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<document id="659D736515438B00C622748AF55C6D70" ID-CLB-Dataset="121760" ID-DOI="10.1093/zoolinnean/zlab105" ID-GBIF-Dataset="83d946c2-fa80-4948-9160-16625c240886" ID-ISSN="0024-4082" ID-Zenodo-Dep="6994599" ID-ZooBank="C06A1F94-AF08-4A21-B1F3-A0865FB1A8DF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660324512590" checkinUser="plazi" docAuthor="Sites, Robert W." docDate="2022" docId="E25E878FFF80FFDDFF34822A0529FDE1" docLanguage="en" docName="zlab105.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Ambrysus Stal 1862" docType="treatment" docVersion="6" lastPageNumber="1259" masterDocId="1E67FFF7FF8DFFD3FFC487270740FFAD" masterDocTitle="Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)" masterLastPageNumber="1286" masterPageNumber="1245" pageNumber="1258" updateTime="1699366499196" updateUser="ExternalLinkService">
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<mods:title id="5BA0BFB9CA06D8F8753635AE091F492A">Phylogeny and revised classification of the saucer bugs (Hemiptera: Nepomorpha: Naucoridae)</mods:title>
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<mods:namePart id="110F9D762DE2ABDA1360DCF5B6B26FE8">Sites, Robert W.</mods:namePart>
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<mods:title id="41ED899E5AD12477F97C71DC60871AE5">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="DC46642C2F5CE5BC0B5D4E981B9181F1">2022</mods:date>
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<subSubSection id="22ED6512FF80FFDEFF34822A05FDFA8B" box="[240,701,1293,1318]" pageId="13" pageNumber="1258" type="nomenclature">
<paragraph id="6A483699FF80FFDEFF34822A05FDFA8B" blockId="13.[240,701,1293,1318]" box="[240,701,1293,1318]" pageId="13" pageNumber="1258">
SUBFAMILY
<taxonomicName id="ADF74D1AFF80FFDEFEB2822A05FDFA8B" ID-CoL="J4X" authority="USINGER, 1941" authorityName="Stal" authorityYear="1862" box="[374,701,1293,1318]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="subFamily" subFamily="Ambrysinae">
AMBRYSINAE
<bibRefCitation id="0E664B68FF80FFDEFDD5822A05FDFA8B" author="Usinger RL" box="[529,701,1293,1318]" pageId="13" pageNumber="1258" pagination="5 - 16" refId="ref25189" refString="Usinger RL. 1941. Key to the subfamilies of Naucoridae with a generic synopsis of the new subfamily Ambrysinae (Hemiptera). Annals of the Entomological Society of America 34: 5 - 16." type="journal article" year="1941">USINGER, 1941</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF80FFDEFEB6821E057CFAFC" box="[370,572,1337,1362]" pageId="13" pageNumber="1258" type="description">
<paragraph id="6A483699FF80FFDEFEB6821E057CFAFC" blockId="13.[370,572,1337,1362]" box="[370,572,1337,1362]" pageId="13" pageNumber="1258">
<figureCitation id="F2CC2A1CFF80FFDEFEB6821E068CFAFC" box="[370,460,1337,1362]" captionStart="Figure 3" captionStartId="8.[145,230,1468,1490]" captionTargetBox="[172,733,195,1428]" captionTargetId="figure-532@8.[172,733,195,1428]" captionTargetPageId="8" captionText="Figure 3. Mouthpart positions: recessed rostrum of Cheirochela thailandana (a); lateral views of Laccocoris spurcus (b) and Australambrysus crenulatus (c). White arrows indicate positions of labrum with respect to the functional anterior margin of the head." figureDoi="http://doi.org/10.5281/zenodo.6994613" httpUri="https://zenodo.org/record/6994613/files/figure.png" pageId="13" pageNumber="1258">FIGS 3C</figureCitation>
,
<figureCitation id="F2CC2A1CFF80FFDEFE1D821E0544FAFC" box="[473,516,1337,1362]" captionStart="Figure 10" captionStartId="14.[144,226,1765,1787]" captionTargetBox="[144,1424,676,1726]" captionTargetId="figure-25@14.[144,1424,676,1726]" captionTargetPageId="14" captionText="Figure 10. Habitus examples of each subfamily: Ambrysinae: Ambrysus signoreti (a), Cheirochelinae: Cheirochela thailandana (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae: Ilyocoris cimicoides (d), Laccocorinae: Laccocoris limicola (e), Limnocorinae: Limnocoris inornatus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Naucoris scutellaris (Thailand) (h)." figureDoi="http://doi.org/10.5281/zenodo.6994631" httpUri="https://zenodo.org/record/6994631/files/figure.png" pageId="13" pageNumber="1258">10A</figureCitation>
,
<figureCitation id="F2CC2A1CFF80FFDEFDD5821E057CFAFC" box="[529,572,1337,1362]" captionStart="Figure 11" captionStartId="15.[162,240,1540,1562]" captionTargetBox="[163,1443,194,1500]" captionTargetId="figure-103@15.[163,1443,194,1500]" captionTargetPageId="15" captionText="Figure 11. Examples of male genital capsule with pygophore, aedeagus and parameres (terga IX and X removed) for each subfamily: Ambrysinae: Ambrysus lunatus (a), Cheirochelinae: Coptocatus kinabalu (b), Cryphocricinae: Cryphocricos hungerfordi (c), Ilyocorinae:Ilyocoris cimicoides (d), Laccocorinae: Laccocoris spurcus (e), Limnocorinae: Limnocoris ovatulus (f), Macrocorinae subfam. nov.: Macrocoris flavicollis (g), Naucorinae: Afronaucoris gen. nov.: Afronaucoris madagascariensis (h)." figureDoi="http://doi.org/10.5281/zenodo.6994635" httpUri="https://zenodo.org/record/6994635/files/figure.png" pageId="13" pageNumber="1258">11A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF80FFDEFF67825705DAFA28" box="[163,666,1391,1413]" pageId="13" pageNumber="1258" type="type_taxon">
<paragraph id="6A483699FF80FFDEFF67825705DAFA28" blockId="13.[163,666,1391,1413]" box="[163,666,1391,1413]" pageId="13" pageNumber="1258">
<emphasis id="5883EA8BFF80FFDEFF6782570673FA28" box="[163,307,1392,1413]" italics="true" pageId="13" pageNumber="1258">
<typeStatus id="B54C883BFF80FFDEFF6782570798FA28" box="[163,216,1392,1413]" pageId="13" pageNumber="1258">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="ADF74D1AFF80FFDEFEFB824805D3FA28" ID-CoL="7QM5Z" authority="Stal, 1862" authorityName="Stal" authorityYear="1862" box="[319,659,1391,1413]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="species" species="signoreti">
<emphasis id="5883EA8BFF80FFDEFEFB8248055AFA28" box="[319,538,1391,1413]" italics="true" pageId="13" pageNumber="1258">Ambrysus signoreti</emphasis>
<bibRefCitation id="0E664B68FF80FFDEFDE5824805D3FA28" author="Stal C" box="[545,659,1391,1413]" pageId="13" pageNumber="1258" pagination="437 - 462" refId="ref25046" refString="Stal C. 1862. Hemiptera Mexicana enumeravit speciesque novas descripsit (continuatio). Entomologische Zeitung Stettiner 23: 437 - 462." type="journal article" year="1862">Stål, 1862</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF80FFDEFF6782830383F9F4" pageId="13" pageNumber="1258" type="discussion">
<paragraph id="6A483699FF80FFDEFF67828304CEFA71" blockId="13.[163,780,1443,1894]" lastBlockId="13.[827,1442,1295,1500]" pageId="13" pageNumber="1258">
<emphasis id="5883EA8BFF80FFDEFF67828306C6FA15" box="[163,390,1443,1465]" italics="true" pageId="13" pageNumber="1258">Taxonomic history:</emphasis>
This subfamily was established by Usinger in 1941, but was downgraded to tribe level and placed in
<taxonomicName id="ADF74D1AFF80FFDEFE5582C60586FA5B" authority="(Popov, 1970)" authorityName="LEACH" authorityYear="1815" baseAuthorityName="Popov" baseAuthorityYear="1970" box="[401,710,1505,1527]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">
Naucorinae (
<bibRefCitation id="0E664B68FF80FFDEFDEB82C605FBFA5B" author="Popov YA" box="[559,699,1505,1526]" pageId="13" pageNumber="1258" pagination="93 - 98" refId="ref23866" refString="Popov YA. 1970. Notes on the classification of the recent Naucoridae (Heteroptera: Nepomorpha). Bulletin of the Polish Academy of Sciences (Biological Sciences) 18: 93 - 98." type="journal article" year="1970">Popov, 1970</bibRefCitation>
)
</taxonomicName>
, then transferred to
<taxonomicName id="ADF74D1AFF80FFDEFEA082D80567F9B8" baseAuthorityName="Reynoso-Velasco &amp; Sites" baseAuthorityYear="2021" box="[356,551,1535,1557]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1284" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
(
<bibRefCitation id="0E664B68FF80FFDEFDF882D907A2F99E" author="Stys P &amp; Jansson A" pageId="13" pageNumber="1258" pagination="1 - 44" refId="ref25103" refString="Stys P, Jansson A. 1988. Check-list of recent family-group and genus-group names of Nepomorpha (Heteroptera) of the world. Acta Entomologica Fennica 50: 1 - 44." type="journal article" year="1988">Štys &amp; Jansson, 1988</bibRefCitation>
) and ultimately elevated and restored to subfamily rank (
<bibRefCitation id="0E664B68FF80FFDEFE77811A079EF9DD" author="Reynoso-Velasco D &amp; Sites RW" pageId="13" pageNumber="1258" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF80FFDEFD04811A079EF9DD" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">Sites, 2021</taxonomicName>
</bibRefCitation>
). Significant recent changes in the generic composition of the subfamily include the transfer of
<taxonomicName id="ADF74D1AFF80FFDEFF0481BE066BF903" authorityName="Stal" authorityYear="1876" box="[192,299,1689,1710]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFF0481BE066BF903" box="[192,299,1689,1710]" italics="true" pageId="13" pageNumber="1258">Pelocoris</emphasis>
</taxonomicName>
in from
<taxonomicName id="ADF74D1AFF80FFDEFE5581BE055FF902" authorityName="LEACH" authorityYear="1815" box="[401,543,1689,1711]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1271" phylum="Arthropoda" rank="subFamily" subFamily="Naucorinae">Naucorinae</taxonomicName>
, and the erection of the genera
<emphasis id="5883EA8BFF80FFDEFEEF819006BAF961" box="[299,506,1719,1740]" italics="true" pageId="13" pageNumber="1258">Australambrysus</emphasis>
Reynoso &amp;
<taxonomicName id="ADF74D1AFF80FFDEFD4C8190044BF961" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[648,779,1719,1741]" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">
<bibRefCitation id="0E664B68FF80FFDEFD4C8190044BF961" author="Sites RW" box="[648,779,1719,1741]" pageId="13" pageNumber="1258" pagination="72 - 94" refId="ref24528" refString="Sites RW. 2021. A review of Cryphocricos Signoret, 1850 (Naucoridae: Cryphocricinae) with descriptions of three new species. Zootaxa 4958: 72 - 94." type="journal article" year="2021">Sites, 2021</bibRefCitation>
</taxonomicName>
and
<emphasis id="5883EA8BFF80FFDEFF1A81F106EBF946" box="[222,427,1750,1771]" italics="true" pageId="13" pageNumber="1258">Maculambrysus</emphasis>
(
<bibRefCitation id="0E664B68FF80FFDEFE0581F1079DF8A7" author="Reynoso-Velasco D &amp; Sites RW" pageId="13" pageNumber="1258" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF80FFDEFD0081F1079DF8A7" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">Sites, 2021</taxonomicName>
</bibRefCitation>
). The subfamily currently includes
<taxonomicName id="ADF74D1AFF80FFDEFD5081D20672F885" authority="Stal, 1862" authorityName="Stal" authorityYear="1862" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFD5081D2044BF8A7" box="[660,779,1781,1802]" italics="true" pageId="13" pageNumber="1258">Ambrysus</emphasis>
<bibRefCitation id="0E664B68FF80FFDEFF6780340672F885" author="Stal C" box="[163,306,1811,1833]" pageId="13" pageNumber="1258" pagination="437 - 462" refId="ref25046" refString="Stal C. 1862. Hemiptera Mexicana enumeravit speciesque novas descripsit (continuatio). Entomologische Zeitung Stettiner 23: 437 - 462." type="journal article" year="1862">Stål, 1862</bibRefCitation>
</taxonomicName>
,
<emphasis id="5883EA8BFF80FFDEFE8F80340575F884" box="[331,565,1811,1833]" italics="true" pageId="13" pageNumber="1258">Australambrysus</emphasis>
,
<taxonomicName id="ADF74D1AFF80FFDEFD8A8033044BF884" authorityName="De Carlo" authorityYear="1963" box="[590,779,1811,1833]" class="Insecta" family="Naucoridae" genus="Carvalhoiella" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFD8A8033044BF884" box="[590,779,1811,1833]" italics="true" pageId="13" pageNumber="1258">Carvalhoiella</emphasis>
</taxonomicName>
D e C a r l o, 1 9 6 3,
<emphasis id="5883EA8BFF80FFDEFEBB8015057FF8E5" box="[383,575,1842,1864]" italics="true" pageId="13" pageNumber="1258">C a t a r a c t o c o r i s</emphasis>
U s i n g e r, 1 9 4 1,
<emphasis id="5883EA8BFF80FFDEFF678076063AF8CB" box="[163,378,1873,1894]" italics="true" pageId="13" pageNumber="1258">H y g r o p e t r o c o r i s</emphasis>
S i t e s, 2 0 1 5,
<emphasis id="5883EA8BFF80FFDEFDF38076044BF8CB" box="[567,779,1872,1894]" italics="true" pageId="13" pageNumber="1258">M a c u l a m b r y s u s</emphasis>
Reynoso &amp;
<taxonomicName id="ADF74D1AFF80FFDEFC1F82280333FA89" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[987,1139,1295,1317]" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">
<bibRefCitation id="0E664B68FF80FFDEFC1F82280333FA89" author="Sites RW" box="[987,1139,1295,1317]" pageId="13" pageNumber="1258" pagination="72 - 94" refId="ref24528" refString="Sites RW. 2021. A review of Cryphocricos Signoret, 1850 (Naucoridae: Cryphocricinae) with descriptions of three new species. Zootaxa 4958: 72 - 94." type="journal article" year="2021">Sites, 2021</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="ADF74D1AFF80FFDEFB4E822802DAFA89" authority="De Carlo" authorityName="De Carlo" authorityYear="1935" box="[1162,1434,1295,1317]" class="Insecta" family="Naucoridae" genus="Melloiella" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFB4E82280252FA89" box="[1162,1298,1295,1316]" italics="true" pageId="13" pageNumber="1258">Melloiella</emphasis>
De Carlo
</taxonomicName>
, 1 9 3 5,
<emphasis id="5883EA8BFF80FFDEFC4B82090346FAEE" box="[911,1030,1325,1347]" italics="true" pageId="13" pageNumber="1258">Pe l o c o r i s</emphasis>
,
<emphasis id="5883EA8BFF80FFDEFBD98209033FFAEE" box="[1053,1151,1326,1347]" italics="true" pageId="13" pageNumber="1258">P i c r o p s</emphasis>
L a R i v e r s, 1 9 5 2 a n d
<taxonomicName id="ADF74D1AFF80FFDEFCFF826B0399FACC" authority="J. Polhemus, 1991" authorityName="J. Polhemus" authorityYear="1991" box="[827,1241,1356,1378]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFCFF826B04BBFACC" box="[827,1019,1356,1377]" italics="true" pageId="13" pageNumber="1258">Procryphocricos</emphasis>
J.
<bibRefCitation id="0E664B68FF80FFDEFBD9826B0399FACC" author="Polhemus JT" box="[1053,1241,1356,1378]" pageId="13" pageNumber="1258" pagination="119 - 123" refId="ref23840" refString="Polhemus JT. 1991. A new and primitive genus of Cryphocricinae (Heteroptera: Naucoridae). Pan-Pacific Entomologist 67: 119 - 123." type="journal article" year="1991">Polhemus, 1991</bibRefCitation>
</taxonomicName>
. Species of all of these genera, except
<taxonomicName id="ADF74D1AFF80FFDEFBFC824C03E3FA2D" authorityName="Stal" authorityYear="1876" box="[1080,1187,1387,1408]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFBFC824C03E3FA2D" box="[1080,1187,1387,1408]" italics="true" pageId="13" pageNumber="1258">Pelocoris</emphasis>
</taxonomicName>
, were included along with
<taxonomicName id="ADF74D1AFF80FFDEFCBD82AE0356FA33" authorityName="Signoret" authorityYear="1850" box="[889,1046,1417,1438]" class="Insecta" family="Naucoridae" genus="Cryphocricos" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFCBD82AE0356FA33" box="[889,1046,1417,1438]" italics="true" pageId="13" pageNumber="1258">Cryphocricos</emphasis>
</taxonomicName>
in the subfamily
<taxonomicName id="ADF74D1AFF80FFDEFB2F82AE02E2FA32" baseAuthorityName="Reynoso-Velasco &amp; Sites" baseAuthorityYear="2021" box="[1259,1442,1417,1439]" class="Insecta" family="Naucoridae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="subFamily" subFamily="Cryphocricinae">Cryphocricinae</taxonomicName>
prior to the revision by
<bibRefCitation id="0E664B68FF80FFDEFBA9828F04C6FA71" author="Reynoso-Velasco D &amp; Sites RW" pageId="13" pageNumber="1258" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF80FFDEFAA1828F04C6FA71" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">Sites (2021)</taxonomicName>
</bibRefCitation>
.
</paragraph>
<paragraph id="6A483699FF80FFDEFCFF81220383F9F4" blockId="13.[827,1442,1541,1625]" pageId="13" pageNumber="1258">
<emphasis id="5883EA8BFF80FFDEFCFF8122036AF9B6" box="[827,1066,1541,1563]" italics="true" pageId="13" pageNumber="1258">Revised taxonomy:</emphasis>
The subfamily was recently revised by
<bibRefCitation id="0E664B68FF80FFDEFC728103025CF994" author="Reynoso-Velasco D &amp; Sites RW" box="[950,1308,1572,1594]" pageId="13" pageNumber="1258" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF80FFDEFB508103025CF994" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1172,1308,1572,1594]" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">Sites (2021)</taxonomicName>
</bibRefCitation>
. No further taxonomic action is proposed here.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF80FFDDFCFF81A5057EFE42" lastPageId="14" lastPageNumber="1259" pageId="13" pageNumber="1258" type="diagnosis">
<paragraph id="6A483699FF80FFDDFCFF81A5057EFE42" blockId="13.[827,1443,1665,1902]" lastBlockId="14.[145,762,197,495]" lastPageId="14" lastPageNumber="1259" pageId="13" pageNumber="1258">
<emphasis id="5883EA8BFF80FFDEFCFF81A504FDF93A" box="[827,957,1666,1687]" italics="true" pageId="13" pageNumber="1258">Diagnosis:</emphasis>
Diagnostic features were provided by
<bibRefCitation id="0E664B68FF80FFDEFCFF818703E5F918" author="Reynoso-Velasco D &amp; Sites RW" box="[827,1189,1696,1718]" pageId="13" pageNumber="1258" pagination="900 - 914" refId="ref23980" refString="Reynoso-Velasco D, Sites RW. 2021. Molecular phylogeny and revised classification of the New World subfamily Cryphocricinae, including the reinstatement of Ambrysinae (Insecta: Hemiptera: Heteroptera: Nepomorpha: Naucoridae). Systematic Entomology 46: 900 - 914." type="journal article" year="2021">
Reynoso-Velasco &amp;
<taxonomicName id="ADF74D1AFF80FFDEFBD8818703E5F918" authority="Sites, 2022" authorityName="Sites" authorityYear="2022" box="[1052,1189,1696,1718]" genus="Sites" pageId="13" pageNumber="1270" rank="genus" status="gen. nov.">Sites (2021)</taxonomicName>
</bibRefCitation>
as follows: (1) labium inserted near the anterior margin of the head; (2) labrum well developed; (3) maxillary plates only slightly produced anteriorly; (4) anterior margin of the pronotum generally with a concavity of different depths, but straight in
<taxonomicName id="ADF74D1AFF80FFDEFC73801E0359F8E3" authorityName="Stal" authorityYear="1876" box="[951,1049,1849,1870]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="1258" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF80FFDEFC73801E0359F8E3" box="[951,1049,1849,1870]" italics="true" pageId="13" pageNumber="1258">Pelocoris</emphasis>
</taxonomicName>
; (5) pronotum with lateral margins smooth or finely crenate; (6) prosternellum concealed by the mesally directed propleural projections, but prosternellum exposed in
<taxonomicName id="ADF74D1AFF83FFDDFE6987C3054FFF54" authorityName="Stal" authorityYear="1876" box="[429,527,228,249]" class="Insecta" family="Naucoridae" genus="Pelocoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFE6987C3054FFF54" box="[429,527,228,249]" italics="true" pageId="14" pageNumber="1259">Pelocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="ADF74D1AFF83FFDDFD8787C305B3FF54" authorityName="J. Polhemus" authorityYear="1991" box="[579,755,228,249]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFD8787C305B3FF54" box="[579,755,228,249]" italics="true" pageId="14" pageNumber="1259">Procryphocricos</emphasis>
</taxonomicName>
; (7) forewing macropterous, but brachyptery also present in
<taxonomicName id="ADF74D1AFF83FFDDFED586060689FE9B" authorityName="J. Polhemus" authorityYear="1991" box="[273,457,289,310]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFED586060689FE9B" box="[273,457,289,310]" italics="true" pageId="14" pageNumber="1259">Procryphocricos</emphasis>
</taxonomicName>
; (8) no sternal abdominal sense organs (hydrostats), except in
<taxonomicName id="ADF74D1AFF83FFDDFDFF866705B2FEF8" authorityName="J. Polhemus" authorityYear="1991" box="[571,754,320,341]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFDFF866705B2FEF8" box="[571,754,320,341]" italics="true" pageId="14" pageNumber="1259">Procryphocricos</emphasis>
</taxonomicName>
; (9) compressible air-bubble respiration, but a plastron could be used by brachypterous
<taxonomicName id="ADF74D1AFF83FFDDFDE9865A05B1FE3F" authorityName="J. Polhemus" authorityYear="1991" box="[557,753,381,402]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFDE9865A05B1FE3F" box="[557,753,381,402]" italics="true" pageId="14" pageNumber="1259">Procryphocricos</emphasis>
</taxonomicName>
; (10) pubescent abdomen (scattered long setae in
<taxonomicName id="ADF74D1AFF83FFDDFF55869D0615FE62" authorityName="J. Polhemus" authorityYear="1991" box="[145,341,442,463]" class="Insecta" family="Naucoridae" genus="Procryphocricos" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="1259" phylum="Arthropoda" rank="genus">
<emphasis id="5883EA8BFF83FFDDFF55869D0615FE62" box="[145,341,442,463]" italics="true" pageId="14" pageNumber="1259">Procryphocricos</emphasis>
</taxonomicName>
); and (11) males with vesica not modified into a long tubular structure.
</paragraph>
</subSubSection>
<subSubSection id="22ED6512FF83FFDDFF55853F0529FDE1" pageId="14" pageNumber="1259" type="discussion">
<paragraph id="6A483699FF83FFDDFF55853F0529FDE1" blockId="14.[145,758,536,588]" pageId="14" pageNumber="1259">
<emphasis id="5883EA8BFF83FFDDFF55853F0661FD80" box="[145,289,536,557]" italics="true" pageId="14" pageNumber="1259">C o m m e n t s:</emphasis>
A l l s p e c i e s i n t h i s s u b f a m i l y a r e distributed exclusively in the New World.
</paragraph>
</subSubSection>
</treatment>
</document>
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