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<document id="833CBBFFFC186B07AA10D31E4A6B85A0" ID-CLB-Dataset="8556" ID-DOI="10.11646/zootaxa.4843.1.1" ID-GBIF-Dataset="a2aef684-44f1-429d-9dd5-dd394539fa36" ID-ISSN="1175-5326" ID-Zenodo-Dep="4405729" ID-ZooBank="57044928-1950-4525-850E-E381A52599A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609419131752" checkinUser="plazi" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="03938025FF8BA91DA7D8B7D2FEA1F98A" docLanguage="en" docName="zootaxa.4843.1.1.pdf" docOrigin="Zootaxa 4843 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole bessonii Forel 1891" docType="key" docVersion="7" lastPageNumber="12" masterDocId="FFAAF85DFF8EA916A74FB608FFC6FFFD" masterDocTitle="The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group" masterLastPageNumber="64" masterPageNumber="1" pageNumber="6" updateTime="1698850390174" updateUser="ExternalLinkService">
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<mods:title id="938FFFF3D49F75BFC314931AAE9CE03D">The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group</mods:title>
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<mods:namePart id="EDA1CB39DBE9B43AA76AAF7C2A2204BE">Salata, Sebastian</mods:namePart>
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<mods:namePart id="22B0618FC87D1AC717020777C35CF7B5">Fisher, Brian L.</mods:namePart>
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<mods:title id="D14AA3E28B974ED87F1AB019DADDCC4F">Zootaxa</mods:title>
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<mods:date id="5B2A39345195E6E4A730F139A334D177">2020</mods:date>
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<treatment id="03938025FF8BA91DA7D8B7D2FEA1F98A" ID-DOI="http://doi.org/10.5281/zenodo.4488375" ID-GBIF-Taxon="177202127" ID-Zenodo-Dep="4488375" LSID="urn:lsid:plazi:treatment:03938025FF8BA91DA7D8B7D2FEA1F98A" httpUri="http://treatment.plazi.org/id/03938025FF8BA91DA7D8B7D2FEA1F98A" lastPageId="11" lastPageNumber="12" pageId="5" pageNumber="6">
<subSubSection id="C32062B8FF8BA913A7D8B7D2FD3AFE08" box="[151,764,474,501]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8B853133FF8BA913A7D8B7D2FD3AFE08" blockId="5.[151,764,474,501]" box="[151,764,474,501]" pageId="5" pageNumber="6">
<emphasis id="B94EED21FF8BA913A7D8B7D2FD3AFE08" bold="true" box="[151,764,474,501]" pageId="5" pageNumber="6">
Synopsis of species of the
<taxonomicName id="4C3A4AB0FF8BA913A698B7D2FD6DFE08" ID-CoL="4FY85" authorityName="Forel" authorityYear="1891" box="[471,683,474,501]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF8BA913A698B7D2FD6DFE08" bold="true" box="[471,683,474,501]" italics="true" pageId="5" pageNumber="6">Pheidole bessonii</emphasis>
</taxonomicName>
group
</emphasis>
</paragraph>
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<subSubSection id="C32062B8FF8BA913A7D8B42BFE64FB5C" pageId="5" pageNumber="6" type="description">
<paragraph id="8B853133FF8BA913A7D8B42BFE13FDC0" blockId="5.[151,612,547,1185]" box="[151,469,547,573]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B42BFEB3FDC0" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,373,547,573]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="antsahabe" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B42BFEB3FDC0" box="[151,373,547,573]" italics="true" pageId="5" pageNumber="6">Pheidole antsahabe</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A633B42BFE13FDC0" bold="true" box="[380,469,547,573]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A633B42BFE13FDC0" box="[380,469,547,573]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B44FFDCCFD9C" blockId="5.[151,612,547,1185]" box="[151,522,583,609]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B44FFE6CFD9C" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,426,583,609]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="atsirakambiaty" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B44FFE6CFD9C" box="[151,426,583,609]" italics="true" pageId="5" pageNumber="6">Pheidole atsirakambiaty</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A6FEB44FFDCCFD9C" bold="true" box="[433,522,583,609]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A6FEB44FFDCCFD9C" box="[433,522,583,609]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B463FE20FD78" blockId="5.[151,612,547,1185]" box="[151,486,619,645]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B463FE20FD78" authority="Forel, 1891" authorityName="Forel" authorityYear="1891" box="[151,486,619,645]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF8BA913A7D8B463FE98FD78" box="[151,350,619,645]" italics="true" pageId="5" pageNumber="6">Pheidole bessonii</emphasis>
<bibRefCitation id="EFAB4CC2FF8BA913A62BB463FE20FD78" author="Forel, A." box="[356,486,619,645]" pageId="5" pageNumber="6" pagination="1 - 280" refId="ref52533" refString="Forel, A. (1891) Histoire naturelle des Hymenopteres. Deuxieme partie: Les Formicides. In: Grandidier, A. (Ed.), Histoire physique, naturelle, et politique de Madagascar. L'Imprimerie Nationale, Paris, pp. 1 - 280." type="book chapter" year="1891">Forel, 1891</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B487FE5BFD54" blockId="5.[151,612,547,1185]" box="[151,413,655,681]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B487FEFBFD54" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,317,655,681]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="clara" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B487FEFBFD54" box="[151,317,655,681]" italics="true" pageId="5" pageNumber="6">Pheidole clara</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A60BB487FE5BFD54" bold="true" box="[324,413,655,681]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A60BB487FE5BFD54" box="[324,413,655,681]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B4BBFDA2FD30" blockId="5.[151,612,547,1185]" box="[151,612,691,717]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B4BBFE34FD30" authority="Forel, 1892" authorityName="Forel" authorityYear="1892" box="[151,498,691,717]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="decollata" status="stat. nov.">
<emphasis id="B94EED21FF8BA913A7D8B4BBFEACFD30" box="[151,362,691,717]" italics="true" pageId="5" pageNumber="6">Pheidole decollata</emphasis>
<bibRefCitation id="EFAB4CC2FF8BA913A63EB4BBFE34FD30" author="Forel, A." box="[369,498,691,717]" pageId="5" pageNumber="6" pagination="516 - 535" refId="ref52583" refString="Forel, A. (1892) Nouvelles especes de Formicides de Madagascar (recoltees par M. Sikora). Premiere serie. Annales de la Societe Entomologique de Belgique, 36, 516 - 535." type="journal article" year="1892">Forel, 1892</bibRefCitation>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A6B6B4BBFDA2FD30" bold="true" box="[505,612,691,717]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A6B6B4BBFDA2FD30" box="[505,612,691,717]" pageId="5" pageNumber="6">stat. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B4DFFE05FD0C" blockId="5.[151,612,547,1185]" box="[151,451,727,753]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B4DFFEA4FD0C" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,354,727,753]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="flammea" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B4DFFEA4FD0C" box="[151,354,727,753]" italics="true" pageId="5" pageNumber="6">Pheidole flammea</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A626B4DFFE05FD0C" bold="true" box="[361,451,727,753]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A626B4DFFE05FD0C" box="[361,451,727,753]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B4F3FE3BFCE8" blockId="5.[151,612,547,1185]" box="[151,509,763,789]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B4F3FE5AFCE8" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,412,763,789]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="flavodepressa" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B4F3FE5AFCE8" box="[151,412,763,789]" italics="true" pageId="5" pageNumber="6">Pheidole flavodepressa</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A6ECB4F3FE3BFCE8" bold="true" box="[419,509,763,789]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A6ECB4F3FE3BFCE8" box="[419,509,763,789]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B517FDC5FCC4" blockId="5.[151,612,547,1185]" box="[151,515,799,825]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B517FDC5FCC4" authority="Emery, 1899" authorityName="Emery" authorityYear="1899" box="[151,515,799,825]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="grallatrix">
<emphasis id="B94EED21FF8BA913A7D8B517FEABFCC4" box="[151,365,799,825]" italics="true" pageId="5" pageNumber="6">Pheidole grallatrix</emphasis>
Emery, 1899
</taxonomicName>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B54BFDC3FCA0" blockId="5.[151,612,547,1185]" box="[151,517,835,861]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B54BFDC3FCA0" authority="Forel, 1892" authorityName="Forel" authorityYear="1892" box="[151,517,835,861]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FF8BA913A7D8B54BFEBBFCA0" box="[151,381,835,861]" italics="true" pageId="5" pageNumber="6">Pheidole madecassa</emphasis>
<bibRefCitation id="EFAB4CC2FF8BA913A6CBB54BFDC3FCA0" author="Forel, A." box="[388,517,835,861]" pageId="5" pageNumber="6" pagination="516 - 535" refId="ref52583" refString="Forel, A. (1892) Nouvelles especes de Formicides de Madagascar (recoltees par M. Sikora). Premiere serie. Annales de la Societe Entomologique de Belgique, 36, 516 - 535." type="journal article" year="1892">Forel, 1892</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B56FFDC2FC7C" blockId="5.[151,612,547,1185]" box="[151,516,871,897]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B56FFE62FC7C" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,420,871,897]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mantadioflava" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B56FFE62FC7C" box="[151,420,871,897]" italics="true" pageId="5" pageNumber="6">Pheidole mantadioflava</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A6E5B56FFDC2FC7C" bold="true" box="[426,516,871,897]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A6E5B56FFDC2FC7C" box="[426,516,871,897]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B583FE5AFC58" blockId="5.[151,612,547,1185]" box="[151,412,907,933]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B583FEFAFC59" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,316,907,933]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maro" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B583FEFAFC59" box="[151,316,907,933]" italics="true" pageId="5" pageNumber="6">Pheidole maro</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A60CB583FE5AFC58" bold="true" box="[323,412,907,933]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A60CB583FE5AFC58" box="[323,412,907,933]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B5A7FE27FC34" blockId="5.[151,612,547,1185]" box="[151,481,943,969]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B5A7FE27FC34" authority="Forel, 1891" authorityName="Forel" authorityYear="1891" box="[151,481,943,969]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="B94EED21FF8BA913A7D8B5A7FE9FFC34" box="[151,345,943,969]" italics="true" pageId="5" pageNumber="6">Pheidole oswaldi</emphasis>
<bibRefCitation id="EFAB4CC2FF8BA913A62FB5A7FE27FC34" author="Forel, A." box="[352,481,943,969]" pageId="5" pageNumber="6" pagination="1 - 280" refId="ref52533" refString="Forel, A. (1891) Histoire naturelle des Hymenopteres. Deuxieme partie: Les Formicides. In: Grandidier, A. (Ed.), Histoire physique, naturelle, et politique de Madagascar. L'Imprimerie Nationale, Paris, pp. 1 - 280." type="book chapter" year="1891">Forel, 1891</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B5DBFE14FC10" blockId="5.[151,612,547,1185]" box="[151,466,979,1005]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B5DBFEB4FC10" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,370,979,1005]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ovalinoda" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B5DBFEB4FC10" box="[151,370,979,1005]" italics="true" pageId="5" pageNumber="6">Pheidole ovalinoda</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A636B5DBFE14FC10" bold="true" box="[377,466,979,1005]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A636B5DBFE14FC10" box="[377,466,979,1005]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B5FFFE6DFBEC" blockId="5.[151,612,547,1185]" box="[151,427,1015,1041]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B5FFFE8DFBEC" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,331,1015,1041]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B5FFFE8DFBEC" box="[151,331,1015,1041]" italics="true" pageId="5" pageNumber="6">Pheidole similis</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A61DB5FFFE6DFBEC" bold="true" box="[338,427,1015,1041]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A61DB5FFFE6DFBEC" box="[338,427,1015,1041]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B213FDD5FBC8" blockId="5.[151,612,547,1185]" box="[151,531,1051,1077]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B213FE74FBC8" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,434,1051,1077]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="tenebrovulgaris" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B213FE74FBC8" box="[151,434,1051,1077]" italics="true" pageId="5" pageNumber="6">Pheidole tenebrovulgaris</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A6F6B213FDD5FBC8" bold="true" box="[441,531,1051,1077]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A6F6B213FDD5FBC8" box="[441,531,1051,1077]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B237FE76FBA4" blockId="5.[151,612,547,1185]" box="[151,432,1087,1113]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B237FE96FBA5" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,336,1087,1113]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="uranus" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B237FE96FBA5" box="[151,336,1087,1113]" italics="true" pageId="5" pageNumber="6">Pheidole uranus</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A619B237FE76FBA4" bold="true" box="[342,432,1087,1113]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A619B237FE76FBA4" box="[342,432,1087,1113]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B26BFE72FB80" blockId="5.[151,612,547,1185]" box="[151,436,1123,1149]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B26BFE95FB81" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,339,1123,1149]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="voreios" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B26BFE95FB81" box="[151,339,1123,1149]" italics="true" pageId="5" pageNumber="6">Pheidole voreios</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A615B26BFE72FB80" bold="true" box="[346,436,1123,1149]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A615B26BFE72FB80" box="[346,436,1123,1149]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B853133FF8BA913A7D8B28FFE64FB5C" blockId="5.[151,612,547,1185]" box="[151,418,1159,1185]" pageId="5" pageNumber="6">
<taxonomicName id="4C3A4AB0FF8BA913A7D8B28FFE84FB5C" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,322,1159,1185]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="zirafy" status="sp. nov.">
<emphasis id="B94EED21FF8BA913A7D8B28FFE84FB5C" box="[151,322,1159,1185]" italics="true" pageId="5" pageNumber="6">Pheidole zirafy</emphasis>
</taxonomicName>
<emphasis id="B94EED21FF8BA913A607B28FFE64FB5C" bold="true" box="[328,418,1159,1185]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A27D505AFF8BA913A607B28FFE64FB5C" box="[328,418,1159,1185]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF8BA913A7D8B2FBFEDCFAF0" box="[151,282,1267,1293]" pageId="5" pageNumber="6" type="multiple">
<paragraph id="8B853133FF8BA913A7D8B2FBFEDCFAF0" blockId="5.[151,282,1267,1293]" box="[151,282,1267,1293]" pageId="5" pageNumber="6">
<emphasis id="B94EED21FF8BA913A7D8B2FBFEDCFAF0" bold="true" box="[151,282,1267,1293]" pageId="5" pageNumber="6">Taxonomy</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF8BA913A7D8B333FD5CF868" pageId="5" pageNumber="6" type="description">
<paragraph id="8B853133FF8BA913A7D8B333FB22FA84" blockId="5.[151,1437,1339,1941]" pageId="5" pageNumber="6">The following repetitive characters occurring in the majority of species have been omitted from the individual species accounts. Unless stated otherwise, the following descriptions apply to all species treated here:</paragraph>
<paragraph id="8B853133FF8BA913A788B38AFF13F940" blockId="5.[151,1437,1339,1941]" pageId="5" pageNumber="6">
<emphasis id="B94EED21FF8BA913A788B38AFE45FA60" bold="true" box="[199,387,1410,1437]" pageId="5" pageNumber="6">Major workers.</emphasis>
Dorsal face of head in lateral view not depressed posteriorly; antennal socket shallow; frontal lobe absent; head in full-face view with distinct posteromedian concavity; antenna 12-segmented with 3-segmented club; masticatory margin of mandible with large, stout apical tooth and preapical tooth, followed by long diastema and then short and crenulate tooth just before rounded basal angle; outer surface of mandible mostly smooth and shining, sometimes with weak and sparse puncta; antennal scrobe absent; promesonotum strongly convex, well above level of propodeum; postpetiole shiny and microreticulate, short with slightly convex dorsum, in dorsal view oval with lateromedian dentate projection, pilosity erect; petiole shiny and microrugulate, peduncle with small horizontal lobes on its basal part, node triangular with rounded top, in rear view node dorsoventrally concave, pilosity erect.
</paragraph>
<paragraph id="8B853133FF8BA913A788B0CEFD5CF868" blockId="5.[151,1437,1339,1941]" pageId="5" pageNumber="6">
<emphasis id="B94EED21FF8BA913A788B0CEFE45F91C" bold="true" box="[199,387,1734,1761]" pageId="5" pageNumber="6">Minor workers.</emphasis>
Antennal socket shallow, surrounded with few indistinct wrinkles that curve posterolaterally, most often with sculptured interspaces; frontal lobe absent; head in full-face view oval, posterior and anterior of compound eye convex; antenna 12-segmented with 3-segmented club; humeral tubercle not developed into projection; clypeus smooth and shiny; anterior margin regularly convex; promesonotum well above level of propodeum; petiole smooth with ventral face slightly convex, node triangular with few erect setae; postpetiole smooth, convex with few erect setae; gaster smooth and shiny.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF88A910A7D8B69FFDBBFF4C" box="[151,637,151,178]" pageId="6" pageNumber="7" type="multiple">
<paragraph id="8B853133FF88A910A7D8B69FFDBBFF4C" blockId="6.[151,637,151,178]" box="[151,637,151,178]" pageId="6" pageNumber="7">
<emphasis id="B94EED21FF88A910A7D8B69FFDBBFF4C" bold="true" box="[151,637,151,178]" pageId="6" pageNumber="7">
Revision of the
<taxonomicName id="4C3A4AB0FF88A910A617B69FFDEAFF4F" authorityName="Forel" authorityYear="1891" box="[344,556,151,178]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF88A910A617B69FFDEAFF4F" bold="true" box="[344,556,151,178]" italics="true" pageId="6" pageNumber="7">Pheidole bessonii</emphasis>
</taxonomicName>
group
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF88A910A7D8B6D7FB33FD30" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8B853133FF88A910A7D8B6D7FB33FD30" blockId="6.[151,1437,223,1150]" pageId="6" pageNumber="7">
<emphasis id="B94EED21FF88A910A7D8B6D7FE09FF04" bold="true" box="[151,463,223,249]" pageId="6" pageNumber="7">Diagnosis. Major worker.</emphasis>
Postpetiole in profile without conspicuous ventral convexity; antennal socket shallow; frontal lobe absent or indistinct; propodeal spine moderate to large, wide to narrow; head in full-face view sub-rectangular, with lateral margins varying from relatively straight to slightly convex, not or slightly widening posteriorly; head in lateral view sub-oval with distinctly convex dorsal and ventral cephalic margins; occipital lobe never with transverse costulae and usually entirely sculptured (except
<taxonomicName id="4C3A4AB0FF88A910A4E1B778FB83FE74" authorityName="Forel" authorityYear="1892" box="[942,1093,367,393]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FF88A910A4E1B778FB83FE74" box="[942,1093,367,393]" italics="true" pageId="6" pageNumber="7">P. madecassa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4AB0FF88A910A31CB778FB6FFE74" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1107,1193,368,393]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FF88A910A31CB778FB6FFE74" box="[1107,1193,368,393]" italics="true" pageId="6" pageNumber="7">P. maro</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4AB0FF88A910A3AFB778FA8CFE74" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1248,1354,368,393]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="uranus">
<emphasis id="B94EED21FF88A910A3AFB778FA8CFE74" box="[1248,1354,368,393]" italics="true" pageId="6" pageNumber="7">P. uranus</emphasis>
</taxonomicName>
); frons entirely sculptured, medially costulate to rugocostulate, lateral sides with usually weaker sculpture; antennal scrobe absent or very indistinct; costulae and rugulae on the rest of head thick to very thin; promesonotum short, angular, and moderately low to very low; mesonotal process usually distinct; postpetiole in dorsal view with lateromedian dentate projection; gaster usually at least partially shagreened.
<emphasis id="B94EED21FF88A910A42FB7F7FBD6FDE4" bold="true" box="[864,1040,511,537]" pageId="6" pageNumber="7">Minor worker.</emphasis>
Postpetiole in profile without conspicuous ventral convexity; antennal socket shallow; frontal lobe absent or indistinct; propodeal spine minute to moderately large, wide to narrow; promesonotum in lateral view never box-like and posterior mesonotum never steep; promesonotal groove present and distinct (except
<taxonomicName id="4C3A4AB0FF88A910A45EB464FC1BFD78" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[785,989,619,645]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tenebrovulgaris">
<emphasis id="B94EED21FF88A910A45EB464FC1BFD78" box="[785,989,619,645]" italics="true" pageId="6" pageNumber="7">P. tenebrovulgaris</emphasis>
</taxonomicName>
); posterior region of head usually elongated, forming short to long neck; nuchal collar present and weakly developed to distinct (except
<taxonomicName id="4C3A4AB0FF88A910A398B498FAABFD54" authorityName="Forel" authorityYear="1892" box="[1239,1389,655,681]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FF88A910A398B498FAABFD54" box="[1239,1389,655,681]" italics="true" pageId="6" pageNumber="7">P. madecassa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4AB0FF88A910A7D8B4BCFF2BFD30" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,237,692,717]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FF88A910A7D8B4BCFF2BFD30" box="[151,237,692,717]" italics="true" pageId="6" pageNumber="7">P. maro</emphasis>
</taxonomicName>
); promesonotum moderately low to very low and moderately long to very long, and arched.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF88A910A788B4DFFABEFB80" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8B853133FF88A910A788B4DFFABEFB80" blockId="6.[151,1437,223,1150]" pageId="6" pageNumber="7">
<emphasis id="B94EED21FF88A910A788B4DFFE89FD0C" bold="true" box="[199,335,727,753]" pageId="6" pageNumber="7">Comments.</emphasis>
The
<taxonomicName id="4C3A4AB0FF88A910A6C7B4D0FE39FD0C" authorityName="Forel" authorityYear="1891" box="[392,511,727,753]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF88A910A6C7B4D0FE39FD0C" box="[392,511,727,753]" italics="true" pageId="6" pageNumber="7">P. bessonii</emphasis>
</taxonomicName>
group consist of species widely distributed on the island. Six species—
<taxonomicName id="4C3A4AB0FF88A910A251B4D0FA53FD0C" authorityName="Forel" authorityYear="1891" box="[1310,1429,727,753]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF88A910A251B4D0FA53FD0C" box="[1310,1429,727,753]" italics="true" pageId="6" pageNumber="7">P. bessonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4AB0FF88A910A7D8B4F4FEDCFCE8" authorityName="Forel" authorityYear="1892" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[151,282,763,789]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="decollata">
<emphasis id="B94EED21FF88A910A7D8B4F4FEDCFCE8" box="[151,282,763,789]" italics="true" pageId="6" pageNumber="7">P. decollata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4AB0FF88A910A667B4F4FE68FCE8" authorityName="Emery" authorityYear="1899" box="[296,430,763,789]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="grallatrix">
<emphasis id="B94EED21FF88A910A667B4F4FE68FCE8" box="[296,430,763,789]" italics="true" pageId="6" pageNumber="7">P. grallatrix</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4AB0FF88A910A6F3B4F4FDD7FCE8" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[444,529,764,789]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FF88A910A6F3B4F4FDD7FCE8" box="[444,529,764,789]" italics="true" pageId="6" pageNumber="7">P. maro</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4AB0FF88A910A550B4F4FD57FCE8" authorityName="Forel" authorityYear="1891" box="[543,657,763,789]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="B94EED21FF88A910A550B4F4FD57FCE8" box="[543,657,763,789]" italics="true" pageId="6" pageNumber="7">P. oswaldi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4AB0FF88A910A588B4F4FC54FCE8" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[711,914,763,789]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tenebrovulgaris">
<emphasis id="B94EED21FF88A910A588B4F4FC54FCE8" box="[711,914,763,789]" italics="true" pageId="6" pageNumber="7">P. tenebrovulgaris</emphasis>
</taxonomicName>
can be considered common. The remaining 12 species have more restricted distributions primarily limited to the northern and northeastern parts of the island. The majority of the major and minor workers of the
<taxonomicName id="4C3A4AB0FF88A910A5E3B54BFCCEFCA0" authorityName="Forel" authorityYear="1891" box="[684,776,835,861]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF88A910A5E3B54BFCCEFCA0" box="[684,776,835,861]" italics="true" pageId="6" pageNumber="7">bessonii</emphasis>
</taxonomicName>
group can be easily separated from other taxa based on the combination of the above-mentioned characters. However, majors of
<taxonomicName id="4C3A4AB0FF88A910A4FFB560FB8FFC7C" authorityName="Forel" authorityYear="1892" box="[944,1097,871,897]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FF88A910A4FFB560FB8FFC7C" box="[944,1097,871,897]" italics="true" pageId="6" pageNumber="7">P. madecassa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4AB0FF88A910A3CCB560FB1DFC7C" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1155,1243,872,897]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FF88A910A3CCB560FB1DFC7C" box="[1155,1243,872,897]" italics="true" pageId="6" pageNumber="7">P. maro</emphasis>
</taxonomicName>
can be confused with major workers of the
<emphasis id="B94EED21FF88A910A68DB583FD97FC58" box="[450,593,907,933]" italics="true" pageId="6" pageNumber="7">longispinosa</emphasis>
group. They can be separated based on distinctly shorter propodeal spines, denser frons sculpture, and brighter body coloration. Additionally, minors of
<taxonomicName id="4C3A4AB0FF88A910A343B5B8FB1CFC34" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1036,1242,943,969]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tenebrovulgaris">
<emphasis id="B94EED21FF88A910A343B5B8FB1CFC34" box="[1036,1242,943,969]" italics="true" pageId="6" pageNumber="7">P. tenebrovulgaris</emphasis>
</taxonomicName>
can be confused with two members of the
<taxonomicName id="4C3A4AB0FF88A910A6F4B5DBFDCBFC10" authorityName="Forel" authorityYear="1891" box="[443,525,979,1005]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sikorae">
<emphasis id="B94EED21FF88A910A6F4B5DBFDCBFC10" box="[443,525,979,1005]" italics="true" pageId="6" pageNumber="7">sikorae</emphasis>
</taxonomicName>
group:
<taxonomicName id="4C3A4AB0FF88A910A529B5DCFC58FC10" authority="Salata &amp; Fisher" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[614,926,979,1005]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="trichotos">
<emphasis id="B94EED21FF88A910A529B5DCFD20FC10" box="[614,742,979,1005]" italics="true" pageId="6" pageNumber="7">P. trichotos</emphasis>
Salata &amp; Fisher
</taxonomicName>
and
<taxonomicName id="4C3A4AB0FF88A910A499B5DCFB6CFC10" authority="Forel." authorityName="Forel." box="[982,1194,979,1005]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="veteratrix">
<emphasis id="B94EED21FF88A910A499B5DCFB99FC10" box="[982,1119,980,1005]" italics="true" pageId="6" pageNumber="7">P. veteratrix</emphasis>
Forel.
</taxonomicName>
<taxonomicName id="4C3A4AB0FF88A910A3FEB5DBFF17FBEC" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tenebrovulgaris">
<emphasis id="B94EED21FF88A910A3FEB5DBFF17FBEC" italics="true" pageId="6" pageNumber="7">Pheidole tenebrovulgaris</emphasis>
</taxonomicName>
can be separated based on larger body size, longer propodeal spines, higher promesonotum, and dark brown body coloration. There is also one member of the
<taxonomicName id="4C3A4AB0FF88A910A5F1B213FCDCFBC8" authorityName="Forel" authorityYear="1891" box="[702,794,1051,1077]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF88A910A5F1B213FCDCFBC8" box="[702,794,1051,1077]" italics="true" pageId="6" pageNumber="7">bessonii</emphasis>
</taxonomicName>
group known from
<collectingCountry id="F32D71A3FF88A910A4B8B214FB9FFBCB" box="[1015,1113,1052,1078]" name="Mayotte" pageId="6" pageNumber="7">Mayotte</collectingCountry>
:
<taxonomicName id="4C3A4AB0FF88A910A32DB214FB0AFBC8" authorityName="Fischer &amp; Fisher" authorityYear="2013" box="[1122,1228,1052,1077]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ragnax">
<emphasis id="B94EED21FF88A910A32DB214FB0AFBC8" box="[1122,1228,1052,1077]" italics="true" pageId="6" pageNumber="7">P. ragnax</emphasis>
</taxonomicName>
. Because its original description is comprehensive and the species is not recorded from
<collectingCountry id="F32D71A3FF88A910A4FFB237FBFFFBA4" box="[944,1081,1087,1113]" name="Madagascar" pageId="6" pageNumber="7">Madagascar</collectingCountry>
, we decided not to include it in our revision. However, we included it in comparative diagnoses within the descriptions of most similar species.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF88A910A7D8B2C7FD82FB14" box="[151,580,1231,1258]" pageId="6" pageNumber="7" type="multiple">
<paragraph id="8B853133FF88A910A7D8B2C7FD82FB14" blockId="6.[151,580,1231,1258]" box="[151,580,1231,1258]" pageId="6" pageNumber="7">
<emphasis id="B94EED21FF88A910A7D8B2C7FD82FB14" bold="true" box="[151,580,1231,1258]" pageId="6" pageNumber="7">
Key to the
<taxonomicName id="4C3A4AB0FF88A910A650B2C7FE34FB17" authorityName="Forel" authorityYear="1891" box="[287,498,1231,1258]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF88A910A650B2C7FE34FB17" bold="true" box="[287,498,1231,1258]" italics="true" pageId="6" pageNumber="7">Pheidole bessonii</emphasis>
</taxonomicName>
group
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF88A91DA7D8B31CFA5BFEC2" lastPageId="11" lastPageNumber="12" pageId="6" pageNumber="7" type="key">
<key id="FFAB6602FF88A91DA7D8B31CFA5BFEC2" lastPageId="11" lastPageNumber="12" pageId="6" pageNumber="7">
<keyStep id="30CE2996FF88A910A7D8B31CFBB3F991" pageId="6" pageNumber="7">
<paragraph id="8B853133FF88A910A7D8B31CFA5AFA7C" blockId="6.[151,1437,1300,2026]" pageId="6" pageNumber="7">
<keyLead id="30CB9206FF88A910A7D8B31CFA5AFA7C" pageId="6" pageNumber="7">
1 Major worker. Head, in full face view, slightly widening posteriorly and occipital lobe mostly smooth
<emphasis id="B94EED21FF88A910A384B31DFB26FAD7" box="[1227,1248,1301,1322]" italics="true" pageId="6" pageNumber="7">or</emphasis>
head not widening posteriorly and occipital lobes with rugae that never arch posterolaterally, body never yellow or orange, head never with short and dense setae (
<figureCitation id="13012DB6FF88A910A6C8B347FE04FA98" box="[391,450,1359,1381]" captionStart="FIGURE 2" captionStartId="7.[151,250,1070,1094]" captionTargetBox="[151,1436,433,1046]" captionTargetId="figure-387@7.[151,1436,433,1047]" captionTargetPageId="7" captionText="FIGURE 2. Major worker, head. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
). Minor worker. Posterior part of head not to slightly elongated but never forming a neck, and nuchal collar absent to weakly developed (
<figureCitation id="13012DB6FF88A910A579B364FDA9FA7F" box="[566,623,1388,1410]" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1940]" captionTargetBox="[152,1433,1160,1893]" captionTargetId="figure-422@7.[151,1436,1155,1893]" captionTargetPageId="7" captionText="FIGURE 3. Minor worker, head. Pheidole decollata, full-face (A), lateral (H). Pheidole madecassa, full-face (B), lateral (I). Pheidole maro, full-face (C), lateral (J). Pheidole oswaldi, full-face (D), lateral (K). Pheidole ovalinoda, full-face (E), lateral (L). Pheidole tenebrovulgaris, full-face (F), lateral (M). Pheidole uranus, full-face (G), lateral (N)." figureDoi="http://doi.org/10.5281/zenodo.4405737" httpUri="https://zenodo.org/record/4405737/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
).................................................................. 2
</keyLead>
</paragraph>
<paragraph id="8B853133FF88A910A7D8B382FBB3F991" blockId="6.[151,1437,1300,2026]" pageId="6" pageNumber="7">
<keyLead id="30CB9206FF88A910A7D8B382FBB3F991" pageId="6" pageNumber="7">
- Major worker. Head not or slightly widening posteriorly, occipital lobe and sometimes frons with rugae or costulae that arch posterolaterally distinctly to indistinctly. If occipital lobes with rugae then body yellow to orange
<emphasis id="B94EED21FF88A910A3D8B3AFFB6AFA41" box="[1175,1196,1447,1468]" italics="true" pageId="6" pageNumber="7">or</emphasis>
brown with lateral sides of head with short, dense setae (
<figureCitation id="13012DB6FF88A910A55CB3CCFD8BFA27" box="[531,589,1476,1498]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
). Minor worker. Posterior part of head elongated and often forming a neck, nuchal collar distinct (
<figureCitation id="13012DB6FF88A910A679B3E9FEB7FA0A" box="[310,369,1505,1527]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
)....................................................................................... 8 Note. Major worker of
<taxonomicName id="4C3A4AB0FF88A910A68FB3F7FDF1F9E9" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[448,567,1535,1556]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FF88A910A68FB3F7FDF1F9E9" box="[448,567,1535,1556]" italics="true" pageId="6" pageNumber="7">P. ovalinoda</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4AB0FF88A910A52AB3F7FD09F9E9" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[613,719,1535,1556]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FF88A910A52AB3F7FD09F9E9" box="[613,719,1535,1556]" italics="true" pageId="6" pageNumber="7">P. flammea</emphasis>
</taxonomicName>
have occipital lobes with distinct rugae and dark body coloration. The best character to distinguish between these taxa is the difference in pilosity on the side of head.
<taxonomicName id="4C3A4AB0FF88A910A32AB014FAE7F9CC" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1125,1313,1564,1585]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FF88A910A32AB014FAE7F9CC" box="[1125,1313,1564,1585]" italics="true" pageId="6" pageNumber="7">Pheidole ovalinoda</emphasis>
</taxonomicName>
has sides of head with short, decumbent to suberect setae while
<taxonomicName id="4C3A4AB0FF88A910A583B032FCF0F9B3" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[716,822,1593,1615]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FF88A910A583B032FCF0F9B3" box="[716,822,1593,1615]" italics="true" pageId="6" pageNumber="7">P. flammea</emphasis>
</taxonomicName>
has sides of head with long, suberect to erect setae. If characters match
<taxonomicName id="4C3A4AB0FF88A910A66DB05FFE5CF991" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[290,410,1623,1644]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FF88A910A66DB05FFE5CF991" box="[290,410,1623,1644]" italics="true" pageId="6" pageNumber="7">P. ovalinoda</emphasis>
</taxonomicName>
then go to couplet #2; if characters match
<taxonomicName id="4C3A4AB0FF88A910A47BB05FFC58F991" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[820,926,1623,1644]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FF88A910A47BB05FFC58F991" box="[820,926,1623,1644]" italics="true" pageId="6" pageNumber="7">P. flammea</emphasis>
</taxonomicName>
then go to couplet #8.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF88A910A7D8B07CFA5AF8E1" pageId="6" pageNumber="7">
<paragraph id="8B853133FF88A910A7D8B07CFA5AF939" blockId="6.[151,1437,1300,2026]" pageId="6" pageNumber="7">
<keyLead id="30CB9206FF88A910A7D8B07CFA5AF939" pageId="6" pageNumber="7">
2 Major worker. Occipital lobes with thick and moderately dense rugae with mostly smooth interspaces, outer hypostomal tooth absent to weakly developed (
<figureCitation id="13012DB6FF88A910A6B7B099FD82F95B" box="[504,580,1681,1703]" captionStart="FIGURE 2" captionStartId="7.[151,250,1070,1094]" captionTargetBox="[151,1436,433,1046]" captionTargetId="figure-387@7.[151,1436,433,1047]" captionTargetPageId="7" captionText="FIGURE 2. Major worker, head. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="6" pageNumber="7">Figs 2E</figureCitation>
,
<figureCitation id="13012DB6FF88A910A501B099FDADF95B" box="[590,619,1681,1702]" captionStart="FIGURE 4" captionStartId="8.[151,250,1039,1063]" captionTargetBox="[187,1400,491,1016]" captionTargetId="figure-388@8.[187,1400,491,1016]" captionTargetPageId="8" captionText="FIGURE 4. Major worker. Pheidole decollata, profile (A). Pheidole madecassa, profile (B), hypostomal tooth (E). Pheidole maro, profile (C), hypostomal tooth (F). Pheidole oswaldi, profile (D). Pheidole ovalinoda, hypostomal tooth (G)." figureDoi="http://doi.org/10.5281/zenodo.4405741" httpUri="https://zenodo.org/record/4405741/files/figure.png" pageId="6" pageNumber="7">4G</figureCitation>
). Minor worker. Petiolar peduncle moderately short and thick and node high, postpetiole short and moderately high (
<figureCitation id="13012DB6FF88A910A554B0A7FDA0F938" box="[539,614,1711,1733]" captionStart="FIGURE 5" captionStartId="8.[151,250,1976,2000]" captionTargetBox="[189,1397,1124,1952]" captionTargetId="figure-433@8.[189,1397,1124,1952]" captionTargetPageId="8" captionText="FIGURE 5. Minor worker, profile. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405743" httpUri="https://zenodo.org/record/4405743/files/figure.png" pageId="6" pageNumber="7">Fig. 5E</figureCitation>
)....................................................
<taxonomicName id="4C3A4AB0FF88A910A3AEB0A7FA5AF939" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1249,1436,1711,1732]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FF88A910A3AEB0A7FA5AF939" box="[1249,1436,1711,1732]" italics="true" pageId="6" pageNumber="7">Pheidole ovalinoda</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF88A910A7D8B0C5FA5AF8E1" blockId="6.[151,1437,1300,2026]" pageId="6" pageNumber="7">
<keyLead id="30CB9206FF88A910A7D8B0C5FA5AF8E1" pageId="6" pageNumber="7">
- Major worker. Occipital lobes mostly smooth or with thin and moderately sparse rugae with interspaces most often sculptured, outer hypostomal tooth present and distinct (
<figureCitation id="13012DB6FF88A910A5C8B0E1FD05F903" box="[647,707,1769,1791]" captionStart="FIGURE 2" captionStartId="7.[151,250,1070,1094]" captionTargetBox="[151,1436,433,1046]" captionTargetId="figure-387@7.[151,1436,433,1047]" captionTargetPageId="7" captionText="FIGURE 2. Major worker, head. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
A–D, F, G, 4E, F). Minor worker. Petiolar peduncle moderately long and thin and node low, postpetiole longer and flat (
<figureCitation id="13012DB6FF88A910A539B10FFD68F8E0" box="[630,686,1799,1821]" captionStart="FIGURE 5" captionStartId="8.[151,250,1976,2000]" captionTargetBox="[189,1397,1124,1952]" captionTargetId="figure-433@8.[189,1397,1124,1952]" captionTargetPageId="8" captionText="FIGURE 5. Minor worker, profile. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405743" httpUri="https://zenodo.org/record/4405743/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
A–D, F, G).................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF88A910A7D8B12CFA5AF831" pageId="6" pageNumber="7">
<paragraph id="8B853133FF88A910A7D8B12CFA5AF889" blockId="6.[151,1437,1300,2026]" pageId="6" pageNumber="7">
<keyLead id="30CB9206FF88A910A7D8B12CFA5AF889" pageId="6" pageNumber="7">
3 Major worker. Occipital lobes with thin rugae and indistinctly punctate to smooth interspaces, side of head with moderately dense, short, appressed to subdecumbent pilosity, body dark brown (
<figureCitation id="13012DB6FF88A910A430B149FC0CF8A5" box="[895,970,1857,1880]" captionStart="FIGURE 2" captionStartId="7.[151,250,1070,1094]" captionTargetBox="[151,1436,433,1046]" captionTargetId="figure-387@7.[151,1436,433,1047]" captionTargetPageId="7" captionText="FIGURE 2. Major worker, head. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="6" pageNumber="7">Fig. 2F</figureCitation>
). Minor worker. Promesonotal groove absent to indistinct, mesosoma entirely punctate, body dark brown. (
<figureCitation id="13012DB6FF88A910A459B157FC9BF888" box="[790,861,1887,1909]" captionStart="FIGURE 5" captionStartId="8.[151,250,1976,2000]" captionTargetBox="[189,1397,1124,1952]" captionTargetId="figure-433@8.[189,1397,1124,1952]" captionTargetPageId="8" captionText="FIGURE 5. Minor worker, profile. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405743" httpUri="https://zenodo.org/record/4405743/files/figure.png" pageId="6" pageNumber="7">Fig. 5F</figureCitation>
).........................
<taxonomicName id="4C3A4AB0FF88A910A3E5B157FA5AF889" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1194,1436,1887,1908]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tenebrovulgaris">
<emphasis id="B94EED21FF88A910A3E5B157FA5AF889" box="[1194,1436,1887,1908]" italics="true" pageId="6" pageNumber="7">Pheidole tenebrovulgaris</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF88A910A7D8B175FA5AF831" blockId="6.[151,1437,1300,2026]" pageId="6" pageNumber="7">
<keyLead id="30CB9206FF88A910A7D8B175FA5AF831" pageId="6" pageNumber="7">
- Major worker. Occipital lobes mostly smooth or occipital lobes punctate with sparse and indistinct rugae, sides of head with dense, long, suberect to erect pilosity and body yellowish orange to brown (
<figureCitation id="13012DB6FF88A910A4E3B191FC22F853" box="[940,996,1945,1967]" captionStart="FIGURE 2" captionStartId="7.[151,250,1070,1094]" captionTargetBox="[151,1436,433,1046]" captionTargetId="figure-387@7.[151,1436,433,1047]" captionTargetPageId="7" captionText="FIGURE 2. Major worker, head. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
A–D, G). Minor worker. Promesonotal groove present and distinct, mesosoma sparsely punctate to smooth, body yellow to yellowish brown (
<figureCitation id="13012DB6FF88A910A320B1BFFB61F830" box="[1135,1191,1975,1997]" captionStart="FIGURE 5" captionStartId="8.[151,250,1976,2000]" captionTargetBox="[189,1397,1124,1952]" captionTargetId="figure-433@8.[189,1397,1124,1952]" captionTargetPageId="8" captionText="FIGURE 5. Minor worker, profile. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405743" httpUri="https://zenodo.org/record/4405743/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
A–D, G)............ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF88A911A7D8B1DCFA5AFF15" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<paragraph id="8B853133FF88A911A7D8B1DCFA5AFF50" blockId="6.[151,1437,1300,2026]" lastBlockId="7.[151,1437,151,408]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<keyLead id="30CB9206FF88A911A7D8B1DCFA5AFF50" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
4 Major workers. Occipital lobes never with reduced sculpture or smooth notches (
<figureCitation id="13012DB6FF88A910A4A5B1DCFB92F817" box="[1002,1108,2004,2026]" captionStart="FIGURE 2" captionStartId="7.[151,250,1070,1094]" captionTargetBox="[151,1436,433,1046]" captionTargetId="figure-387@7.[151,1436,433,1047]" captionTargetPageId="7" captionText="FIGURE 2. Major worker, head. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="6" pageNumber="7">Fig. 2A, D</figureCitation>
). Minor workers. Posterior part of head slightly elongated, mesosoma mostly punctate (
<figureCitation id="13012DB6FF89A911A592B69FFC43FF50" box="[733,901,151,173]" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1940]" captionTargetBox="[152,1433,1160,1893]" captionTargetId="figure-422@7.[151,1436,1155,1893]" captionTargetPageId="7" captionText="FIGURE 3. Minor worker, head. Pheidole decollata, full-face (A), lateral (H). Pheidole madecassa, full-face (B), lateral (I). Pheidole maro, full-face (C), lateral (J). Pheidole oswaldi, full-face (D), lateral (K). Pheidole ovalinoda, full-face (E), lateral (L). Pheidole tenebrovulgaris, full-face (F), lateral (M). Pheidole uranus, full-face (G), lateral (N)." figureDoi="http://doi.org/10.5281/zenodo.4405737" httpUri="https://zenodo.org/record/4405737/files/figure.png" pageId="7" pageNumber="8">Figs 3A, D, H, K</figureCitation>
,
<figureCitation id="13012DB6FF89A911A4C0B69FFC0AFF50" box="[911,972,151,173]" captionStart="FIGURE 5" captionStartId="8.[151,250,1976,2000]" captionTargetBox="[189,1397,1124,1952]" captionTargetId="figure-433@8.[189,1397,1124,1952]" captionTargetPageId="8" captionText="FIGURE 5. Minor worker, profile. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405743" httpUri="https://zenodo.org/record/4405743/files/figure.png" pageId="7" pageNumber="8">5A, D</figureCitation>
)..................................... 5
</keyLead>
</paragraph>
<paragraph id="8B853133FF89A911A7D8B6BDFA5AFF15" blockId="7.[151,1437,151,408]" pageId="7" pageNumber="8">
<keyLead id="30CB9206FF89A911A7D8B6BDFA5AFF15" pageId="7" pageNumber="8">
- Major worker. Occipital lobes entirely or mostly smooth (
<figureCitation id="13012DB6FF89A911A445B6BDFC4DFF37" box="[778,907,181,203]" captionStart="FIGURE 2" captionStartId="7.[151,250,1070,1094]" captionTargetBox="[151,1436,433,1046]" captionTargetId="figure-387@7.[151,1436,433,1047]" captionTargetPageId="7" captionText="FIGURE 2. Major worker, head. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="7" pageNumber="8">Fig. 2B, C, G</figureCitation>
). Minor worker. Posterior part of head never elongated and mesosoma mostly or entirely smooth (
<figureCitation id="13012DB6FF89A911A535B6DAFC8EFF15" box="[634,840,210,232]" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1940]" captionTargetBox="[152,1433,1160,1893]" captionTargetId="figure-422@7.[151,1436,1155,1893]" captionTargetPageId="7" captionText="FIGURE 3. Minor worker, head. Pheidole decollata, full-face (A), lateral (H). Pheidole madecassa, full-face (B), lateral (I). Pheidole maro, full-face (C), lateral (J). Pheidole oswaldi, full-face (D), lateral (K). Pheidole ovalinoda, full-face (E), lateral (L). Pheidole tenebrovulgaris, full-face (F), lateral (M). Pheidole uranus, full-face (G), lateral (N)." figureDoi="http://doi.org/10.5281/zenodo.4405737" httpUri="https://zenodo.org/record/4405737/files/figure.png" pageId="7" pageNumber="8">Figs 3B, C, G, I, J, N</figureCitation>
,
<figureCitation id="13012DB6FF89A911A41CB6DAFC6EFF1A" box="[851,936,210,232]" captionStart="FIGURE 5" captionStartId="8.[151,250,1976,2000]" captionTargetBox="[189,1397,1124,1952]" captionTargetId="figure-433@8.[189,1397,1124,1952]" captionTargetPageId="8" captionText="FIGURE 5. Minor worker, profile. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405743" httpUri="https://zenodo.org/record/4405743/files/figure.png" pageId="7" pageNumber="8">5B, C, G</figureCitation>
)....................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF89A911A7D8B6E7FA5AFE6A" pageId="7" pageNumber="8">
<paragraph id="8B853133FF89A911A7D8B6E7FA5BFEC2" blockId="7.[151,1437,151,408]" pageId="7" pageNumber="8">
<keyLead id="30CB9206FF89A911A7D8B6E7FA5BFEC2" pageId="7" pageNumber="8">
5 Major worker. Promesonotum in lateral view moderately long, angular, and very low (
<figureCitation id="13012DB6FF89A911A36CB6E7FBB6FEFB" box="[1059,1136,239,262]" captionStart="FIGURE 4" captionStartId="8.[151,250,1039,1063]" captionTargetBox="[187,1400,491,1016]" captionTargetId="figure-388@8.[187,1400,491,1016]" captionTargetPageId="8" captionText="FIGURE 4. Major worker. Pheidole decollata, profile (A). Pheidole madecassa, profile (B), hypostomal tooth (E). Pheidole maro, profile (C), hypostomal tooth (F). Pheidole oswaldi, profile (D). Pheidole ovalinoda, hypostomal tooth (G)." figureDoi="http://doi.org/10.5281/zenodo.4405741" httpUri="https://zenodo.org/record/4405741/files/figure.png" pageId="7" pageNumber="8">Fig. 4D</figureCitation>
), inner hypostomal tooth small (
<figureCitation id="13012DB6FF89A911A7A6B705FE84FEDE" box="[233,322,269,291]" captionStart="FIGURE 27" captionStartId="60.[151,250,1604,1628]" captionTargetBox="[151,1436,1137,1581]" captionTargetId="figure-432@60.[151,1436,1137,1581]" captionTargetPageId="60" captionText="FIGURE 27. Major worker, hypostomal tooth. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole decollata (E). Pheidole flammea (F). Pheidole flavodepressa (G). Pheidole grallatrix (H). Pheidole madecassa (I). Pheidole mantadioflava (J). Pheidole maro (K). Pheidole oswaldi (L). Pheidole ovalinoda (M). Pheidole similis (N). Pheidole tenebrovulgaris (O). Pheidole uranus (P). Pheidole voreios (Q). Pheidole zirafy (R)." figureDoi="http://doi.org/10.5281/zenodo.4405803" httpUri="https://zenodo.org/record/4405803/files/figure.png" pageId="7" pageNumber="8">Fig. 27L</figureCitation>
). Minor worker. Occiput slightly elongated, promesonotum moderately long, angular, and very low (
<figureCitation id="13012DB6FF89A911A269B705FA5EFEDF" box="[1318,1432,269,291]" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1940]" captionTargetBox="[152,1433,1160,1893]" captionTargetId="figure-422@7.[151,1436,1155,1893]" captionTargetPageId="7" captionText="FIGURE 3. Minor worker, head. Pheidole decollata, full-face (A), lateral (H). Pheidole madecassa, full-face (B), lateral (I). Pheidole maro, full-face (C), lateral (J). Pheidole oswaldi, full-face (D), lateral (K). Pheidole ovalinoda, full-face (E), lateral (L). Pheidole tenebrovulgaris, full-face (F), lateral (M). Pheidole uranus, full-face (G), lateral (N)." figureDoi="http://doi.org/10.5281/zenodo.4405737" httpUri="https://zenodo.org/record/4405737/files/figure.png" pageId="7" pageNumber="8">Figs 3D, K</figureCitation>
,
<figureCitation id="13012DB6FF89A911A7ADB722FEC4FEBD" box="[226,258,298,320]" captionStart="FIGURE 5" captionStartId="8.[151,250,1976,2000]" captionTargetBox="[189,1397,1124,1952]" captionTargetId="figure-433@8.[189,1397,1124,1952]" captionTargetPageId="8" captionText="FIGURE 5. Minor worker, profile. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405743" httpUri="https://zenodo.org/record/4405743/files/figure.png" pageId="7" pageNumber="8">5D</figureCitation>
)...................................................................................
<taxonomicName id="4C3A4AB0FF89A911A3B9B722FA5BFEC2" ID-CoL="4FYHF" authorityName="Forel" authorityYear="1891" box="[1270,1437,298,319]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="B94EED21FF89A911A3B9B722FA5BFEC2" box="[1270,1437,298,319]" italics="true" pageId="7" pageNumber="8">Pheidole oswaldi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF89A911A7D8B740FA5AFE6A" blockId="7.[151,1437,151,408]" pageId="7" pageNumber="8">
<keyLead id="30CB9206FF89A911A7D8B740FA5AFE6A" pageId="7" pageNumber="8">
- Major worker. Promesonotum in lateral view short, angular, and moderately low (
<figureCitation id="13012DB6FF89A911A4A0B74FFBFAFEA3" box="[1007,1084,327,350]" captionStart="FIGURE 4" captionStartId="8.[151,250,1039,1063]" captionTargetBox="[187,1400,491,1016]" captionTargetId="figure-388@8.[187,1400,491,1016]" captionTargetPageId="8" captionText="FIGURE 4. Major worker. Pheidole decollata, profile (A). Pheidole madecassa, profile (B), hypostomal tooth (E). Pheidole maro, profile (C), hypostomal tooth (F). Pheidole oswaldi, profile (D). Pheidole ovalinoda, hypostomal tooth (G)." figureDoi="http://doi.org/10.5281/zenodo.4405741" httpUri="https://zenodo.org/record/4405741/files/figure.png" pageId="7" pageNumber="8">Fig. 4A</figureCitation>
), inner hypostomal tooth moderately large (
<figureCitation id="13012DB6FF89A911A651B76DFEB2FE86" box="[286,372,357,379]" captionStart="FIGURE 27" captionStartId="60.[151,250,1604,1628]" captionTargetBox="[151,1436,1137,1581]" captionTargetId="figure-432@60.[151,1436,1137,1581]" captionTargetPageId="60" captionText="FIGURE 27. Major worker, hypostomal tooth. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole decollata (E). Pheidole flammea (F). Pheidole flavodepressa (G). Pheidole grallatrix (H). Pheidole madecassa (I). Pheidole mantadioflava (J). Pheidole maro (K). Pheidole oswaldi (L). Pheidole ovalinoda (M). Pheidole similis (N). Pheidole tenebrovulgaris (O). Pheidole uranus (P). Pheidole voreios (Q). Pheidole zirafy (R)." figureDoi="http://doi.org/10.5281/zenodo.4405803" httpUri="https://zenodo.org/record/4405803/files/figure.png" pageId="7" pageNumber="8">Fig. 27E</figureCitation>
). Minor worker. Occiput not elongated, promesonotum moderately long, angular, and moderately low (
<figureCitation id="13012DB6FF89A911A206B76DFF33FE65" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1940]" captionTargetBox="[152,1433,1160,1893]" captionTargetId="figure-422@7.[151,1436,1155,1893]" captionTargetPageId="7" captionText="FIGURE 3. Minor worker, head. Pheidole decollata, full-face (A), lateral (H). Pheidole madecassa, full-face (B), lateral (I). Pheidole maro, full-face (C), lateral (J). Pheidole oswaldi, full-face (D), lateral (K). Pheidole ovalinoda, full-face (E), lateral (L). Pheidole tenebrovulgaris, full-face (F), lateral (M). Pheidole uranus, full-face (G), lateral (N)." figureDoi="http://doi.org/10.5281/zenodo.4405737" httpUri="https://zenodo.org/record/4405737/files/figure.png" pageId="7" pageNumber="8">Figs 3A, H</figureCitation>
,
<figureCitation id="13012DB6FF89A911A7B0B78AFED9FE65" box="[255,287,386,408]" captionStart="FIGURE 5" captionStartId="8.[151,250,1976,2000]" captionTargetBox="[189,1397,1124,1952]" captionTargetId="figure-433@8.[189,1397,1124,1952]" captionTargetPageId="8" captionText="FIGURE 5. Minor worker, profile. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405743" httpUri="https://zenodo.org/record/4405743/files/figure.png" pageId="7" pageNumber="8">5A</figureCitation>
)................................................................................
<taxonomicName id="4C3A4AB0FF89A911A3A7B78AFA5AFE6A" authorityName="Forel" authorityYear="1892" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[1256,1436,386,407]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="decollata">
<emphasis id="B94EED21FF89A911A3A7B78AFA5AFE6A" box="[1256,1436,386,407]" italics="true" pageId="7" pageNumber="8">Pheidole decollata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF4561BBFF89A911A7D8B226FC5EFB96" ID-DOI="http://doi.org/10.5281/zenodo.4405735" ID-Zenodo-Dep="4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1070,1094]" targetBox="[151,1436,433,1046]" targetPageId="7">
<paragraph id="8B853133FF89A911A7D8B226FC5EFB96" blockId="7.[151,1437,1070,1131]" pageId="7" pageNumber="8">
<emphasis id="B94EED21FF89A911A7D8B226FED2FBBA" bold="true" box="[151,276,1070,1095]" pageId="7" pageNumber="8">FIGURE 2.</emphasis>
Major worker, head.
<taxonomicName id="4C3A4AB0FF89A911A6BDB227FD77FBBA" authorityName="Forel" authorityYear="1892" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[498,689,1071,1095]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="decollata">
<emphasis id="B94EED21FF89A911A6BDB227FD77FBBA" box="[498,689,1071,1095]" italics="true" pageId="7" pageNumber="8">Pheidole decollata</emphasis>
</taxonomicName>
(A).
<taxonomicName id="4C3A4AB0FF89A911A5A6B227FC7FFBBA" ID-CoL="4FYFF" authorityName="Forel" authorityYear="1892" box="[745,953,1071,1095]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FF89A911A5A6B227FC7FFBBA" box="[745,953,1071,1095]" italics="true" pageId="7" pageNumber="8">Pheidole madecassa</emphasis>
</taxonomicName>
(B).
<taxonomicName id="4C3A4AB0FF89A911A4A0B227FB40FBBB" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1007,1158,1071,1095]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FF89A911A4A0B227FB40FBBB" box="[1007,1158,1071,1095]" italics="true" pageId="7" pageNumber="8">Pheidole maro</emphasis>
</taxonomicName>
(C).
<taxonomicName id="4C3A4AB0FF89A911A3F3B227FAAAFBBA" ID-CoL="4FYHF" authorityName="Forel" authorityYear="1891" box="[1212,1388,1071,1095]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="B94EED21FF89A911A3F3B227FAAAFBBA" box="[1212,1388,1071,1095]" italics="true" pageId="7" pageNumber="8">Pheidole oswaldi</emphasis>
</taxonomicName>
(D).
<taxonomicName id="4C3A4AB0FF89A911A7D8B25BFE9BFB96" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,349,1107,1131]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FF89A911A7D8B25BFE9BFB96" box="[151,349,1107,1131]" italics="true" pageId="7" pageNumber="8">Pheidole ovalinoda</emphasis>
</taxonomicName>
(E).
<taxonomicName id="4C3A4AB0FF89A911A6DFB25BFD56FB96" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[400,656,1107,1131]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tenebrovulgaris">
<emphasis id="B94EED21FF89A911A6DFB25BFD56FB96" box="[400,656,1107,1131]" italics="true" pageId="7" pageNumber="8">Pheidole tenebrovulgaris</emphasis>
</taxonomicName>
(F).
<taxonomicName id="4C3A4AB0FF89A911A58DB25BFCAEFB97" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[706,872,1107,1131]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="uranus">
<emphasis id="B94EED21FF89A911A58DB25BFCAEFB97" box="[706,872,1107,1131]" italics="true" pageId="7" pageNumber="8">Pheidole uranus</emphasis>
</taxonomicName>
(G).
</paragraph>
</caption>
<caption id="DF4561BBFF89A911A7D8B174FB47F820" ID-DOI="http://doi.org/10.5281/zenodo.4405737" ID-Zenodo-Dep="4405737" httpUri="https://zenodo.org/record/4405737/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1916,1940]" targetBox="[152,1433,1160,1893]" targetPageId="7">
<paragraph id="8B853133FF89A911A7D8B174FB47F820" blockId="7.[151,1436,1916,2013]" pageId="7" pageNumber="8">
<emphasis id="B94EED21FF89A911A7D8B174FED2F869" bold="true" box="[151,276,1916,1940]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
Minor worker, head.
<taxonomicName id="4C3A4AB0FF89A911A6BAB175FD73F868" authorityName="Forel" authorityYear="1892" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[501,693,1917,1941]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="decollata">
<emphasis id="B94EED21FF89A911A6BAB175FD73F868" box="[501,693,1917,1941]" italics="true" pageId="7" pageNumber="8">Pheidole decollata</emphasis>
</taxonomicName>
, full-face (A), lateral (H).
<taxonomicName id="4C3A4AB0FF89A911A485B175FB5CF868" ID-CoL="4FYFF" authorityName="Forel" authorityYear="1892" box="[970,1178,1917,1941]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FF89A911A485B175FB5CF868" box="[970,1178,1917,1941]" italics="true" pageId="7" pageNumber="8">Pheidole madecassa</emphasis>
</taxonomicName>
, full-face (B), lateral (I).
<taxonomicName id="4C3A4AB0FF89A911A7D8B1A9FEE8F845" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,302,1953,1977]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FF89A911A7D8B1A9FEE8F845" box="[151,302,1953,1977]" italics="true" pageId="7" pageNumber="8">Pheidole maro</emphasis>
</taxonomicName>
, full-face (C), lateral (J).
<taxonomicName id="4C3A4AB0FF89A911A577B1A9FD2EF844" ID-CoL="4FYHF" authorityName="Forel" authorityYear="1891" box="[568,744,1953,1977]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="B94EED21FF89A911A577B1A9FD2EF844" box="[568,744,1953,1977]" italics="true" pageId="7" pageNumber="8">Pheidole oswaldi</emphasis>
</taxonomicName>
, full-face (D), lateral (K).
<taxonomicName id="4C3A4AB0FF89A911A4B3B1A9FB05F844" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1020,1219,1953,1977]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FF89A911A4B3B1A9FB05F844" box="[1020,1219,1953,1977]" italics="true" pageId="7" pageNumber="8">Pheidole ovalinoda</emphasis>
</taxonomicName>
, full-face (E), lateral (L).
<taxonomicName id="4C3A4AB0FF89A911A78BB1CDFE02F820" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[196,452,1989,2013]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tenebrovulgaris">
<emphasis id="B94EED21FF89A911A78BB1CDFE02F820" box="[196,452,1989,2013]" italics="true" pageId="7" pageNumber="8">Pheidole tenebrovulgaris</emphasis>
</taxonomicName>
, full-face (F), lateral (M).
<taxonomicName id="4C3A4AB0FF89A911A59DB1CDFCBFF821" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[722,889,1989,2013]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="uranus">
<emphasis id="B94EED21FF89A911A59DB1CDFCBFF821" box="[722,889,1989,2013]" italics="true" pageId="7" pageNumber="8">Pheidole uranus</emphasis>
</taxonomicName>
, full-face (G), lateral (N).
</paragraph>
</caption>
<keyStep id="30CE2996FF86A91EA7D8B69FFA5AFEDF" pageId="8" pageNumber="9">
<paragraph id="8B853133FF86A91EA7D8B69FFA5BFF15" blockId="8.[151,1437,151,467]" pageId="8" pageNumber="9">
<keyLead id="30CB9206FF86A91EA7D8B69FFA5BFF15" pageId="8" pageNumber="9">
6 Major worker. Lateral frons densely rugose-reticulate, occipital lobes mostly smooth, posteriorly with sparse and weakly developed rugae (
<figureCitation id="13012DB6FF86A91EA639B6BDFE06FF37" box="[374,448,181,203]" captionStart="FIGURE 2" captionStartId="7.[151,250,1070,1094]" captionTargetBox="[151,1436,433,1046]" captionTargetId="figure-387@7.[151,1436,433,1047]" captionTargetPageId="7" captionText="FIGURE 2. Major worker, head. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="8" pageNumber="9">Fig. 2G</figureCitation>
). Minor worker. Occiput slightly elongated; nuchal collar weakly developed (
<figureCitation id="13012DB6FF86A91EA3E2B6BDFADEFF37" box="[1197,1304,181,203]" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1940]" captionTargetBox="[152,1433,1160,1893]" captionTargetId="figure-422@7.[151,1436,1155,1893]" captionTargetPageId="7" captionText="FIGURE 3. Minor worker, head. Pheidole decollata, full-face (A), lateral (H). Pheidole madecassa, full-face (B), lateral (I). Pheidole maro, full-face (C), lateral (J). Pheidole oswaldi, full-face (D), lateral (K). Pheidole ovalinoda, full-face (E), lateral (L). Pheidole tenebrovulgaris, full-face (F), lateral (M). Pheidole uranus, full-face (G), lateral (N)." figureDoi="http://doi.org/10.5281/zenodo.4405737" httpUri="https://zenodo.org/record/4405737/files/figure.png" pageId="8" pageNumber="9">Fig. 3G, N</figureCitation>
)................................................................................................
<taxonomicName id="4C3A4AB0FF86A91EA3B1B6DAFA5BFF15" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1278,1437,210,232]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="uranus">
<emphasis id="B94EED21FF86A91EA3B1B6DAFA5BFF15" box="[1278,1437,210,232]" italics="true" pageId="8" pageNumber="9">Pheidole uranus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF86A91EA7D8B6F8FA5AFEDF" blockId="8.[151,1437,151,467]" pageId="8" pageNumber="9">
<keyLead id="30CB9206FF86A91EA7D8B6F8FA5AFEDF" pageId="8" pageNumber="9">
- Major worker. Frons laterally smooth with sparse rugulae, occipital lobes smooth, sometimes with weakly developed rugopuncta (
<figureCitation id="13012DB6FF86A91EA660B705FE51FEDE" box="[303,407,269,291]" captionStart="FIGURE 2" captionStartId="7.[151,250,1070,1094]" captionTargetBox="[151,1436,433,1046]" captionTargetId="figure-387@7.[151,1436,433,1047]" captionTargetPageId="7" captionText="FIGURE 2. Major worker, head. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="8" pageNumber="9">Fig. 2B, C</figureCitation>
). Minor worker. Occiput not elongated, nuchal collar absent (
<figureCitation id="13012DB6FF86A91EA4A9B705FBB0FEDF" box="[998,1142,269,291]" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1940]" captionTargetBox="[152,1433,1160,1893]" captionTargetId="figure-422@7.[151,1436,1155,1893]" captionTargetPageId="7" captionText="FIGURE 3. Minor worker, head. Pheidole decollata, full-face (A), lateral (H). Pheidole madecassa, full-face (B), lateral (I). Pheidole maro, full-face (C), lateral (J). Pheidole oswaldi, full-face (D), lateral (K). Pheidole ovalinoda, full-face (E), lateral (L). Pheidole tenebrovulgaris, full-face (F), lateral (M). Pheidole uranus, full-face (G), lateral (N)." figureDoi="http://doi.org/10.5281/zenodo.4405737" httpUri="https://zenodo.org/record/4405737/files/figure.png" pageId="8" pageNumber="9">Fig. 3B, C, I, J</figureCitation>
)...................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF86A91EA7D8B723FA5AFE2F" pageId="8" pageNumber="9">
<paragraph id="8B853133FF86A91EA7D8B723FA5AFE87" blockId="8.[151,1437,151,467]" pageId="8" pageNumber="9">
<keyLead id="30CB9206FF86A91EA7D8B723FA5AFE87" pageId="8" pageNumber="9">
7 Major worker. Antennal scrobe with sparse and very thin rugulae with indistinctly punctate interspaces, mesosoma weakly microreticulate, inner hypostomal tooth large, body reddish brown (
<figureCitation id="13012DB6FF86A91EA422B74FFC7AFEA0" box="[877,956,327,349]" captionStart="FIGURE 2" captionStartId="7.[151,250,1070,1094]" captionTargetBox="[151,1436,433,1046]" captionTargetId="figure-387@7.[151,1436,433,1047]" captionTargetPageId="7" captionText="FIGURE 2. Major worker, head. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="8" pageNumber="9">Figs 2C</figureCitation>
,
<figureCitation id="13012DB6FF86A91EA489B740FC38FEA0" box="[966,1022,328,349]" captionStart="FIGURE 4" captionStartId="8.[151,250,1039,1063]" captionTargetBox="[187,1400,491,1016]" captionTargetId="figure-388@8.[187,1400,491,1016]" captionTargetPageId="8" captionText="FIGURE 4. Major worker. Pheidole decollata, profile (A). Pheidole madecassa, profile (B), hypostomal tooth (E). Pheidole maro, profile (C), hypostomal tooth (F). Pheidole oswaldi, profile (D). Pheidole ovalinoda, hypostomal tooth (G)." figureDoi="http://doi.org/10.5281/zenodo.4405741" httpUri="https://zenodo.org/record/4405741/files/figure.png" pageId="8" pageNumber="9">4C, F</figureCitation>
). Minor worker. Frons and mesosoma with very sparse, weakly developed puncta (
<figureCitation id="13012DB6FF86A91EA514B76DFD6FFE87" box="[603,681,357,379]" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1940]" captionTargetBox="[152,1433,1160,1893]" captionTargetId="figure-422@7.[151,1436,1155,1893]" captionTargetPageId="7" captionText="FIGURE 3. Minor worker, head. Pheidole decollata, full-face (A), lateral (H). Pheidole madecassa, full-face (B), lateral (I). Pheidole maro, full-face (C), lateral (J). Pheidole oswaldi, full-face (D), lateral (K). Pheidole ovalinoda, full-face (E), lateral (L). Pheidole tenebrovulgaris, full-face (F), lateral (M). Pheidole uranus, full-face (G), lateral (N)." figureDoi="http://doi.org/10.5281/zenodo.4405737" httpUri="https://zenodo.org/record/4405737/files/figure.png" pageId="8" pageNumber="9">Figs 3C</figureCitation>
,
<figureCitation id="13012DB6FF86A91EA5FBB76DFD14FE86" box="[692,722,357,379]" captionStart="FIGURE 5" captionStartId="8.[151,250,1976,2000]" captionTargetBox="[189,1397,1124,1952]" captionTargetId="figure-433@8.[189,1397,1124,1952]" captionTargetPageId="8" captionText="FIGURE 5. Minor worker, profile. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405743" httpUri="https://zenodo.org/record/4405743/files/figure.png" pageId="8" pageNumber="9">5C</figureCitation>
)..............................................
<taxonomicName id="4C3A4AB0FF86A91EA240B76DFA5AFE87" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1295,1436,357,378]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FF86A91EA240B76DFA5AFE87" box="[1295,1436,357,378]" italics="true" pageId="8" pageNumber="9">Pheidole maro</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF86A91EA7D8B78BFA5AFE2F" blockId="8.[151,1437,151,467]" pageId="8" pageNumber="9">
<keyLead id="30CB9206FF86A91EA7D8B78BFA5AFE2F" pageId="8" pageNumber="9">
- Major worker. Antennal scrobe with sparse, very thin rugulae with smooth interspaces, mesosoma smooth with very indistinct and thin rugulae, inner hypostomal tooth small, body yellow to orange (
<figureCitation id="13012DB6FF86A91EA4D6B797FC2EFE48" box="[921,1000,415,437]" captionStart="FIGURE 2" captionStartId="7.[151,250,1070,1094]" captionTargetBox="[151,1436,433,1046]" captionTargetId="figure-387@7.[151,1436,433,1047]" captionTargetPageId="7" captionText="FIGURE 2. Major worker, head. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405735" httpUri="https://zenodo.org/record/4405735/files/figure.png" pageId="8" pageNumber="9">Figs 2B</figureCitation>
,
<figureCitation id="13012DB6FF86A91EA4BCB7A8FBEBFE48" box="[1011,1069,416,437]" captionStart="FIGURE 4" captionStartId="8.[151,250,1039,1063]" captionTargetBox="[187,1400,491,1016]" captionTargetId="figure-388@8.[187,1400,491,1016]" captionTargetPageId="8" captionText="FIGURE 4. Major worker. Pheidole decollata, profile (A). Pheidole madecassa, profile (B), hypostomal tooth (E). Pheidole maro, profile (C), hypostomal tooth (F). Pheidole oswaldi, profile (D). Pheidole ovalinoda, hypostomal tooth (G)." figureDoi="http://doi.org/10.5281/zenodo.4405741" httpUri="https://zenodo.org/record/4405741/files/figure.png" pageId="8" pageNumber="9">4B, E</figureCitation>
). Minor worker. Frons and mesosoma smooth, only anterolateral frons with weakly developed puncta (
<figureCitation id="13012DB6FF86A91EA403B7B5FC5CFE2F" box="[844,922,445,467]" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1940]" captionTargetBox="[152,1433,1160,1893]" captionTargetId="figure-422@7.[151,1436,1155,1893]" captionTargetPageId="7" captionText="FIGURE 3. Minor worker, head. Pheidole decollata, full-face (A), lateral (H). Pheidole madecassa, full-face (B), lateral (I). Pheidole maro, full-face (C), lateral (J). Pheidole oswaldi, full-face (D), lateral (K). Pheidole ovalinoda, full-face (E), lateral (L). Pheidole tenebrovulgaris, full-face (F), lateral (M). Pheidole uranus, full-face (G), lateral (N)." figureDoi="http://doi.org/10.5281/zenodo.4405737" httpUri="https://zenodo.org/record/4405737/files/figure.png" pageId="8" pageNumber="9">Figs 3B</figureCitation>
,
<figureCitation id="13012DB6FF86A91EA4EBB7B5FC05FE2E" box="[932,963,445,467]" captionStart="FIGURE 5" captionStartId="8.[151,250,1976,2000]" captionTargetBox="[189,1397,1124,1952]" captionTargetId="figure-433@8.[189,1397,1124,1952]" captionTargetPageId="8" captionText="FIGURE 5. Minor worker, profile. Pheidole decollata (A). Pheidole madecassa (B). Pheidole maro (C). Pheidole oswaldi (D). Pheidole ovalinoda (E). Pheidole tenebrovulgaris (F). Pheidole uranus (G)." figureDoi="http://doi.org/10.5281/zenodo.4405743" httpUri="https://zenodo.org/record/4405743/files/figure.png" pageId="8" pageNumber="9">5B</figureCitation>
).....................
<taxonomicName id="4C3A4AB0FF86A91EA397B7B5FA5AFE2F" ID-CoL="4FYFF" authorityName="Forel" authorityYear="1892" box="[1240,1436,445,466]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FF86A91EA397B7B5FA5AFE2F" box="[1240,1436,445,466]" italics="true" pageId="8" pageNumber="9">Pheidole madecassa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF4561BBFF86A91EA7D8B207FADDFBB1" ID-DOI="http://doi.org/10.5281/zenodo.4405741" ID-Zenodo-Dep="4405741" httpUri="https://zenodo.org/record/4405741/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1039,1063]" targetBox="[187,1400,491,1016]" targetPageId="8">
<paragraph id="8B853133FF86A91EA7D8B207FADDFBB1" blockId="8.[151,1436,1039,1100]" pageId="8" pageNumber="9">
<emphasis id="B94EED21FF86A91EA7D8B207FED2FBDA" bold="true" box="[151,276,1039,1063]" pageId="8" pageNumber="9">FIGURE 4.</emphasis>
Major worker.
<taxonomicName id="4C3A4AB0FF86A91EA6F9B218FDB0FBD5" authorityName="Forel" authorityYear="1892" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[438,630,1040,1064]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="decollata">
<emphasis id="B94EED21FF86A91EA6F9B218FDB0FBD5" box="[438,630,1040,1064]" italics="true" pageId="8" pageNumber="9">Pheidole decollata</emphasis>
</taxonomicName>
, profile (A).
<taxonomicName id="4C3A4AB0FF86A91EA44FB218FC17FBD5" ID-CoL="4FYFF" authorityName="Forel" authorityYear="1892" box="[768,977,1040,1064]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FF86A91EA44FB218FC17FBD5" box="[768,977,1040,1064]" italics="true" pageId="8" pageNumber="9">Pheidole madecassa</emphasis>
</taxonomicName>
, profile (B), hypostomal tooth (E).
<taxonomicName id="4C3A4AB0FF86A91EA20DB218FF0AFBB6" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FF86A91EA20DB218FF0AFBB6" italics="true" pageId="8" pageNumber="9">Pheidole maro</emphasis>
</taxonomicName>
, profile (C), hypostomal tooth (F).
<taxonomicName id="4C3A4AB0FF86A91EA57CB23CFD25FBB1" ID-CoL="4FYHF" authorityName="Forel" authorityYear="1891" box="[563,739,1076,1100]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="B94EED21FF86A91EA57CB23CFD25FBB1" box="[563,739,1076,1100]" italics="true" pageId="8" pageNumber="9">Pheidole oswaldi</emphasis>
</taxonomicName>
, profile (D).
<taxonomicName id="4C3A4AB0FF86A91EA426B23CFBE8FBB1" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[873,1070,1076,1100]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FF86A91EA426B23CFBE8FBB1" box="[873,1070,1076,1100]" italics="true" pageId="8" pageNumber="9">Pheidole ovalinoda</emphasis>
</taxonomicName>
, hypostomal tooth (G).
</paragraph>
</caption>
<caption id="DF4561BBFF86A91EA7D8B1B0FC5EF809" ID-DOI="http://doi.org/10.5281/zenodo.4405743" ID-Zenodo-Dep="4405743" httpUri="https://zenodo.org/record/4405743/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1976,2000]" targetBox="[189,1397,1124,1952]" targetPageId="8">
<paragraph id="8B853133FF86A91EA7D8B1B0FC5EF809" blockId="8.[151,1437,1976,2036]" pageId="8" pageNumber="9">
<emphasis id="B94EED21FF86A91EA7D8B1B0FED4F82D" bold="true" box="[151,274,1976,2000]" pageId="8" pageNumber="9">FIGURE 5.</emphasis>
Minor worker, profile.
<taxonomicName id="4C3A4AB0FF86A91EA54FB1B0FD78F82D" authorityName="Forel" authorityYear="1892" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[512,702,1976,2000]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="decollata">
<emphasis id="B94EED21FF86A91EA54FB1B0FD78F82D" box="[512,702,1976,2000]" italics="true" pageId="8" pageNumber="9">Pheidole decollata</emphasis>
</taxonomicName>
(A).
<taxonomicName id="4C3A4AB0FF86A91EA5BCB1B0FC04F82D" ID-CoL="4FYFF" authorityName="Forel" authorityYear="1892" box="[755,962,1976,2000]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="madecassa">
<emphasis id="B94EED21FF86A91EA5BCB1B0FC04F82D" box="[755,962,1976,2000]" italics="true" pageId="8" pageNumber="9">Pheidole madecassa</emphasis>
</taxonomicName>
(B).
<taxonomicName id="4C3A4AB0FF86A91EA4B9B1B0FB4DF82D" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1014,1163,1976,2000]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="maro">
<emphasis id="B94EED21FF86A91EA4B9B1B0FB4DF82D" box="[1014,1163,1976,2000]" italics="true" pageId="8" pageNumber="9">Pheidole maro</emphasis>
</taxonomicName>
(C).
<taxonomicName id="4C3A4AB0FF86A91EA3F1B1B0FAABF82D" ID-CoL="4FYHF" authorityName="Forel" authorityYear="1891" box="[1214,1389,1976,2000]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="oswaldi">
<emphasis id="B94EED21FF86A91EA3F1B1B0FAABF82D" box="[1214,1389,1976,2000]" italics="true" pageId="8" pageNumber="9">Pheidole oswaldi</emphasis>
</taxonomicName>
(D).
<taxonomicName id="4C3A4AB0FF86A91EA7D8B1D4FE9BF809" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,349,2012,2036]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ovalinoda">
<emphasis id="B94EED21FF86A91EA7D8B1D4FE9BF809" box="[151,349,2012,2036]" italics="true" pageId="8" pageNumber="9">Pheidole ovalinoda</emphasis>
</taxonomicName>
(E).
<taxonomicName id="4C3A4AB0FF86A91EA6DFB1D4FD56F809" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[400,656,2012,2036]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tenebrovulgaris">
<emphasis id="B94EED21FF86A91EA6DFB1D4FD56F809" box="[400,656,2012,2036]" italics="true" pageId="8" pageNumber="9">Pheidole tenebrovulgaris</emphasis>
</taxonomicName>
(F).
<taxonomicName id="4C3A4AB0FF86A91EA58DB1D4FCAEF809" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[706,872,2012,2036]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="uranus">
<emphasis id="B94EED21FF86A91EA58DB1D4FCAEF809" box="[706,872,2012,2036]" italics="true" pageId="8" pageNumber="9">Pheidole uranus</emphasis>
</taxonomicName>
(G).
</paragraph>
</caption>
<keyStep id="30CE2996FF87A91FA7D8B690FA5AFF37" pageId="9" pageNumber="10">
<paragraph id="8B853133FF87A91FA7D8B690FA5AFF50" blockId="9.[151,1438,151,555]" box="[151,1436,151,173]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B690FA5AFF50" box="[151,1436,151,173]" pageId="9" pageNumber="10">8 At least minor worker yellow to orange.................................................................... 9</keyLead>
</paragraph>
<paragraph id="8B853133FF87A91FA7D8B6BDFA5AFF37" blockId="9.[151,1438,151,555]" box="[151,1436,181,203]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B6BDFA5AFF37" box="[151,1436,181,203]" pageId="9" pageNumber="10">- Minor and major worker bright brown to brownish black..................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF87A91FA7D8B6DAFA5AFEC2" pageId="9" pageNumber="10">
<paragraph id="8B853133FF87A91FA7D8B6DAFA5AFEF9" blockId="9.[151,1438,151,555]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B6DAFA5AFEF9" pageId="9" pageNumber="10">
9 Major worker. Head elongated, posteromedian concavity very deep, CI&gt; 112.0, HL&gt;
<quantity id="4CC29CD6FF87A91FA36AB6DAFBA9FF15" box="[1061,1135,210,232]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="10" unit="mm" value="1.8">1.8 mm</quantity>
(
<figureCitation id="13012DB6FF87A91FA332B6DAFB01FF15" box="[1149,1223,210,232]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="9" pageNumber="10">Fig. 6F</figureCitation>
). Minor worker. Head with moderately long and narrow neck (
<figureCitation id="13012DB6FF87A91FA52DB6E7FD00FEFB" box="[610,710,239,262]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="9" pageNumber="10">Fig. 7F, Q</figureCitation>
)........................................
<taxonomicName id="4C3A4AB0FF87A91FA3F3B6E7FA5AFEF9" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1212,1436,239,260]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="flavodepressa">
<emphasis id="B94EED21FF87A91FA3F3B6E7FA5AFEF9" box="[1212,1436,239,260]" italics="true" pageId="9" pageNumber="10">Pheidole flavodepressa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF87A91FA7D8B705FA5AFEC2" blockId="9.[151,1438,151,555]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B705FA5AFEC2" pageId="9" pageNumber="10">
- Major worker. Head not elongated, posteromedian concavity deep, CI &lt;112.0, HL &lt;
<quantity id="4CC29CD6FF87A91FA35FB705FB9FFEDF" box="[1040,1113,269,290]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="10" unit="mm" value="1.8">1.8 mm</quantity>
(
<figureCitation id="13012DB6FF87A91FA329B705FAC6FEDF" box="[1126,1280,269,291]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="9" pageNumber="10">Fig. 6A, D, H, I</figureCitation>
). Minor worker. Head with short and moderately thick neck (
<figureCitation id="13012DB6FF87A91FA5C3B722FC52FEBD" box="[652,916,298,320]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="9" pageNumber="10">Fig. 7A, D, H, I, L, O, S, T</figureCitation>
)........................................ 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF87A91FA7D8B740FA5AFE48" pageId="9" pageNumber="10">
<paragraph id="8B853133FF87A91FA7D8B740FA5BFE87" blockId="9.[151,1438,151,555]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B740FA5BFE87" pageId="9" pageNumber="10">
10 Major worker. Body brown, costulae on posterior frons arching posterolaterally (
<figureCitation id="13012DB6FF87A91FA4A4B74FFBFEFEA0" box="[1003,1080,327,349]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="9" pageNumber="10">Fig. 6A</figureCitation>
). Minor worker. Promesonotum with weakly developed network of sparse microreticulae (
<figureCitation id="13012DB6FF87A91FA591B76DFCEAFE86" box="[734,812,357,379]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="9" pageNumber="10">Fig. 8A</figureCitation>
)...................................
<taxonomicName id="4C3A4AB0FF87A91FA391B76DFA5BFE87" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1246,1437,357,378]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="antsahabe">
<emphasis id="B94EED21FF87A91FA391B76DFA5BFE87" box="[1246,1437,357,378]" italics="true" pageId="9" pageNumber="10">Pheidole antsahabe</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF87A91FA7D8B78BFA5AFE48" blockId="9.[151,1438,151,555]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B78BFA5AFE48" pageId="9" pageNumber="10">
- Major worker. Body yellow to orange, costulae on posterior frons never arching posterolaterally (
<figureCitation id="13012DB6FF87A91FA3CBB78AFAC6FE65" box="[1156,1280,386,408]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="9" pageNumber="10">Fig. 6D, H, I</figureCitation>
). Minor worker. Promesonotum smooth or distinctly punctate (
<figureCitation id="13012DB6FF87A91FA5D1B797FCDDFE48" box="[670,795,415,437]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="9" pageNumber="10">Fig. 8D, H, I</figureCitation>
).................................................. 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF87A91FA7D8B7B5FA5AFDD7" pageId="9" pageNumber="10">
<paragraph id="8B853133FF87A91FA7D8B7B5FA5BFE12" blockId="9.[151,1438,151,555]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B7B5FA5BFE12" pageId="9" pageNumber="10">
11 Major worker. Head with lateral sides relatively straight, frons laterally with thick rugocostulae and rugulate interspaces (
<figureCitation id="13012DB6FF87A91FA23AB7B5FF3EFE0D" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="9" pageNumber="10">Fig. 6I</figureCitation>
). Minor worker. Promesonotum smooth, frons mostly smooth (
<figureCitation id="13012DB6FF87A91FA418B7D2FC5BFE0D" box="[855,925,474,496]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="9" pageNumber="10">Figs 7I</figureCitation>
,
<figureCitation id="13012DB6FF87A91FA4E8B7D2FC78FE0D" box="[935,958,474,496]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="9" pageNumber="10">8I</figureCitation>
)..........................
<taxonomicName id="4C3A4AB0FF87A91FA24DB7D2FA5BFE12" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1282,1437,474,495]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B94EED21FF87A91FA24DB7D2FA5BFE12" box="[1282,1437,474,495]" italics="true" pageId="9" pageNumber="10">Pheidole similis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF87A91FA7D8B7F0FA5AFDD7" blockId="9.[151,1438,151,555]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B7F0FA5AFDD7" pageId="9" pageNumber="10">
- Major worker. Head with lateral sides slightly convex, frons laterally sparsely rugopunctate (
<figureCitation id="13012DB6FF87A91FA300B7FFFB71FDF0" box="[1103,1207,503,525]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="9" pageNumber="10">Fig. 6D, H</figureCitation>
). Minor worker. Promesonotum punctate, frons weakly punctate (
<figureCitation id="13012DB6FF87A91FA537B41DFD23FDD7" box="[632,741,533,555]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="9" pageNumber="10">Figs 7D, H</figureCitation>
,
<figureCitation id="13012DB6FF87A91FA5BFB41DFCEBFDD7" box="[752,813,533,554]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="9" pageNumber="10">8D, H</figureCitation>
)................................................. 12
</keyLead>
</paragraph>
</keyStep>
<caption id="DF4561BBFF87A91FA7D8B3FEFD99F9AB" ID-DOI="http://doi.org/10.5281/zenodo.4405745" ID-Zenodo-Dep="4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1526,1550]" targetBox="[151,1436,579,1502]" targetPageId="9">
<paragraph id="8B853133FF87A91FA7D8B3FEFD99F9AB" blockId="9.[151,1436,1526,1622]" pageId="9" pageNumber="10">
<emphasis id="B94EED21FF87A91FA7D8B3FEFED5F9F3" bold="true" box="[151,275,1526,1550]" pageId="9" pageNumber="10">FIGURE 6.</emphasis>
Major worker, head.
<taxonomicName id="4C3A4AB0FF87A91FA6A0B3FEFD7EF9F3" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[495,696,1526,1550]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="antsahabe">
<emphasis id="B94EED21FF87A91FA6A0B3FEFD7EF9F3" box="[495,696,1526,1550]" italics="true" pageId="9" pageNumber="10">Pheidole antsahabe</emphasis>
</taxonomicName>
(A).
<taxonomicName id="4C3A4AB0FF87A91FA5A0B3FEFC21F9F3" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[751,999,1526,1550]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="atsirakambiaty">
<emphasis id="B94EED21FF87A91FA5A0B3FEFC21F9F3" box="[751,999,1526,1550]" italics="true" pageId="9" pageNumber="10">Pheidole atsirakambiaty</emphasis>
</taxonomicName>
(B).
<taxonomicName id="4C3A4AB0FF87A91FA352B3FEFB17F9F3" ID-CoL="4FY85" authorityName="Forel" authorityYear="1891" box="[1053,1233,1526,1550]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF87A91FA352B3FEFB17F9F3" box="[1053,1233,1526,1550]" italics="true" pageId="9" pageNumber="10">Pheidole bessonii</emphasis>
</taxonomicName>
(C).
<taxonomicName id="4C3A4AB0FF87A91FA249B3FEFA5AF9F3" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1286,1436,1526,1550]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="clara">
<emphasis id="B94EED21FF87A91FA249B3FEFA5AF9F3" box="[1286,1436,1526,1550]" italics="true" pageId="9" pageNumber="10">Pheidole clara</emphasis>
</taxonomicName>
(D).
<taxonomicName id="4C3A4AB0FF87A91FA787B012FE46F9CF" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[200,384,1562,1586]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FF87A91FA787B012FE46F9CF" box="[200,384,1562,1586]" italics="true" pageId="9" pageNumber="10">Pheidole flammea</emphasis>
</taxonomicName>
(E).
<taxonomicName id="4C3A4AB0FF87A91FA6FAB012FD64F9CF" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[437,674,1562,1586]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="flavodepressa">
<emphasis id="B94EED21FF87A91FA6FAB012FD64F9CF" box="[437,674,1562,1586]" italics="true" pageId="9" pageNumber="10">Pheidole flavodepressa</emphasis>
</taxonomicName>
(F).
<taxonomicName id="4C3A4AB0FF87A91FA59AB012FC51F9CF" ID-CoL="4FYCW" authorityName="Emery" authorityYear="1899" box="[725,919,1562,1586]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="grallatrix">
<emphasis id="B94EED21FF87A91FA59AB012FC51F9CF" box="[725,919,1562,1586]" italics="true" pageId="9" pageNumber="10">Pheidole grallatrix</emphasis>
</taxonomicName>
(G).
<taxonomicName id="4C3A4AB0FF87A91FA481B012FB07F9CF" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[974,1217,1562,1586]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mantadioflava">
<emphasis id="B94EED21FF87A91FA481B012FB07F9CF" box="[974,1217,1562,1586]" italics="true" pageId="9" pageNumber="10">Pheidole mantadioflava</emphasis>
</taxonomicName>
(H).
<taxonomicName id="4C3A4AB0FF87A91FA3B6B012FA5AF9CF" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1273,1436,1562,1586]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B94EED21FF87A91FA3B6B012FA5AF9CF" box="[1273,1436,1562,1586]" italics="true" pageId="9" pageNumber="10">Pheidole similis</emphasis>
</taxonomicName>
(I).
<taxonomicName id="4C3A4AB0FF87A91FA7F2B036FEAEF9AB" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[189,360,1598,1622]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="voreios">
<emphasis id="B94EED21FF87A91FA7F2B036FEAEF9AB" box="[189,360,1598,1622]" italics="true" pageId="9" pageNumber="10">Pheidole voreios</emphasis>
</taxonomicName>
(J).
<taxonomicName id="4C3A4AB0FF87A91FA6DAB036FDE9F9AB" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[405,559,1598,1622]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="zirafy">
<emphasis id="B94EED21FF87A91FA6DAB036FDE9F9AB" box="[405,559,1598,1622]" italics="true" pageId="9" pageNumber="10">Pheidole zirafy</emphasis>
</taxonomicName>
(K).
</paragraph>
</caption>
<keyStep id="30CE2996FF87A91FA7D8B089FA5BF8D4" pageId="9" pageNumber="10">
<paragraph id="8B853133FF87A91FA7D8B089FA5BF92C" blockId="9.[151,1437,1665,2039]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B089FA5BF92C" pageId="9" pageNumber="10">
12 Major worker. Antennal scrobes very indistinct, mostly punctate with additional network of thin and sparse rugae, anterior mesonotum placed slightly higher than pronotum (
<figureCitation id="13012DB6FF87A91FA5E5B096FD3FF949" box="[682,761,1694,1716]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="9" pageNumber="10">Figs 6H</figureCitation>
,
<figureCitation id="13012DB6FF87A91FA44CB097FCE2F948" box="[771,804,1695,1717]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="9" pageNumber="10">8N</figureCitation>
). Minor worker. Frons with sparse puncta, propodeal spines small with wide base (
<figureCitation id="13012DB6FF87A91FA630B0B4FE08F92C" box="[383,462,1724,1746]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="9" pageNumber="10">Figs 7H</figureCitation>
,
<figureCitation id="13012DB6FF87A91FA697B0B4FE3FF92F" box="[472,505,1724,1746]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="9" pageNumber="10">8H</figureCitation>
).........................................................
<taxonomicName id="4C3A4AB0FF87A91FA3F9B0B4FA5BF92C" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1206,1437,1724,1745]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mantadioflava">
<emphasis id="B94EED21FF87A91FA3F9B0B4FA5BF92C" box="[1206,1437,1724,1745]" italics="true" pageId="9" pageNumber="10">Pheidole mantadioflava</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF87A91FA7D8B0D2FA5BF8D4" blockId="9.[151,1437,1665,2039]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B0D2FA5BF8D4" pageId="9" pageNumber="10">
- Major worker. Antennal scrobes absent, frons laterally with thick and dense rugae with distinctly punctate interspaces, anterior mesonotum placed lower than pronotum (
<figureCitation id="13012DB6FF87A91FA53AB0FEFD03F8F1" box="[629,709,1782,1804]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="9" pageNumber="10">Figs 6D</figureCitation>
,
<figureCitation id="13012DB6FF87A91FA580B0FFFD2BF8F0" box="[719,749,1783,1805]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="9" pageNumber="10">8L</figureCitation>
). Minor worker. Only anterolateral frons with sparse puncta, propodeal spines small and thin (
<figureCitation id="13012DB6FF87A91FA6F6B11CFDCEF8D4" box="[441,520,1812,1834]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="9" pageNumber="10">Figs 7D</figureCitation>
,
<figureCitation id="13012DB6FF87A91FA55CB11CFDF4F8D7" box="[531,562,1812,1834]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="9" pageNumber="10">8D</figureCitation>
)............................................................
<taxonomicName id="4C3A4AB0FF87A91FA241B11CFA5BF8D4" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1294,1437,1812,1833]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="clara">
<emphasis id="B94EED21FF87A91FA241B11CFA5BF8D4" box="[1294,1437,1812,1833]" italics="true" pageId="9" pageNumber="10">Pheidole clara</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF87A91FA7D8B139FA5AF824" pageId="9" pageNumber="10">
<paragraph id="8B853133FF87A91FA7D8B139FA5BF87C" blockId="9.[151,1437,1665,2039]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B139FA5BF87C" pageId="9" pageNumber="10">
13 Major worker. Occipital lobes with thick, sparse rugae with smooth interspaces, frons laterally with thick, dense rugae with smooth to indistinctly rugulate interspaces (
<figureCitation id="13012DB6FF87A91FA5C0B146FD1DF899" box="[655,731,1870,1892]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="9" pageNumber="10">Fig. 6E</figureCitation>
). Minor worker. Head with short and moderately narrow neck (
<figureCitation id="13012DB6FF87A91FA202B146FF34F87C" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="9" pageNumber="10">Fig. 7E, P</figureCitation>
)....................................................................................
<taxonomicName id="4C3A4AB0FF87A91FA3A1B164FA5BF87C" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1262,1437,1900,1921]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FF87A91FA3A1B164FA5BF87C" box="[1262,1437,1900,1921]" italics="true" pageId="9" pageNumber="10">Pheidole flammea</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF87A91FA7D8B182FA5AF824" blockId="9.[151,1437,1665,2039]" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91FA7D8B182FA5AF824" pageId="9" pageNumber="10">
- Major worker. Occipital lobes with very thin, dense rugae, most often with sculptured interspaces, frons laterally with thin, dense rugae with distinctly sculptured interspaces (
<figureCitation id="13012DB6FF87A91FA584B1AEFC47F841" box="[715,897,1958,1980]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="9" pageNumber="10">Fig. 6B, C, G, J, K</figureCitation>
). Minor worker. Head without a neck or with a long and narrow neck (
<figureCitation id="13012DB6FF87A91FA629B1CCFD76F824" box="[358,688,1988,2010]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="9" pageNumber="10">Fig. 7B, C, G, J, K, M, N, R, U, V</figureCitation>
)........................................................... 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF87A91CA7D8B1E9FA5AFEF8" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<paragraph id="8B853133FF87A91CA7D8B1E9FA5AFF36" blockId="9.[151,1437,1665,2039]" lastBlockId="10.[151,1437,151,408]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<keyLead id="30CB9206FF87A91CA7D8B1E9FA5AFF36" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
14 Major worker. Head elongated, posteromedian concavity very deep, HL&gt;
<quantity id="4CC29CD6FF87A91FA4FFB1E9FC3AF80A" box="[944,1020,2017,2039]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="10" unit="mm" value="2.4">2.4 mm</quantity>
, CI&gt; 110.0 or head not elongated and posteromedian concavity deep and mesosoma mostly smooth (
<figureCitation id="13012DB6FF84A91CA45CB69FFCB8FF50" box="[787,894,151,173]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="10" pageNumber="11">Figs 6G, K</figureCitation>
,
<figureCitation id="13012DB6FF84A91CA4C8B690FC0EFF53" box="[903,968,152,174]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="10" pageNumber="11">8M, O</figureCitation>
). Minor worker. Head with long and narrow neck (
<figureCitation id="13012DB6FF84A91CA7A6B6BDFE4FFF37" box="[233,393,181,203]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="10" pageNumber="11">Fig. 7G, K, R, V</figureCitation>
).................................................................................... 15
</keyLead>
</paragraph>
<paragraph id="8B853133FF84A91CA7D8B6DBFA5AFEF8" blockId="10.[151,1437,151,408]" pageId="10" pageNumber="11">
<keyLead id="30CB9206FF84A91CA7D8B6DBFA5AFEF8" pageId="10" pageNumber="11">
- Major worker. Head not elongated, posteromedian concavity deep, and mesosoma entirely sculptured, HL &lt;
<quantity id="4CC29CD6FF84A91CA3AAB6DAFAE9FF15" box="[1253,1327,210,232]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="11" unit="mm" value="2.4">2.4 mm</quantity>
, CI &lt;110.0 (
<figureCitation id="13012DB6FF84A91CA7A6B6E7FEAEFEF8" box="[233,360,239,261]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="10" pageNumber="11">Figs 6B, C, J</figureCitation>
,
<figureCitation id="13012DB6FF84A91CA63CB6F8FE59FEFB" box="[371,415,240,262]" captionStart="FIGURE 10" captionStartId="15.[151,250,1956,1980]" captionTargetBox="[151,1436,453,1933]" captionTargetId="figure-137@15.[151,1436,453,1933]" captionTargetPageId="15" captionText="FIGURE 10. Pheidole atsirakambiaty, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0491661) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0491660)." figureDoi="http://doi.org/10.5281/zenodo.4405757" httpUri="https://zenodo.org/record/4405757/files/figure.png" pageId="10" pageNumber="11">10D</figureCitation>
). Minor worker. Head without neck. (
<figureCitation id="13012DB6FF84A91CA446B6E7FC24FEF8" box="[777,994,239,261]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="10" pageNumber="11">Fig. 7B, C, J, M, N, U</figureCitation>
).................................. 16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF84A91CA7D8B705FA5AFE6A" pageId="10" pageNumber="11">
<paragraph id="8B853133FF84A91CA7D8B705FA5BFEA0" blockId="10.[151,1437,151,408]" pageId="10" pageNumber="11">
<keyLead id="30CB9206FF84A91CA7D8B705FA5BFEA0" pageId="10" pageNumber="11">
15 Major worker. Head elongated, posteromedian concavity very deep, and mesosoma entirely sculptured (
<figureCitation id="13012DB6FF84A91CA39DB705FAE5FEDE" box="[1234,1315,269,291]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="10" pageNumber="11">Figs 6K</figureCitation>
,
<figureCitation id="13012DB6FF84A91CA261B705FA88FEDE" box="[1326,1358,269,291]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="10" pageNumber="11">8O</figureCitation>
). Minor worker. Neck very long, promesonotal and propodeal dorsum mostly smooth with sparse, transverse rugae, propodeal side and mesopleuron densely rugoreticulate (
<figureCitation id="13012DB6FF84A91CA50AB74FFD6BFEA0" box="[581,685,327,349]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="10" pageNumber="11">Figs 7K, V</figureCitation>
,
<figureCitation id="13012DB6FF84A91CA5F6B740FD1CFEA3" box="[697,730,328,350]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="10" pageNumber="11">8K</figureCitation>
).............................................
<taxonomicName id="4C3A4AB0FF84A91CA245B74FFA5BFEA0" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1290,1437,327,349]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="zirafy">
<emphasis id="B94EED21FF84A91CA245B74FFA5BFEA0" box="[1290,1437,327,349]" italics="true" pageId="10" pageNumber="11">Pheidole zirafy</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF84A91CA7D8B76DFA5AFE6A" blockId="10.[151,1437,151,408]" pageId="10" pageNumber="11">
<keyLead id="30CB9206FF84A91CA7D8B76DFA5AFE6A" pageId="10" pageNumber="11">
- Major worker. Head not elongated and posteromedian concavity deep, mesosoma mostly smooth (
<figureCitation id="13012DB6FF84A91CA3C9B76DFB15FE87" box="[1158,1235,357,379]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="10" pageNumber="11">Figs 6G</figureCitation>
,
<figureCitation id="13012DB6FF84A91CA391B76DFAC4FE86" box="[1246,1282,357,379]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="10" pageNumber="11">8M</figureCitation>
). Minor worker. Neck long, mesosoma smooth (
<figureCitation id="13012DB6FF84A91CA55FB78AFDBAFE65" box="[528,636,386,408]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="10" pageNumber="11">Figs 7G, R</figureCitation>
,
<figureCitation id="13012DB6FF84A91CA5C9B78AFD65FE6A" box="[646,675,386,407]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="10" pageNumber="11">8G</figureCitation>
)..............................................
<taxonomicName id="4C3A4AB0FF84A91CA3AAB78AFA5AFE6A" ID-CoL="4FYCW" authorityName="Emery" authorityYear="1899" box="[1253,1436,386,407]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="grallatrix">
<emphasis id="B94EED21FF84A91CA3AAB78AFA5AFE6A" box="[1253,1436,386,407]" italics="true" pageId="10" pageNumber="11">Pheidole grallatrix</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="DF4561BBFF84A91CA7D8B026FE1AF92A" ID-DOI="http://doi.org/10.5281/zenodo.4405749" ID-Zenodo-Dep="4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,250,1582,1606]" targetBox="[151,1436,433,1559]" targetPageId="10">
<paragraph id="8B853133FF84A91CA7D8B026FE1AF92A" blockId="10.[151,1437,1582,1751]" pageId="10" pageNumber="11">
<emphasis id="B94EED21FF84A91CA7D8B026FED5F9BA" bold="true" box="[151,275,1582,1607]" pageId="10" pageNumber="11">FIGURE 7.</emphasis>
Minor worker, head.
<taxonomicName id="4C3A4AB0FF84A91CA6BFB026FD7EF9BB" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[496,696,1582,1606]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="antsahabe">
<emphasis id="B94EED21FF84A91CA6BFB026FD7EF9BB" box="[496,696,1582,1606]" italics="true" pageId="10" pageNumber="11">Pheidole antsahabe</emphasis>
</taxonomicName>
, full-face (A), profile (L).
<taxonomicName id="4C3A4AB0FF84A91CA486B026FB07F9BB" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[969,1217,1582,1606]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="atsirakambiaty">
<emphasis id="B94EED21FF84A91CA486B026FB07F9BB" box="[969,1217,1582,1606]" italics="true" pageId="10" pageNumber="11">Pheidole atsirakambiaty</emphasis>
</taxonomicName>
, full-face (B), profile (M).
<taxonomicName id="4C3A4AB0FF84A91CA782B05AFE44F997" ID-CoL="4FY85" authorityName="Forel" authorityYear="1891" box="[205,386,1618,1642]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF84A91CA782B05AFE44F997" box="[205,386,1618,1642]" italics="true" pageId="10" pageNumber="11">Pheidole bessonii</emphasis>
</taxonomicName>
, full-face (C), profile (N).
<taxonomicName id="4C3A4AB0FF84A91CA5D3B05AFCF2F997" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[668,820,1618,1642]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="clara">
<emphasis id="B94EED21FF84A91CA5D3B05AFCF2F997" box="[668,820,1618,1642]" italics="true" pageId="10" pageNumber="11">Pheidole clara</emphasis>
</taxonomicName>
, full-face (D), profile (O).
<taxonomicName id="4C3A4AB0FF84A91CA31FB05AFACFF997" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1104,1289,1618,1642]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FF84A91CA31FB05AFACFF997" box="[1104,1289,1618,1642]" italics="true" pageId="10" pageNumber="11">Pheidole flammea</emphasis>
</taxonomicName>
, full-face (E), profile (P).
<taxonomicName id="4C3A4AB0FF84A91CA643B07EFE3EF973" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[268,504,1654,1678]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="flavodepressa">
<emphasis id="B94EED21FF84A91CA643B07EFE3EF973" box="[268,504,1654,1678]" italics="true" pageId="10" pageNumber="11">Pheidole flavodepressa</emphasis>
</taxonomicName>
, full-face (F), profile (Q).
<taxonomicName id="4C3A4AB0FF84A91CA44BB07EFC03F973" ID-CoL="4FYCW" authorityName="Emery" authorityYear="1899" box="[772,965,1654,1678]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="grallatrix">
<emphasis id="B94EED21FF84A91CA44BB07EFC03F973" box="[772,965,1654,1678]" italics="true" pageId="10" pageNumber="11">Pheidole grallatrix</emphasis>
</taxonomicName>
, full-face (G), profile (R).
<taxonomicName id="4C3A4AB0FF84A91CA39BB07EFF0CF94F" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mantadioflava">
<emphasis id="B94EED21FF84A91CA39BB07EFF0CF94F" italics="true" pageId="10" pageNumber="11">Pheidole mantadioflava</emphasis>
</taxonomicName>
, full-face (H), profile (S).
<taxonomicName id="4C3A4AB0FF84A91CA693B092FDB9F94F" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[476,639,1690,1714]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B94EED21FF84A91CA693B092FDB9F94F" box="[476,639,1690,1714]" italics="true" pageId="10" pageNumber="11">Pheidole similis</emphasis>
</taxonomicName>
, full-face (I), profile (T).
<taxonomicName id="4C3A4AB0FF84A91CA4C6B092FBF2F94F" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[905,1076,1690,1714]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="voreios">
<emphasis id="B94EED21FF84A91CA4C6B092FBF2F94F" box="[905,1076,1690,1714]" italics="true" pageId="10" pageNumber="11">Pheidole voreios</emphasis>
</taxonomicName>
, full-face (J), profile (U).
<taxonomicName id="4C3A4AB0FF84A91CA20DB092FF17F92B" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="zirafy">
<emphasis id="B94EED21FF84A91CA20DB092FF17F92B" italics="true" pageId="10" pageNumber="11">Pheidole zirafy</emphasis>
</taxonomicName>
, full-face (K), profile (V).
</paragraph>
</caption>
<keyStep id="30CE2996FF84A91CA7D8B109FA5AF854" pageId="10" pageNumber="11">
<paragraph id="8B853133FF84A91CA7D8B109FA5AF8AC" blockId="10.[151,1437,1793,2021]" pageId="10" pageNumber="11">
<keyLead id="30CB9206FF84A91CA7D8B109FA5AF8AC" pageId="10" pageNumber="11">
16 Major worker. Body black, frons and occipital lobes with moderately thick, moderately dense costulae that arch posterolaterally, interspaces between costulae smooth to indistinctly rugopunctate (
<figureCitation id="13012DB6FF84A91CA4CBB117FC09F8C8" box="[900,975,1823,1845]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="10" pageNumber="11">Fig. 6B</figureCitation>
). Minor worker. Body black, frons punctate with few rugae, mesosoma distinctly punctate (
<figureCitation id="13012DB6FF84A91CA539B134FD02F8AF" box="[630,708,1852,1874]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="10" pageNumber="11">Figs 7B</figureCitation>
,
<figureCitation id="13012DB6FF84A91CA581B134FD2BF8AF" box="[718,749,1852,1874]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="10" pageNumber="11">8B</figureCitation>
)....................................
<taxonomicName id="4C3A4AB0FF84A91CA3FEB134FA5AF8AC" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1201,1436,1852,1873]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="atsirakambiaty">
<emphasis id="B94EED21FF84A91CA3FEB134FA5AF8AC" box="[1201,1436,1852,1873]" italics="true" pageId="10" pageNumber="11">Pheidole atsirakambiaty</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF84A91CA7D8B152FA5AF854" blockId="10.[151,1437,1793,2021]" pageId="10" pageNumber="11">
<keyLead id="30CB9206FF84A91CA7D8B152FA5AF854" pageId="10" pageNumber="11">
- Major worker. Body brown, at least frons laterally more rugulate, frons and occipital lobes with distinctly punctate to rugopunctate interspaces (
<figureCitation id="13012DB6FF84A91CA6FFB17FFDD7F871" box="[432,529,1911,1933]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="10" pageNumber="11">Fig. 6C, J</figureCitation>
). Minor worker. Body yellowish brown to blackish brown, frons usually with strongly reduced sculpture and with smooth notches, mesosoma with smooth notches (
<figureCitation id="13012DB6FF84A91CA435B19CFC18F857" box="[890,990,1940,1962]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="10" pageNumber="11">Figs 7C, J</figureCitation>
,
<figureCitation id="13012DB6FF84A91CA4A7B19CFBDAF857" box="[1000,1052,1940,1962]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="10" pageNumber="11">8C, J</figureCitation>
)............................. 17
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CE2996FF84A91DA7D8B1BAFA5BFEC2" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<paragraph id="8B853133FF84A91DA7D8B1BAFA5AFF37" blockId="10.[151,1437,1793,2021]" lastBlockId="11.[151,1437,151,320]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<keyLead id="30CB9206FF84A91DA7D8B1BAFA5AFF37" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
17 Major worker. Occipital lobes with moderately thin and sparse rugulae that arch posterolaterally, interspaces between rugulae punctate, frons laterally with sparse costulae with distinctly punctate interspaces (
<figureCitation id="13012DB6FF84A91CA4B4B1C7FB87F818" box="[1019,1089,1999,2021]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="10" pageNumber="11">Fig. 6J</figureCitation>
). Minor worker. Occiput elongated, mesosoma microreticulate, promesonotum with sparse microreticulae, promesonotal dorsum mostly to entirely smooth, propodeal spines small and narrow (
<figureCitation id="13012DB6FF85A91DA54AB6BDFD8BFF37" box="[517,589,181,203]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="11" pageNumber="12">Figs 7J</figureCitation>
,
<figureCitation id="13012DB6FF85A91DA518B6BDFDA8FF36" box="[599,622,181,203]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="11" pageNumber="12">8J</figureCitation>
).....................................................
<taxonomicName id="4C3A4AB0FF85A91DA3B4B6BDFA5AFF37" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1275,1436,181,202]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="voreios">
<emphasis id="B94EED21FF85A91DA3B4B6BDFA5AFF37" box="[1275,1436,181,202]" italics="true" pageId="11" pageNumber="12">Pheidole voreios</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B853133FF85A91DA7D8B6DBFA5BFEC2" blockId="11.[151,1437,151,320]" pageId="11" pageNumber="12">
<keyLead id="30CB9206FF85A91DA7D8B6DBFA5BFEC2" pageId="11" pageNumber="12">
- Major worker. Occipital lobes punctate, sometimes with thin and very sparse rugulae that arch posterolaterally, frons laterally with sparser, thinner rugocostulae with distinctly punctate interspaces (
<figureCitation id="13012DB6FF85A91DA4C5B6E7FC10FEF8" box="[906,982,239,261]" captionStart="FIGURE 6" captionStartId="9.[151,250,1526,1550]" captionTargetBox="[151,1436,579,1502]" captionTargetId="figure-583@9.[151,1436,579,1502]" captionTargetPageId="9" captionText="FIGURE 6. Major worker, head. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole flammea (E). Pheidole flavodepressa (F). Pheidole grallatrix (G). Pheidole mantadioflava (H). Pheidole similis (I). Pheidole voreios (J). Pheidole zirafy (K)." figureDoi="http://doi.org/10.5281/zenodo.4405745" httpUri="https://zenodo.org/record/4405745/files/figure.png" pageId="11" pageNumber="12">Fig. 6C</figureCitation>
). Minor worker. Occiput not elongated or elongated very slightly, promesonotum mostly smooth, lateral mesosoma sparsely microreticulate, propodeal spines very small with relatively wide base (
<figureCitation id="13012DB6FF85A91DA6E0B722FE38FEC2" box="[431,510,298,320]" captionStart="FIGURE 7" captionStartId="10.[151,250,1582,1606]" captionTargetBox="[151,1436,433,1559]" captionTargetId="figure-387@10.[151,1436,433,1559]" captionTargetPageId="10" captionText="FIGURE 7. Minor worker, head. Pheidole antsahabe, full-face (A), profile (L). Pheidole atsirakambiaty, full-face (B), profile (M). Pheidole bessonii, full-face (C), profile (N). Pheidole clara, full-face (D), profile (O). Pheidole flammea, full-face (E), profile (P). Pheidole flavodepressa, full-face (F), profile (Q). Pheidole grallatrix, full-face (G), profile (R). Pheidole mantadioflava, full-face (H), profile (S). Pheidole similis, full-face (I), profile (T). Pheidole voreios, full-face (J), profile (U). Pheidole zirafy, full-face (K), profile (V)." figureDoi="http://doi.org/10.5281/zenodo.4405749" httpUri="https://zenodo.org/record/4405749/files/figure.png" pageId="11" pageNumber="12">Figs 7C</figureCitation>
,
<figureCitation id="13012DB6FF85A91DA548B722FDE0FEBD" box="[519,550,298,320]" captionStart="FIGURE 8" captionStartId="11.[151,250,1378,1402]" captionTargetBox="[151,1436,345,1355]" captionTargetId="figure-211@11.[151,1436,345,1355]" captionTargetPageId="11" captionText="FIGURE 8. Profile. Pheidole antsahabe, minor worker (A). Pheidole atsirakambiaty, minor worker (B). Pheidole bessonii, minor worker (C). Pheidole clara, minor worker (D), major worker (L). Pheidole flammea, minor worker (E). Pheidole flavodepressa, minor worker (F). Pheidole grallatrix, minor worker (G), major worker (M). Pheidole mantadioflava, minor worker (H), major worker (N). Pheidole similis, minor worker (I). Pheidole voreios, minor worker. (J). Pheidole zirafy, minor worker (K), major worker (O)." figureDoi="http://doi.org/10.5281/zenodo.4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="11" pageNumber="12">8C</figureCitation>
)..........................................................
<taxonomicName id="4C3A4AB0FF85A91DA3BDB722FA5BFEC2" ID-CoL="4FY85" authorityName="Forel" authorityYear="1891" box="[1266,1437,298,319]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF85A91DA3BDB722FA5BFEC2" box="[1266,1437,298,319]" italics="true" pageId="11" pageNumber="12">Pheidole bessonii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<caption id="DF4561BBFF85A91DA7D8B36AFE46F9F6" ID-DOI="http://doi.org/10.5281/zenodo.4405751" ID-Zenodo-Dep="4405751" httpUri="https://zenodo.org/record/4405751/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1378,1402]" targetBox="[151,1436,345,1355]" targetPageId="11">
<paragraph id="8B853133FF85A91DA7D8B36AFE46F9F6" blockId="11.[151,1436,1378,1547]" pageId="11" pageNumber="12">
<emphasis id="B94EED21FF85A91DA7D8B36AFED3FA87" bold="true" box="[151,277,1378,1402]" pageId="11" pageNumber="12">FIGURE 8.</emphasis>
Profile.
<taxonomicName id="4C3A4AB0FF85A91DA63EB36BFDFAFA86" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[369,572,1379,1403]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="antsahabe">
<emphasis id="B94EED21FF85A91DA63EB36BFDFAFA86" box="[369,572,1379,1403]" italics="true" pageId="11" pageNumber="12">Pheidole antsahabe</emphasis>
</taxonomicName>
, minor worker (A).
<taxonomicName id="4C3A4AB0FF85A91DA45DB36BFBCAFA86" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[786,1036,1379,1403]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="atsirakambiaty">
<emphasis id="B94EED21FF85A91DA45DB36BFBCAFA86" box="[786,1036,1379,1403]" italics="true" pageId="11" pageNumber="12">Pheidole atsirakambiaty</emphasis>
</taxonomicName>
, minor worker (B).
<taxonomicName id="4C3A4AB0FF85A91DA3AFB36BFA50FA86" authorityName="Forel" authorityYear="1891" box="[1248,1430,1379,1403]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bessonii">
<emphasis id="B94EED21FF85A91DA3AFB36BFA50FA86" box="[1248,1430,1379,1403]" italics="true" pageId="11" pageNumber="12">Pheidole bessonii</emphasis>
</taxonomicName>
, minor worker (C).
<taxonomicName id="4C3A4AB0FF85A91DA61BB38FFE2EFA62" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[340,488,1415,1439]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="clara">
<emphasis id="B94EED21FF85A91DA61BB38FFE2EFA62" box="[340,488,1415,1439]" italics="true" pageId="11" pageNumber="12">Pheidole clara</emphasis>
</taxonomicName>
, minor worker (D), major worker (L).
<taxonomicName id="4C3A4AB0FF85A91DA423B38FFBE5FA62" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[876,1059,1415,1439]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="flammea">
<emphasis id="B94EED21FF85A91DA423B38FFBE5FA62" box="[876,1059,1415,1439]" italics="true" pageId="11" pageNumber="12">Pheidole flammea</emphasis>
</taxonomicName>
, minor worker (E).
<taxonomicName id="4C3A4AB0FF85A91DA3A5B38FFF1EFA3F" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="flavodepressa">
<emphasis id="B94EED21FF85A91DA3A5B38FFF1EFA3F" italics="true" pageId="11" pageNumber="12">Pheidole flavodepressa</emphasis>
</taxonomicName>
, minor worker (F).
<taxonomicName id="4C3A4AB0FF85A91DA6E5B3A3FDABFA3E" authorityName="Emery" authorityYear="1899" box="[426,621,1451,1475]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="grallatrix">
<emphasis id="B94EED21FF85A91DA6E5B3A3FDABFA3E" box="[426,621,1451,1475]" italics="true" pageId="11" pageNumber="12">Pheidole grallatrix</emphasis>
</taxonomicName>
, minor worker (G), major worker (M).
<taxonomicName id="4C3A4AB0FF85A91DA343B3A3FAC7FA3E" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1036,1281,1451,1475]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mantadioflava">
<emphasis id="B94EED21FF85A91DA343B3A3FAC7FA3E" box="[1036,1281,1451,1475]" italics="true" pageId="11" pageNumber="12">Pheidole mantadioflava</emphasis>
</taxonomicName>
, minor worker (H), major worker (N).
<taxonomicName id="4C3A4AB0FF85A91DA6C6B3C7FDEAFA1A" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[393,556,1487,1511]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B94EED21FF85A91DA6C6B3C7FDEAFA1A" box="[393,556,1487,1511]" italics="true" pageId="11" pageNumber="12">Pheidole similis</emphasis>
</taxonomicName>
, minor worker (I).
<taxonomicName id="4C3A4AB0FF85A91DA5BDB3C7FC5BFA1B" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[754,925,1487,1511]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="voreios">
<emphasis id="B94EED21FF85A91DA5BDB3C7FC5BFA1B" box="[754,925,1487,1511]" italics="true" pageId="11" pageNumber="12">Pheidole voreios</emphasis>
</taxonomicName>
, minor worker. (J).
<taxonomicName id="4C3A4AB0FF85A91DA326B3C7FAC2FA1A" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1129,1284,1487,1511]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="zirafy">
<emphasis id="B94EED21FF85A91DA326B3C7FAC2FA1A" box="[1129,1284,1487,1511]" italics="true" pageId="11" pageNumber="12">Pheidole zirafy</emphasis>
</taxonomicName>
, minor worker (K), major worker (O).
</paragraph>
</caption>
<subSubSection id="C32062B8FF85A91DA7D8B054FEA1F98A" box="[151,359,1628,1655]" pageId="11" pageNumber="12" type="multiple">
<paragraph id="8B853133FF85A91DA7D8B054FEA1F98A" blockId="11.[151,359,1628,1655]" box="[151,359,1628,1655]" pageId="11" pageNumber="12">
<emphasis id="B94EED21FF85A91DA7D8B054FEA1F98A" bold="true" box="[151,359,1628,1655]" pageId="11" pageNumber="12">Species accounts</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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