnit="m" metricValue="1.43" pageId="35" pageNumber="36" unit="km" value="14.3">14.3 km</quantity>
57°
<location id="8EE567E8FFADA935A408B58EFC17FC5D" LSID="urn:lsid:plazi:treatment:03938025FFADA933A7D8B7BEFDC7FC58:8EE567E8FFADA935A408B58EFC17FC5D" box="[839,977,902,928]" country="Madagascar" municipality="Foret Ambatovy" name="Moramanga" pageId="35" pageNumber="36">Moramanga</location>
, -18.85083 48.32,
<quantity id="4CC29CD6FFADA935A3D0B58DFB32FC5D" box="[1183,1268,901,928]" metricMagnitude="3" metricUnit="m" metricValue="1.075" pageId="35" pageNumber="36" unit="m" value="1075.0">
<elevation id="0017D600FFADA935A3D0B58DFB32FC5D" box="[1183,1268,901,928]" metricMagnitude="3" metricUnit="m" metricValue="1.075" pageId="35" pageNumber="36" unit="m" value="1075.0">1075 m</elevation>
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,
<collectorName id="26CF54E5FFADA935A3B0B58EFA44FC62" box="[1279,1410,901,928]" pageId="35" pageNumber="36">B. L. Fisher</collectorName>
<emphasis id="B94EED21FFADA935A2C7B58EFF6BFC3E" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. leg. (
<collectionCode id="ED2BA9F6FFADA935A7BBB5A1FE85FC3E" box="[244,323,937,963]" pageId="35" pageNumber="36">CASC</collectionCode>
)
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;
<materialsCitation id="3B523B6EFFADA935A618B5A1FBEFFC3E" ID-GBIF-Occurrence="3027954367" box="[343,1065,937,964]" country="Madagascar" location="Camp Vohitsivalana" municipality="Reserve Betampona" pageId="35" pageNumber="36" specimenCount="2">
<specimenCount id="9D3CFABAFFADA935A618B5A1FE47FC3E" box="[343,385,937,963]" pageId="35" pageNumber="36" type="generic">2w.</specimenCount>
, 1s.,
<collectingMunicipality id="6BE1AB49FFADA935A6F4B5A2FD58FC39" box="[443,670,938,964]" pageId="35" pageNumber="36">Reserve Betampona</collectingMunicipality>
,
<location id="8EE567E8FFADA935A5E5B5A1FC49FC3E" LSID="urn:lsid:plazi:treatment:03938025FFADA933A7D8B7BEFDC7FC58:8EE567E8FFADA935A5E5B5A1FC49FC3E" box="[682,911,937,963]" country="Madagascar" municipality="Reserve Betampona" name="Camp Vohitsivalana" pageId="35" pageNumber="36">Camp Vohitsivalana</location>
,
<quantity id="4CC29CD6FFADA935A4D5B5A1FC30FC3E" box="[922,1014,937,963]" metricMagnitude="4" metricUnit="m" metricValue="3.71" pageId="35" pageNumber="36" unit="km" value="37.1">37.1 km</quantity>
338
</materialsCitation>
°
<materialsCitation id="3B523B6EFFADA935A376B5A1FDA0FC1A" ID-GBIF-Occurrence="3027954554" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="520" latitude="-17.88667" location="Toamasina" longLatPrecision="5" longitude="49.2025" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Toamasina">
<collectingRegion id="49FEFFD1FFADA935A376B5A1FB73FC3E" box="[1081,1205,937,963]" country="Madagascar" name="Toamasina" pageId="35" pageNumber="36">Toamasina</collectingRegion>
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<geoCoordinate id="EE0E57F4FFADA935A275B5A1FA5EFC39" box="[1338,1432,937,964]" degrees="49.2025" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="5" value="49.2025">49.2025</geoCoordinate>
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<quantity id="4CC29CD6FFADA935A7D8B5C5FF22FC15" box="[151,228,973,1000]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="35" pageNumber="36" unit="m" value="520.0">
<elevation id="0017D600FFADA935A7D8B5C5FF22FC15" box="[151,228,973,1000]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="35" pageNumber="36" unit="m" value="520.0">520 m</elevation>
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,
<collectorName id="26CF54E5FFADA935A7BBB5C6FE47FC1A" box="[244,385,973,1000]" pageId="35" pageNumber="36">B. L. Fisher</collectorName>
<emphasis id="B94EED21FFADA935A6C3B5C6FE07FC1A" box="[396,449,973,999]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. leg. (
<collectionCode id="ED2BA9F6FFADA935A541B5C5FD9BFC1A" box="[526,605,973,999]" pageId="35" pageNumber="36">CASC</collectionCode>
)
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;
<materialsCitation id="3B523B6EFFADA935A53AB5C5FD71FBF6" ID-GBIF-Occurrence="3027954620" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="475" latitude="-16.76912" location="Sandrangato river" longLatPrecision="1" longitude="49.26704" municipality="Reserve Speciale Ambatovaky" pageId="35" pageNumber="36" specimenCount="1">
<specimenCount id="9D3CFABAFFADA935A53AB5C5FD58FC1A" box="[629,670,973,999]" pageId="35" pageNumber="36" type="generic">1w.</specimenCount>
, 2s.,
<collectingMunicipality id="6BE1AB49FFADA935A5AFB5C5FBF9FC1A" box="[736,1087,973,999]" pageId="35" pageNumber="36">Réserve Spéciale Ambatovaky</collectingMunicipality>
,
<location id="8EE567E8FFADA935A301B5C5FADDFC1A" LSID="urn:lsid:plazi:treatment:03938025FFADA933A7D8B7BEFDC7FC58:8EE567E8FFADA935A301B5C5FADDFC1A" box="[1102,1307,973,999]" country="Madagascar" latitude="-16.76912" longLatPrecision="1" longitude="49.26704" municipality="Reserve Speciale Ambatovaky" name="Sandrangato river" pageId="35" pageNumber="36">Sandrangato river</location>
,
<geoCoordinate id="EE0E57F4FFADA935A265B5C5FA5AFC1A" box="[1322,1436,973,999]" degrees="16.76912" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="1" value="-16.76912">-16.76912</geoCoordinate>
<geoCoordinate id="EE0E57F4FFADA935A7D8B5F9FEC5FBF6" box="[151,259,1009,1035]" degrees="49.26704" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="1" value="49.26704">49.26704</geoCoordinate>
,
<quantity id="4CC29CD6FFADA935A642B5F9FE90FBF1" box="[269,342,1009,1036]" metricMagnitude="2" metricUnit="m" metricValue="4.75" pageId="35" pageNumber="36" unit="m" value="475.0">
<elevation id="0017D600FFADA935A642B5F9FE90FBF1" box="[269,342,1009,1036]" metricMagnitude="2" metricUnit="m" metricValue="4.75" pageId="35" pageNumber="36" unit="m" value="475.0">475 m</elevation>
</quantity>
,
<collectorName id="26CF54E5FFADA935A62EB5FAFE23FBF6" box="[353,485,1009,1036]" pageId="35" pageNumber="36">B. L. Fisher</collectorName>
<emphasis id="B94EED21FFADA935A6A4B5FAFDDAFBF6" box="[491,540,1009,1035]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. leg. (
<collectionCode id="ED2BA9F6FFADA935A52FB5F9FD69FBF6" box="[608,687,1009,1035]" pageId="35" pageNumber="36">CASC</collectionCode>
)
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<materialsCitation id="3B523B6EFFADA935A58DB5F9FD78FBD2" ID-GBIF-Occurrence="3027954675" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="480" latitude="-16.80561" location="Sandrangato river" longLatPrecision="1" longitude="49.29507" municipality="Reserve Speciale Ambatovaky" pageId="35" pageNumber="36" specimenCount="3">
<specimenCount id="9D3CFABAFFADA935A58DB5F9FD2AFBF6" box="[706,748,1009,1035]" pageId="35" pageNumber="36" type="generic">3w.</specimenCount>
,
<collectingMunicipality id="6BE1AB49FFADA935A5B8B5F9FB8BFBF6" box="[759,1101,1009,1035]" pageId="35" pageNumber="36">Réserve Spéciale Ambatovaky</collectingMunicipality>
,
<location id="8EE567E8FFADA935A318B5F9FAE6FBF6" LSID="urn:lsid:plazi:treatment:03938025FFADA933A7D8B7BEFDC7FC58:8EE567E8FFADA935A318B5F9FAE6FBF6" box="[1111,1312,1009,1035]" country="Madagascar" latitude="-16.80561" longLatPrecision="1" longitude="49.29507" municipality="Reserve Speciale Ambatovaky" name="Sandrangato river" pageId="35" pageNumber="36">Sandrangato river</location>
,
<geoCoordinate id="EE0E57F4FFADA935A265B5F9FA5AFBF1" box="[1322,1436,1009,1036]" degrees="16.80561" direction="south" orientation="latitude" pageId="35" pageNumber="36" precision="1" value="-16.80561">-16.80561</geoCoordinate>
<geoCoordinate id="EE0E57F4FFADA935A7D8B21DFEC5FBCD" box="[151,259,1045,1072]" degrees="49.29507" direction="east" orientation="longitude" pageId="35" pageNumber="36" precision="1" value="49.29507">49.29507</geoCoordinate>
,
<quantity id="4CC29CD6FFADA935A641B21DFE91FBD2" box="[270,343,1045,1071]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="35" pageNumber="36" unit="m" value="480.0">
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,
<collectorName id="26CF54E5FFADA935A62CB21DFE2FFBD2" box="[355,489,1045,1071]" pageId="35" pageNumber="36">B. L. Fisher</collectorName>
<emphasis id="B94EED21FFADA935A6BFB21EFDE7FBD2" box="[496,545,1045,1071]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. leg. (
<collectionCode id="ED2BA9F6FFADA935A528B21DFD70FBD2" box="[615,694,1045,1071]" pageId="35" pageNumber="36">CASC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B853133FFADA935A788B230FAD6FBAF" blockId="35.[151,1437,613,2037]" box="[199,1296,1080,1107]" pageId="35" pageNumber="36">
<emphasis id="B94EED21FFADA935A788B230FE60FBAE" bold="true" box="[199,422,1080,1107]" pageId="35" pageNumber="36">Geographic range.</emphasis>
<materialsCitation id="3B523B6EFFADA935A6E2B230FCBBFBAF" ID-GBIF-Occurrence="3027954517" box="[429,893,1080,1106]" country="Madagascar" location="Antsiranana" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Antsiranana">
<collectingCountry id="F32D71A3FFADA935A6E2B230FDF0FBAF" box="[429,566,1080,1106]" name="Madagascar" pageId="35" pageNumber="36">Madagascar</collectingCountry>
, from Andapa in
<collectingRegion id="49FEFFD1FFADA935A5B9B230FCBBFBAF" box="[758,893,1080,1106]" country="Madagascar" name="Antsiranana" pageId="35" pageNumber="36">Antsiranana</collectingRegion>
</materialsCitation>
south to Moramanga in
<collectingRegion id="49FEFFD1FFADA935A3C0B230FACAFBAF" box="[1167,1292,1080,1106]" country="Madagascar" name="Toamasina" pageId="35" pageNumber="36">Toamasina</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFADA935A788B254FABFFA21" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="8B853133FFADA935A788B254FABFFA21" blockId="35.[151,1437,613,2037]" pageId="35" pageNumber="36">
<emphasis id="B94EED21FFADA935A788B254FDC3FB8B" bold="true" box="[199,517,1116,1142]" pageId="35" pageNumber="36">Diagnosis. Major workers.</emphasis>
Head in full-face view sub-rectangular, slightly widening posteriorly, with lateral margins slightly convex; side of head with dense, long, subdecumbent to suberect pilosity; antennal scrobes very indistinct with sparse and very thin rugocostulae; interspaces between rugocostulae smooth; frons with sparse and thin costulae; interspaces between costulae smooth; occipital lobes smooth; scape, when laid back, not reaching the midlength of head; inner hypostomal tooth indistinct, tooth-like, closely spaced, and directed posteriorly; outer hypostomal tooth lobe-like, high, and moderately wide; median tooth absent; promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process distinct and tubercle-like; mesosoma smooth with very indistinct, and thin rugulae; gaster indistinctly shagreened; body yellow to orange.
<emphasis id="B94EED21FFADA935A24DB35EFF04FA69" bold="true" pageId="35" pageNumber="36">Minor workers.</emphasis>
Occiput not elongated; nuchal collar absent; head sculpture smooth; scape, when laid back, exceeding the posterior head margin by one-fifth of its length; promesonotum moderately low, moderately short, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines indistinct; mesosoma smooth; body yellow.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFADA933A788B3EDFE1FFDC0" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36" type="description">
<paragraph id="8B853133FFADA935A788B3EDFB62F996" blockId="35.[151,1437,613,2037]" pageId="35" pageNumber="36">
<emphasis id="B94EED21FFADA935A788B3EDFDD8FA02" bold="true" box="[199,542,1509,1535]" pageId="35" pageNumber="36">Description. Major workers.</emphasis>
Measurements (n=6): HL: 1.872.38 (2.15); HW: 1.882.35 (2.07); SL: 0.94 0.99 (0.96); EL: 0.220.26 (0.23); WL: 1.451.67 (1.55); PSL: 0.20.3 (0.24); MTL: 1.071.21 (1.14); PNW: 0.690.8 (0.74); PTW: 0.180.28 (0.23); PPW: 0.550.73 (0.63); CI: 99.7107.9 (104.0); SI: 39.952.4 (46.8); PSLI: 9.713.0 (11.2); PPI: 32.941.1 (37.4); PNI: 34.236.6 (35.7); MTI: 51.461.6 (55.3).
</paragraph>
<paragraph id="8B853133FFADA932A788B07CFE47FE98" blockId="35.[151,1437,613,2037]" lastBlockId="36.[151,1436,151,357]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="B94EED21FFADA935A788B07CFEC8F973" bold="true" box="[199,270,1652,1678]" pageId="35" pageNumber="36">Head.</emphasis>
In full-face view sub-rectangular, slightly widening posteriorly, with lateral sides slightly convex (
<figureCitation id="13012DB6FFADA935A220B07CFF0FF94F" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="35" pageNumber="36">Fig. 17B</figureCitation>
). In lateral view sub-oval, not depressed posteriorly; ventral and dorsal margins convex; inner hypostomal tooth visible. Side of head with dense, long, subdecumbent to suberect pilosity; head dorsum with relatively dense, long, suberect to erect pilosity. Antennal scrobes very indistinct; with sparse and very thin rugocostulae; interspaces between rugocostulae smooth. Frons with sparse and thin costulae; interspaces between costulae smooth. Occipital lobes smooth. Gena with moderately dense and thin costulae; interspaces between costulae smooth. Sides posterolateral from eyes smooth; anterior parts indistinctly shagreened. Center of clypeus shiny and smooth, lateral sides with distinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, not reaching the midlength of head; pilosity subdecumbent to erect (
<figureCitation id="13012DB6FFADA935A67BB1BEFE79F82C" box="[308,447,1974,2001]" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="35" pageNumber="36">Fig. 17B, D</figureCitation>
). Inner hypostomal tooth indistinct, tooth-like, closely spaced, and directed posteriorly; outer hypostomal tooth lobe-like, high, and moderately wide; median tooth absent (
<figureCitation id="13012DB6FFADA935A373B1D2FB5FF808" box="[1084,1177,2010,2037]" captionStart="FIGURE 27" captionStartId="60.[151,250,1604,1628]" captionTargetBox="[151,1436,1137,1581]" captionTargetId="figure-432@60.[151,1436,1137,1581]" captionTargetPageId="60" captionText="FIGURE 27. Major worker, hypostomal tooth. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole decollata (E). Pheidole flammea (F). Pheidole flavodepressa (G). Pheidole grallatrix (H). Pheidole madecassa (I). Pheidole mantadioflava (J). Pheidole maro (K). Pheidole oswaldi (L). Pheidole ovalinoda (M). Pheidole similis (N). Pheidole tenebrovulgaris (O). Pheidole uranus (P). Pheidole voreios (Q). Pheidole zirafy (R)." figureDoi="http://doi.org/10.5281/zenodo.4405803" httpUri="https://zenodo.org/record/4405803/files/figure.png" pageId="35" pageNumber="36">Fig. 27I</figureCitation>
).
<emphasis id="B94EED21FFADA935A3E3B1D3FAF7F808" bold="true" box="[1196,1329,2011,2037]" pageId="35" pageNumber="36">Mesosoma.</emphasis>
In lateral view, promesonotum short, angular, and moderately low; posterior mesonotum moderately steep; mesonotal process distinct and tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, with wide base and acute top; humeral tubercle weakly produced (
<figureCitation id="13012DB6FFAAA932A4D9B6D7FBC7FF04" box="[918,1025,223,249]" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="36" pageNumber="37">Fig. 17D</figureCitation>
). Surface mostly smooth, with very indistinct, and thin rugulae. Pilosity moderately dense, long, and erect (
<figureCitation id="13012DB6FFAAA932A4FDB70BFBF1FEE0" box="[946,1079,259,285]" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="36" pageNumber="37">Fig. 17D, F</figureCitation>
).
<emphasis id="B94EED21FFAAA932A304B70AFB67FEE1" bold="true" box="[1099,1185,258,284]" pageId="36" pageNumber="37">Gaster.</emphasis>
Shiny and indistinctly shagreened; pilosity moderately dense, long, and erect (
<figureCitation id="13012DB6FFAAA932A44EB72FFC42FEBC" box="[769,900,295,321]" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="36" pageNumber="37">Fig. 17D, F</figureCitation>
).
<emphasis id="B94EED21FFAAA932A4D7B72EFC27FEBD" bold="true" box="[920,993,294,320]" pageId="36" pageNumber="37">Color.</emphasis>
Yellow to orange, gaster and mandibles darker (
<figureCitation id="13012DB6FFAAA932A7A1B743FEB2FE98" box="[238,372,331,357]" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="36" pageNumber="37">Fig. 17D, F</figureCitation>
).
</paragraph>
<caption id="DF4561BBFFAAA932A7D8B1ABFB8EF822" ID-DOI="http://doi.org/10.5281/zenodo.4405775" ID-Zenodo-Dep="4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1955,1979]" targetBox="[151,1436,381,1931]" targetPageId="36">
<paragraph id="8B853133FFAAA932A7D8B1ABFB8EF822" blockId="36.[151,1436,1955,2015]" pageId="36" pageNumber="37">
<emphasis id="B94EED21FFAAA932A7D8B1ABFEE7F846" bold="true" box="[151,289,1955,1979]" pageId="36" pageNumber="37">FIGURE 17.</emphasis>
<taxonomicName id="4C3A4AB0FFAAA932A666B1ABFE3CF846" authorityName="Forel" authorityYear="1892" box="[297,506,1955,1979]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FFAAA932A666B1ABFE3CF846" box="[297,506,1955,1979]" italics="true" pageId="36" pageNumber="37">Pheidole madecassa</emphasis>
</taxonomicName>
, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255).
</paragraph>
</caption>
<paragraph id="8B853133FFABA933A788B69EFB93FEE0" blockId="37.[151,1437,150,933]" pageId="37" pageNumber="38">
<emphasis id="B94EED21FFABA933A788B69EFDDEFF4D" bold="true" box="[199,536,150,176]" pageId="37" pageNumber="38">Description. Minor workers.</emphasis>
Measurements (n=7): HL: 0.680.77 (0.73); HW: 0.60.66 (0.63); SL: 0.850.96 (0.9); EL: 0.120.14 (0.13); WL: 0.861.04 (0.96); PSL: 0.090.11 (0.1); MTL: 0.680.89 (0.78); PNW: 0.370.42 (0.4); PTW: 0.10.11 (0.1); PPW: 0.170.2 (0.18); CI: 113.3126.3 (117.0); SI: 132.1153.6 (143.4); PSLI: 12.9 15.3 (13.9); PPI: 53.363.3 (58.1); PNI: 61.365.1 (63.3); MTI: 110.5144.6 (124.6).
</paragraph>
<paragraph id="8B853133FFABA933A788B72EFA5AFDE4" blockId="37.[151,1437,150,933]" pageId="37" pageNumber="38">
<emphasis id="B94EED21FFABA933A788B72EFEC8FEBD" bold="true" box="[199,270,294,320]" pageId="37" pageNumber="38">Head.</emphasis>
In full-face view oval, posterior region not elongated; nuchal collar absent (
<figureCitation id="13012DB6FFABA933A3C8B72EFB35FEBD" box="[1159,1267,294,320]" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="37" pageNumber="38">Fig. 17A</figureCitation>
). Pilosity relatively sparse, moderately long, subdecumbent to erect. Sculpture smooth; anterolateral frons with weakly developed puncta. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by one-fifth of its length; pilosity dense, subdecumbent to erect (
<figureCitation id="13012DB6FFABA933A220B79AFF2AFE2D" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="37" pageNumber="38">Fig. 17A, C</figureCitation>
).
<emphasis id="B94EED21FFABA933A64EB7BEFE41FE2D" bold="true" box="[257,391,438,464]" pageId="37" pageNumber="38">Mesosoma.</emphasis>
In lateral view, promesonotum moderately low, moderately short, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines weakly developed and triangular (
<figureCitation id="13012DB6FFABA933A3F8B7D2FADBFE09" box="[1207,1309,474,500]" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="37" pageNumber="38">Fig. 17C</figureCitation>
). Sculpture smooth. Pilosity very sparse, long, and erect (
<figureCitation id="13012DB6FFABA933A5EBB7F6FCE8FDE4" box="[676,814,510,537]" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="37" pageNumber="38">Fig. 17C, E</figureCitation>
).
<emphasis id="B94EED21FFABA933A40CB7F6FC5EFDE5" bold="true" box="[835,920,510,536]" pageId="37" pageNumber="38">Gaster.</emphasis>
With sparse and erect pilosity (
<figureCitation id="13012DB6FFABA933A249B7F6FA56FDE4" box="[1286,1424,510,537]" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="37" pageNumber="38">Fig. 17C, E</figureCitation>
).
</paragraph>
<paragraph id="8B853133FFABA933A7D8B42AFE1FFDC0" blockId="37.[151,1437,150,933]" box="[151,473,546,573]" pageId="37" pageNumber="38">
<emphasis id="B94EED21FFABA933A7D8B42AFF26FDC1" bold="true" box="[151,224,546,572]" pageId="37" pageNumber="38">Color.</emphasis>
Yellow (
<figureCitation id="13012DB6FFABA933A609B42AFE0AFDC0" box="[326,460,546,573]" captionStart="FIGURE 17" captionStartId="36.[151,250,1955,1979]" captionTargetBox="[151,1436,381,1931]" captionTargetId="figure-116@36.[151,1436,381,1931]" captionTargetPageId="36" captionText="FIGURE 17. Pheidole madecassa, full-face view (A), profile (C), and dorsal view (E) of minor worker (CASENT0162735) and full-face view (B), profile (D), and dorsal view (F) of major worker (CASENT0923255)." figureDoi="http://doi.org/10.5281/zenodo.4405775" httpUri="https://zenodo.org/record/4405775/files/figure.png" pageId="37" pageNumber="38">Fig. 17C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFABA933A788B44EFE51FD55" pageId="37" pageNumber="38" type="biology_ecology">
<paragraph id="8B853133FFABA933A788B44EFE51FD55" blockId="37.[151,1437,150,933]" pageId="37" pageNumber="38">
<emphasis id="B94EED21FFABA933A788B44EFEE1FD9D" bold="true" box="[199,295,582,608]" pageId="37" pageNumber="38">Biology.</emphasis>
The species was collected between
<quantity id="4CC29CD6FFABA933A5F6B44EFC80FD9C" box="[697,838,582,609]" metricMagnitude="2" metricUnit="m" metricValue="8.375" metricValueMax="12.0" metricValueMin="4.75" pageId="37" pageNumber="38" unit="m" value="837.5" valueMax="1200.0" valueMin="475.0">4751200 m</quantity>
elevation, in rainforest and montane rainforest. Nests were located in rotten logs, dead branches above the ground, and root mats. Workers were collected from sifted litter and on low vegetation.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FFABA933A788B4BAFDC7FC58" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8B853133FFABA933A788B4BAFDC7FC58" blockId="37.[151,1437,150,933]" pageId="37" pageNumber="38">
<emphasis id="B94EED21FFABA933A788B4BAFE89FD31" bold="true" box="[199,335,690,716]" pageId="37" pageNumber="38">Comments.</emphasis>
<taxonomicName id="4C3A4AB0FFABA933A618B4BAFDFBFD31" authorityName="Forel" authorityYear="1892" box="[343,573,690,716]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FFABA933A618B4BAFDFBFD31" box="[343,573,690,716]" italics="true" pageId="37" pageNumber="38">Pheidole madecassa</emphasis>
</taxonomicName>
is most similar to sympatric
<emphasis id="B94EED21FFABA933A4CBB4BBFC24FD31" box="[900,994,691,716]" italics="true" pageId="37" pageNumber="38">
<taxonomicName id="4C3A4AB0FFABA933A4CBB4BBFC18FD31" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[900,990,691,716]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="maro">P. maro</taxonomicName>
.
</emphasis>
Major workers of
<taxonomicName id="4C3A4AB0FFABA933A3FAB4BBFA8AFD31" authorityName="Forel" authorityYear="1892" box="[1205,1356,690,716]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FFABA933A3FAB4BBFA8AFD31" box="[1205,1356,690,716]" italics="true" pageId="37" pageNumber="38">P. madecassa</emphasis>
</taxonomicName>
can be separated by smooth interspaces between rugocostulae on antennal scrobes, mostly smooth mesosoma with very indistinct and thin rugulae, small inner hypostomal tooth, and yellow to orange body. Majors of
<taxonomicName id="4C3A4AB0FFABA933A38AB4F3FADCFCE9" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1221,1306,763,788]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FFABA933A38AB4F3FADCFCE9" box="[1221,1306,763,788]" italics="true" pageId="37" pageNumber="38">P. maro</emphasis>
</taxonomicName>
have indistinctly punctate interspaces between rugocostulae on antennal scrobes, sculptured mesosoma, large inner hypostomal tooth, and reddish-brown body. Minor workers of
<taxonomicName id="4C3A4AB0FFABA933A5B9B54BFC4AFCA1" authorityName="Forel" authorityYear="1892" box="[758,908,834,860]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FFABA933A5B9B54BFC4AFCA1" box="[758,908,834,860]" italics="true" pageId="37" pageNumber="38">P. madecassa</emphasis>
</taxonomicName>
can be distinguished by smooth mesosoma and mostly smooth frons with weakly punctuated anterolateral areas. In contrast, minors of
<taxonomicName id="4C3A4AB0FFABA933A3CFB56FFB1EFC7D" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1152,1240,871,896]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FFABA933A3CFB56FFB1EFC7D" box="[1152,1240,871,896]" italics="true" pageId="37" pageNumber="38">P. maro</emphasis>
</taxonomicName>
have indistinctly sculptured frons and mesosoma.
</paragraph>
</subSubSection>
</treatment>
</document>