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<mods:title id="35BA81E1E610C33A0867EFA7E831238F">Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae)</mods:title>
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<taxonomicName id="8FEA4D01D847FFAAFF017590FE24FDBA" ID-CoL="8X7DL" authority="King, 1866" authorityName="King. A" authorityYear="1866" box="[151,384,919,944]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D847FFAAFF017590FE24FDBA" bold="true" box="[151,384,919,944]" pageId="5" pageNumber="6">
<emphasis id="7A9EEA90D847FFAAFF017590FEA1FDBA" bold="true" box="[151,261,920,944]" italics="true" pageId="5" pageNumber="6">EUPINES</emphasis>
<bibRefCitation id="2C7B4B73D847FFAAFE9D759FFE24FDBA" author="King, R. L." box="[267,384,919,944]" pageId="5" pageNumber="6" pagination="299 - 315" refId="ref58602" refString="King, R. L. (1866) On the Pselaphidae of Australia. No. IV. Transactions Entomological Society of New South Wales, 1, 299 - 315." type="journal article" year="1866">King, 1866</bibRefCitation>
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<paragraph id="48553682D847FFAAFF0175B2FF5BFDDE" blockId="5.[151,384,919,980]" box="[151,255,954,980]" pageId="5" pageNumber="6">
<figureCitation id="D0D12A07D847FFAAFF0175B2FF5BFDDE" box="[151,255,954,980]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="6.[151,250,1798,1823]" captionStartId-1="7.[151,250,1676,1701]" captionTargetBox-0="[161,1420,353,1766]" captionTargetBox-1="[162,1432,187,1639]" captionTargetId-0="figure@6.[151,1435,345,1773]" captionTargetId-1="figure@7.[151,1435,181,1651]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 2. Morphology of Eupines King (based on E. (B.) crassicornides). A) Head, in dorsal view. B) Same, in lateral view. C) Same, in ventral view. D) Mouth parts. E) Mandibles and labrum. F) Female antenna. G) Male antenna. H) Pronotum. I) Same, in lateral view. J) Prothorax. K) Meso- and metaventrite. Abbreviations: pc—postantennal cavities, gf—gular foveae, gr—gular ridge, omc—ocular-mandibular carinae, l—lacinia, lb—labrum, lp—labial palp, m—mentum, lmcf—lateral mesocoxal foveae, lmtf—lateral metaventral foveae, lpcf—lateral procoxal foveae, lmvf—lateral mesoventral foveae, mmsf—median mesoventral foveae, ppf—prepectal foveae, vf—vertexal foveae. Scale bars: 0.2 mm, except D, E, H, I, J = 0.1 mm." captionText-1="FIGURE 3. Morphology of Eupines King. A) Tergites 15. B) Tergite 1. C) Tergite 2. D) Tergite 3. E) Tergite 4. F) Tergite 5. G) Ventrites 26. H) Ventrites 46. I) Ventrite 2. J) Ventrite 3. K) Ventrite 4. L) Ventrite 5. M) Ventrite 6. N) Elytra, in dorsal view. Scale bars: 0.1 mm, except A, N = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3821178" figureDoi-1="http://doi.org/10.5281/zenodo.3821180" httpUri-0="https://zenodo.org/record/3821178/files/figure.png" httpUri-1="https://zenodo.org/record/3821180/files/figure.png" pageId="5" pageNumber="6">Figs. 23</figureCitation>
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<taxonomicName id="8FEA4D01D847FFAAFF0175F7FE87FA1D" ID-CoL="8X7DL" authority="King" authorityName="King. A" authorityYear="1866" box="[151,291,1022,1047]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D847FFAAFF0175F7FF4DFA1C" box="[151,233,1023,1046]" italics="true" pageId="5" pageNumber="6">Eupines</emphasis>
King
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(subgenus of
<taxonomicName id="8FEA4D01D847FFAAFE2575F7FDC9FA1D" authority="Kugelann" authorityName="Kugelann" authorityYear="1794" box="[435,621,1022,1047]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D847FFAAFE2575F7FDA4FA1C" box="[435,512,1023,1046]" italics="true" pageId="5" pageNumber="6">Bryaxis</emphasis>
Kugelann
</taxonomicName>
), 1866: 309. Elevated to genus by
<treatmentCitation id="C94B1093D847FFAAFC4675F7FB24FA1D" author="Broun, T." box="[976,1152,1022,1047]" page="759" pageId="5" pageNumber="6" year="1886">
<bibRefCitation id="2C7B4B73D847FFAAFC4675F7FB24FA1D" author="Broun, T." box="[976,1152,1022,1047]" pageId="5" pageNumber="6" pagination="745 - 973" refId="ref57925" refString="Broun, T. (1886) Manual of the New Zealand Coleoptera. Parts III &amp; IV. Colonial Museum and Geological Survey Department, Wellington, 228 pp. [pp. I-XVII + 745 - 973]" type="book chapter" year="1886">Broun 1886: 759</bibRefCitation>
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<typeStatus id="97518820D847FFAAFB1D75F7FB64FA1D" box="[1163,1216,1023,1047]" pageId="5" pageNumber="6">Type</typeStatus>
species:
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<taxonomicName id="8FEA4D01D847FFAAFAB675F7FEE6FA3F" authority="King" authorityName="King" authorityYear="1864" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="clavatula">
<emphasis id="7A9EEA90D847FFAAFAB675F7FEA2FA3F" italics="true" pageId="5" pageNumber="6">Bryaxis clavatula</emphasis>
King
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, subsequent designation by
<treatmentCitation id="C94B1093D847FFAAFDF77214FCB4FA3F" author="Jeannel, R." box="[609,784,1052,1077]" page="84" pageId="5" pageNumber="6" year="1952">
<bibRefCitation id="2C7B4B73D847FFAAFDF77214FCB4FA3F" author="Jeannel, R." box="[609,784,1052,1077]" pageId="5" pageNumber="6" pagination="69 - 113" refId="ref58578" refString="Jeannel, R. (1952) Pselaphides de Saigon. Revue Francaise d'Entomologie, 19 (2), 69 - 113." type="journal article" year="1952">Jeannel 1952: 84</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="C94B1093D847FFAAFC8D7214FC73FA3F" author="Raffray, A." box="[795,983,1052,1077]" page="258" pageId="5" pageNumber="6" year="1897">
<bibRefCitation id="2C7B4B73D847FFAAFC8D7214FC73FA3F" author="Raffray, A." box="[795,983,1052,1077]" pageId="5" pageNumber="6" pagination="227 - 284" refId="ref59120" refString="Raffray, A. (1897) Nouvelles etudes sur les Pselaphides et les Clavigerides. Annales de la Societe entomologique de France, 65, 227 - 284." type="journal article" year="1897">Raffray 1897: 258</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="C94B1093D847FFAAFC747215FBEFFA3F" author="Raffray, A." box="[994,1099,1053,1077]" page="171" pageId="5" pageNumber="6" year="1900">
<bibRefCitation id="2C7B4B73D847FFAAFC747215FBEFFA3F" author="Raffray, A." box="[994,1099,1053,1077]" pageId="5" pageNumber="6" pagination="131 - 249" refId="ref59150" refString="Raffray, A. (1900) Australian Pselaphidae. Proceedings of the Linnean Society of New South Wales, 25, 131 - 249." type="journal article" year="1900">1900: 171</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="C94B1093D847FFAAFBC07215FB64FA3F" author="Raffray, A." box="[1110,1216,1052,1077]" page="206" pageId="5" pageNumber="6" year="1908">
<bibRefCitation id="2C7B4B73D847FFAAFBC07215FB64FA3F" author="Raffray, A." box="[1110,1216,1052,1077]" pageId="5" pageNumber="6" pagination="1 - 484" refId="ref59206" refString="Raffray, A. (1908) Coleoptera fam. Pselaphidae. In: Wytsman, P. (Ed.), Genera insectorum. 64 th Fascicule. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 484, 489 pls." type="book chapter" year="1908">1908: 206</bibRefCitation>
</treatmentCitation>
;
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<bibRefCitation id="2C7B4B73D847FFAAFB5D7214FA3CFA3F" author="Chandler, D. S." box="[1227,1432,1052,1077]" pageId="5" pageNumber="6" refId="ref58398" refString="Chandler, D. S. (2001) Biology, morphology, and systematics of the ant-like litter beetle genera of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology International. Entomology International, Associated Publishers, Gainesville, Florida, VIII + 560 pp." type="book" year="2001">Chandler 2001: 320</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="C94B1093D847FFAAFF517234FDABFA5E" author="Nomura, S. &amp; Leschen, R. A. B." box="[199,527,1083,1108]" page="253" pageId="5" pageNumber="6" year="2006">
<bibRefCitation id="2C7B4B73D847FFAAFF517234FDABFA5E" author="Nomura, S. &amp; Leschen, R. A. B." box="[199,527,1083,1108]" pageId="5" pageNumber="6" pagination="239 - 272" refId="ref58967" refString="Nomura, S. &amp; Leschen, R. A. B. (2006) Faunistic review on the pselaphine species known from New Zealand (Insecta, Coleoptera, Staphylinidae). National Science Museum Monographs, 34, 239 - 272." type="journal article" year="2006">Nomura and Leschen 2006: 253</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<treatmentCitation id="C94B1093D847FFAAFF017253FE0FFA79" author="Reitter, E." box="[151,427,1114,1139]" page="166" pageId="5" pageNumber="6" year="1880">
<taxonomicName id="8FEA4D01D847FFAAFF017253FE0FFA79" authority="Reitter, 1880: 166" authorityName="Reitter" authorityPageNumber="166" authorityYear="1880" box="[151,427,1114,1139]" class="Insecta" family="Staphylinidae" genus="Byraxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D847FFAAFF017253FF40FA78" box="[151,228,1115,1138]" italics="true" pageId="5" pageNumber="6">Byraxis</emphasis>
<bibRefCitation id="2C7B4B73D847FFAAFF7B7252FE0FFA79" author="Reitter, E." box="[237,427,1114,1139]" pageId="5" pageNumber="6" pagination="165 - 183" refId="ref59347" refString="Reitter, E. (1880) Beitrage zur Kaferfauna von Neu-Zeeland. Verhandlungen des Naturforschenden Vereines in Brunn, 18, 165 - 183." type="journal article" year="1880">Reitter, 1880: 166</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="97518820D847FFAAFE207253FE4CFA79" box="[438,488,1115,1139]" pageId="5" pageNumber="6">Type</typeStatus>
species:
<taxonomicName id="8FEA4D01D847FFAAFDDF7253FCF3FA79" ID-CoL="9LDFL" authority="Reitter" authorityName="Reitter" authorityYear="1880" box="[585,855,1114,1139]" class="Insecta" family="Staphylinidae" genus="Byraxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="monstrosa">
<emphasis id="7A9EEA90D847FFAAFDDF7253FCACFA78" box="[585,776,1115,1138]" italics="true" pageId="5" pageNumber="6">Byraxis monstrosa</emphasis>
Reitter
</taxonomicName>
, by monotypy. Reduction to subgenus by
<treatmentCitation id="C94B1093D847FFAAFA9B7252FF54FA9B" author="Raffray, A." page="202" pageId="5" pageNumber="6" year="1904">
<bibRefCitation id="2C7B4B73D847FFAAFA9B7252FF54FA9B" author="Raffray, A." pageId="5" pageNumber="6" pagination="1 - 400" refId="ref59176" refString="Raffray, A. (1904) Genera et catalogue des Pselaphides [continued]. Annales de la Societe entomologique de France, 72, 1 - 400." type="journal article" year="1904">Raffray 1904: 202</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="48553682D847FFAAFF017291FE4FFAC5" blockId="5.[151,1437,1022,2043]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="68FA11ACD847FFAAFF017291FE4FFAC5" pageId="5" pageNumber="6">
<treatmentCitation id="C94B1093D847FFAAFF017291FE1AFABA" author="Raffray, A." box="[151,446,1175,1200]" page="123" pageId="5" pageNumber="6" year="1890">
<taxonomicName id="8FEA4D01D847FFAAFF017291FE1AFABA" ID-CoL="9LFYZ" authority="Raffray, 1890: 123" authorityName="Raffray" authorityPageNumber="123" authorityYear="1890" box="[151,446,1175,1200]" class="Insecta" family="Staphylinidae" genus="Patranus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D847FFAAFF017291FF50FABA" box="[151,244,1177,1200]" italics="true" pageId="5" pageNumber="6">Patranus</emphasis>
<bibRefCitation id="2C7B4B73D847FFAAFF6D729FFE1AFABA" author="Raffray, A." box="[251,446,1175,1200]" pageId="5" pageNumber="6" pagination="81 - 172" refId="ref59092" refString="Raffray, A. (1890) Etude sur les Pselaphides. V. Tableaux synoptiques-Notes et synonymie. Revue d'Entomologie, 9, 81 - 172." type="journal article" year="1890">Raffray, 1890: 123</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="97518820D847FFAAFE5E7290FE5EFABA" box="[456,506,1176,1200]" pageId="5" pageNumber="6">Type</typeStatus>
species:
<taxonomicName id="8FEA4D01D847FFAAFDCF7290FCFDFABA" ID-CoL="59SNN" authority="Schaufuss" authorityName="Schaufuss" box="[601,857,1175,1200]" class="Insecta" family="Staphylinidae" genus="Tychus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="politus">
<emphasis id="7A9EEA90D847FFAAFDCF7290FD4FFABA" box="[601,747,1176,1200]" italics="true" pageId="5" pageNumber="6">Tychus politus</emphasis>
Schaufuss
</taxonomicName>
(=
<taxonomicName id="8FEA4D01D847FFAAFCE17291FB06FABA" authority="Schaufuss" authorityName="Schaufuss" box="[887,1186,1175,1200]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="globulifer">
<emphasis id="7A9EEA90D847FFAAFCE17291FB8BFABA" box="[887,1071,1176,1200]" italics="true" pageId="5" pageNumber="6">Bryaxis globulifer</emphasis>
Schaufuss
</taxonomicName>
), by monotypy. Synonymized by
<treatmentCitation id="C94B1093D847FFAAFEBD72BEFE43FAC5" author="Raffray, A." box="[299,487,1206,1231]" page="171" pageId="5" pageNumber="6" year="1900">
<bibRefCitation id="2C7B4B73D847FFAAFEBD72BEFE43FAC5" author="Raffray, A." box="[299,487,1206,1231]" pageId="5" pageNumber="6" pagination="131 - 249" refId="ref59150" refString="Raffray, A. (1900) Australian Pselaphidae. Proceedings of the Linnean Society of New South Wales, 25, 131 - 249." type="journal article" year="1900">Raffray 1900: 171</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="48553682D847FFAAFF0172DDFC4DFB06" blockId="5.[151,1437,1022,2043]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="68FA11ACD847FFAAFF0172DDFC4DFB06" pageId="5" pageNumber="6">
<treatmentCitation id="C94B1093D847FFAAFF0172DDFE1EFAE7" author="Raffray, A." box="[151,442,1236,1261]" page="125" pageId="5" pageNumber="6" year="1890">
<taxonomicName id="8FEA4D01D847FFAAFF0172DDFE1EFAE7" ID-CoL="8XK4D" authority="Raffray, 1890: 125" authorityName="Raffray" authorityPageNumber="125" authorityYear="1890" box="[151,442,1236,1261]" class="Insecta" family="Staphylinidae" genus="Brabaxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D847FFAAFF0172DDFF56FAE7" box="[151,242,1237,1261]" italics="true" pageId="5" pageNumber="6">Brabaxis</emphasis>
<bibRefCitation id="2C7B4B73D847FFAAFF6F72DCFE1EFAE7" author="Raffray, A." box="[249,442,1236,1261]" pageId="5" pageNumber="6" pagination="81 - 172" refId="ref59092" refString="Raffray, A. (1890) Etude sur les Pselaphides. V. Tableaux synoptiques-Notes et synonymie. Revue d'Entomologie, 9, 81 - 172." type="journal article" year="1890">Raffray, 1890: 125</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="97518820D847FFAAFE5572DEFE51FAE4" box="[451,501,1238,1262]" pageId="5" pageNumber="6">Type</typeStatus>
species:
<taxonomicName id="8FEA4D01D847FFAAFDC472DEFCD1FAE7" authority="Schaufuss" authorityName="Schaufuss" box="[594,885,1236,1261]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="7A9EEA90D847FFAAFDC472DEFCACFAE7" box="[594,776,1238,1261]" italics="true" pageId="5" pageNumber="6">Bryaxis siamensis</emphasis>
Schaufuss
</taxonomicName>
(=
<taxonomicName id="8FEA4D01D847FFAAFC0472DEFB73FAE7" authority="Motschulsky" authorityName="Motschulsky" authorityYear="1851" box="[914,1239,1236,1261]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sphaerica">
<emphasis id="7A9EEA90D847FFAAFC0472DEFBEDFAE7" box="[914,1097,1237,1261]" italics="true" pageId="5" pageNumber="6">Bryaxis sphaerica</emphasis>
Motschulsky
</taxonomicName>
), subsequent designation by
<treatmentCitation id="C94B1093D847FFAAFEBB72FCFDC9FB06" author="Newton, A. F. &amp; Chandler, D. S." box="[301,621,1267,1292]" page="45" pageId="5" pageNumber="6" year="1989">
<bibRefCitation id="2C7B4B73D847FFAAFEBB72FCFDC9FB06" author="Newton, A. F. &amp; Chandler, D. S." box="[301,621,1267,1292]" pageId="5" pageNumber="6" pagination="1 - 93" refId="ref58915" refString="Newton, A. F. &amp; Chandler, D. S. (1989) World catalog of the genera of Pselaphidae (Coleoptera). Fieldiana Zoology, 53, I-IV + 1 - 93. https: // doi. org / 10.5962 / bhl. title. 3209" type="journal article" year="1989">Newton and Chandler 1989: 45</bibRefCitation>
</treatmentCitation>
. Synonymized by
<treatmentCitation id="C94B1093D847FFAAFCBF72FBFC41FB06" author="Raffray, A." box="[809,997,1267,1292]" page="258" pageId="5" pageNumber="6" year="1897">
<bibRefCitation id="2C7B4B73D847FFAAFCBF72FBFC41FB06" author="Raffray, A." box="[809,997,1267,1292]" pageId="5" pageNumber="6" pagination="227 - 284" refId="ref59120" refString="Raffray, A. (1897) Nouvelles etudes sur les Pselaphides et les Clavigerides. Annales de la Societe entomologique de France, 65, 227 - 284." type="journal article" year="1897">Raffray 1897: 258</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="48553682D847FFAAFF01731AFDD8FB45" blockId="5.[151,1437,1022,2043]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="68FA11ACD847FFAAFF01731AFDD8FB45" pageId="5" pageNumber="6">
<treatmentCitation id="C94B1093D847FFAAFF01731AFE1AFB21" author="Raffray, A." box="[151,446,1298,1323]" page="168" pageId="5" pageNumber="6" year="1900">
<taxonomicName id="8FEA4D01D847FFAAFF01731AFE1AFB21" ID-CoL="9LCBW" authority="Raffray, 1900: 168" authorityName="Raffray" authorityPageNumber="168" authorityYear="1900" box="[151,446,1298,1323]" class="Insecta" family="Staphylinidae" genus="Eupinoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D847FFAAFF01731AFF5EFB20" box="[151,250,1298,1322]" italics="true" pageId="5" pageNumber="6">Eupinoda</emphasis>
<bibRefCitation id="2C7B4B73D847FFAAFF69731AFE1AFB21" author="Raffray, A." box="[255,446,1298,1323]" pageId="5" pageNumber="6" pagination="131 - 249" refId="ref59150" refString="Raffray, A. (1900) Australian Pselaphidae. Proceedings of the Linnean Society of New South Wales, 25, 131 - 249." type="journal article" year="1900">Raffray, 1900: 168</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="97518820D847FFAAFE50731BFE5CFB21" box="[454,504,1299,1323]" pageId="5" pageNumber="6">Type</typeStatus>
species:
<taxonomicName id="8FEA4D01D847FFAAFDC5731AFCE2FB21" authority="Raffray" authorityName="Raffray" authorityYear="1900" box="[595,838,1298,1323]" class="Insecta" family="Staphylinidae" genus="Eupinoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="leana">
<emphasis id="7A9EEA90D847FFAAFDC5731AFD57FB20" box="[595,755,1298,1322]" italics="true" pageId="5" pageNumber="6">Eupinoda leana</emphasis>
Raffray
</taxonomicName>
, subsequent designation by
<treatmentCitation id="C94B1093D847FFAAFBC8731BFAACFB21" author="Lucas, R." box="[1118,1288,1299,1323]" page="284" pageId="5" pageNumber="6" year="1920">
<bibRefCitation id="2C7B4B73D847FFAAFBC8731BFAACFB21" author="Lucas, R." box="[1118,1288,1299,1323]" pageId="5" pageNumber="6" refId="ref58726" refString="Lucas, R. (1920) Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Vol. 84. Archiv fur Naturgeschichte, Berlin, XXXI + 696 pp." type="book" year="1920">Lucas 1920: 284</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="C94B1093D847FFAAFA86731AFF54FB45" author="Raffray, A." page="217" pageId="5" pageNumber="6" year="1908">
<bibRefCitation id="2C7B4B73D847FFAAFA86731AFF54FB45" author="Raffray, A." pageId="5" pageNumber="6" pagination="1 - 484" refId="ref59206" refString="Raffray, A. (1908) Coleoptera fam. Pselaphidae. In: Wytsman, P. (Ed.), Genera insectorum. 64 th Fascicule. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 484, 489 pls." type="book chapter" year="1908">Raffray 1908: 217</bibRefCitation>
</treatmentCitation>
. Synonymized by
<treatmentCitation id="C94B1093D847FFAAFE3D733EFDD3FB45" author="Chandler, D. S." box="[427,631,1334,1359]" page="320" pageId="5" pageNumber="6" year="2001">
<bibRefCitation id="2C7B4B73D847FFAAFE3D733EFDD3FB45" author="Chandler, D. S." box="[427,631,1334,1359]" pageId="5" pageNumber="6" refId="ref58398" refString="Chandler, D. S. (2001) Biology, morphology, and systematics of the ant-like litter beetle genera of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology International. Entomology International, Associated Publishers, Gainesville, Florida, VIII + 560 pp." type="book" year="2001">Chandler 2001: 320</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="00F06509D847FFAAFF017350FC4FF82D" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="48553682D847FFAAFF017350FC4FF82D" blockId="5.[151,1437,1022,2043]" pageId="5" pageNumber="6">
<emphasis id="7A9EEA90D847FFAAFF017350FEB6FB79" bold="true" box="[151,274,1368,1395]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Head with asetose vertexal foveae small or lacking; lacking frontal fovea; antennal clubs 14 segment- ed; antennal tubercles moderately developed to lacking; postantennal notches small or lacking; postantennal cavities present from vaguely visible to as long as antennal tubercles; ventrolateral margin beneath eyes often rounded; median gular ridge narrow and low, barely visible in lateral view. Pronotum with lateral antebasal foveae punctiform or lacking; lacking median antebasal fovea. Elytra lacking basal foveae and discal stria. Abdominal tergite 1 lacking mediobasal foveae and discal carinae; ventrite 2 lacking mediobasal foveae.
</paragraph>
</subSubSection>
<subSubSection id="00F06509D847FFA7FF517038FAFAFD1D" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="description">
<paragraph id="48553682D847FFA9FF517038FD55FEDD" blockId="5.[151,1437,1022,2043]" lastBlockId="6.[151,1437,153,323]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="7A9EEA90D847FFAAFF517038FEC9F841" bold="true" box="[199,365,1584,1611]" pageId="5" pageNumber="6">Redescription</emphasis>
[based on
<collectingCountry id="30FD7612D847FFAAFE7E7039FDDAF841" box="[488,638,1585,1611]" name="New Zealand" pageId="5" pageNumber="6">New Zealand</collectingCountry>
members of
<taxonomicName id="8FEA4D01D847FFAAFC84703AFCC8F841" authorityName="King. A" authorityYear="1866" box="[786,876,1586,1611]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D847FFAAFC84703AFCC8F841" box="[786,876,1586,1611]" italics="true" pageId="5" pageNumber="6">Eupines</emphasis>
</taxonomicName>
]. Body length
<quantity id="8F129B67D847FFAAFB827039FB1EF841" box="[1044,1210,1585,1611]" metricMagnitude="-3" metricUnit="m" metricValue="1.665" metricValueMax="2.1" metricValueMin="1.23" pageId="5" pageNumber="6" unit="mm" value="1.665" valueMax="2.1" valueMin="1.23">1.232.10 mm</quantity>
, generally short and compact in dorsal view; body color reddish brown to dark brown and rarely black; head and pronotum smooth, elytra and abdomen micropunctate or smooth; lacking setae or with sparsely scattered short decumbent setae on pronotum, elytra and abdomen, average length longer than length of eyes. Head with asetose vertexal foveae (
<figureCitation id="D0D12A07D847FFAAFAF97095FF1FF8D1" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1823]" captionTargetBox="[161,1420,353,1766]" captionTargetId="figure@6.[151,1435,345,1773]" captionTargetPageId="6" captionText="FIGURE 2. Morphology of Eupines King (based on E. (B.) crassicornides). A) Head, in dorsal view. B) Same, in lateral view. C) Same, in ventral view. D) Mouth parts. E) Mandibles and labrum. F) Female antenna. G) Male antenna. H) Pronotum. I) Same, in lateral view. J) Prothorax. K) Meso- and metaventrite. Abbreviations: pc—postantennal cavities, gf—gular foveae, gr—gular ridge, omc—ocular-mandibular carinae, l—lacinia, lb—labrum, lp—labial palp, m—mentum, lmcf—lateral mesocoxal foveae, lmtf—lateral metaventral foveae, lpcf—lateral procoxal foveae, lmvf—lateral mesoventral foveae, mmsf—median mesoventral foveae, ppf—prepectal foveae, vf—vertexal foveae. Scale bars: 0.2 mm, except D, E, H, I, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821178" httpUri="https://zenodo.org/record/3821178/files/figure.png" pageId="5" pageNumber="6">Fig. 2A</figureCitation>
) small or lacking, often punctiform; lacking frontal fovea; frontal rostrum often moderately developed; antennal tubercles moderately developed to lacking; postantennal notches distinctively margined to lacking; postantennal cavity (
<figureCitation id="D0D12A07D847FFAAFF7C7101FE92F929" box="[234,310,1801,1827]" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1823]" captionTargetBox="[161,1420,353,1766]" captionTargetId="figure@6.[151,1435,345,1773]" captionTargetPageId="6" captionText="FIGURE 2. Morphology of Eupines King (based on E. (B.) crassicornides). A) Head, in dorsal view. B) Same, in lateral view. C) Same, in ventral view. D) Mouth parts. E) Mandibles and labrum. F) Female antenna. G) Male antenna. H) Pronotum. I) Same, in lateral view. J) Prothorax. K) Meso- and metaventrite. Abbreviations: pc—postantennal cavities, gf—gular foveae, gr—gular ridge, omc—ocular-mandibular carinae, l—lacinia, lb—labrum, lp—labial palp, m—mentum, lmcf—lateral mesocoxal foveae, lmtf—lateral metaventral foveae, lpcf—lateral procoxal foveae, lmvf—lateral mesoventral foveae, mmsf—median mesoventral foveae, ppf—prepectal foveae, vf—vertexal foveae. Scale bars: 0.2 mm, except D, E, H, I, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821178" httpUri="https://zenodo.org/record/3821178/files/figure.png" pageId="5" pageNumber="6">Figs. 2</figureCitation>
AB) extend from dorsal side of head to lateral side of antennal insertions, size vary from vaguely visible to as long as antennal tubercles; eyes often well developed and prominent; antennae (
<figureCitation id="D0D12A07D847FFAAFBEE7125FB60F94D" box="[1144,1220,1837,1863]" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1823]" captionTargetBox="[161,1420,353,1766]" captionTargetId="figure@6.[151,1435,345,1773]" captionTargetPageId="6" captionText="FIGURE 2. Morphology of Eupines King (based on E. (B.) crassicornides). A) Head, in dorsal view. B) Same, in lateral view. C) Same, in ventral view. D) Mouth parts. E) Mandibles and labrum. F) Female antenna. G) Male antenna. H) Pronotum. I) Same, in lateral view. J) Prothorax. K) Meso- and metaventrite. Abbreviations: pc—postantennal cavities, gf—gular foveae, gr—gular ridge, omc—ocular-mandibular carinae, l—lacinia, lb—labrum, lp—labial palp, m—mentum, lmcf—lateral mesocoxal foveae, lmtf—lateral metaventral foveae, lpcf—lateral procoxal foveae, lmvf—lateral mesoventral foveae, mmsf—median mesoventral foveae, ppf—prepectal foveae, vf—vertexal foveae. Scale bars: 0.2 mm, except D, E, H, I, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821178" httpUri="https://zenodo.org/record/3821178/files/figure.png" pageId="5" pageNumber="6">Figs. 2</figureCitation>
FG) 9-segmented, 10-segmented or 11-segmented in males and 11-segmented in females, antennal clubs 14 segmented, with scape and pedicel elongate; ocular-mandibular carinae (
<figureCitation id="D0D12A07D847FFAAFD5A717DFC80F985" box="[716,804,1909,1935]" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1823]" captionTargetBox="[161,1420,353,1766]" captionTargetId="figure@6.[151,1435,345,1773]" captionTargetPageId="6" captionText="FIGURE 2. Morphology of Eupines King (based on E. (B.) crassicornides). A) Head, in dorsal view. B) Same, in lateral view. C) Same, in ventral view. D) Mouth parts. E) Mandibles and labrum. F) Female antenna. G) Male antenna. H) Pronotum. I) Same, in lateral view. J) Prothorax. K) Meso- and metaventrite. Abbreviations: pc—postantennal cavities, gf—gular foveae, gr—gular ridge, omc—ocular-mandibular carinae, l—lacinia, lb—labrum, lp—labial palp, m—mentum, lmcf—lateral mesocoxal foveae, lmtf—lateral metaventral foveae, lpcf—lateral procoxal foveae, lmvf—lateral mesoventral foveae, mmsf—median mesoventral foveae, ppf—prepectal foveae, vf—vertexal foveae. Scale bars: 0.2 mm, except D, E, H, I, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821178" httpUri="https://zenodo.org/record/3821178/files/figure.png" pageId="5" pageNumber="6">Fig. 2B</figureCitation>
) present; ventrolateral margin rounded beneath eyes or with short basal carinae; median gular ridge (
<figureCitation id="D0D12A07D847FFAAFD187191FD47F9B9" box="[654,739,1945,1971]" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1823]" captionTargetBox="[161,1420,353,1766]" captionTargetId="figure@6.[151,1435,345,1773]" captionTargetPageId="6" captionText="FIGURE 2. Morphology of Eupines King (based on E. (B.) crassicornides). A) Head, in dorsal view. B) Same, in lateral view. C) Same, in ventral view. D) Mouth parts. E) Mandibles and labrum. F) Female antenna. G) Male antenna. H) Pronotum. I) Same, in lateral view. J) Prothorax. K) Meso- and metaventrite. Abbreviations: pc—postantennal cavities, gf—gular foveae, gr—gular ridge, omc—ocular-mandibular carinae, l—lacinia, lb—labrum, lp—labial palp, m—mentum, lmcf—lateral mesocoxal foveae, lmtf—lateral metaventral foveae, lpcf—lateral procoxal foveae, lmvf—lateral mesoventral foveae, mmsf—median mesoventral foveae, ppf—prepectal foveae, vf—vertexal foveae. Scale bars: 0.2 mm, except D, E, H, I, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821178" httpUri="https://zenodo.org/record/3821178/files/figure.png" pageId="5" pageNumber="6">Fig. 2C</figureCitation>
) narrow and low, barely visible in lateral view, curved posteriorly to asetose gular foveae (
<figureCitation id="D0D12A07D847FFAAFE4171B5FD8BF9DD" box="[471,559,1981,2007]" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1823]" captionTargetBox="[161,1420,353,1766]" captionTargetId="figure@6.[151,1435,345,1773]" captionTargetPageId="6" captionText="FIGURE 2. Morphology of Eupines King (based on E. (B.) crassicornides). A) Head, in dorsal view. B) Same, in lateral view. C) Same, in ventral view. D) Mouth parts. E) Mandibles and labrum. F) Female antenna. G) Male antenna. H) Pronotum. I) Same, in lateral view. J) Prothorax. K) Meso- and metaventrite. Abbreviations: pc—postantennal cavities, gf—gular foveae, gr—gular ridge, omc—ocular-mandibular carinae, l—lacinia, lb—labrum, lp—labial palp, m—mentum, lmcf—lateral mesocoxal foveae, lmtf—lateral metaventral foveae, lpcf—lateral procoxal foveae, lmvf—lateral mesoventral foveae, mmsf—median mesoventral foveae, ppf—prepectal foveae, vf—vertexal foveae. Scale bars: 0.2 mm, except D, E, H, I, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821178" httpUri="https://zenodo.org/record/3821178/files/figure.png" pageId="5" pageNumber="6">Fig. 2C</figureCitation>
); mouth (
<figureCitation id="D0D12A07D847FFAAFD0F71B5FD41F9DD" box="[665,741,1981,2007]" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1823]" captionTargetBox="[161,1420,353,1766]" captionTargetId="figure@6.[151,1435,345,1773]" captionTargetPageId="6" captionText="FIGURE 2. Morphology of Eupines King (based on E. (B.) crassicornides). A) Head, in dorsal view. B) Same, in lateral view. C) Same, in ventral view. D) Mouth parts. E) Mandibles and labrum. F) Female antenna. G) Male antenna. H) Pronotum. I) Same, in lateral view. J) Prothorax. K) Meso- and metaventrite. Abbreviations: pc—postantennal cavities, gf—gular foveae, gr—gular ridge, omc—ocular-mandibular carinae, l—lacinia, lb—labrum, lp—labial palp, m—mentum, lmcf—lateral mesocoxal foveae, lmtf—lateral metaventral foveae, lpcf—lateral procoxal foveae, lmvf—lateral mesoventral foveae, mmsf—median mesoventral foveae, ppf—prepectal foveae, vf—vertexal foveae. Scale bars: 0.2 mm, except D, E, H, I, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821178" httpUri="https://zenodo.org/record/3821178/files/figure.png" pageId="5" pageNumber="6">Figs. 2</figureCitation>
DE) with labial palp truncate at apex, with pair of long apical setae, mentum widest at middle, lacinia well developed, galea fused with lacinia, maxillary palpi with segment 2 slender and clavate, segment 3 trapezoidal, segment 4 large and fusiform, sensory area with stout apical setae, mandible with pair of subapical setae, labrum trapezoidal.
</paragraph>
<paragraph id="48553682D844FFA9FF5176E9FEF8FF49" blockId="6.[151,1437,153,323]" pageId="6" pageNumber="7">
Pronotum about as long as wide; lacking median antebasal fovea; lateral antebasal foveae punctiform or lacking; lacking antebasal sulcus and median longitudinal sulcus; setose lateral procoxal foveae (
<figureCitation id="D0D12A07D844FFA9FB01770DFB41FF15" box="[1175,1253,261,287]" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1823]" captionTargetBox="[161,1420,353,1766]" captionTargetId="figure@6.[151,1435,345,1773]" captionTargetPageId="6" captionText="FIGURE 2. Morphology of Eupines King (based on E. (B.) crassicornides). A) Head, in dorsal view. B) Same, in lateral view. C) Same, in ventral view. D) Mouth parts. E) Mandibles and labrum. F) Female antenna. G) Male antenna. H) Pronotum. I) Same, in lateral view. J) Prothorax. K) Meso- and metaventrite. Abbreviations: pc—postantennal cavities, gf—gular foveae, gr—gular ridge, omc—ocular-mandibular carinae, l—lacinia, lb—labrum, lp—labial palp, m—mentum, lmcf—lateral mesocoxal foveae, lmtf—lateral metaventral foveae, lpcf—lateral procoxal foveae, lmvf—lateral mesoventral foveae, mmsf—median mesoventral foveae, ppf—prepectal foveae, vf—vertexal foveae. Scale bars: 0.2 mm, except D, E, H, I, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821178" httpUri="https://zenodo.org/record/3821178/files/figure.png" pageId="6" pageNumber="7">Fig. 2J</figureCitation>
) small and often widely separated.
</paragraph>
<caption id="1C95660AD844FFA9FF01710EFAC4F9D8" ID-DOI="http://doi.org/10.5281/zenodo.3821178" ID-Zenodo-Dep="3821178" httpUri="https://zenodo.org/record/3821178/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,250,1798,1823]" targetBox="[161,1420,353,1766]" targetPageId="6">
<paragraph id="48553682D844FFA9FF01710EFAC4F9D8" blockId="6.[151,1437,1798,2003]" pageId="6" pageNumber="7">
<emphasis id="7A9EEA90D844FFA9FF01710EFEB7F915" bold="true" box="[151,275,1798,1823]" pageId="6" pageNumber="7">FIGURE 2.</emphasis>
Morphology of
<taxonomicName id="8FEA4D01D844FFA9FE2C7100FDE1F915" authority="King" authorityName="King. A" authorityYear="1866" box="[442,581,1798,1823]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D844FFA9FE2C7100FDA8F915" box="[442,524,1800,1823]" italics="true" pageId="6" pageNumber="7">Eupines</emphasis>
King
</taxonomicName>
(based on
<taxonomicName id="8FEA4D01D844FFA9FD257100FC34F915" authorityName="Newton" authorityYear="2017" box="[691,912,1799,1823]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="10" phylum="Arthropoda" rank="species" species="crassicornides" subGenus="Byraxis">
<emphasis id="7A9EEA90D844FFA9FD257100FD66F915" box="[691,706,1800,1823]" italics="true" pageId="6" pageNumber="7">E</emphasis>
. (
<emphasis id="7A9EEA90D844FFA9FD417100FD42F915" box="[727,742,1800,1823]" italics="true" pageId="6" pageNumber="7">B</emphasis>
.)
<emphasis id="7A9EEA90D844FFA9FD6A710FFC34F915" box="[764,912,1799,1823]" italics="true" pageId="6" pageNumber="7">crassicornides</emphasis>
</taxonomicName>
).
<emphasis id="7A9EEA90D844FFA9FC33710EFC1BF915" bold="true" box="[933,959,1798,1823]" pageId="6" pageNumber="7">A)</emphasis>
Head, in dorsal view.
<emphasis id="7A9EEA90D844FFA9FB34710EFB18F915" bold="true" box="[1186,1212,1798,1823]" pageId="6" pageNumber="7">B)</emphasis>
Same, in lateral view.
<emphasis id="7A9EEA90D844FFA9FF017122FF16F949" bold="true" box="[151,178,1834,1859]" pageId="6" pageNumber="7">C)</emphasis>
Same, in ventral view.
<emphasis id="7A9EEA90D844FFA9FE317122FE65F949" bold="true" box="[423,449,1834,1859]" pageId="6" pageNumber="7">D)</emphasis>
Mouth parts.
<emphasis id="7A9EEA90D844FFA9FDC27122FDC9F949" bold="true" box="[596,621,1834,1859]" pageId="6" pageNumber="7">E)</emphasis>
Mandibles and labrum.
<emphasis id="7A9EEA90D844FFA9FCFF7122FC26F949" bold="true" box="[873,898,1834,1859]" pageId="6" pageNumber="7">F)</emphasis>
Female antenna.
<emphasis id="7A9EEA90D844FFA9FBAE7122FBF0F949" bold="true" box="[1080,1108,1834,1859]" pageId="6" pageNumber="7">G)</emphasis>
Male antenna.
<emphasis id="7A9EEA90D844FFA9FB627122FAB4F949" bold="true" box="[1268,1296,1834,1859]" pageId="6" pageNumber="7">H)</emphasis>
Pronotum.
<emphasis id="7A9EEA90D844FFA9FA1C7122FA38F949" bold="true" box="[1418,1436,1834,1859]" pageId="6" pageNumber="7">I)</emphasis>
Same, in lateral view.
<emphasis id="7A9EEA90D844FFA9FEE97146FE30F96D" bold="true" box="[383,404,1870,1895]" pageId="6" pageNumber="7">J)</emphasis>
Prothorax.
<emphasis id="7A9EEA90D844FFA9FD997146FD8EF96D" bold="true" box="[527,554,1870,1895]" pageId="6" pageNumber="7">K)</emphasis>
Meso- and metaventrite. Abbreviations: pc—postantennal cavities, gf—gular foveae, gr—gular ridge, omc—ocular-mandibular carinae, l—lacinia, lb—labrum, lp—labial palp, m—mentum, lmcf—lateral mesocoxal foveae, lmtf—lateral metaventral foveae, lpcf—lateral procoxal foveae, lmvf—lateral mesoventral foveae, mmsf—me-dian mesoventral foveae, ppf—prepectal foveae, vf—vertexal foveae. Scale bars: 0.2 mm, except D, E, H, I, J = 0.1 mm.
</paragraph>
</caption>
<caption id="1C95660AD845FFA8FF017084FD2EF8E7" ID-DOI="http://doi.org/10.5281/zenodo.3821180" ID-Zenodo-Dep="3821180" httpUri="https://zenodo.org/record/3821180/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1676,1701]" targetBox="[162,1432,187,1639]" targetPageId="7">
<paragraph id="48553682D845FFA8FF017084FD2EF8E7" blockId="7.[151,1436,1676,1773]" pageId="7" pageNumber="8">
<emphasis id="7A9EEA90D845FFA8FF017084FEB7F8AF" bold="true" box="[151,275,1676,1701]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
Morphology of
<taxonomicName id="8FEA4D01D845FFA8FE2D7086FDC2F8AF" authority="King. A" authorityName="King. A" authorityYear="1866" box="[443,614,1676,1701]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D845FFA8FE2D7086FDA9F8AF" box="[443,525,1678,1701]" italics="true" pageId="7" pageNumber="8">Eupines</emphasis>
King.
<emphasis id="7A9EEA90D845FFA8FDC57084FDC2F8AF" bold="true" box="[595,614,1676,1701]" pageId="7" pageNumber="8">A</emphasis>
</taxonomicName>
<emphasis id="7A9EEA90D845FFA8FDF17084FDCAF8AF" bold="true" box="[615,622,1676,1701]" pageId="7" pageNumber="8">)</emphasis>
Tergites 15.
<emphasis id="7A9EEA90D845FFA8FD697084FCBDF8AF" bold="true" box="[767,793,1676,1701]" pageId="7" pageNumber="8">B)</emphasis>
Tergite 1.
<emphasis id="7A9EEA90D845FFA8FC107084FC05F8AF" bold="true" box="[902,929,1676,1701]" pageId="7" pageNumber="8">C)</emphasis>
Tergite 2.
<emphasis id="7A9EEA90D845FFA8FB987084FB8CF8AF" bold="true" box="[1038,1064,1676,1701]" pageId="7" pageNumber="8">D)</emphasis>
Tergite 3.
<emphasis id="7A9EEA90D845FFA8FB007084FB0BF8AF" bold="true" box="[1174,1199,1676,1701]" pageId="7" pageNumber="8">E)</emphasis>
Tergite 4.
<emphasis id="7A9EEA90D845FFA8FA8B7084FA92F8AF" bold="true" box="[1309,1334,1676,1701]" pageId="7" pageNumber="8">F)</emphasis>
Tergite 5.
<emphasis id="7A9EEA90D845FFA8FF0170B8FF17F8C3" bold="true" box="[151,179,1712,1737]" pageId="7" pageNumber="8">G)</emphasis>
Ventrites 26.
<emphasis id="7A9EEA90D845FFA8FED870B8FECEF8C3" bold="true" box="[334,362,1712,1737]" pageId="7" pageNumber="8">H)</emphasis>
Ventrites 46.
<emphasis id="7A9EEA90D845FFA8FD9370B8FDB3F8C3" bold="true" box="[517,535,1712,1737]" pageId="7" pageNumber="8">I)</emphasis>
Ventrite 2.
<emphasis id="7A9EEA90D845FFA8FD1870B8FD07F8C3" bold="true" box="[654,675,1712,1737]" pageId="7" pageNumber="8">J)</emphasis>
Ventrite 3.
<emphasis id="7A9EEA90D845FFA8FC8D70B8FC93F8C3" bold="true" box="[795,823,1712,1737]" pageId="7" pageNumber="8">K)</emphasis>
Ventrite 4.
<emphasis id="7A9EEA90D845FFA8FC3870B8FC6CF8C3" bold="true" box="[942,968,1712,1737]" pageId="7" pageNumber="8">L)</emphasis>
Ventrite 5.
<emphasis id="7A9EEA90D845FFA8FBA970B8FBFBF8C3" bold="true" box="[1087,1119,1712,1737]" pageId="7" pageNumber="8">M)</emphasis>
Ventrite 6.
<emphasis id="7A9EEA90D845FFA8FB4170B8FB55F8C3" bold="true" box="[1239,1265,1712,1737]" pageId="7" pageNumber="8">N)</emphasis>
Elytra, in dorsal view. Scale bars: 0.1 mm, except A, N = 0.2 mm.
</paragraph>
</caption>
<paragraph id="48553682D845FFA8FF517113FB14F953" blockId="7.[151,1437,1819,2026]" pageId="7" pageNumber="8">
Elytra (
<figureCitation id="D0D12A07D845FFA8FE887113FEDDF93C" box="[286,377,1819,1846]" captionStart="FIGURE 3" captionStartId="7.[151,250,1676,1701]" captionTargetBox="[162,1432,187,1639]" captionTargetId="figure@7.[151,1435,181,1651]" captionTargetPageId="7" captionText="FIGURE 3. Morphology of Eupines King. A) Tergites 15. B) Tergite 1. C) Tergite 2. D) Tergite 3. E) Tergite 4. F) Tergite 5. G) Ventrites 26. H) Ventrites 46. I) Ventrite 2. J) Ventrite 3. K) Ventrite 4. L) Ventrite 5. M) Ventrite 6. N) Elytra, in dorsal view. Scale bars: 0.1 mm, except A, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3821180" httpUri="https://zenodo.org/record/3821180/files/figure.png" pageId="7" pageNumber="8">Fig. 3N</figureCitation>
) slightly wider than long, longer than abdomen in dorsal view, with posterior margin often rounded; each elytron lacking basal foveae and discal carinae; sutural stria distinct to lacking.
</paragraph>
<paragraph id="48553682D845FFA8FF51716BFAFEF9CF" blockId="7.[151,1437,1819,2026]" pageId="7" pageNumber="8">
Mesoventrite (
<figureCitation id="D0D12A07D845FFA8FEFC716BFE66F974" box="[362,450,1891,1918]" captionStart="FIGURE 2" captionStartId="6.[151,250,1798,1823]" captionTargetBox="[161,1420,353,1766]" captionTargetId="figure@6.[151,1435,345,1773]" captionTargetPageId="6" captionText="FIGURE 2. Morphology of Eupines King (based on E. (B.) crassicornides). A) Head, in dorsal view. B) Same, in lateral view. C) Same, in ventral view. D) Mouth parts. E) Mandibles and labrum. F) Female antenna. G) Male antenna. H) Pronotum. I) Same, in lateral view. J) Prothorax. K) Meso- and metaventrite. Abbreviations: pc—postantennal cavities, gf—gular foveae, gr—gular ridge, omc—ocular-mandibular carinae, l—lacinia, lb—labrum, lp—labial palp, m—mentum, lmcf—lateral mesocoxal foveae, lmtf—lateral metaventral foveae, lpcf—lateral procoxal foveae, lmvf—lateral mesoventral foveae, mmsf—median mesoventral foveae, ppf—prepectal foveae, vf—vertexal foveae. Scale bars: 0.2 mm, except D, E, H, I, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821178" httpUri="https://zenodo.org/record/3821178/files/figure.png" pageId="7" pageNumber="8">Fig. 2K</figureCitation>
) with asetose prepectal foveae vaguely visible; setose median mesoventral foveae bifurcate at apex, often forming triangular cavity; pair of setose lateral mesoventral foveae large and nearly meet internally; setose lateral mesocoxal foveae often small. Metaventrite with setose lateral metaventral foveae small.
</paragraph>
<paragraph id="48553682D845FFA7FF5171C7FE67FF15" blockId="7.[151,1437,1819,2026]" lastBlockId="8.[151,1437,153,1295]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Abdomen (
<figureCitation id="D0D12A07D845FFA8FED371C7FE72F9E0" box="[325,470,1999,2026]" captionStart="FIGURE 3" captionStartId="7.[151,250,1676,1701]" captionTargetBox="[162,1432,187,1639]" captionTargetId="figure@7.[151,1435,181,1651]" captionTargetPageId="7" captionText="FIGURE 3. Morphology of Eupines King. A) Tergites 15. B) Tergite 1. C) Tergite 2. D) Tergite 3. E) Tergite 4. F) Tergite 5. G) Ventrites 26. H) Ventrites 46. I) Ventrite 2. J) Ventrite 3. K) Ventrite 4. L) Ventrite 5. M) Ventrite 6. N) Elytra, in dorsal view. Scale bars: 0.1 mm, except A, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3821180" httpUri="https://zenodo.org/record/3821180/files/figure.png" pageId="7" pageNumber="8">Figs. 3A, 3G</figureCitation>
) convex, with tergite 1 about 1.5 × to 2 × longer than tergite 2; tergite 1 (
<figureCitation id="D0D12A07D845FFA8FA9071C7FAFFF9E0" box="[1286,1371,1999,2026]" captionStart="FIGURE 3" captionStartId="7.[151,250,1676,1701]" captionTargetBox="[162,1432,187,1639]" captionTargetId="figure@7.[151,1435,181,1651]" captionTargetPageId="7" captionText="FIGURE 3. Morphology of Eupines King. A) Tergites 15. B) Tergite 1. C) Tergite 2. D) Tergite 3. E) Tergite 4. F) Tergite 5. G) Ventrites 26. H) Ventrites 46. I) Ventrite 2. J) Ventrite 3. K) Ventrite 4. L) Ventrite 5. M) Ventrite 6. N) Elytra, in dorsal view. Scale bars: 0.1 mm, except A, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3821180" httpUri="https://zenodo.org/record/3821180/files/figure.png" pageId="7" pageNumber="8">Fig. 3B</figureCitation>
) with basolateral foveae vaguely visible; lacking mediobasal fovea, discal carinae and mediobasal impression; tergites 24 (
<figureCitation id="D0D12A07D84AFFA7FF4776B5FEB8FEDD" box="[209,284,189,215]" captionStart="FIGURE 3" captionStartId="7.[151,250,1676,1701]" captionTargetBox="[162,1432,187,1639]" captionTargetId="figure@7.[151,1435,181,1651]" captionTargetPageId="7" captionText="FIGURE 3. Morphology of Eupines King. A) Tergites 15. B) Tergite 1. C) Tergite 2. D) Tergite 3. E) Tergite 4. F) Tergite 5. G) Ventrites 26. H) Ventrites 46. I) Ventrite 2. J) Ventrite 3. K) Ventrite 4. L) Ventrite 5. M) Ventrite 6. N) Elytra, in dorsal view. Scale bars: 0.1 mm, except A, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3821180" httpUri="https://zenodo.org/record/3821180/files/figure.png" pageId="8" pageNumber="9">Figs. 3</figureCitation>
CE) lacking basolateral foveae; paratergites visible on tergites 13 and carinate on tergite 4. Abdominal ventrite 2 with basolateral foveae (
<figureCitation id="D0D12A07D84AFFA7FDD776E9FD28FEF1" box="[577,652,225,251]" captionStart="FIGURE 3" captionStartId="7.[151,250,1676,1701]" captionTargetBox="[162,1432,187,1639]" captionTargetId="figure@7.[151,1435,181,1651]" captionTargetPageId="7" captionText="FIGURE 3. Morphology of Eupines King. A) Tergites 15. B) Tergite 1. C) Tergite 2. D) Tergite 3. E) Tergite 4. F) Tergite 5. G) Ventrites 26. H) Ventrites 46. I) Ventrite 2. J) Ventrite 3. K) Ventrite 4. L) Ventrite 5. M) Ventrite 6. N) Elytra, in dorsal view. Scale bars: 0.1 mm, except A, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3821180" httpUri="https://zenodo.org/record/3821180/files/figure.png" pageId="8" pageNumber="9">Fig. 3I</figureCitation>
) vaguely visible, lacking mediobasal fovea, ventrites 35 (
<figureCitation id="D0D12A07D84AFFA7FA8A76E9FAC3FEF1" box="[1308,1383,225,251]" captionStart="FIGURE 3" captionStartId="7.[151,250,1676,1701]" captionTargetBox="[162,1432,187,1639]" captionTargetId="figure@7.[151,1435,181,1651]" captionTargetPageId="7" captionText="FIGURE 3. Morphology of Eupines King. A) Tergites 15. B) Tergite 1. C) Tergite 2. D) Tergite 3. E) Tergite 4. F) Tergite 5. G) Ventrites 26. H) Ventrites 46. I) Ventrite 2. J) Ventrite 3. K) Ventrite 4. L) Ventrite 5. M) Ventrite 6. N) Elytra, in dorsal view. Scale bars: 0.1 mm, except A, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3821180" httpUri="https://zenodo.org/record/3821180/files/figure.png" pageId="8" pageNumber="9">Figs. 3</figureCitation>
JL) lacking basolateral foveae.
</paragraph>
<paragraph id="48553682D84AFFA7FF517721FD70FF6D" blockId="8.[151,1437,153,1295]" pageId="8" pageNumber="9">Legs with protarsomeres 2 and 3 subequal in length, mesotarsomere 2 slightly longer than mesotarsomere 3, metatarsomere 2 about as long as metatarsomere 3.</paragraph>
<paragraph id="48553682D84AFFA7FF517779FE18FFD9" blockId="8.[151,1437,153,1295]" pageId="8" pageNumber="9">Males with antennae, protrochanters, protibiae, mesotibiae, ventrite 2 or ventrite 6 modified. Aedeagus with well-developed and often symmetrical parameres; median lobe often with two sclerites; basal bulb relatively small; lacking dorsal diaphragm.</paragraph>
<paragraph id="48553682D84AFFA7FF5177D4FAFAFD1D" blockId="8.[151,1437,153,1295]" pageId="8" pageNumber="9">
<emphasis id="7A9EEA90D84AFFA7FF5177D4FED6FFFD" bold="true" box="[199,370,476,503]" pageId="8" pageNumber="9">Relationships:</emphasis>
Within the
<collectingCountry id="30FD7612D84AFFA7FE6477D5FD23FFFD" box="[498,647,477,503]" name="New Zealand" pageId="8" pageNumber="9">New Zealand</collectingCountry>
fauna, members of
<taxonomicName id="8FEA4D01D84AFFA7FCF777D6FC1FFFFD" authorityName="King. A" authorityYear="1866" box="[865,955,478,503]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FCF777D6FC1FFFFD" box="[865,955,478,503]" italics="true" pageId="8" pageNumber="9">Eupines</emphasis>
</taxonomicName>
are most similar to those of
<taxonomicName id="8FEA4D01D84AFFA7FB6577D5FEB7FC11" authority="Park and Pearce" authorityName="Park and Pearce" class="Insecta" family="Staphylinidae" genus="Simkinion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FB6577D5FAC7FFFD" box="[1267,1379,477,503]" italics="true" pageId="8" pageNumber="9">Simkinion</emphasis>
Park and Pearce
</taxonomicName>
based on combination of the following characers: pronotum lacking a median antebasal fovea, tergite 1 lacking the mediobasal impression and ventrites 45 lacking the basolateral foveae. Members of
<taxonomicName id="8FEA4D01D84AFFA7FB64742EFAE8FC35" authorityName="King. A" authorityYear="1866" box="[1266,1356,550,575]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FB64742EFAE8FC35" box="[1266,1356,550,575]" italics="true" pageId="8" pageNumber="9">Eupines</emphasis>
</taxonomicName>
can be readily separated from those of
<taxonomicName id="8FEA4D01D84AFFA7FD947441FDD6FC69" box="[514,626,585,611]" class="Insecta" family="Staphylinidae" genus="Simkinion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FD947441FDD6FC69" box="[514,626,585,611]" italics="true" pageId="8" pageNumber="9">Simkinion</emphasis>
</taxonomicName>
by the absence of basal elytral foveae and the low median gular ridge, which is prominent in the latters. Members of
<taxonomicName id="8FEA4D01D84AFFA7FD0D7466FD51FC8D" authorityName="King. A" authorityYear="1866" box="[667,757,622,647]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FD0D7466FD51FC8D" box="[667,757,622,647]" italics="true" pageId="8" pageNumber="9">Eupines</emphasis>
</taxonomicName>
are also similar to that of
<taxonomicName id="8FEA4D01D84AFFA7FB817465FB1AFC8D" authority="Oke" authorityName="Oke, Nomura &amp; Leschen" authorityYear="2006" box="[1047,1214,621,647]" class="Insecta" family="Staphylinidae" genus="Eupinolus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FB817465FB2DFC8D" box="[1047,1161,621,647]" italics="true" pageId="8" pageNumber="9">Eupinolus</emphasis>
Oke
</taxonomicName>
in lacking the basal elytral foveae and the short abdomen. They may be separated by the presence of a small median antebasal fovea and the punctate pronotal base in members of
<taxonomicName id="8FEA4D01D84AFFA7FDFC74BDFD78FCC5" authorityName="Oke, Nomura &amp; Leschen" authorityYear="2006" box="[618,732,693,719]" class="Insecta" family="Staphylinidae" genus="Eupinolus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FDFC74BDFD78FCC5" box="[618,732,693,719]" italics="true" pageId="8" pageNumber="9">Eupinolus</emphasis>
</taxonomicName>
, which are lacking in members of
<taxonomicName id="8FEA4D01D84AFFA7FBC974BEFB1DFCC5" authorityName="King. A" authorityYear="1866" box="[1119,1209,694,719]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FBC974BEFB1DFCC5" box="[1119,1209,694,719]" italics="true" pageId="8" pageNumber="9">Eupines</emphasis>
</taxonomicName>
. In
<collectingCountry id="30FD7612D84AFFA7FB7574BDFAE9FCC5" box="[1251,1357,693,719]" name="Australia" pageId="8" pageNumber="9">Australia</collectingCountry>
,
<bibRefCitation id="2C7B4B73D84AFFA7FACE74BDFEB1FCF9" author="Chandler, D. S." pageId="8" pageNumber="9" refId="ref58398" refString="Chandler, D. S. (2001) Biology, morphology, and systematics of the ant-like litter beetle genera of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology International. Entomology International, Associated Publishers, Gainesville, Florida, VIII + 560 pp." type="book" year="2001">Chandler (2001)</bibRefCitation>
stated that
<taxonomicName id="8FEA4D01D84AFFA7FE0174D2FE55FCF9" authorityName="King. A" authorityYear="1866" box="[407,497,730,755]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FE0174D2FE55FCF9" box="[407,497,730,755]" italics="true" pageId="8" pageNumber="9">Eupines</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="8FEA4D01D84AFFA7FD4C74D1FC1DFCF9" authority="Chandler" authorityName="Chandler" authorityYear="2001" box="[730,953,729,755]" class="Insecta" family="Staphylinidae" genus="Storeyella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FD4C74D1FCEFFCF9" box="[730,843,729,755]" italics="true" pageId="8" pageNumber="9">Storeyella</emphasis>
Chandler
</taxonomicName>
and
<taxonomicName id="8FEA4D01D84AFFA7FC6674D1FB7CFCF9" authority="Chandler" authorityName="Chandler" authorityYear="2001" box="[1008,1240,729,755]" class="Insecta" family="Staphylinidae" genus="Wataranka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FC6674D1FBCEFCF9" box="[1008,1130,729,755]" italics="true" pageId="8" pageNumber="9">Wataranka</emphasis>
Chandler
</taxonomicName>
but can be easily differentiated from both by the smooth basal pronotal area and the lack of a median antebasal pronotal fovea.
</paragraph>
</subSubSection>
<subSubSection id="00F06509D84AFFA7FF517528FBF2FB05" pageId="8" pageNumber="9" type="discussion">
<paragraph id="48553682D84AFFA7FF517528FC6FFAA9" blockId="8.[151,1437,153,1295]" pageId="8" pageNumber="9">
<emphasis id="7A9EEA90D84AFFA7FF517528FEEBFD31" bold="true" box="[199,335,800,827]" pageId="8" pageNumber="9">Comments.</emphasis>
The lack of the basal elytral foveae, the median antebasal pronotal fovea and the basolateral foveae on tergites and ventrites, suggests that
<taxonomicName id="8FEA4D01D84AFFA7FD15754EFD79FD55" authorityName="King. A" authorityYear="1866" box="[643,733,838,863]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FD15754EFD79FD55" box="[643,733,838,863]" italics="true" pageId="8" pageNumber="9">Eupines</emphasis>
</taxonomicName>
may be more derived than all other goniacerite genera in
<collectingCountry id="30FD7612D84AFFA7FAF1754DFF51FD89" name="New Zealand" pageId="8" pageNumber="9">New Zealand</collectingCountry>
. The subgenus
<taxonomicName id="8FEA4D01D84AFFA7FE357562FE5DFD89" authorityName="Reitter" authorityYear="1880" box="[419,505,874,899]" class="Insecta" family="Staphylinidae" genus="Byraxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FE357562FE5DFD89" box="[419,505,874,899]" italics="true" pageId="8" pageNumber="9">Byraxis</emphasis>
</taxonomicName>
is characterized by males possessing 10 antennomeres, and males of the nominate
<taxonomicName id="8FEA4D01D84AFFA7FF017586FF55FDAD" authorityName="King. A" authorityYear="1866" box="[151,241,910,935]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FF017586FF55FDAD" box="[151,241,910,935]" italics="true" pageId="8" pageNumber="9">Eupines</emphasis>
</taxonomicName>
possess 11 antennomeres: females of both subgenera have 11-segmented antennae.
<bibRefCitation id="2C7B4B73D84AFFA7FB2C7585FF40FDC6" author="Nomura, S. &amp; Leschen, R. A. B." pageId="8" pageNumber="9" pagination="239 - 272" refId="ref58967" refString="Nomura, S. &amp; Leschen, R. A. B. (2006) Faunistic review on the pselaphine species known from New Zealand (Insecta, Coleoptera, Staphylinidae). National Science Museum Monographs, 34, 239 - 272." type="journal article" year="2006">Nomura &amp; Leschen (2006)</bibRefCitation>
, in their faunistic review of New Zeland Pselaphinae, listed nine species under the nominate subgenus in
<collectingCountry id="30FD7612D84AFFA7FF0175DDFE97FDE5" box="[151,307,981,1007]" name="New Zealand" pageId="8" pageNumber="9">New Zealand</collectingCountry>
. In the following treatment of this subgenus by
<bibRefCitation id="2C7B4B73D84AFFA7FCC875DDFBD6FDE5" author="Shen, J. - W. &amp; Leschen, R. A. B." box="[862,1138,981,1007]" pageId="8" pageNumber="9" pagination="255 - 302" refId="ref59638" refString="Shen, J. - W. &amp; Leschen, R. A. B. (2019) Catalogue and type designations for New Zealand Goniaceritae (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4614 (2), 255 - 302. https: // doi. org / 10.11646 / zootaxa. 4614.2.2" type="journal article" year="2019">Shen &amp; Leschen (2019)</bibRefCitation>
, three species were found based solely on female
<typeStatus id="97518820D84AFFA7FE3775F1FE7BFA19" box="[417,479,1017,1043]" pageId="8" pageNumber="9">types</typeStatus>
, and one was transferred to the subgenus
<taxonomicName id="8FEA4D01D84AFFA7FC2175F2FBA9FA19" authorityName="Reitter" authorityYear="1880" box="[951,1037,1018,1043]" class="Insecta" family="Staphylinidae" genus="Byraxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FC2175F2FBA9FA19" box="[951,1037,1018,1043]" italics="true" pageId="8" pageNumber="9">Byraxis</emphasis>
</taxonomicName>
. During work of the nominate subgenus in this paper, five species are revealed not members of
<taxonomicName id="8FEA4D01D84AFFA7FCF47216FC18FA3D" authorityName="King. A" authorityYear="1866" box="[866,956,1054,1079]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FCF47216FC18FA3D" box="[866,956,1054,1079]" italics="true" pageId="8" pageNumber="9">Eupines</emphasis>
</taxonomicName>
, and one new species is described. Up to date, the subgenus
<taxonomicName id="8FEA4D01D84AFFA7FEFB724AFE63FA51" authorityName="King. A" authorityYear="1866" box="[365,455,1090,1115]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FEFB724AFE63FA51" box="[365,455,1090,1115]" italics="true" pageId="8" pageNumber="9">Eupines</emphasis>
</taxonomicName>
comprises only one species in
<collectingCountry id="30FD7612D84AFFA7FCB37249FC1AFA51" box="[805,958,1089,1115]" name="New Zealand" pageId="8" pageNumber="9">New Zealand</collectingCountry>
, with all remaining species placed into the diverse subgenus
<taxonomicName id="8FEA4D01D84AFFA7FECB726EFE17FA75" authorityName="Reitter" authorityYear="1880" box="[349,435,1126,1151]" class="Insecta" family="Staphylinidae" genus="Byraxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FECB726EFE17FA75" box="[349,435,1126,1151]" italics="true" pageId="8" pageNumber="9">Byraxis</emphasis>
</taxonomicName>
. In
<collectingCountry id="30FD7612D84AFFA7FE4A726DFDE7FA75" box="[476,579,1125,1151]" name="Australia" pageId="8" pageNumber="9">Australia</collectingCountry>
it is the reverse, with the subgenus
<taxonomicName id="8FEA4D01D84AFFA7FC5B726EFB83FA75" authorityName="King. A" authorityYear="1866" box="[973,1063,1126,1151]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="7A9EEA90D84AFFA7FC5B726EFB83FA75" box="[973,1063,1126,1151]" italics="true" pageId="8" pageNumber="9">Eupines</emphasis>
</taxonomicName>
being more diverse, possibly due to incorrect subgeneric placements or insufficient study (
<bibRefCitation id="2C7B4B73D84AFFA7FC857281FC1BFAA9" author="Chandler, D. S." box="[787,959,1161,1187]" pageId="8" pageNumber="9" refId="ref58398" refString="Chandler, D. S. (2001) Biology, morphology, and systematics of the ant-like litter beetle genera of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology International. Entomology International, Associated Publishers, Gainesville, Florida, VIII + 560 pp." type="book" year="2001">Chandler 2001</bibRefCitation>
).
</paragraph>
<paragraph id="48553682D84AFFA7FF5172A5FBF2FB05" blockId="8.[151,1437,153,1295]" pageId="8" pageNumber="9">
Three species are excluded from this revision (
<taxonomicName id="8FEA4D01D84AFFA7FD7572A6FC70FAC2" authority="(Sharp)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[739,980,1197,1224]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="grata" subGenus="Eupines">
<emphasis id="7A9EEA90D84AFFA7FD7572A6FD5FFACD" box="[739,763,1198,1223]" italics="true" pageId="8" pageNumber="9">E.</emphasis>
(
<emphasis id="7A9EEA90D84AFFA7FC9B72A6FC82FACD" box="[781,806,1198,1223]" italics="true" pageId="8" pageNumber="9">E.</emphasis>
)
<emphasis id="7A9EEA90D84AFFA7FCAE72A6FCD1FACD" box="[824,885,1198,1223]" italics="true" pageId="8" pageNumber="9">grata</emphasis>
(Sharp)
</taxonomicName>
,
<taxonomicName id="8FEA4D01D84AFFA7FC7472A6FB08FACD" authority="Broun" authorityName="Broun" authorityYear="1921" box="[994,1196,1197,1223]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="clemens">
<emphasis id="7A9EEA90D84AFFA7FC7472A6FBF9FACD" box="[994,1117,1197,1223]" italics="true" pageId="8" pageNumber="9">E. clemens</emphasis>
Broun
</taxonomicName>
and
<taxonomicName id="8FEA4D01D84AFFA7FB7172A6FF58FAE1" authority="(Broun))" authorityName="" baseAuthorityName="Broun" baseAuthorityYear="1893" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sternalis" subGenus="Eupines">
<emphasis id="7A9EEA90D84AFFA7FB7172A6FB5BFACD" box="[1255,1279,1198,1223]" italics="true" pageId="8" pageNumber="9">E.</emphasis>
(
<emphasis id="7A9EEA90D84AFFA7FA8772A6FA8EFACD" box="[1297,1322,1198,1223]" italics="true" pageId="8" pageNumber="9">E.</emphasis>
)
<emphasis id="7A9EEA90D84AFFA7FAAA72A5FA38FACD" box="[1340,1436,1197,1223]" italics="true" pageId="8" pageNumber="9">sternalis</emphasis>
(Broun))
</taxonomicName>
. These are being described in a separate manuscript as a new genus due to the presence of the shallow antebasal pronotal sulcus, short discal elytral striae, and different genitalic characters.
</paragraph>
</subSubSection>
</treatment>
</document>