treatments-xml/data/AF/FA/2F/AFFA2F3D7128607210CEFA2DA6835501.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.412.7282" ID-GBIF-Dataset="d3df7824-b26c-4825-b90d-161c868ab486" ID-PMC="PMC4042694" ID-Pensoft-Pub="1313-2970-412-9" ID-PubMed="24899860" ID-ZBK="442595E8D2014E59AC9F5FD0AD4580E1" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-412-9" ModsDocOrigin="ZooKeys 412" ModsDocTitle="Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant &amp; Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae)" checkinTime="1451245848117" checkinUser="pensoft" docAuthor="Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Bourdon, Caroline, Hammond, H. E. James, Pohl, Greg R. &amp; Godin, Benoit" docDate="2014" docId="AFFA2F3D7128607210CEFA2DA6835501" docLanguage="en" docName="ZooKeys 412: 9-40" docOrigin="ZooKeys 412" docSource="http://dx.doi.org/10.3897/zookeys.412.7282" docTitle="Gnathusa Fenyes" docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="FFC6FFDF7A63FFE1BF16424FFF8BE33F" masterDocTitle="Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant &amp; Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae)" masterLastPageNumber="40" masterPageNumber="9" pageNumber="12" updateTime="1668158569973" updateUser="ExternalLinkService">
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<mods:title>Review of Canadian species of the genera Gnathusa Fenyes, Mniusa Mulsant &amp; Rey and Ocyusa Kraatz (Coleoptera, Staphylinidae, Aleocharinae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Klimaszewski, Jan</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Webster, Reginald P.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Langor, David W.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bourdon, Caroline</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hammond, H. E. James</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pohl, Greg R.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Godin, Benoit</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>412</mods:number>
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<mods:start>9</mods:start>
<mods:end>40</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.412.7282</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.412.7282</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-412-9</mods:identifier>
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<treatment ID-GBIF-Taxon="152053255" LSID="urn:lsid:plazi:treatment:AFFA2F3D7128607210CEFA2DA6835501" httpUri="http://treatment.plazi.org/id/AFFA2F3D7128607210CEFA2DA6835501" lastPageId="4" lastPageNumber="12" pageId="3" pageNumber="12">
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<paragraph pageId="3" pageNumber="12">
<taxonomicName LSID="http://species-id.net/wiki/Gnathusa" authority="Fenyes" class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa" order="Coleoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">Gnathusa Fenyes</taxonomicName>
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<paragraph pageId="3" pageNumber="12">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa" order="Coleoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">Gnathusa</taxonomicName>
<bibRefCitation author="Fenyes, A" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="24" pageNumber="33" pagination="197 - 199" title="Two new species of Aleocharinae from California." volume="11" year="1909">Fenyes 1909</bibRefCitation>
: 197. Type species: 
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa eva" order="Coleoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="eva">Gnathusa eva</taxonomicName>
Fenyes.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="12" type="diagnosis">
<paragraph pageId="3" pageNumber="12">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="12">
Body dark brown to black, compact and robust, subparallel (Figs 1a, 2a, b, 3a, 4a), length 3.5-4.3 mm; integument with distinct meshed microsculpture; head large with mandibles extremely long and sickle-shaped, apices very slender, each crossing the other in resting position, each mandible bearing long spine or tooth (Figs 1h, i, 2i, j, 3h, i, 4h, i); infraorbital carina strong and complete; ligula more or less deeply bilobed (Figs 1l, 2m, 3l, 4l); last article of maxillary palpus needle-shaped (Figs 1k, 2l, 3k, 4k); frontal suture of head absent; anterior margin of mesosternum with short V-shaped basal carina, remaining mesosternum uncarinated; mesosternal process long, triangular basally and then narrowly produced and extending to about 
<normalizedToken originalValue="¾">3/4</normalizedToken>
length of mesosternal cavities, metasternal process triangular in shape and short, isthmus short; median lobe of aedeagus with large crista apicalis, internal sac structures variable in shape (Figs 1b, 2c, 3b, 4b), paramere with narrowly elongate apical lobe bearing four macrosetae; spermatheca pipe-shaped with spherical or tubular capsule and elongate narrow stem (Figs 1e, 2d, 3e, 4e).
</paragraph>
<caption pageId="3" pageNumber="12">
<paragraph pageId="3" pageNumber="12">
Figure 1. 
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa eva" order="Coleoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="eva">Gnathusa eva</taxonomicName>
Fenyes: 1a habitus 1b median lobe of aedeagus in lateral view 1c male tergite VIII 1d male sternite VIII 1e spermatheca in lateral view 1f female tergite VIII 1g female sternite VIII 1h left mandible 1i right mandible 1j labrum 1k maxilla 1l menthum, labial palpi and ligula. Habitus scale bar = 1.0 mm; other scale bars = 0.2 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="12">
<paragraph pageId="3" pageNumber="12">
Figure 2. 
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa tenuicornis" order="Coleoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="tenuicornis">Gnathusa tenuicornis</taxonomicName>
Fenyes: 2a habitus, based on female from New Brunswick 2b habitus based on male from Alberta 2c median lobe of aedeagus in lateral view 2d spermatheca in lateral view 2e male tergite VIII 2f male sternite VIII 2g female tergite VIII 2h female sternite VIII 2i left mandible 2j right mandible 2k labrum 2l maxilla 2m menthum, labial palpi and ligula. Habitus scale bar = 1.0 mm; other scale bars = 0.2 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="12">
<paragraph pageId="3" pageNumber="12">
Figure 3. 
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa caribou" order="Coleoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="caribou">Gnathusa caribou</taxonomicName>
Lohse: 3a habitus 3b median lobe of aedeagus in lateral view 3c male tergite VIII 3d male sternite VIII 3e spermatheca in lateral view 3f female tergite VIII 3g female sternite VIII 3h left mandible 3i right mandible 3j labrum 3k maxilla 3l menthum, labial palpi and ligula. Habitus scale bar = 1.0 mm; other scale bars = 0.2 mm.
</paragraph>
</caption>
<caption pageId="3" pageNumber="12">
<paragraph pageId="3" pageNumber="12">
Figure 4. 
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa alfacaribou" order="Coleoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="alfacaribou">Gnathusa alfacaribou</taxonomicName>
Klimaszewski and Langor: 4a habitus 4b median lobe of aedeagus in lateral view, partially reconstructed below broken line in Fig. 4b, and above broken line in Fig. 4c, 4d; based on the holotype 4c male tergite VIII partially reconstructed above broken line, based on the holotype 4d male sternite VIII partially reconstructed above broken line, based on the holotype 4e spermatheca in lateral view 4f female tergite VIII 4g female sternite VIII 4h left mandible 4i right mandible 4j labrum 4k maxilla 4l menthum, labial palpi and ligula. Habitus scale bar = 1.0 mm; other scale bars = 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="13" pageId="3" pageNumber="12" type="key to canadian species of gnathusa">
<paragraph pageId="3" pageNumber="12">
Key to Canadian species of 
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa" order="Coleoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="genus">Gnathusa</taxonomicName>
</paragraph>
<paragraph pageId="3" pageNumber="12">New provincial and territorial records are indicated in boldface font.</paragraph>
<paragraph lastPageId="4" lastPageNumber="13" pageId="3" pageNumber="12">
<table lastPageId="4" lastPageNumber="13" pageId="3" pageNumber="12">
<tr pageId="3" pageNumber="12">
<td colspan="1" pageId="3" pageNumber="12" rowspan="1">Fig. 1jFig. 1aFig. 1b, e</td>
<td colspan="1" pageId="3" pageNumber="12" rowspan="1">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa eva" order="Coleoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="eva">Gnathusa eva</taxonomicName>
Fenyes
</td>
</tr>
<tr pageId="3" pageNumber="12">
<td colspan="1" pageId="3" pageNumber="12" rowspan="1">Figs 2k3j4jFigs 2a, b3a4aFigs 2c, d3b, e4b, e</td>
<td colspan="1" pageId="3" pageNumber="12" rowspan="1">2</td>
</tr>
<tr pageId="3" pageNumber="12">
<td colspan="1" pageId="3" pageNumber="12" rowspan="1">Fig. 2a, bFigs 2c, d</td>
<td colspan="1" pageId="3" pageNumber="12" rowspan="1">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa tenuicornis" order="Coleoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="tenuicornis">Gnathusa tenuicornis</taxonomicName>
FenyesABORNB
</td>
</tr>
<tr pageId="3" pageNumber="12">
<td colspan="1" pageId="3" pageNumber="12" rowspan="1">Figs 3a4aFigs 3b, e4b, e</td>
<td colspan="1" pageId="3" pageNumber="12" rowspan="1">3</td>
</tr>
<tr pageId="3" pageNumber="12">
<td colspan="1" pageId="3" pageNumber="12" rowspan="1">Fig. 4aFig. 4b, e</td>
<td colspan="1" pageId="3" pageNumber="12" rowspan="1">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa alfacaribou" order="Coleoptera" pageId="3" pageNumber="12" phylum="Arthropoda" rank="species" species="alfacaribou">Gnathusa alfacaribou</taxonomicName>
Klimaszewski &amp; Langor
</td>
</tr>
<tr pageId="4" pageNumber="13">
<td colspan="1" pageId="4" pageNumber="13" rowspan="1">
<pageBreakToken pageId="4" pageNumber="13" start="start">Fig</pageBreakToken>
. 3aFigs 3b, e 
<bibRefCitation author="Lohse, GA" journalOrPublisher="The Coleopterists Bulletin" pageId="24" pageNumber="33" pagination="121 - 202" title="Revision of Arctic Aleocharinae of North America (Coleoptera: Staphylinidae)." volume="44" year="1990">Lohse et al. 1990</bibRefCitation>
</td>
<td colspan="1" pageId="4" pageNumber="13" rowspan="1">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnathusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gnathusa caribou" order="Coleoptera" pageId="4" pageNumber="13" phylum="Arthropoda" rank="species" species="caribou">Gnathusa caribou</taxonomicName>
Lohse
</td>
</tr>
</table>
</paragraph>
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</treatment>
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