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<document id="74790E98015C719AE85C173B40DD043F" ID-CLB-Dataset="7681" ID-DOI="10.11646/zootaxa.4942.1.1" ID-GBIF-Dataset="114135c4-23cb-444b-8cff-75d9f43aad6a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4596025" ID-ZooBank="CDF398FE-B0F6-40E6-967D-FB857C1565BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615447261711" checkinUser="plazi" docAuthor="Yasunaga, Tomohide, Duwal, Ram Keshari &amp; Nakatani, Yukinobu" docDate="2021" docId="A16ADB37EB110C6BFF47C1C27E67FF59" docLanguage="en" docName="zootaxa.4942.1.1.pdf" docOrigin="Zootaxa 4942 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pilophorus pullulus Poppius 1915" docType="treatment" docVersion="8" lastPageNumber="21" masterDocId="5D53A34FEB050C7EFFD0C03C7D09FF8C" masterDocTitle="Reclassification of the plant bug genus Pilophorus in Japan and key to the genera and species of Japanese Pilophorini (Hemiptera: Heteroptera: Miridae Phylinae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="21" updateTime="1698933027681" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5B3AE8F12319936A375739FF051F7371">Reclassification of the plant bug genus Pilophorus in Japan and key to the genera and species of Japanese Pilophorini (Hemiptera: Heteroptera: Miridae Phylinae)</mods:title>
</mods:titleInfo>
<mods:name id="E221C49B945A237CA06488308939E771" type="personal">
<mods:role id="78F2141568DD23F0FF971C86E7EB3232">
<mods:roleTerm id="32F310D765544EABC88A6D05A6C6FEB0">Author</mods:roleTerm>
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<mods:namePart id="8216AA1CB111F5BA9E7980385C68BBE3">Yasunaga, Tomohide</mods:namePart>
<mods:affiliation id="F82160D6A0C1A5576C4C0A83DEC0D9E5">Research Associate, Division of Invertebrate Zoology, American Museum of Natural History, Central Park at 79 th Street, New York, NY 10024, c / o Nameshi 2 - 33 - 2, Nagasaki 852 - 8061, Japan.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="B34A9259A3840F7F36224E745180891A">Duwal, Ram Keshari</mods:namePart>
<mods:affiliation id="FC3C71034553CAC4EA443CD1DF5EB1C9">Visiting Researcher, Agriculture &amp; Agri-Food Canada Environmental Health, K. W. Neatby: Bldg # 20, 960 Carling Avenue, Central Experimental Farm, Ottawa Ontario, Canada K 1 A 0 C 6.</mods:affiliation>
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</mods:role>
<mods:namePart id="780551D78C357F5A637CA461A3D15522">Nakatani, Yukinobu</mods:namePart>
<mods:affiliation id="D741488FC95B64C7A5889E3F87780B20">Division of Informatics and Inventory, Institute for Agro-Environmental Sciences, NARO, Kannondai 3 - 1 - 3, Tsukuba, Ibaraki 305 - 8604, Japan.</mods:affiliation>
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<mods:title id="731018BB6C4273EB0901FAA25EFB3688">Zootaxa</mods:title>
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<mods:date id="54C36D99E91A39F1ABF12F9DE580037F">2021</mods:date>
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<mods:number id="363C464D6A7C9FAE425F83CC85B24CD9">4942</mods:number>
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<treatment id="A16ADB37EB110C6BFF47C1C27E67FF59" ID-DOI="http://doi.org/10.5281/zenodo.4618380" ID-GBIF-Taxon="180588578" ID-Zenodo-Dep="4618380" LSID="urn:lsid:plazi:treatment:A16ADB37EB110C6BFF47C1C27E67FF59" httpUri="http://treatment.plazi.org/id/A16ADB37EB110C6BFF47C1C27E67FF59" lastPageId="21" lastPageNumber="21" pageId="20" pageNumber="21">
<subSubSection id="61D939AAEB110C6AFF47C1C27F94FD95" box="[151,669,510,537]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="297C6A21EB110C6AFF47C1C27F94FD95" blockId="20.[151,669,510,575]" box="[151,669,510,537]" pageId="20" pageNumber="21">
<heading id="7234DD4DEB110C6AFF47C1C27F94FD95" bold="true" box="[151,669,510,537]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="1BB7B633EB110C6AFF47C1C27F94FD95" bold="true" box="[151,669,510,537]" pageId="20" pageNumber="21">
<taxonomicName id="EEC311A2EB110C6AFF47C1C27F37FD94" ID-CoL="6VK4K" authority="Poppius, 1915" authorityName="Poppius" authorityYear="1915" box="[151,574,510,537]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pullulus">
<emphasis id="1BB7B633EB110C6AFF47C1C27C8EFD95" bold="true" box="[151,391,510,537]" italics="true" pageId="20" pageNumber="21">Pilophorus pullulus</emphasis>
Poppius, 1915
</taxonomicName>
sp. rev.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61D939AAEB110C6AFF47C21F7CB9FDB1" box="[151,432,547,575]" pageId="20" pageNumber="21" type="description">
<paragraph id="297C6A21EB110C6AFF47C21F7CB9FDB1" blockId="20.[151,669,510,575]" box="[151,432,547,575]" pageId="20" pageNumber="21">
(
<figureCitation id="B1F876A4EB110C6AFF4FC21F7DF1FDB3" box="[159,248,547,575]" captionStart="FIGURE 1" captionStartId="7.[151,250,1918,1942]" captionTargetBox="[174,1436,428,1895]" captionTargetId="figure-146@7.[151,1436,417,1895]" captionTargetPageId="7" captionText="FIGURE 1. Habitus images of Pilophorus spp. A. P. choii ♂. B. P. maeharai, holotype ♂. C. P. niger ♀. D–E. P. pullulus, holotype ♀ (Taiwan). F–G. P. pullulus ♂ (Amami-Oshima Is., Japan)." figureDoi="http://doi.org/10.5281/zenodo.4596027" httpUri="https://zenodo.org/record/4596027/files/figure.png" pageId="20" pageNumber="21">Figs 1D</figureCitation>
̅G, 9P̅S, 14O)
</paragraph>
</subSubSection>
<subSubSection id="61D939AAEB110C6AFF47C25B7C11FD30" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="297C6A21EB110C6AFF47C25B7C11FD30" blockId="20.[151,1436,615,700]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="09D34D0FEB110C6AFF47C25B7895FD12" pageId="20" pageNumber="21">
<taxonomicName id="EEC311A2EB110C6AFF47C25B7F92FDF3" ID-CoL="6VK4K" authority="Poppius, 1914: 238" authorityName="Poppius" authorityPageNumber="238" authorityYear="1914" box="[151,667,615,639]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pullulus">
<treatmentCitation id="A8624C30EB110C6AFF47C25B7F25FDF3" author="Poppius, B." box="[151,556,615,639]" page="238" pageId="20" pageNumber="21" year="1914">
<emphasis id="1BB7B633EB110C6AFF47C25B7C69FDF3" box="[151,352,615,639]" italics="true" pageId="20" pageNumber="21">Pilophorus pullulus</emphasis>
<bibRefCitation id="4D5217D0EB110C6AFEB6C25B7F25FDF3" author="Poppius, B." box="[358,556,615,639]" pageId="20" pageNumber="21" pagination="237 - 254" refId="ref25114" refString="Poppius, B. (1914) Ubersicht der Pilophorus - Arten nebst Beschreibung verwandter Gattungen (Hem. Het.). Annales de la Societe Entomologique de Belgique, 58, 237 - 254." type="journal article" year="1914">Poppius, 1914: 238</bibRefCitation>
</treatmentCitation>
, 1915a: 64
</taxonomicName>
(n. sp.) [synonymized with
<taxonomicName id="EEC311A2EB110C6AFC69C25479B0FDF3" ID-CoL="6VJT2" authority="(Distant, 1909)" baseAuthorityName="Distant" baseAuthorityYear="1909" box="[953,1209,615,639]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="typicus">
<emphasis id="1BB7B633EB110C6AFC69C2547913FDF3" box="[953,1050,616,639]" italics="true" pageId="20" pageNumber="21">P. typicus</emphasis>
(Distant, 1909)
</taxonomicName>
by
<bibRefCitation id="4D5217D0EB110C6AFB0FC25B7864FDF3" author="Schuh, R. T." box="[1247,1389,615,639]" pageId="20" pageNumber="21" pagination="1 - 476" refId="ref25151" refString="Schuh, R. T. (1984) Revision of the Phylinae (Hemiptera, Miridae) of the Indo-Pacific. Bulletin of the American Museum of Natural History, 177, 1 - 476." type="journal article" year="1984">Schuh (1984)</bibRefCitation>
].
<taxonomicName id="EEC311A2EB110C6AFF47C2BA7F01FD12" ID-CoL="6VJT2" authority="(Distant, 1909)" baseAuthorityName="Distant" baseAuthorityYear="1909" box="[151,520,646,670]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="typicus">
<emphasis id="1BB7B633EB110C6AFF47C2BA7C52FD11" box="[151,347,646,670]" italics="true" pageId="20" pageNumber="21">Pilophorus typicus</emphasis>
(Distant, 1909)
</taxonomicName>
:
<treatmentCitation id="A8624C30EB110C6AFDC6C2BA7FCDFD12" author="Schuh, R. T." box="[534,708,646,670]" page="71" pageId="20" pageNumber="21" year="1984">
<bibRefCitation id="4D5217D0EB110C6AFDC6C2BA7FCDFD12" author="Schuh, R. T." box="[534,708,646,670]" pageId="20" pageNumber="21" pagination="1 - 476" refId="ref25151" refString="Schuh, R. T. (1984) Revision of the Phylinae (Hemiptera, Miridae) of the Indo-Pacific. Bulletin of the American Museum of Natural History, 177, 1 - 476." type="journal article" year="1984">Schuh, 1984: 71</bibRefCitation>
</treatmentCitation>
(n. syn.);
<bibRefCitation id="4D5217D0EB110C6AFCEAC2BA7EB7FD12" author="Schuh, R. T." box="[826,958,646,670]" pageId="20" pageNumber="21" refId="ref25329" refString="Schuh, R. T. (2002 - 2013) On-line Systematic Catalog of Plant Bugs (Insecta: Heteroptera: Miridae). Available from: http: // research. amnh. org / pbi / catalog / (accessed 5 December 2020)" type="url" year="2002">Schuh, 2002</bibRefCitation>
–2013, online catalog;
<bibRefCitation id="4D5217D0EB110C6AFB62C2BA7847FD12" author="Aukema, B." box="[1202,1358,646,670]" pageId="20" pageNumber="21" refId="ref24010" refString="Aukema, B. (2018) Catalogue of the Palaearctic Heteroptera (searchable database). Available from: https: // catpalhet. linnaeus. naturalis. nl / (accessed 5 December 2020)" type="url" year="2018">Aukema, 2018</bibRefCitation>
, online
</treatmentCitationGroup>
catalog.
</paragraph>
</subSubSection>
<subSubSection id="61D939AAEB110C6AFF47C2D67C64FBE1" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="297C6A21EB110C6AFF47C2D67E84FC1A" blockId="20.[151,1437,746,2034]" pageId="20" pageNumber="21">
<emphasis id="1BB7B633EB110C6AFF47C2D67C88FC88" bold="true" box="[151,385,746,772]" pageId="20" pageNumber="21">Material examined.</emphasis>
<materialsCitation id="99AB607CEB110C6AFE59C2D7795BFC89" ID-GBIF-Occurrence="3053984320" box="[393,1106,746,773]" collectionCode="SDEI" collectorName="Aug, H" country="Taiwan" location="City" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" stateProvince="Tainan" typeStatus="holotype">
<typeStatus id="F678D483EB110C6AFE59C2D77CF8FC89" box="[393,497,747,773]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
(
<specimenCount id="3FC5A1A8EB110C6AFDD5C2D77F11FC89" box="[517,536,747,773]" pageId="20" pageNumber="21" type="female">♀</specimenCount>
).
<emphasis id="1BB7B633EB110C6AFDE0C2D67FA3FC88" bold="true" box="[560,682,746,772]" pageId="20" pageNumber="21">
<collectingCountry id="51D42AB1EB110C6AFDE0C2D67FAFFC88" box="[560,678,746,772]" name="Taiwan" pageId="20" pageNumber="21">TAIWAN</collectingCountry>
:
</emphasis>
<collectingRegion id="EB07A4C3EB110C6AFD62C2D77FF7FC89" box="[690,766,747,773]" country="Taiwan" name="Tainan" pageId="20" pageNumber="21">Tainan</collectingRegion>
<location id="2C1C3CFAEB110C6AFCD5C2D77E3FFC89" LSID="urn:lsid:plazi:treatment:A16ADB37EB110C6BFF47C1C27E67FF59:2C1C3CFAEB110C6AFCD5C2D77E3FFC89" box="[773,822,747,773]" country="Taiwan" name="City" pageId="20" pageNumber="21" stateProvince="Tainan">City</location>
, 7
<collectorName id="84360FF7EB110C6AFC87C2D77EA4FC89" box="[855,941,747,773]" pageId="20" pageNumber="21">Aug, H</collectorName>
. Sauter (
<collectionCode id="4FD2F2E4EB110C6AFBC0C2D7795BFC89" box="[1040,1106,747,773]" pageId="20" pageNumber="21">SDEI</collectionCode>
</materialsCitation>
, images available on website (http://twinsecttype.nmns.edu.tw/specimen/?id=NMNS-SDEI-00202).
<materialsCitation id="99AB607CEB110C6AFC69C3327E80FC1A" ID-GBIF-Occurrence="3053984310" collectingDate="1954-07-26" collectingDateMax="1960-04-04" collectingDateMin="1954-07-26" collectionCode="KUEC, AMNH" collectorName="S. Miyamoto" country="Japan" county="Ryukyus" latitude="28.2843" location="Santaro-toge" longLatPrecision="6" longitude="129.4215" municipality="Amami-Oshima Island" pageId="20" pageNumber="21" specimenCode="PBI 00380672" specimenCount="1">
<emphasis id="1BB7B633EB110C6AFC69C3327815FCA4" bold="true" box="[953,1308,782,808]" pageId="20" pageNumber="21">
Additional material.
<collectingCountry id="51D42AB1EB110C6AFB12C3327815FCA4" box="[1218,1308,782,808]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
</emphasis>
:
<collectingCounty id="C01D12ADEB110C6AFAE3C3337891FCA5" box="[1331,1432,783,809]" pageId="20" pageNumber="21">Ryukyus</collectingCounty>
,
<collectingMunicipality id="C918F05BEB110C6AFF47C30F7C96FCC1" box="[151,415,819,845]" pageId="20" pageNumber="21">Amami-Oshima Island</collectingMunicipality>
,
<location id="2C1C3CFAEB110C6AFE60C30F7F36FCC1" LSID="urn:lsid:plazi:treatment:A16ADB37EB110C6BFF47C1C27E67FF59:2C1C3CFAEB110C6AFE60C30F7F36FCC1" box="[432,575,819,845]" country="Japan" county="Ryukyus" latitude="28.2843" longLatPrecision="6" longitude="129.4215" municipality="Amami-Oshima Island" name="Santaro-toge" pageId="20" pageNumber="21">Santaro-toge</location>
[= current Sumiyo Town, Kamiya,
<geoCoordinate id="4CF70CE6EB110C6AFC20C30F7947FCC1" box="[1008,1102,819,845]" degrees="28.2843" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="5" value="28.2843">28.2843</geoCoordinate>
,
<geoCoordinate id="4CF70CE6EB110C6AFB8EC30E79C4FCC1" box="[1118,1229,818,845]" degrees="129.4215" direction="east" orientation="longitude" pageId="20" pageNumber="21" precision="5" value="129.4215">129.4215</geoCoordinate>
],
<date id="5D7D4CE1EB110C6AFB34C30F787CFCC1" box="[1252,1397,818,845]" pageId="20" pageNumber="21" value="1954-07-26">
<collectingDate id="4D39B509EB110C6AFB34C30F787CFCC1" box="[1252,1397,818,845]" pageId="20" pageNumber="21" value="1954-07-26">26 Jul 1954</collectingDate>
</date>
,
<collectorName id="84360FF7EB110C6AFA56C30F7C05FCFD" pageId="20" pageNumber="21">S. Miyamoto</collectorName>
&amp;
<collectorName id="84360FF7EB110C6AFEE9C36B7CDDFCFD" box="[313,468,855,881]" pageId="20" pageNumber="21">Y. Hirashima</collectorName>
, 1 ♂ (
<collectionCode id="4FD2F2E4EB110C6AFDF6C36B7F7FFCFD" box="[550,630,855,881]" country="Japan" name="Kyushu University Entomology Collection" pageId="20" pageNumber="21">KUEC</collectionCode>
) (
<collectionCode id="4FD2F2E4EB110C6AFD41C36B7FE1FCFD" box="[657,744,855,881]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="20" pageNumber="21">AMNH</collectionCode>
_
<specimenCode id="7965C25AEB110C6AFD26C36B7EAAFCFE" box="[758,931,855,882]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="20" pageNumber="21">PBI 00380672</specimenCode>
);
<location id="2C1C3CFAEB110C6AFC69C36B79C8FCFD" LSID="urn:lsid:plazi:treatment:A16ADB37EB110C6BFF47C1C27E67FF59:2C1C3CFAEB110C6AFC69C36B79C8FCFD" box="[953,1217,855,881]" country="Japan" county="Ryukyus" latitude="28.2843" longLatPrecision="6" longitude="129.4215" municipality="Amami-Oshima Island" name="Amami-Oshima Island" pageId="20" pageNumber="21">Amami-Oshima Island</location>
,
<collectorName id="84360FF7EB110C6AFB01C36B7891FCFD" box="[1233,1432,855,881]" pageId="20" pageNumber="21">Shinmura-Yuwan</collectorName>
, 28.24, 129.33,
<date id="5D7D4CE1EB110C6AFEEFC3477CC9FC19" box="[319,448,891,917]" pageId="20" pageNumber="21" value="1960-04-04">
<collectingDate id="4D39B509EB110C6AFEEFC3477CC9FC19" box="[319,448,891,917]" pageId="20" pageNumber="21" value="1960-04-04">4 Apr 1960</collectingDate>
</date>
,
<collectorName id="84360FF7EB110C6AFE1BC3477F68FC19" box="[459,609,891,917]" pageId="20" pageNumber="21">S. Miyamoto</collectorName>
, 1 ♂ (
<collectionCode id="4FD2F2E4EB110C6AFD74C3477FFDFC19" box="[676,756,891,917]" country="Japan" name="Kyushu University Entomology Collection" pageId="20" pageNumber="21">KUEC</collectionCode>
) (00380673)
</materialsCitation>
.
</paragraph>
<paragraph id="297C6A21EB110C6AFF17C3A27C64FBE1" blockId="20.[151,1437,746,2034]" pageId="20" pageNumber="21">
<emphasis id="1BB7B633EB110C6AFF17C3A27C57FC34" bold="true" box="[199,350,926,952]" pageId="20" pageNumber="21">Rediagnosis.</emphasis>
Closely allied to
<taxonomicName id="EEC311A2EB110C6AFDF1C3A37F81FC35" authority="Yasunaga &amp; Duwal &amp; Nakatani, 2020" authorityName="Yasunaga &amp; Duwal &amp; Nakatani" authorityYear="2020" box="[545,648,927,953]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hyotan" status="sp. nov.">
<emphasis id="1BB7B633EB110C6AFDF1C3A37F81FC35" box="[545,648,927,953]" italics="true" pageId="20" pageNumber="21">P. hyotan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00840B48EB110C6AFD5FC3A37FC3FC35" box="[655,714,927,953]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="EEC311A2EB110C6AFD2FC3A37E60FC34" baseAuthorityName="Distant" baseAuthorityYear="1909" box="[767,873,927,952]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="typicus">
<emphasis id="1BB7B633EB110C6AFD2FC3A37E60FC34" box="[767,873,927,952]" italics="true" pageId="20" pageNumber="21">P. typicus</emphasis>
</taxonomicName>
, to which this species is superficially very similar, but
<taxonomicName id="EEC311A2EB110C6AFF14C3FF7C32FC51" authorityName="Poppius" authorityYear="1914" box="[196,315,963,989]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pullulus">
<emphasis id="1BB7B633EB110C6AFF14C3FF7C32FC51" box="[196,315,963,989]" italics="true" pageId="20" pageNumber="21">P. pullulus</emphasis>
</taxonomicName>
can be distinguished by the following characters: Smaller size (e.g., total body length ±
<quantity id="EE3BC7C4EB110C6AFAE4C3FE7899FC51" box="[1332,1424,962,989]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="mm" value="2.5">2.5 mm</quantity>
); brownish dorsum; dispersed patches of silvery setae on posterior corium; sparsely distributed claval golden-brown setae; less splayed out left paramere with weakly produced lateral margin of sensory lobe; and rather thickened median process of endosoma. The external structure of a female (
<typeStatus id="F678D483EB110C6AFCB8C4137EC4FBC6" box="[872,973,1071,1098]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
) of
<taxonomicName id="EEC311A2EB110C6AFC28C4137964FBC5" authorityName="Poppius" authorityYear="1914" box="[1016,1133,1071,1097]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pullulus">
<emphasis id="1BB7B633EB110C6AFC28C4137964FBC5" box="[1016,1133,1071,1097]" italics="true" pageId="20" pageNumber="21">P. pullulus</emphasis>
</taxonomicName>
(
<figureCitation id="B1F876A4EB110C6AFBABC41379B1FBC5" box="[1147,1208,1071,1097]" captionStart="FIGURE 1" captionStartId="7.[151,250,1918,1942]" captionTargetBox="[174,1436,428,1895]" captionTargetId="figure-146@7.[151,1436,417,1895]" captionTargetPageId="7" captionText="FIGURE 1. Habitus images of Pilophorus spp. A. P. choii ♂. B. P. maeharai, holotype ♂. C. P. niger ♀. D–E. P. pullulus, holotype ♀ (Taiwan). F–G. P. pullulus ♂ (Amami-Oshima Is., Japan)." figureDoi="http://doi.org/10.5281/zenodo.4596027" httpUri="https://zenodo.org/record/4596027/files/figure.png" pageId="20" pageNumber="21">Fig. 1</figureCitation>
D–E) was described by Poppius (1915).
</paragraph>
</subSubSection>
<subSubSection id="61D939AAEB110C6AFF17C44A7FA5F895" pageId="20" pageNumber="21" type="description">
<paragraph id="297C6A21EB110C6AFF17C44A7FDEF95D" blockId="20.[151,1437,746,2034]" pageId="20" pageNumber="21">
<emphasis id="1BB7B633EB110C6AFF17C44A7C50FB1C" bold="true" box="[199,345,1142,1168]" pageId="20" pageNumber="21">Description.</emphasis>
<emphasis id="1BB7B633EB110C6AFE8EC44B7C91FB1D" box="[350,408,1143,1169]" italics="true" pageId="20" pageNumber="21">Male</emphasis>
: Macropterous; body generally castaneous to dark reddish brown, antlike (HCR 0.77); dorsal surface shining, with sparsely distributed, simple, reclining setae and scattered patches of scale-like setae (
<figureCitation id="B1F876A4EB110C6AFAEAC4A77887FB39" box="[1338,1422,1179,1205]" captionStart="FIGURE 1" captionStartId="7.[151,250,1918,1942]" captionTargetBox="[174,1436,428,1895]" captionTargetId="figure-146@7.[151,1436,417,1895]" captionTargetPageId="7" captionText="FIGURE 1. Habitus images of Pilophorus spp. A. P. choii ♂. B. P. maeharai, holotype ♂. C. P. niger ♀. D–E. P. pullulus, holotype ♀ (Taiwan). F–G. P. pullulus ♂ (Amami-Oshima Is., Japan)." figureDoi="http://doi.org/10.5281/zenodo.4596027" httpUri="https://zenodo.org/record/4596027/files/figure.png" pageId="20" pageNumber="21">Fig. 1F</figureCitation>
). Head shiny castaneous, slightly wider than high, narrowly margined along basal vertex; eye small; vertex wider than an eye in dorsal view; head below eyes tinged with red. Antenna dark brown, generally short; segment I, basal 2/3 of II and almost whole IV creamy brown; segment II somewhat clavate, with apical part about as thick as profemur; segments III and IV filiform. Labium shiny reddish brown, exceeding base of metacoxa; apical 3/4 of segment III and basal half of segment IV creamy brown. Pronotum shiny fuscous, polished, with uniformly distributed, simple, semierect setae (
<figureCitation id="B1F876A4EB110C6AFE7CC54F7F1CFA01" box="[428,533,1395,1421]" captionStart="FIGURE 14" captionStartId="28.[151,250,1888,1912]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-27@28.[151,1436,181,1864]" captionTargetPageId="28" captionText="FIGURE 14. Scanning electron micrographs for Japanese Pilophorus spp., males (except for E, L, P: ♀). A–D. P. choii. E–K. P. erraticus. L–N. P. pseudoperplexus. O–Q. P. miyamotoi. A, E. Left-dorsolateral habitus. B, I, N. Apex of pygophore, left lateral view. C–D, J–K, Q. Endosoma (vesica).A, E. Dorsal habitus. F. Thoracic pleura, left lateral view. H, L, M, O, P. Lateral habitus. G. Pygophore, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4596067" httpUri="https://zenodo.org/record/4596067/files/figure.png" pageId="20" pageNumber="21">Fig. 14O</figureCitation>
); scutellum shiny fuscous, with roundly clustered scale-like setae at each angle; pleura dark brown, with clustered scale-like setae on mesepimeron (
<figureCitation id="B1F876A4EB110C6AFC57C5AB7ED6FA3D" box="[903,991,1431,1457]" captionStart="FIGURE 1" captionStartId="7.[151,250,1918,1942]" captionTargetBox="[174,1436,428,1895]" captionTargetId="figure-146@7.[151,1436,417,1895]" captionTargetPageId="7" captionText="FIGURE 1. Habitus images of Pilophorus spp. A. P. choii ♂. B. P. maeharai, holotype ♂. C. P. niger ♀. D–E. P. pullulus, holotype ♀ (Taiwan). F–G. P. pullulus ♂ (Amami-Oshima Is., Japan)." figureDoi="http://doi.org/10.5281/zenodo.4596027" httpUri="https://zenodo.org/record/4596027/files/figure.png" pageId="20" pageNumber="21">Fig. 1G</figureCitation>
). Hemelytron almost totally castaneous, somewhat matte, with sparsely distributed, simple, reclining setae, and clustered scale-like at subapical clavus and basal 1/3 of corium; median band of scale-like setae interrupted, separated into two clusters (but excluding old specimens, presumably rubbed off); membrane pale smoky brown, with a narrow, pale, semitransparent part along apical margin of cuneus. Coxae creamy yellow, except for entirely dark brown mesocoxa; profemur, and all tibiae and tarsi creamy yellow; meso- and metafemora and basal parts of meso- and metatibiae reddish or dark brown; tarsomeres III darkened. Abdomen shiny dark brown. Male genitalia (
<figureCitation id="B1F876A4EB110C6AFBC2C653795EF905" box="[1042,1111,1647,1673]" captionStart="FIGURE 9" captionStartId="21.[151,250,1881,1905]" captionTargetBox="[161,1424,249,1850]" captionTargetId="figure-47@21.[151,1436,237,1858]" captionTargetPageId="21" captionText="FIGURE 9. Male genitalia of Pilophorus spp. A–E, I–K. P. hyotan n. sp. (from Nagasaki, Japan: A–C, K; Solkhumbu, Nepal: D–E, I–J). F–H, L–N. P. typicus (Nakhon Nayok, Thailand: F–G; India: H, L–M; Nagasaki: N). O. P. miyamotoi (Kochi Pref., Japan). P–S. P. pullulus (Amami-Oshima Is.). A– B, D, F–G, O–P. Endosoma (vesica). C, E, H, S. Phallotheca. I–N, Q–R. Parameres." figureDoi="http://doi.org/10.5281/zenodo.4596053" httpUri="https://zenodo.org/record/4596053/files/figure.png" pageId="20" pageNumber="21">Fig. 9</figureCitation>
P–S): Left paramere narrow, not sprayed-out (
<figureCitation id="B1F876A4EB110C6AFE87C6AF7CBAF921" box="[343,435,1683,1709]" captionStart="FIGURE 9" captionStartId="21.[151,250,1881,1905]" captionTargetBox="[161,1424,249,1850]" captionTargetId="figure-47@21.[151,1436,237,1858]" captionTargetPageId="21" captionText="FIGURE 9. Male genitalia of Pilophorus spp. A–E, I–K. P. hyotan n. sp. (from Nagasaki, Japan: A–C, K; Solkhumbu, Nepal: D–E, I–J). F–H, L–N. P. typicus (Nakhon Nayok, Thailand: F–G; India: H, L–M; Nagasaki: N). O. P. miyamotoi (Kochi Pref., Japan). P–S. P. pullulus (Amami-Oshima Is.). A– B, D, F–G, O–P. Endosoma (vesica). C, E, H, S. Phallotheca. I–N, Q–R. Parameres." figureDoi="http://doi.org/10.5281/zenodo.4596053" httpUri="https://zenodo.org/record/4596053/files/figure.png" pageId="20" pageNumber="21">Fig. 9Q</figureCitation>
); endosoma nearly J-shaped, with simple, rather thick median process (
<figureCitation id="B1F876A4EB110C6AFB0CC6AF783BF921" box="[1244,1330,1683,1709]" captionStart="FIGURE 9" captionStartId="21.[151,250,1881,1905]" captionTargetBox="[161,1424,249,1850]" captionTargetId="figure-47@21.[151,1436,237,1858]" captionTargetPageId="21" captionText="FIGURE 9. Male genitalia of Pilophorus spp. A–E, I–K. P. hyotan n. sp. (from Nagasaki, Japan: A–C, K; Solkhumbu, Nepal: D–E, I–J). F–H, L–N. P. typicus (Nakhon Nayok, Thailand: F–G; India: H, L–M; Nagasaki: N). O. P. miyamotoi (Kochi Pref., Japan). P–S. P. pullulus (Amami-Oshima Is.). A– B, D, F–G, O–P. Endosoma (vesica). C, E, H, S. Phallotheca. I–N, Q–R. Parameres." figureDoi="http://doi.org/10.5281/zenodo.4596053" httpUri="https://zenodo.org/record/4596053/files/figure.png" pageId="20" pageNumber="21">Fig. 9P</figureCitation>
); apex of phallotheca with a small, pointed process (
<figureCitation id="B1F876A4EB110C6AFDA4C68B7FC3F95D" box="[628,714,1719,1745]" captionStart="FIGURE 9" captionStartId="21.[151,250,1881,1905]" captionTargetBox="[161,1424,249,1850]" captionTargetId="figure-47@21.[151,1436,237,1858]" captionTargetPageId="21" captionText="FIGURE 9. Male genitalia of Pilophorus spp. A–E, I–K. P. hyotan n. sp. (from Nagasaki, Japan: A–C, K; Solkhumbu, Nepal: D–E, I–J). F–H, L–N. P. typicus (Nakhon Nayok, Thailand: F–G; India: H, L–M; Nagasaki: N). O. P. miyamotoi (Kochi Pref., Japan). P–S. P. pullulus (Amami-Oshima Is.). A– B, D, F–G, O–P. Endosoma (vesica). C, E, H, S. Phallotheca. I–N, Q–R. Parameres." figureDoi="http://doi.org/10.5281/zenodo.4596053" httpUri="https://zenodo.org/record/4596053/files/figure.png" pageId="20" pageNumber="21">Fig. 9S</figureCitation>
).
</paragraph>
<paragraph id="297C6A21EB110C6AFF17C6E77FA5F895" blockId="20.[151,1437,746,2034]" pageId="20" pageNumber="21">
<emphasis id="1BB7B633EB110C6AFF17C6E77C77F979" bold="true" box="[199,382,1755,1781]" pageId="20" pageNumber="21">Measurements.</emphasis>
See
<tableCitation id="64415F9AEB110C6AFE62C6E77F0EF979" box="[434,519,1755,1781]" captionStart="TABLE 2" captionStartId="10.[151,239,440,464]" captionText="TABLE 2. Measurements for eight Japanese Pilophorus species." pageId="20" pageNumber="21">Table 2</tableCitation>
. According to the original description of the
<typeStatus id="F678D483EB110C6AFC27C6E77950F979" box="[1015,1113,1755,1781]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
female (
<figureCitation id="B1F876A4EB110C6AFB6AC6E77819F979" box="[1210,1296,1755,1781]" captionStart="FIGURE 1" captionStartId="7.[151,250,1918,1942]" captionTargetBox="[174,1436,428,1895]" captionTargetId="figure-146@7.[151,1436,417,1895]" captionTargetPageId="7" captionText="FIGURE 1. Habitus images of Pilophorus spp. A. P. choii ♂. B. P. maeharai, holotype ♂. C. P. niger ♀. D–E. P. pullulus, holotype ♀ (Taiwan). F–G. P. pullulus ♂ (Amami-Oshima Is., Japan)." figureDoi="http://doi.org/10.5281/zenodo.4596027" httpUri="https://zenodo.org/record/4596027/files/figure.png" pageId="20" pageNumber="21">Fig. 1D</figureCitation>
) by Poppius (1915), body length
<quantity id="EE3BC7C4EB110C6AFEACC6C37CDBF895" box="[380,466,1791,1817]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="20" pageNumber="21" unit="mm" value="2.2">2.2 mm</quantity>
and width
<quantity id="EE3BC7C4EB110C6AFD9FC6C37FA1F895" box="[591,680,1791,1817]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="21" unit="mm" value="0.8">0.8 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="61D939AAEB110C6AFF17C71E7CADF8B1" box="[199,420,1826,1853]" pageId="20" pageNumber="21" type="biology_ecology">
<paragraph id="297C6A21EB110C6AFF17C71E7CADF8B1" blockId="20.[151,1437,746,2034]" box="[199,420,1826,1853]" pageId="20" pageNumber="21">
<emphasis id="1BB7B633EB110C6AFF17C71E7C2EF8B0" bold="true" box="[199,295,1826,1852]" pageId="20" pageNumber="21">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="61D939AAEB110C6BFF17C77A7E67FF59" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="discussion">
<paragraph id="297C6A21EB110C6BFF17C77A7E67FF59" blockId="20.[151,1437,746,2034]" lastBlockId="21.[151,1437,151,213]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="1BB7B633EB110C6AFF17C77A7C45F8EC" bold="true" box="[199,332,1862,1888]" pageId="20" pageNumber="21">Discussion.</emphasis>
This species was described by Poppius (1915), based on a single female specimen collected in
<collectingRegion id="EB07A4C3EB110C6AFF47C7577DEAF809" box="[151,227,1899,1925]" country="Taiwan" name="Tainan" pageId="20" pageNumber="21">Tainan</collectingRegion>
City, southern
<collectingCountry id="51D42AB1EB110C6AFE40C7577CEBF809" box="[400,482,1899,1925]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
(
<figureCitation id="B1F876A4EB110C6AFE22C7577F41F80B" box="[498,584,1899,1927]" captionStart="FIGURE 1" captionStartId="7.[151,250,1918,1942]" captionTargetBox="[174,1436,428,1895]" captionTargetId="figure-146@7.[151,1436,417,1895]" captionTargetPageId="7" captionText="FIGURE 1. Habitus images of Pilophorus spp. A. P. choii ♂. B. P. maeharai, holotype ♂. C. P. niger ♀. D–E. P. pullulus, holotype ♀ (Taiwan). F–G. P. pullulus ♂ (Amami-Oshima Is., Japan)." figureDoi="http://doi.org/10.5281/zenodo.4596027" httpUri="https://zenodo.org/record/4596027/files/figure.png" pageId="20" pageNumber="21">Fig. 1D</figureCitation>
̅E).
<bibRefCitation id="4D5217D0EB110C6AFD52C7577E13F809" author="Schuh, R. T." box="[642,794,1899,1925]" pageId="20" pageNumber="21" pagination="1 - 476" refId="ref25151" refString="Schuh, R. T. (1984) Revision of the Phylinae (Hemiptera, Miridae) of the Indo-Pacific. Bulletin of the American Museum of Natural History, 177, 1 - 476." type="journal article" year="1984">Schuh (1984)</bibRefCitation>
synonymized
<taxonomicName id="EEC311A2EB110C6AFC6FC757793CF809" authorityName="Poppius" authorityYear="1914" box="[959,1077,1899,1925]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pullulus">
<emphasis id="1BB7B633EB110C6AFC6FC757793CF809" box="[959,1077,1899,1925]" italics="true" pageId="20" pageNumber="21">P. pullulus</emphasis>
</taxonomicName>
with
<taxonomicName id="EEC311A2EB110C6AFBA7C75779EBF808" baseAuthorityName="Distant" baseAuthorityYear="1909" box="[1143,1250,1899,1924]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="typicus">
<emphasis id="1BB7B633EB110C6AFBA7C75779EBF808" box="[1143,1250,1899,1924]" italics="true" pageId="20" pageNumber="21">P. typicus</emphasis>
</taxonomicName>
, as the
<typeStatus id="F678D483EB110C6AFAEAC7577895F809" box="[1338,1436,1899,1925]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
had been damaged since original description (now only left clavus remaining as in
<figureCitation id="B1F876A4EB110C6AFBB2C7B379B2F825" box="[1122,1211,1935,1961]" captionStart="FIGURE 1" captionStartId="7.[151,250,1918,1942]" captionTargetBox="[174,1436,428,1895]" captionTargetId="figure-146@7.[151,1436,417,1895]" captionTargetPageId="7" captionText="FIGURE 1. Habitus images of Pilophorus spp. A. P. choii ♂. B. P. maeharai, holotype ♂. C. P. niger ♀. D–E. P. pullulus, holotype ♀ (Taiwan). F–G. P. pullulus ♂ (Amami-Oshima Is., Japan)." figureDoi="http://doi.org/10.5281/zenodo.4596027" httpUri="https://zenodo.org/record/4596027/files/figure.png" pageId="20" pageNumber="21">Fig. 1D</figureCitation>
and coloration and vestiture pattern of entire hemelytra depending only on the original description—hemelytron brown, matte, with whitish band of scale-like setae on posterior corium and clavus). Judging from the description of Poppius and remaining body parts of the
<typeStatus id="F678D483EB100C6BFE06C0AB7F32FF3D" box="[470,571,151,177]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
, we conclude that two male specimens, collected from Amami-Oshima more than six decades ago, are in all likelihood identical to
<taxonomicName id="EEC311A2EB100C6BFD22C0877E6EFF59" authorityName="Poppius" authorityYear="1914" box="[754,871,187,213]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pullulus">
<emphasis id="1BB7B633EB100C6BFD22C0877E6EFF59" box="[754,871,187,213]" italics="true" pageId="21" pageNumber="22">P. pullulus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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