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<document id="E59C2D002EA1229C80FDE38DEE74D800" ID-CLB-Dataset="100956" ID-DOI="10.5281/zenodo.6867065" ID-GBIF-Dataset="3d520847-5163-4b5c-87bf-2cdceb781098" ID-ISBN="978-84-96553-89-7" ID-Zenodo-Dep="6867065" IM.illustrations_requiresApprovalFor="plazi" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1658256599114" checkinUser="jonas" docAuthor="Russell A. Mittermeier, Anthony B. Rylands &amp; Don E. Wilson" docDate="2013" docId="CE199B17FF9AFF9FFF2164CFFEC4F787" docLanguage="en" docName="hbmw_3_Cercopithecidae_0550.pdf.imf" docOrigin="Handbook of the Mammals of the World – Volume 3 Primates, Barcelona: Lynx Edicions" docTitle="Presbytis sumatrana" docType="treatment" docVersion="7" lastPageNumber="719" masterDocId="3220E36FFFC1FFC4FF596510FFB7FFFC" masterDocTitle="Cercopithecidae" masterLastPageNumber="755" masterPageNumber="550" pageNumber="719" updateTime="1699359875441" updateUser="ExternalLinkService">
<mods:mods id="81D2D0EA08BF296733645A9E9DFE8096" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0A3C63C8533097C866D8841B301A2E01">
<mods:title id="C93E2BD52DC34B73EFF013C8D7774BC5">Cercopithecidae</mods:title>
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<mods:namePart id="CC1E28F2D442107D647C50C1F8A7EDB9">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="9D3C8CC43A186AB3DF8EC920D99BE793">Anthony B. Rylands</mods:namePart>
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<mods:namePart id="1D90B0B2954B5DB0EE5CB73F18D763CC">Don E. Wilson</mods:namePart>
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<mods:dateIssued id="D72C144719A8850D41A61057AEB98AFC">2013</mods:dateIssued>
<mods:dateOther id="626EC734C70C08A9EBBECAC46B202182" type="pubDate">2013-03-31</mods:dateOther>
<mods:publisher id="CA56C0A533088465FF7BF4068A99CAD7">Lynx Edicions</mods:publisher>
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<mods:placeTerm id="E182D94D2BCBFF0854156842FFA22F21">Barcelona</mods:placeTerm>
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<mods:titleInfo id="656659385EE16C89211684E9F0D48895">
<mods:title id="0EC224BE26B2058A7ADAA35BA30756AD">Handbook of the Mammals of the World – Volume 3 Primates</mods:title>
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<mods:classification id="782D120109E7A66A0802E9D033913B2A">book chapter</mods:classification>
<mods:identifier id="E18B35B1C96C3C1BE7DB1650746D03F6" type="CLB-Dataset">100956</mods:identifier>
<mods:identifier id="D4A6820275C3D625F5F00A5DB13CA8CD" type="DOI">10.5281/zenodo.6867065</mods:identifier>
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<treatment id="CE199B17FF9AFF9FFF2164CFFEC4F787" ID-DOI="http://doi.org/10.5281/zenodo.6863398" ID-GBIF-Taxon="197824749" ID-Zenodo-Dep="6863398" LSID="urn:lsid:plazi:treatment:CE199B17FF9AFF9FFF2164CFFEC4F787" httpUri="http://treatment.plazi.org/id/CE199B17FF9AFF9FFF2164CFFEC4F787" lastPageNumber="719" pageId="91" pageNumber="719">
<subSubSection id="0EAA798AFF9AFF9FFF2164CFFF7FFDF9" box="[120,200,479,517]" pageId="91" pageNumber="719" type="multiple">
<paragraph id="460F2A01FF9AFF9FFF2164CFFF7FFDF9" blockId="91.[116,1142,479,562]" box="[120,200,479,517]" pageId="91" pageNumber="719">
<heading id="1D479D6DFF9AFF9FFF2164CFFF7FFDF9" box="[120,200,479,517]" pageId="91" pageNumber="719">
<figureCitation id="DE8B3684FF9AFF9FFF2164CFFF7FFDF9" box="[120,200,479,517]" captionStart="Plate 47: Cercopithecidae" captionStartId="87.[100,130,3411,3432]" captionTargetBox="[13,2745,15,3644]" captionTargetPageId="86" captionText="5. 5- 6 kg ( females ). The Sabah Grizzled Languris gray above and below, except white belly and inner surfaces of limbs, contrasting with black on hands and feet. Crown is gray, with a tall black crest on back of head and two frontal whorls. Facial skin is red- dish, with bluish lips and a pair of distinct black spots on each side of the face, between eyes and mouth — a pattern sometimes seen in more contrastingly colored individuals of Hose’s Langur ( Presbytis hoset (null), 6. [MISSING], 7. [MISSING], 8. [MISSING], 9. [MISSING], 10. [MISSING], 11. [MISSING], 12. [MISSING], 13. [MISSING], 14. [MISSING], 15. [MISSING], 16. [MISSING], 17. [MISSING], 18. [MISSING], 19. [MISSING], 20. [MISSING], 21. [MISSING], 22. [MISSING], 23. [MISSING], 24. [MISSING], 25. [MISSING], 26. [MISSING], 27. [MISSING], 28. [MISSING], 29. [MISSING], 30. [MISSING], 31. [MISSING], 32. [MISSING], 33. [MISSING], 34. [MISSING], 35. [MISSING], 36. [MISSING], 37. [MISSING], 38. [MISSING], 39. [MISSING], 40. [MISSING], 41. [MISSING], 42. [MISSING], 43. [MISSING], 44. [MISSING], 45. [MISSING], 46. [MISSING], 47. [MISSING], 48. [MISSING], 49. [MISSING], 50. [MISSING], 51. [MISSING], 52. [MISSING], 53. [MISSING], 54. [MISSING], 55. [MISSING], 56. [MISSING], 57. [MISSING], 58. [MISSING], 59. [MISSING], 60. [MISSING], 61. [MISSING], 62. [MISSING], 63. [MISSING], 64. [MISSING], 65. [MISSING], 66. [MISSING], 67. [MISSING], 68. [MISSING], 69. [MISSING], 70. [MISSING], 71. [MISSING], 72. [MISSING], 73. [MISSING], 74. [MISSING], 75. [MISSING], 76. [MISSING], 77. [MISSING], 78. [MISSING], 79. [MISSING], 80. [MISSING], 81. [MISSING], 82. [MISSING], 83. [MISSING], 84. [MISSING], 85. [MISSING], 86. [MISSING], 87. [MISSING], 88. [MISSING], 89. [MISSING], 90. [MISSING], 91. [MISSING], 92. [MISSING], 93. [MISSING], 94. [MISSING], 95. [MISSING], 96. [MISSING], 97. [MISSING], 98. [MISSING], 99. [MISSING], 100. [MISSING], 101. [MISSING], 102. [MISSING], 103. [MISSING], 105. Javan Langur (Presbytis comata), 106. Banded Langur (Presbytis femoralis), 107. Cross-marked Langur (Presbytis chrysomelas), 108. White-fronted Langur (Presbytis frontata), 109. Hose’s Langur (Presbytis hosei), 110. Miller’s Langur (Presbytis canicrus), 111. Sabah Grizzled Langur (Presbytis sabana), 112. Black-crested Sumatran Langur (Presbytis melalophos), 113. Black-and-white Langur (Presbytis bicolor), 114. Black Sumatran Langur (Presbytis sumatrana), 115. Mitered Langur (Presbytis mitrata)" figureDoi="http://doi.org/10.5281/zenodo.6867463" httpUri="https://zenodo.org/record/6867463/files/figure.png" pageId="91" pageNumber="719">114.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFF8064CFFD7CFDF9" box="[217,715,479,517]" pageId="91" pageNumber="719" type="vernacular_names">
<paragraph id="460F2A01FF9AFF9FFF8064CFFD7CFDF9" blockId="91.[116,1142,479,562]" box="[217,715,479,517]" pageId="91" pageNumber="719">
<heading id="1D479D6DFF9AFF9FFF8064CFFD7CFDF9" box="[217,715,479,517]" pageId="91" pageNumber="719">
<vernacularName id="C8B35A2FFF9AFF9FFF8064CFFD7CFDF9" box="[217,715,479,517]" pageId="91" pageNumber="719">Black Sumatran Langur</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFDA564CFFBC3FDF9" box="[764,1140,479,517]" pageId="91" pageNumber="719" type="nomenclature">
<paragraph id="460F2A01FF9AFF9FFDA564CFFBC3FDF9" blockId="91.[116,1142,479,562]" box="[764,1140,479,517]" pageId="91" pageNumber="719">
<heading id="1D479D6DFF9AFF9FFDA564CFFBC3FDF9" box="[764,1140,479,517]" pageId="91" pageNumber="719">
<taxonomicName id="81B05182FF9AFF9FFDA564CFFBC3FDF9" ID-CoL="4MBBM" baseAuthorityName="S. Miller &amp; Schlegel" baseAuthorityYear="1841" box="[764,1140,479,517]" class="Mammalia" family="Cercopithecidae" genus="Presbytis" kingdom="Animalia" order="Primates" pageId="91" pageNumber="719" phylum="Chordata" rank="species" species="sumatrana">
<emphasis id="74C4F613FF9AFF9FFDA564CFFBC3FDF9" box="[764,1140,479,517]" italics="true" pageId="91" pageNumber="719">Presbytis sumatrana</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFF2C6707FBDEFDD0" box="[117,1129,535,556]" pageId="91" pageNumber="719" type="vernacular_names">
<paragraph id="460F2A01FF9AFF9FFF2C6707FBDEFDD0" blockId="91.[116,1142,479,562]" box="[117,1129,535,556]" pageId="91" pageNumber="719">
<heading id="1D479D6DFF9AFF9FFF2C6707FBDEFDD0" box="[117,1129,535,556]" pageId="91" pageNumber="719">
<emphasis id="74C4F613FF9AFF9FFF2C6707FF75FDD0" bold="true" box="[117,194,535,556]" pageId="91" pageNumber="719">French:</emphasis>
<vernacularName id="C8B35A2FFF9AFF9FFF956707FE3EFDD0" box="[204,393,535,556]" pageId="91" pageNumber="719">Langur de Sumatra</vernacularName>
/
<emphasis id="74C4F613FF9AFF9FFEC66707FE4DFDD0" bold="true" box="[415,506,535,556]" pageId="91" pageNumber="719">German:</emphasis>
<vernacularName id="C8B35A2FFF9AFF9FFD5A6707FD46FDD0" box="[515,753,535,556]" pageId="91" pageNumber="719">Schwarzer Kammlangur</vernacularName>
/
<emphasis id="74C4F613FF9AFF9FFC5F6707FCD6FDD0" bold="true" box="[774,865,535,556]" pageId="91" pageNumber="719">Spanish:</emphasis>
<vernacularName id="C8B35A2FFF9AFF9FFC326707FBDEFDD0" box="[875,1129,535,556]" pageId="91" pageNumber="719">Langur negro de Sumatra</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFD84674EFBF6FD5E" pageId="91" pageNumber="719" type="reference_group">
<paragraph id="460F2A01FF9AFF9FFD84674EFBF6FD5E" blockId="91.[732,1320,606,1029]" pageId="91" pageNumber="719">
<emphasis id="74C4F613FF9AFF9FFD84674EFCCEFD87" bold="true" box="[733,889,606,635]" pageId="91" pageNumber="719">Taxonomy.</emphasis>
<taxonomicName id="81B05182FF9AFF9FFCCD674EFB8BFD5E" authority="S. Miller &amp; Schlegel, 1841" authorityName="S. Miller &amp; Schlegel" authorityYear="1841" class="Mammalia" family="Cercopithecidae" genus="Semnopithecus" kingdom="Animalia" order="Primates" pageId="91" pageNumber="719" phylum="Chordata" rank="species" species="sumatranus">Semnopithecus sumatranus S. Miller &amp; Schlegel, 1841</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFB086791FB3CFD36" pageId="91" pageNumber="719" type="materials_examined">
<paragraph id="460F2A01FF9AFF9FFB086791FB3CFD36" blockId="91.[732,1320,606,1029]" pageId="91" pageNumber="719">
<materialsCitation id="F6D8205CFF9AFF9FFB086791FB3CFD36" ID-GBIF-Occurrence="3864421334" pageId="91" pageNumber="719">“Mt. Ophir” (= G. Talamau), north of Padang.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFD8767C0FEA7FBD6" pageId="91" pageNumber="719" type="discussion">
<paragraph id="460F2A01FF9AFF9FFD8767C0FEA7FBD6" blockId="91.[732,1320,606,1029]" lastBlockId="91.[112,1321,1033,2171]" pageId="91" pageNumber="719">
<taxonomicName id="81B05182FF9AFF9FFD8767C0FC2EFD0D" box="[734,921,720,753]" class="Mammalia" family="Cercopithecidae" genus="Presbytis" kingdom="Animalia" order="Primates" pageId="91" pageNumber="719" phylum="Chordata" rank="species" species="sumatrana">P. sumatrana</taxonomicName>
was formerly considered a subspecies of
<taxonomicName id="81B05182FF9AFF9FFC8D67E8FB3EFCE5" authorityName="Raffles" authorityYear="1821" box="[980,1161,760,793]" class="Mammalia" family="Cercopithecidae" genus="Presbytis" kingdom="Animalia" order="Primates" pageId="91" pageNumber="719" phylum="Chordata" rank="species" species="melalophos">P. melalophos</taxonomicName>
. Northern populations are darker and blacker than those in the south, which are more brownish, and could be separated as the subspecies margae. It is separated from
<taxonomicName id="81B05182FF9AFF9FFBC06685FA9FFC4A" baseAuthorityName="Martin" baseAuthorityYear="1838" box="[1177,1320,917,950]" class="Mammalia" family="Cercopithecidae" genus="Presbytis" kingdom="Animalia" order="Primates" pageId="91" pageNumber="719" phylum="Chordata" rank="species" species="femoralis">P. femoralis</taxonomicName>
percura by the Rokan River and from
<taxonomicName id="81B05182FF9AFF9FFB8466ADFC86FBF9" baseAuthorityName="Muller &amp; Schlegel" baseAuthorityYear="1841" class="Mammalia" family="Cercopithecidae" genus="Presbytis" kingdom="Animalia" order="Primates" pageId="91" pageNumber="719" phylum="Chordata" rank="species" species="siamensis">P. siamensis</taxonomicName>
paenulata by the Barumun River. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFF2A6122FCC4FB5D" pageId="91" pageNumber="719" type="distribution">
<caption id="12CF7A89FF9AFF9FFF2A6122FCC4FB5D" ID-DOI="http://doi.org/10.5281/zenodo.6867333" ID-Zenodo-Dep="6867333" httpUri="https://zenodo.org/record/6867333/files/figure.png" inLine="true" pageId="91" pageNumber="719" targetBox="[119,701,615,1022]" targetPageId="91">
<paragraph id="460F2A01FF9AFF9FFF2A6122FCC4FB5D" blockId="91.[112,1321,1033,2171]" pageId="91" pageNumber="719">
<emphasis id="74C4F613FF9AFF9FFF2A6122FE94FBAF" bold="true" box="[115,291,1074,1107]" pageId="91" pageNumber="719">Distribution.</emphasis>
W &amp; NC Sumatra (N highlands and W coast S of the Simpang Kiri River, N of Gunung Talamau, and on the E coast from the Wampu River to the Barumun River, and SE to the Rokan River), and Batu Is (Pini I).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFF2A61BCFE60FA20" pageId="91" pageNumber="719" type="description">
<paragraph id="460F2A01FF9AFF9FFF2A61BCFE60FA20" blockId="91.[112,1321,1033,2171]" pageId="91" pageNumber="719">
<emphasis id="74C4F613FF9AFF9FFF2A61BCFEC7FB35" bold="true" box="[115,368,1196,1225]" pageId="91" pageNumber="719">Descriptive notes.</emphasis>
Head—body 33-35 cm (males) and 34-39 cm (females), tail 64— 72 cm (males) and 66-84 cm (females); weight 4.9-7.2 kg (males) and 5.1-8.5 kg (females). Upperparts of the Black Sumatran Langur are dark gray-brown to nearly black, including dorsal surface of tail. Hands, feet, and outer limbs are black; throat, underparts, underside oftail, and inner surfaces of limbs as far as wrists and ankles are white, sharply set off from dark color of upperparts. Crest is gray to dark brown, and sometimes indistinctly formed. Facial skin is mostly bluish, flesh colored around the mouth, but lips are black.
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFF2A60F4FB1EF9F9" box="[115,1193,1508,1541]" pageId="91" pageNumber="719" type="biology_ecology">
<paragraph id="460F2A01FF9AFF9FFF2A60F4FB1EF9F9" blockId="91.[112,1321,1033,2171]" box="[115,1193,1508,1541]" pageId="91" pageNumber="719">
<emphasis id="74C4F613FF9AFF9FFF2A60F4FF56F9F9" bold="true" box="[115,225,1508,1541]" pageId="91" pageNumber="719">Habitat.</emphasis>
[Lowland and hill rainforest, old growth forest, and secondary forest.
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFF2A631BFC7EF9AF" pageId="91" pageNumber="719" type="food_feeding">
<paragraph id="460F2A01FF9AFF9FFF2A631BFC7EF9AF" blockId="91.[112,1321,1033,2171]" pageId="91" pageNumber="719">
<emphasis id="74C4F613FF9AFF9FFF2A631BFEC1F9D0" bold="true" box="[115,374,1547,1580]" pageId="91" pageNumber="719">Food and Feeding.</emphasis>
The diet of the Black Sumatran Langur probably consists largely of young leaves, unripe fruits, flowers, mature leaves, and seeds.
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFF2B6349FC01F986" box="[114,950,1625,1658]" pageId="91" pageNumber="719" type="breeding">
<paragraph id="460F2A01FF9AFF9FFF2B6349FC01F986" blockId="91.[112,1321,1033,2171]" box="[114,950,1625,1658]" pageId="91" pageNumber="719">
<emphasis id="74C4F613FF9AFF9FFF2B6349FF4FF986" bold="true" box="[114,248,1625,1658]" pageId="91" pageNumber="719">Breeding.</emphasis>
There is no information available for this species.
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFF296395FBF0F95E" box="[112,1095,1669,1698]" pageId="91" pageNumber="719" type="activity">
<paragraph id="460F2A01FF9AFF9FFF296395FBF0F95E" blockId="91.[112,1321,1033,2171]" box="[112,1095,1669,1698]" pageId="91" pageNumber="719">
<emphasis id="74C4F613FF9AFF9FFF296395FEECF95E" bold="true" box="[112,347,1669,1698]" pageId="91" pageNumber="719">Activity patterns.</emphasis>
The Black Sumatran Langur is diurnal and arboreal.
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFF2B63B8FD41F8E4" pageId="91" pageNumber="719" type="biology_ecology">
<paragraph id="460F2A01FF9AFF9FFF2B63B8FD41F8E4" blockId="91.[112,1321,1033,2171]" pageId="91" pageNumber="719">
<emphasis id="74C4F613FF9AFF9FFF2B63B8FC88F935" bold="true" box="[114,831,1704,1737]" pageId="91" pageNumber="719">Movements, Home range and Social organization.</emphasis>
The Black Sumatran Langur has never been studied in the wild. Like other species of
<taxonomicName id="81B05182FF9AFF9FFC1763C0FC04F90D" authorityName="Eschscholtz" authorityYear="1821" box="[846,947,1744,1777]" class="Mammalia" family="Cercopithecidae" genus="Presbytis" kingdom="Animalia" order="Primates" pageId="91" pageNumber="719" phylum="Chordata" rank="genus">Presbytis</taxonomicName>
,it likely lives in small onemale groups and defends exclusive territories.
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFF2B620FFD97F7FF" pageId="91" pageNumber="719" type="conservation">
<paragraph id="460F2A01FF9AFF9FFF2B620FFD97F7FF" blockId="91.[112,1321,1033,2171]" pageId="91" pageNumber="719">
<emphasis id="74C4F613FF9AFF9FFF2B620FFE66F8BC" bold="true" box="[114,465,1823,1856]" pageId="91" pageNumber="719">Status and Conservation.</emphasis>
CITES Appendix II. Classified as Endangered on The IUCN Red List (as
<taxonomicName id="81B05182FF9AFF9FFE416256FE76F89B" authorityName="Raffles" authorityYear="1821" box="[280,449,1862,1895]" class="Mammalia" family="Cercopithecidae" genus="Presbytis" kingdom="Animalia" order="Primates" pageId="91" pageNumber="719" phylum="Chordata" rank="species" species="melalophos">P. melalophos</taxonomicName>
<taxonomicName id="81B05182FF9AFF9FFE926256FDDFF89B" authorityName="S. Miller &amp; Schlegel" authorityYear="1841" box="[459,616,1862,1895]" class="Mammalia" family="Cercopithecidae" genus="Semnopithecus" kingdom="Animalia" order="Primates" pageId="91" pageNumber="719" phylum="Chordata" rank="species" species="sumatranus">sumatranus</taxonomicName>
). The Black Sumatran Langur is protected under Indonesian law. About 50% of the forests in its range have been degraded over the past 30 years. Remaining forest is a fragmented mosaic among commercial plantations. The illegal pet trade is a further threat to the Black Sumatran Langur. It occurs in Batang Gadis National Park.
</paragraph>
</subSubSection>
<subSubSection id="0EAA798AFF9AFF9FFF2B6D04FEC4F787" pageId="91" pageNumber="719" type="bibRefCitation_list">
<paragraph id="460F2A01FF9AFF9FFF2B6D04FEC4F787" blockId="91.[112,1321,1033,2171]" pageId="91" pageNumber="719">
<emphasis id="74C4F613FF9AFF9FFF2B6D04FEBBF7D1" bold="true" box="[114,268,2068,2093]" pageId="91" pageNumber="719">Bibliography.</emphasis>
Aimi &amp; Bakar (1992, 1996), Davies et al. (1988), Fleagle (1977), Groves (2001), Kawamura (1984), Meyer, Hodges et al. (2012), Meyer, Rinaldi et al. (2011), Mukhtar et al. (1990), Nijman &amp; Manullang (2008), Zingeser (1970).
</paragraph>
</subSubSection>
</treatment>
</document>
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