treatments-xml/data/7F/09/EC/7F09EC44FF9BA405FF3FD456FA4DAA9D.xml

<document id="6B02379285351388E1D4A00ECAA3A7B1" ID-DOI="10.5281/zenodo.211197" ID-GBIF-Dataset="14afa521-5f4c-4690-a8f3-71080067f2c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="211197" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460500196774" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Lai, Joelle C. Y." docDate="2012" docId="7F09EC44FF9BA405FF3FD456FA4DAA9D" docLanguage="en" docName="zt03393p065.pdf" docOrigin="Zootaxa 3393" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Calappa karenae Ng &amp; Lai, 2012, sp. nov." docType="treatment" docVersion="10" lastPageNumber="64" masterDocId="8330943CFF9BA402FFA8D320FFCBAE02" masterDocTitle="Calappa karenae, a new species of box crab from Guam (Crustacea: Decapoda: Brachyura: Calappidae)" masterLastPageNumber="65" masterPageNumber="57" originalUpdateDomain="Boston" originalUpdateTime="1544534532577" originalUpdateUser="gglab" pageNumber="57" updateTime="1698311602805" updateUser="plazi">
<mods:mods id="4C53D6D63960DBB176389729FF6289B9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F4F0122FB8F22B4F30D5D4993F722F0B">
<mods:title id="3E965E0E79220DE45776DBC4DA886110">Calappa karenae, a new species of box crab from Guam (Crustacea: Decapoda: Brachyura: Calappidae)</mods:title>
</mods:titleInfo>
<mods:name id="F32B46D78B10BBBCD012F5A716175188" type="personal">
<mods:role id="6E5CA1E788A37EF569D2FBC03437E962">
<mods:roleTerm id="19C9B60941051B6EB8099A7D57A7507C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3004FB06383743C3D08AC3BE57AB51A3">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="CB531A229C0EC2E3BD4F4B0C156F8A56" type="personal">
<mods:role id="4497F881E6A801546F00DC4D9B971D6A">
<mods:roleTerm id="2A575CFE11CB255B69E6C9C50CBA5B3E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="255630664A301FD4E873D2523CF90D2C">Lai, Joelle C. Y.</mods:namePart>
</mods:name>
<mods:typeOfResource id="E6D7E97A8D9ACC63EE9E74A7002B53C2">text</mods:typeOfResource>
<mods:relatedItem id="D25F08316E24241E48785056E9609AA9" type="host">
<mods:titleInfo id="3E548D7BD65DA85F803186DD02CEB8F3">
<mods:title id="8FC7B7B55F2BC44A75BB35E661A9EB1F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1D0E01B783E3D30B727AD33C2FC40725">
<mods:date id="FBADA404026AC11CCE03D75DB203FEF3">2012</mods:date>
<mods:detail id="81BE85B158F51F24D3F5E95DE2006832" type="volume">
<mods:number id="19B432FB56A3F3FA5B9B05E727B41EC7">3393</mods:number>
</mods:detail>
<mods:extent id="31A95D3C66B1A6D1C20D2553E308A201" unit="page">
<mods:start id="CAC2D525127F14825EAED14008B85B67">57</mods:start>
<mods:end id="27D81D6F7F6F302CDEC5FFCC37DC1F0D">65</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9AC9CA6F6E2DAC136B0C4AA49C99F508">journal article</mods:classification>
<mods:identifier id="06E1EC2649EC03BB88D1BF7F87FD6876" type="DOI">10.5281/zenodo.211197</mods:identifier>
<mods:identifier id="6C0ABE9026DB9447A2D53BD93397CDD5" type="GBIF-Dataset">14afa521-5f4c-4690-a8f3-71080067f2c5</mods:identifier>
<mods:identifier id="AE7FB987744B0093E8EC13214703E184" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1D48F009C6B294AEB04DC830C26D20D5" type="Zenodo-Dep">211197</mods:identifier>
</mods:mods>
<treatment id="7F09EC44FF9BA405FF3FD456FA4DAA9D" ID-DOI="http://doi.org/10.5281/zenodo.6170014" ID-GBIF-Taxon="119521100" ID-Zenodo-Dep="6170014" LSID="urn:lsid:plazi:treatment:7F09EC44FF9BA405FF3FD456FA4DAA9D" httpUri="http://treatment.plazi.org/id/7F09EC44FF9BA405FF3FD456FA4DAA9D" lastPageId="7" lastPageNumber="64" pageId="0" pageNumber="57">
<subSubSection id="BFBA0ED9FF9BA402FF3FD456FED9A9B2" pageId="0" pageNumber="57" type="nomenclature">
<paragraph id="F71F5D52FF9BA402FF3FD456FE02A992" blockId="0.[151,457,1910,1968]" box="[151,457,1910,1936]" pageId="0" pageNumber="57">
<heading id="AC57EA3EFF9BA402FF3FD456FE02A992" bold="true" box="[151,457,1910,1936]" fontSize="11" level="1" pageId="0" pageNumber="57" reason="1">
<emphasis id="C5D48140FF9BA402FF3FD456FE02A992" bold="true" box="[151,457,1910,1936]" pageId="0" pageNumber="57">
<taxonomicName id="30A026D1FF9BA402FF3FD456FEAEA992" ID-CoL="7T362" box="[151,357,1910,1936]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9BA402FF3FD456FEAEA992" bold="true" box="[151,357,1910,1936]" italics="true" pageId="0" pageNumber="57">Calappa karenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEE73C3BFF9BA402FEC4D456FE02A992" box="[364,457,1910,1936]" pageId="0" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F71F5D52FF9BA402FF3FD4B8FED9A9B2" blockId="0.[151,457,1910,1968]" box="[151,274,1944,1968]" pageId="0" pageNumber="57">
(
<figureCitation id="6F9B41D7FF9BA402FF37D4B8FEC3A9B2" box="[159,264,1944,1968]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1979,2002]" captionStartId-1="3.[151,250,1959,1982]" captionStartId-2="4.[151,250,1934,1957]" captionStartId-3="5.[151,250,1162,1185]" captionStartId-4="6.[151,250,1912,1935]" captionTargetBox-0="[151,1435,1065,1958]" captionTargetBox-1="[203,1384,193,1936]" captionTargetBox-2="[224,1364,193,1899]" captionTargetBox-3="[151,1435,193,1140]" captionTargetBox-4="[213,1370,204,1858]" captionTargetId-0="figure@2.[151,1436,1055,1958]" captionTargetId-1="figure@3.[203,1386,193,1937]" captionTargetId-2="figure@4.[224,1366,193,1900]" captionTargetId-3="figure@5.[151,1436,193,1141]" captionTargetId-4="figure@6.[200,1387,193,1891]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." captionText-1="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." captionText-2="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." captionText-3="FIGURE 4. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, outer view of right chela, carpus and merus; B, outer view of left chela; C, dorsal view of merus and carpus of left cheliped; D, frontal view showing third maxillipeds, front, orbit, antennules and antennae." captionText-4="FIGURE 5. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, left basal antennal segment; B, C, left G 1; D, E, distal part of left G 1; F, left G 2; G, distal part of left G 2. Scale = 1.0 mm." httpUri-0="https://zenodo.org/record/211198/files/figure.png" httpUri-1="https://zenodo.org/record/211199/files/figure.png" httpUri-2="https://zenodo.org/record/211200/files/figure.png" httpUri-3="https://zenodo.org/record/211201/files/figure.png" httpUri-4="https://zenodo.org/record/211202/files/figure.png" pageId="0" pageNumber="57">Figs. 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BFBA0ED9FF9BA403FF3FD4FBFABFAEF4" lastPageId="1" lastPageNumber="58" pageId="0" pageNumber="57" type="reference_group">
<paragraph id="F71F5D52FF9BA403FF3FD4FBFABFAEF4" blockId="0.[151,1216,2011,2033]" lastBlockId="1.[151,1436,151,2007]" lastPageId="1" lastPageNumber="58" pageId="0" pageNumber="57">
<taxonomicName id="30A026D1FF9BA402FF3FD4FBFF25A9F3" ID-CoL="7NRJ9" box="[151,238,2011,2033]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="C5D48140FF9BA402FF3FD4FBFF25A9F3" box="[151,238,2011,2033]" italics="true" pageId="0" pageNumber="57">Calappa</emphasis>
</taxonomicName>
sp. 1 (
<taxonomicName id="30A026D1FF9BA402FE9AD4FBFDF3A9F3" authority="Galil, 1997" authorityName="Galil" authorityYear="1997" box="[306,568,2011,2033]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="sebastiani">
aff. 
<emphasis id="C5D48140FF9BA402FEF3D4FBFE09A9F3" box="[347,450,2011,2033]" italics="true" pageId="0" pageNumber="57">sebastiani</emphasis>
<bibRefCitation id="933120A3FF9BA402FE60D4FBFDF3A9F3" author="Galil" box="[456,568,2011,2033]" pageId="0" pageNumber="57" refString="Galil, B. S. (1997). Crustacea Decapoda: A revision of the Indo-Pacific species of the genus Calappa Weber, 1795 (Calappidae) In A. Crosnier (ed.) Resultats des Campagnes MUSORTORM, Volume 18. Memoires du Museum national d'Histoire naturelle, Paris, 176, 271 - 335." type="journal article" year="1997">Galil, 1997</bibRefCitation>
</taxonomicName>
)— 
<bibRefCitation id="933120A3FF9BA402FDF2D4FBFCDDA9F3" author="Paulay" box="[602,790,2011,2033]" pageId="0" pageNumber="57" refString="Paulay, G., Kropp, R., Ng, P. K. L. &amp; Eldredge, L. G. (2003) The crustaceans and pycnogonids of the Mariana Islands. Micronesica, 35 - 36, 456 - 513." type="book chapter" year="2003">
Paulay 
<emphasis id="C5D48140FF9BA402FD0FD4FCFD11A9F3" box="[679,730,2011,2033]" italics="true" pageId="0" pageNumber="57">et al.</emphasis>
2003
</bibRefCitation>
: 496 [not 
<taxonomicName id="30A026D1FF9BA402FCD3D4FBFB71A9F3" authority="Galil, 1997" authorityName="Galil" authorityYear="1997" box="[891,1210,2011,2033]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="sebastiani">
<emphasis id="C5D48140FF9BA402FCD3D4FBFB8BA9F3" box="[891,1088,2011,2033]" italics="true" pageId="0" pageNumber="57">Calappa sebastiani</emphasis>
<bibRefCitation id="933120A3FF9BA402FBEED4FBFB71A9F3" author="Galil" box="[1094,1210,2011,2033]" pageId="0" pageNumber="57" refString="Galil, B. S. (1997). Crustacea Decapoda: A revision of the Indo-Pacific species of the genus Calappa Weber, 1795 (Calappidae) In A. Crosnier (ed.) Resultats des Campagnes MUSORTORM, Volume 18. Memoires du Museum national d'Histoire naturelle, Paris, 176, 271 - 335." type="journal article" year="1997">Galil, 1997</bibRefCitation>
</taxonomicName>
] 
<materialsCitation id="47C8570FFF9AA403FF3FD3B7FC6EAED1" ID-GBIF-Occurrence="1948792852" collectingDate="1986-04-10" collectionCode="USNM" collectorName="E. Smith &amp; L. Eldredge" country="Guam" elevation="-183" location="Haputo" pageId="1" pageNumber="58" specimenCode="USNM 1150290" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="C5D48140FF9AA403FF3FD3B7FE4FAEB2" bold="true" box="[151,388,151,176]" pageId="1" pageNumber="58">Material examined.</emphasis>
<typeStatus id="281BE3F0FF9AA403FE26D3B7FE30AEAD" box="[398,507,151,175]" pageId="1" pageNumber="58" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="E1A696DBFF9AA403FDA1D3B7FDF4AEAD" box="[521,575,151,175]" pageId="1" pageNumber="58" type="male">male</specimenCount>
(180.4 by 
<quantity id="3058F0B7FF9AA403FD6CD3B7FCE5AEAD" box="[708,814,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="9.59" pageId="1" pageNumber="58" unit="mm" value="95.9">95.9 mm</quantity>
) (USNM 
<specimenCode id="A706F529FF9AA403FC0DD3B7FBC1AEAD" box="[933,1034,151,175]" pageId="1" pageNumber="58">1150290</specimenCode>
), Haputo, 
<collectingCountry id="8FB71DC2FF9AA403FB2DD3B7FB06AEAD" box="[1157,1229,151,175]" name="Guam" pageId="1" pageNumber="58" value="Guam">Guam</collectingCountry>
, in crab-trap, 100 fathoms (= 
<quantity id="3058F0B7FF9AA403FE8AD39BFEBAAED1" box="[290,369,187,211]" metricMagnitude="2" metricUnit="m" metricValue="1.83" pageId="1" pageNumber="58" unit="m" value="183.0">183 m</quantity>
), coll. 
<collectorName id="5A553884FF9AA403FE6FD39CFD2CAED1" box="[455,743,187,211]" pageId="1" pageNumber="58">E. Smith &amp; L. Eldredge</collectorName>
, 
<collectingDate id="935A827AFF9AA403FD5FD39BFC69AED1" box="[759,930,187,211]" pageId="1" pageNumber="58" value="1986-04-10">
<date id="831E7B92FF9AA403FD5FD39BFC69AED1" box="[759,930,187,211]" pageId="1" pageNumber="58" value="1986-04-10">10 April 1986</date>
</collectingDate>
.
</materialsCitation>
<materialsCitation id="47C8570FFF9AA403FC19D39CFABFAEF4" ID-GBIF-Occurrence="1948792851" collectingDate="1986-04-10" collectionCode="USNM" collectorName="E. Smith &amp; L. Eldredge" country="Guam" elevation="-183" location="Haputo" pageId="1" pageNumber="58" specimenCode="USNM 1150291" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="281BE3F0FF9AA403FC19D39CFBDCAED6" box="[945,1047,188,212]" pageId="1" pageNumber="58" type="paratype">Paratype</typeStatus>
: 
<specimenCount id="E1A696DBFF9AA403FB8ED39BFBBDAED1" box="[1062,1142,187,211]" pageId="1" pageNumber="58" type="male">1 male</specimenCount>
(CL 
<quantity id="3058F0B7FF9AA403FB13D39BFAECAED1" box="[1211,1319,187,211]" metricMagnitude="-2" metricUnit="m" metricValue="8.959999999999999" pageId="1" pageNumber="58" unit="mm" value="89.6">89.6 mm</quantity>
, damaged carapace; dimensions estimated from 
<figureCitation id="6F9B41D7FF9AA403FD96D3FEFDB4AEF4" box="[574,639,222,246]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2002]" captionTargetBox="[151,1435,1065,1958]" captionTargetId="figure@2.[151,1436,1055,1958]" captionTargetPageId="2" captionText="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="1" pageNumber="58">Fig. 1</figureCitation>
, 197.0 mm by 95.0 mm) (USNM 
<specimenCode id="A706F529FF9AA403FC56D3FEFBA8AEF4" box="[1022,1123,222,246]" pageId="1" pageNumber="58">1150291</specimenCode>
), same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BFBA0ED9FF9AA403FF6FD221FD57AFEE" pageId="1" pageNumber="58" type="diagnosis">
<paragraph id="F71F5D52FF9AA403FF6FD221FD57AFEE" blockId="1.[151,1436,151,2007]" pageId="1" pageNumber="58">
<emphasis id="C5D48140FF9AA403FF6FD221FE8AAF18" bold="true" box="[199,321,257,282]" pageId="1" pageNumber="58">Diagnosis.</emphasis>
Carapace much broader than long, width to length ratio 1.88–2.07. Anterior two-thirds of dorsal surface with large, round submammillate tubercles; posterior one-third with numerous low small granules (
<figureCitation id="6F9B41D7FF9AA403FAFAD204FA5EAF3E" box="[1362,1429,292,316]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2002]" captionTargetBox="[151,1435,1065,1958]" captionTargetId="figure@2.[151,1436,1055,1958]" captionTargetPageId="2" captionText="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="1" pageNumber="58">Fig. 1</figureCitation>
, 
<figureCitation id="6F9B41D7FF9AA403FF3FD267FF63AF5D" box="[151,168,327,351]" captionStart="FIGURE 2" captionStartId="3.[151,250,1959,1982]" captionTargetBox="[203,1384,193,1936]" captionTargetId="figure@3.[203,1386,193,1937]" captionTargetPageId="3" captionText="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="1" pageNumber="58">2</figureCitation>
). Pterygostomian lobe with broad inner oblique longitudinal groove (
<figureCitation id="6F9B41D7FF9AA403FC56D267FB81AF5D" box="[1022,1098,327,351]" captionStart="FIGURE 4" captionStartId="5.[151,250,1162,1185]" captionTargetBox="[151,1435,193,1140]" captionTargetId="figure@5.[151,1436,193,1141]" captionTargetPageId="5" captionText="FIGURE 4. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, outer view of right chela, carpus and merus; B, outer view of left chela; C, dorsal view of merus and carpus of left cheliped; D, frontal view showing third maxillipeds, front, orbit, antennules and antennae." httpUri="https://zenodo.org/record/211201/files/figure.png" pageId="1" pageNumber="58">Fig. 4</figureCitation>
D). Posterolateral margin of clypeiform expansion with 8 triangular teeth (
<figureCitation id="6F9B41D7FF9AA403FD3ED24BFD2FAF81" box="[662,740,363,387]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2002]" captionTargetBox="[151,1435,1065,1958]" captionTargetId="figure@2.[151,1436,1055,1958]" captionTargetPageId="2" captionText="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="1" pageNumber="58">Figs. 1</figureCitation>
, 
<figureCitation id="6F9B41D7FF9AA403FD59D24BFCC9AF81" box="[753,770,363,387]" captionStart="FIGURE 2" captionStartId="3.[151,250,1959,1982]" captionTargetBox="[203,1384,193,1936]" captionTargetId="figure@3.[203,1386,193,1937]" captionTargetPageId="3" captionText="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="1" pageNumber="58">2</figureCitation>
). Outer margin of merus of cheliped with 4 lobes (
<figureCitation id="6F9B41D7FF9AA403FA9FD24BFAB0AF81" box="[1335,1403,363,387]" captionStart="FIGURE 4" captionStartId="5.[151,250,1162,1185]" captionTargetBox="[151,1435,193,1140]" captionTargetId="figure@5.[151,1436,193,1141]" captionTargetPageId="5" captionText="FIGURE 4. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, outer view of right chela, carpus and merus; B, outer view of left chela; C, dorsal view of merus and carpus of left cheliped; D, frontal view showing third maxillipeds, front, orbit, antennules and antennae." httpUri="https://zenodo.org/record/211201/files/figure.png" pageId="1" pageNumber="58">Fig. 4</figureCitation>
C). Chela with 6 high, lamelliform teeth on dorsal margin (
<figureCitation id="6F9B41D7FF9AA403FCB4D2AEFCA1AFA4" box="[796,874,398,422]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Figs. 3</figureCitation>
A, 4A, B). Male thoracic sternites 4–6 relatively broad (
<figureCitation id="6F9B41D7FF9AA403FF4ED291FEE2AFCB" box="[230,297,433,457]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Fig. 3</figureCitation>
B, C). Male abdomen relatively broad; somite 6 subrectangular, slightly wider than long (
<figureCitation id="6F9B41D7FF9AA403FABDD291FA93AFCB" box="[1301,1368,433,457]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Fig. 3</figureCitation>
B, C). G1 C-shaped; tip truncated (
<figureCitation id="6F9B41D7FF9AA403FE7CD2F4FDDCAFEE" box="[468,535,468,492]" captionStart="FIGURE 5" captionStartId="6.[151,250,1912,1935]" captionTargetBox="[213,1370,204,1858]" captionTargetId="figure@6.[200,1387,193,1891]" captionTargetPageId="6" captionText="FIGURE 5. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, left basal antennal segment; B, C, left G 1; D, E, distal part of left G 1; F, left G 2; G, distal part of left G 2. Scale = 1.0 mm." httpUri="https://zenodo.org/record/211202/files/figure.png" pageId="1" pageNumber="58">Fig. 5</figureCitation>
B, C, D, E).
</paragraph>
</subSubSection>
<subSubSection id="BFBA0ED9FF9AA400FF6FD2D7FC33AC31" lastPageId="2" lastPageNumber="59" pageId="1" pageNumber="58" type="description">
<paragraph id="F71F5D52FF9AA403FF6FD2D7FBD8AB14" blockId="1.[151,1436,151,2007]" pageId="1" pageNumber="58">
<emphasis id="C5D48140FF9AA403FF6FD2D7FE92AC12" bold="true" box="[199,345,503,528]" pageId="1" pageNumber="58">Description.</emphasis>
Carapace transversely ovate, much broader than long, width to length ratio 1.88 (
<typeStatus id="281BE3F0FF9AA403FB58D2D7FA93AC0D" box="[1264,1368,503,527]" pageId="1" pageNumber="58" type="holotype">holotype</typeStatus>
), 2.07 (estimated from photograph of 
<typeStatus id="281BE3F0FF9AA403FDA2D13CFDA1AC36" box="[522,618,540,564]" pageId="1" pageNumber="58" type="paratype">paratype</typeStatus>
when intact) (
<figureCitation id="6F9B41D7FF9AA403FCB4D13BFCA4AC31" box="[796,879,539,563]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2002]" captionTargetBox="[151,1435,1065,1958]" captionTargetId="figure@2.[151,1436,1055,1958]" captionTargetPageId="2" captionText="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="1" pageNumber="58">Figs. 1</figureCitation>
, 
<figureCitation id="6F9B41D7FF9AA403FC2AD13BFC58AC31" box="[898,915,539,563]" captionStart="FIGURE 2" captionStartId="3.[151,250,1959,1982]" captionTargetBox="[203,1384,193,1936]" captionTargetId="figure@3.[203,1386,193,1937]" captionTargetPageId="3" captionText="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="1" pageNumber="58">2</figureCitation>
). Dorsal surface convex, anterior two-thirds covered with large, round submammillate tubercles, resembling low pustules, surface between tubercles finely granular; posterior third with numerous low small granules, short transverse striae, becoming more prominent, relatively larger towards posterior margin (
<figureCitation id="6F9B41D7FF9AA403FD2DD1A4FD1EAC9E" box="[645,725,644,668]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2002]" captionTargetBox="[151,1435,1065,1958]" captionTargetId="figure@2.[151,1436,1055,1958]" captionTargetPageId="2" captionText="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="1" pageNumber="58">Figs. 1</figureCitation>
, 
<figureCitation id="6F9B41D7FF9AA403FD4DD1A4FD3DAC9E" box="[741,758,644,668]" captionStart="FIGURE 2" captionStartId="3.[151,250,1959,1982]" captionTargetBox="[203,1384,193,1936]" captionTargetId="figure@3.[203,1386,193,1937]" captionTargetPageId="3" captionText="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="1" pageNumber="58">2</figureCitation>
). Gastric region convex, with median part highest point of carapace, surface with low granules, tubercles; longitudinal gastro-cardiac grooves relatively deep, extending to just before posterior margin (
<figureCitation id="6F9B41D7FF9AA403FE4CD1EBFDFFACE1" box="[484,564,715,739]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2002]" captionTargetBox="[151,1435,1065,1958]" captionTargetId="figure@2.[151,1436,1055,1958]" captionTargetPageId="2" captionText="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="1" pageNumber="58">Figs. 1</figureCitation>
, 
<figureCitation id="6F9B41D7FF9AA403FDEBD1EBFD9FACE1" box="[579,596,715,739]" captionStart="FIGURE 2" captionStartId="3.[151,250,1959,1982]" captionTargetBox="[203,1384,193,1936]" captionTargetId="figure@3.[203,1386,193,1937]" captionTargetPageId="3" captionText="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="1" pageNumber="58">2</figureCitation>
). Frontal margin distinctly bifurcated, with deep, broad U-shaped median cleft separating triangular teeth, separated from supraorbital margin by broad cleft (
<figureCitation id="6F9B41D7FF9AA403FBE2D1CEFB44AD04" box="[1098,1167,750,774]" captionStart="FIGURE 2" captionStartId="3.[151,250,1959,1982]" captionTargetBox="[203,1384,193,1936]" captionTargetId="figure@3.[203,1386,193,1937]" captionTargetPageId="3" captionText="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="1" pageNumber="58">Fig. 2</figureCitation>
A). Supraorbital margin uneven, granular, with 2 narrow longitudinal fissures on outer half (
<figureCitation id="6F9B41D7FF9AA403FC13D031FBC8AD2B" box="[955,1027,785,809]" captionStart="FIGURE 2" captionStartId="3.[151,250,1959,1982]" captionTargetBox="[203,1384,193,1936]" captionTargetId="figure@3.[203,1386,193,1937]" captionTargetPageId="3" captionText="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="1" pageNumber="58">Fig. 2</figureCitation>
A). Eyes folded obliquely, external orbital tooth low, tubercular (
<figureCitation id="6F9B41D7FF9AA403FE77D014FDE9AD4E" box="[479,546,820,844]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Fig. 3</figureCitation>
A). Suborbital margin broad, dentiform, appearing sublobiform, with 3 low but distinct teeth, separated from external orbital angle by deep fissure (
<figureCitation id="6F9B41D7FF9AA403FC69D078FBC3AD72" box="[961,1032,856,880]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Fig. 3</figureCitation>
A). Pterygostomian lobe at end of posterior margin of epistome with relatively broad inner oblique longitudinal groove (
<figureCitation id="6F9B41D7FF9AA403FBD7D05BFB0EAD91" box="[1151,1221,891,915]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Fig. 3</figureCitation>
A). Basal antennal article subquadrate, cusp-like, with transverse median ridge; proximal margin with prominent granules, distal margin deeply notched (
<figureCitation id="6F9B41D7FF9AA403FE0CD0E1FE3BADDB" box="[420,496,961,985]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Figs. 3</figureCitation>
A, 5A). Longitudinal epistomial septum appears anteriorly complete (
<figureCitation id="6F9B41D7FF9AA403FB53D0E1FA8DADDB" box="[1275,1350,961,985]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Figs. 3</figureCitation>
A, 4D). Proepistome rounded (
<figureCitation id="6F9B41D7FF9AA403FE08D0C4FE23ADFE" box="[416,488,996,1020]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Fig. 3</figureCitation>
A). Third maxilliped with margins of merus densely setose, distal margin cleft, anterolateral angle auriculiform (
<figureCitation id="6F9B41D7FF9AA403FDA0D728FD86AA22" box="[520,589,1032,1056]" captionStart="FIGURE 4" captionStartId="5.[151,250,1162,1185]" captionTargetBox="[151,1435,193,1140]" captionTargetId="figure@5.[151,1436,193,1141]" captionTargetPageId="5" captionText="FIGURE 4. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, outer view of right chela, carpus and merus; B, outer view of left chela; C, dorsal view of merus and carpus of left cheliped; D, frontal view showing third maxillipeds, front, orbit, antennules and antennae." httpUri="https://zenodo.org/record/211201/files/figure.png" pageId="1" pageNumber="58">Fig. 4</figureCitation>
D). Anterolateral margin arcuate, with 12–15 blunt, rounded unevenly sized tubercles, first tubercle distinct, subsequent ones very low, not easily discernible; last 5 or 6 progressively larger, more dentiform. Posterolateral margin of clypeiform expansion well developed, margins densely setose but not obscuring margin, with 8 triangular teeth, each tooth with denticulate margins, longitudinal median ridge (very strong in last 3 posterior teeth); first 4 teeth symmetrical, directed obliquely anteriorly; fifth tooth slightly asymmetrical, posterior part slightly longer, directed laterally; sixth tooth largest, broadest, asymmetrical, inner margin twice length of outer margin, directed posteriorly; last 2 teeth progressively smaller; posterior carapace margin gently concave, granular, not flanked by teeth but low lobes (
<figureCitation id="6F9B41D7FF9AA403FC32D7DEFC2CAB14" box="[922,999,1278,1302]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2002]" captionTargetBox="[151,1435,1065,1958]" captionTargetId="figure@2.[151,1436,1055,1958]" captionTargetPageId="2" captionText="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="1" pageNumber="58">Figs. 1</figureCitation>
, 
<figureCitation id="6F9B41D7FF9AA403FC5DD7DEFBCEAB14" box="[1013,1029,1278,1302]" captionStart="FIGURE 2" captionStartId="3.[151,250,1959,1982]" captionTargetBox="[203,1384,193,1936]" captionTargetId="figure@3.[203,1386,193,1937]" captionTargetPageId="3" captionText="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="1" pageNumber="58">2</figureCitation>
).
</paragraph>
<paragraph id="F71F5D52FF9AA403FF61D601FCF5A8E2" blockId="1.[151,1436,151,2007]" pageId="1" pageNumber="58">
Chelipeds asymmetrical, right larger (
<figureCitation id="6F9B41D7FF9AA403FDDBD601FD0AAB3B" box="[627,705,1313,1337]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2002]" captionTargetBox="[151,1435,1065,1958]" captionTargetId="figure@2.[151,1436,1055,1958]" captionTargetPageId="2" captionText="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="1" pageNumber="58">Figs. 1</figureCitation>
, 
<figureCitation id="6F9B41D7FF9AA403FD78D601FD15AB3B" box="[720,734,1313,1337]" captionStart="FIGURE 2" captionStartId="3.[151,250,1959,1982]" captionTargetBox="[203,1384,193,1936]" captionTargetId="figure@3.[203,1386,193,1937]" captionTargetPageId="3" captionText="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="1" pageNumber="58">2</figureCitation>
A, B, 3A). Outer margin of merus with 4 lobes, first lobe very low, distalmost largest, sharpest; tip of lobes 3, 4 with sharp tubercle distally (
<figureCitation id="6F9B41D7FF9AA403FC57D664FB88AB5E" box="[1023,1091,1348,1372]" captionStart="FIGURE 4" captionStartId="5.[151,250,1162,1185]" captionTargetBox="[151,1435,193,1140]" captionTargetId="figure@5.[151,1436,193,1141]" captionTargetPageId="5" captionText="FIGURE 4. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, outer view of right chela, carpus and merus; B, outer view of left chela; C, dorsal view of merus and carpus of left cheliped; D, frontal view showing third maxillipeds, front, orbit, antennules and antennae." httpUri="https://zenodo.org/record/211201/files/figure.png" pageId="1" pageNumber="58">Fig. 4</figureCitation>
C). Carpus subtriangular, outer surface granular with scattered low tubercles (
<figureCitation id="6F9B41D7FF9AA403FD08D648FD2EAB82" box="[672,741,1384,1408]" captionStart="FIGURE 4" captionStartId="5.[151,250,1162,1185]" captionTargetBox="[151,1435,193,1140]" captionTargetId="figure@5.[151,1436,193,1141]" captionTargetPageId="5" captionText="FIGURE 4. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, outer view of right chela, carpus and merus; B, outer view of left chela; C, dorsal view of merus and carpus of left cheliped; D, frontal view showing third maxillipeds, front, orbit, antennules and antennae." httpUri="https://zenodo.org/record/211201/files/figure.png" pageId="1" pageNumber="58">Fig. 4</figureCitation>
B, C). Chela high, dorsal margin with prominent crest, with 6 high, lamelliform teeth, first smaller than second, directed slightly anteriorly; next 5 teeth vertical, acutely triangular, progressively smaller; base of crest with 2 small teeth; outer surface of manus with numerous granules, more prominent becoming denser, more prominent towards ventral margin with scattered large rounded tubercles; granules relatively more prominent in smaller chela; ventral margin lined with prominent row of granules, with prominent triangular tooth on proximal edge of ventral margin (
<figureCitation id="6F9B41D7FF9AA403FC84D538FCB1A832" box="[812,890,1560,1584]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Figs. 3</figureCitation>
A, 4A-C). Dactylus of major chela with distinct basal curved cutting tooth, rest of margin with several low teeth; pollex with 3 molariform teeth, first tooth largest, subsequent teeth small. Dactylus of minor chela slender, curved, proximal two-thirds granular, cutting margin lined with denticles; pollex relatively stouter, surface granular, proximal cutting margin lined with denticles, distal part more blade-like (
<figureCitation id="6F9B41D7FF9AA403FEFFD584FE50A8BE" box="[343,411,1700,1724]" captionStart="FIGURE 4" captionStartId="5.[151,250,1162,1185]" captionTargetBox="[151,1435,193,1140]" captionTargetId="figure@5.[151,1436,193,1141]" captionTargetPageId="5" captionText="FIGURE 4. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, outer view of right chela, carpus and merus; B, outer view of left chela; C, dorsal view of merus and carpus of left cheliped; D, frontal view showing third maxillipeds, front, orbit, antennules and antennae." httpUri="https://zenodo.org/record/211201/files/figure.png" pageId="1" pageNumber="58">Fig. 4</figureCitation>
A, B). Ambulatory legs slender, surfaces smooth; first leg longest; merus, carpus, propodus laterally flattened; dactylus styliform, gently curved (
<figureCitation id="6F9B41D7FF9AA403FD44D5E8FCE6A8E2" box="[748,813,1736,1760]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2002]" captionTargetBox="[151,1435,1065,1958]" captionTargetId="figure@2.[151,1436,1055,1958]" captionTargetPageId="2" captionText="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="1" pageNumber="58">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="F71F5D52FF9AA403FF61D5CBFC96A96E" blockId="1.[151,1436,151,2007]" pageId="1" pageNumber="58">
Thoracic sternum narrow; sternites 1, 2 completely fused; separated from sternite 3 by convex shallow groove; sternites 3, 4 fused but median sutures still visible, medially interrupted with longitudinal depression (
<figureCitation id="6F9B41D7FF9AA403FA9BD42EFAB3A924" box="[1331,1400,1806,1830]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Fig. 3</figureCitation>
B); male thoracic sternites 4–6 relatively low, broad, surface covered with low, flattened granules (
<figureCitation id="6F9B41D7FF9AA403FB15D411FB34A94B" box="[1213,1279,1841,1865]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Fig. 3</figureCitation>
B, C). Sternoabdominal cavity almost reaching thoracic suture 3/4 (
<figureCitation id="6F9B41D7FF9AA403FD50D474FCF1A96E" box="[760,826,1876,1900]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="1" pageNumber="58">Fig. 3</figureCitation>
B).
</paragraph>
<paragraph id="F71F5D52FF9AA400FF61D458FDFEAF1B" blockId="1.[151,1436,151,2007]" lastBlockId="2.[151,1436,151,1028]" lastPageId="2" lastPageNumber="59" pageId="1" pageNumber="58">
Abdomen relatively broad; somite 1 longitudinally narrow with lateral parts expanded, anterior margin distinctly granular; somite 2 with lateral parts expanded to form subauriculiform process, surface prominently granular; somites 3–5 fused but median sutures just visible as shallow broad grooves, present as clefts laterally; somite 3 sub-rectangular, outer margin appears lobiform, granular, scalloped; somite 4 trapezoidal, lateral margins sinuous; somite 5 rectangular, lateral margins concave, distal margin granular; somite 6 subrectangular, slightly wider than long, lateral margins concave; telson acutely triangular, converging gradually to sharp tip; lateral margins gently concave (
<figureCitation id="6F9B41D7FF99A400FE07D221FE39AF1B" box="[431,498,257,281]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="2" pageNumber="59">Fig. 3</figureCitation>
B, C).
</paragraph>
<paragraph id="F71F5D52FF99A400FF61D204FB2CAF5D" blockId="2.[151,1436,151,1028]" pageId="2" pageNumber="59">
G1 relatively stout, slightly sinuous (
<figureCitation id="6F9B41D7FF99A400FDD1D204FD74AF3E" box="[633,703,292,316]" captionStart="FIGURE 5" captionStartId="6.[151,250,1912,1935]" captionTargetBox="[213,1370,204,1858]" captionTargetId="figure@6.[200,1387,193,1891]" captionTargetPageId="6" captionText="FIGURE 5. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, left basal antennal segment; B, C, left G 1; D, E, distal part of left G 1; F, left G 2; G, distal part of left G 2. Scale = 1.0 mm." httpUri="https://zenodo.org/record/211202/files/figure.png" pageId="2" pageNumber="59">Fig. 5</figureCitation>
B, C); tip truncated, distal surface lined with numerous minute spinules (
<figureCitation id="6F9B41D7FF99A400FEAAD267FE8FAF5D" box="[258,324,327,351]" captionStart="FIGURE 5" captionStartId="6.[151,250,1912,1935]" captionTargetBox="[213,1370,204,1858]" captionTargetId="figure@6.[200,1387,193,1891]" captionTargetPageId="6" captionText="FIGURE 5. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, left basal antennal segment; B, C, left G 1; D, E, distal part of left G 1; F, left G 2; G, distal part of left G 2. Scale = 1.0 mm." httpUri="https://zenodo.org/record/211202/files/figure.png" pageId="2" pageNumber="59">Fig. 5</figureCitation>
D, E). G2 subequal in length to G1, slender, curved, distal tip rounded (
<figureCitation id="6F9B41D7FF99A400FBCCD267FB6DAF5D" box="[1124,1190,327,351]" captionStart="FIGURE 5" captionStartId="6.[151,250,1912,1935]" captionTargetBox="[213,1370,204,1858]" captionTargetId="figure@6.[200,1387,193,1891]" captionTargetPageId="6" captionText="FIGURE 5. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, left basal antennal segment; B, C, left G 1; D, E, distal part of left G 1; F, left G 2; G, distal part of left G 2. Scale = 1.0 mm." httpUri="https://zenodo.org/record/211202/files/figure.png" pageId="2" pageNumber="59">Fig. 5</figureCitation>
F, G).
</paragraph>
<paragraph id="F71F5D52FF99A400FF6FD24AFC33AC31" blockId="2.[151,1436,151,1028]" pageId="2" pageNumber="59">
<emphasis id="C5D48140FF99A400FF6FD24AFEEBAF81" bold="true" box="[199,288,362,387]" pageId="2" pageNumber="59">Colour.</emphasis>
On the basis of the colour photograph by Roy Kropp (
<figureCitation id="6F9B41D7FF99A400FC35D24BFC2BAF81" box="[925,992,363,387]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2002]" captionTargetBox="[151,1435,1065,1958]" captionTargetId="figure@2.[151,1436,1055,1958]" captionTargetPageId="2" captionText="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="2" pageNumber="59">Fig. 1</figureCitation>
), the live colouration of 
<taxonomicName id="30A026D1FF99A400FAACD24CFA4BAF81" box="[1284,1408,363,387]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="karenae">
<emphasis id="C5D48140FF99A400FAACD24CFA4BAF81" box="[1284,1408,363,387]" italics="true" pageId="2" pageNumber="59">C. karenae</emphasis>
</taxonomicName>
is striking. It ranges from orange-red anteriorly to a light salmon pink in the posterior part of the carapace, with the round, submammillate tubercles in the middle third of the carapace scarlet red and bright yellow at the apex, resembling small pustules. Tips of movable fingers are dark brown, as are the tips of propodus on its ambulatory legs. The inner surface of the cheliped carpus is red, and the inner surface of its manus bears numerous small red spots. The preserved specimens have lost their colour and are now uniformly beige.
</paragraph>
</subSubSection>
<subSubSection id="BFBA0ED9FF99A400FF6FD11EFAF4AC7B" pageId="2" pageNumber="59" type="etymology">
<paragraph id="F71F5D52FF99A400FF6FD11EFAF4AC7B" blockId="2.[151,1436,151,1028]" pageId="2" pageNumber="59">
<emphasis id="C5D48140FF99A400FF6FD11EFE85AC54" bold="true" box="[199,334,574,598]" pageId="2" pageNumber="59">Etymology.</emphasis>
The authors are delighted to dedicate this distinctive new species to Ms Karen Reed, a custodian to the priceless crustacean collections at USNM, who helped us eventually locate these elusive specimens.
</paragraph>
</subSubSection>
<subSubSection id="BFBA0ED9FF99A400FF6FD1A5FC94AA06" pageId="2" pageNumber="59" type="discussion">
<paragraph id="F71F5D52FF99A400FF6FD1A5FC94AA06" blockId="2.[151,1436,151,1028]" pageId="2" pageNumber="59">
<emphasis id="C5D48140FF99A400FF6FD1A5FEF0AC9F" bold="true" box="[199,315,645,669]" pageId="2" pageNumber="59">Remarks.</emphasis>
A label associated with the present specimens, written in 1979 by the late Austin Williams notes that the species “keys to 
<taxonomicName id="30A026D1FF99A400FE1CD189FDDEACC3" box="[436,533,681,705]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="hepatica">
<emphasis id="C5D48140FF99A400FE1CD189FDDEACC3" box="[436,533,681,705]" italics="true" pageId="2" pageNumber="59">hepatica</emphasis>
</taxonomicName>
in Sakai 1976 but cannot be that”. The general features of 
<taxonomicName id="30A026D1FF99A400FB14D189FAF3ACC3" box="[1212,1336,681,705]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF99A400FB14D189FAF3ACC3" box="[1212,1336,681,705]" italics="true" pageId="2" pageNumber="59">C. karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF99A400FAE9D189FA57ACC3" bold="true" box="[1345,1436,681,705]" pageId="2" pageNumber="59">
<taxonomicNameLabel id="DEE73C3BFF99A400FAE9D189FA57ACC3" box="[1345,1436,681,705]" pageId="2" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
superficially resemble 
<taxonomicName id="30A026D1FF99A400FE3ED1EDFD2BACE6" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[406,736,716,741]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="hepatica">
<emphasis id="C5D48140FF99A400FE3ED1EDFDD3ACE7" box="[406,536,717,741]" italics="true" pageId="2" pageNumber="59">C. hepatica</emphasis>
(
<bibRefCitation id="933120A3FF99A400FD8ED1ECFD13ACE6" author="Linnaeus" box="[550,728,716,740]" pageId="2" pageNumber="59" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum caracteribus, differentiis, synonymis, locis. Ed. 10, 1, L. Salvii, Holmiae, iv + 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
, but the differences are marked. 
<taxonomicName id="30A026D1FF99A400FBE6D1EDFADCACE7" box="[1102,1303,717,741]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="hepatica">
<emphasis id="C5D48140FF99A400FBE6D1EDFADCACE7" box="[1102,1303,717,741]" italics="true" pageId="2" pageNumber="59">Calappa hepatica</emphasis>
</taxonomicName>
has a green to purplish-grey carapace, never red. The front does not project so far anteriorly, there are no mammary-like tubercles on the carapace, and most importantly, the carapace is much less broad, with the width to length ratio of about 1.6 to 1.7 (see 
<bibRefCitation id="933120A3FF99A400FED6D018FE33AD52" author="Galil" box="[382,504,824,848]" pageId="2" pageNumber="59" refString="Galil, B. S. (1997). Crustacea Decapoda: A revision of the Indo-Pacific species of the genus Calappa Weber, 1795 (Calappidae) In A. Crosnier (ed.) Resultats des Campagnes MUSORTORM, Volume 18. Memoires du Museum national d'Histoire naturelle, Paris, 176, 271 - 335." type="journal article" year="1997">Galil 1997</bibRefCitation>
). The basal article of the antenna is also completely different (
<bibRefCitation id="933120A3FF99A400FB03D018FA44AD52" author="Galil" box="[1195,1423,824,848]" pageId="2" pageNumber="59" refString="Galil, B. S. (1997). Crustacea Decapoda: A revision of the Indo-Pacific species of the genus Calappa Weber, 1795 (Calappidae) In A. Crosnier (ed.) Resultats des Campagnes MUSORTORM, Volume 18. Memoires du Museum national d'Histoire naturelle, Paris, 176, 271 - 335." type="journal article" year="1997" yearSuffix="a">Galil 1997: Fig. 14a</bibRefCitation>
). The carapace of 
<taxonomicName id="30A026D1FF99A400FEFAD07DFE06AD77" box="[338,461,861,885]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF99A400FEFAD07DFE06AD77" box="[338,461,861,885]" italics="true" pageId="2" pageNumber="59">C. karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF99A400FE7CD07DFDE5AD77" bold="true" box="[468,558,861,885]" pageId="2" pageNumber="59">
<taxonomicNameLabel id="DEE73C3BFF99A400FE7CD07DFDE5AD77" box="[468,558,861,885]" pageId="2" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is wider than any other 
<taxonomicName id="30A026D1FF99A400FCE8D07DFC6BAD77" box="[832,928,861,885]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="C5D48140FF99A400FCE8D07DFC6BAD77" box="[832,928,861,885]" italics="true" pageId="2" pageNumber="59">Calappa</emphasis>
</taxonomicName>
species in the Indo-West Pacific: 
<taxonomicName id="30A026D1FF99A400FA88D07DFE9DAD9A" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="calappa">
<emphasis id="C5D48140FF99A400FA88D07DFA57AD77" box="[1312,1436,861,885]" italics="true" pageId="2" pageNumber="59">C. calappa</emphasis>
(
<bibRefCitation id="933120A3FF99A400FF36D0A0FE85AD9A" author="Linnaeus" box="[158,334,896,920]" pageId="2" pageNumber="59" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum caracteribus, differentiis, synonymis, locis. Ed. 10, 1, L. Salvii, Holmiae, iv + 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
the next widest species, has a width to length ratio of 1.7 to 1.8 (
<bibRefCitation id="933120A3FF99A400FB98D0A0FB66AD9A" author="Galil" box="[1072,1197,896,920]" pageId="2" pageNumber="59" refString="Galil, B. S. (1997). Crustacea Decapoda: A revision of the Indo-Pacific species of the genus Calappa Weber, 1795 (Calappidae) In A. Crosnier (ed.) Resultats des Campagnes MUSORTORM, Volume 18. Memoires du Museum national d'Histoire naturelle, Paris, 176, 271 - 335." type="journal article" year="1997">Galil 1997</bibRefCitation>
). The carapace width to length ratios of the 
<typeStatus id="281BE3F0FF99A400FE0DD084FDCCADBE" box="[421,519,932,956]" pageId="2" pageNumber="59" type="holotype">holotype</typeStatus>
and 
<typeStatus id="281BE3F0FF99A400FDEFD085FD6CADBF" box="[583,679,933,957]" pageId="2" pageNumber="59" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="30A026D1FF99A400FD49D085FC94ADBF" box="[737,863,933,957]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF99A400FD49D085FC94ADBF" box="[737,863,933,957]" italics="true" pageId="2" pageNumber="59">C. karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF99A400FCC4D085FC01ADBF" bold="true" box="[876,970,933,957]" pageId="2" pageNumber="59">
<taxonomicNameLabel id="DEE73C3BFF99A400FCC4D085FC01ADBF" box="[876,970,933,957]" pageId="2" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is 1.88 and 2.07. respectively 
<taxonomicName id="30A026D1FF99A400FA94D085FE67ADE2" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="C5D48140FF99A400FA94D085FF32ADE3" italics="true" pageId="2" pageNumber="59">Calappa japonica</emphasis>
<bibRefCitation id="933120A3FF99A400FEA9D0E8FE67ADE2" author="Ortmann" box="[257,428,968,992]" pageId="2" pageNumber="59" refString="Ortmann, A. E. (1892) Die Decapoda-Krebe des Strassburger Museum. Theil 5, Die Abtheilungen Hippidea, Dromiidea und Oxystomata, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum sufbewahrten Formen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588." type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
</taxonomicName>
has been reported to attain a larger carapace length (see 
<bibRefCitation id="933120A3FF99A400FB9BD0E8FB2FADE2" author="Lai" box="[1075,1252,968,993]" pageId="2" pageNumber="59" refString="Lai, J. C. Y. &amp; Ng, P. K. L. (2006) A new species of Calappa Weber, 1795 (Crustacea: Decapoda: Calappidae) from East and South Africa. Zootaxa, 1358, 39 - 48." type="journal article" year="2006">Lai &amp; Ng 2006</bibRefCitation>
; Ng 
<emphasis id="C5D48140FF99A400FAB2D0EAFA9FADE3" box="[1306,1364,969,993]" italics="true" pageId="2" pageNumber="59">et al.</emphasis>
2011) but its width to length ratio is approximately 
<quantity id="3058F0B7FF99A400FD39D0CCFD1AAA06" box="[657,721,1004,1028]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" pageId="2" pageNumber="59" unit="in" value="1.4">1.4 in</quantity>
adult males.
</paragraph>
</subSubSection>
<caption id="A3DF0DDAFF99A400FF3FD49BFF14A9F2" httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="2" pageNumber="59" targetBox="[151,1435,1065,1958]" targetPageId="2">
<paragraph id="F71F5D52FF99A400FF3FD49BFF14A9F2" blockId="2.[151,1437,1979,2032]" pageId="2" pageNumber="59">
<emphasis id="C5D48140FF99A400FF3FD49BFED9A9D3" bold="true" box="[151,274,1979,2002]" pageId="2" pageNumber="59">FIGURE 1.</emphasis>
<taxonomicName id="30A026D1FF99A400FEB0D49CFE0DA9D0" box="[280,454,1980,2002]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF99A400FEB0D49CFE0DA9D0" box="[280,454,1980,2002]" italics="true" pageId="2" pageNumber="59">Calappa karenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEE73C3BFF99A400FE64D49CFDD6A9D3" box="[460,541,1980,2001]" pageId="2" pageNumber="59" rank="species">
<emphasis id="C5D48140FF99A400FE64D49CFDD6A9D3" bold="true" box="[460,541,1980,2001]" pageId="2" pageNumber="59">sp. nov.</emphasis>
</taxonomicNameLabel>
Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp.
</paragraph>
</caption>
<caption id="A3DF0DDAFF98A401FF3FD487FD35A9DE" httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="3" pageNumber="60" targetBox="[203,1384,193,1936]" targetPageId="3">
<paragraph id="F71F5D52FF98A401FF3FD487FD35A9DE" blockId="3.[151,1436,1959,2012]" pageId="3" pageNumber="60">
<emphasis id="C5D48140FF98A401FF3FD487FEDEA9BF" bold="true" box="[151,277,1959,1982]" pageId="3" pageNumber="60">FIGURE 2.</emphasis>
<taxonomicName id="30A026D1FF98A401FEB6D487FE05A9BF" box="[286,462,1959,1981]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF98A401FEB6D487FE05A9BF" box="[286,462,1959,1981]" italics="true" pageId="3" pageNumber="60">Calappa karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF98A401FE70D488FDE1A9BF" bold="true" box="[472,554,1960,1981]" pageId="3" pageNumber="60">
<taxonomicNameLabel id="DEE73C3BFF98A401FE70D488FDE1A9BF" box="[472,554,1960,1981]" pageId="3" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace.
</paragraph>
</caption>
<caption id="A3DF0DDAFF9FA406FF3FD4AEFD37A9E0" httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="4" pageNumber="61" targetBox="[224,1364,193,1899]" targetPageId="4">
<paragraph id="F71F5D52FF9FA406FF3FD4AEFD37A9E0" blockId="4.[151,1437,1934,2018]" pageId="4" pageNumber="61">
<emphasis id="C5D48140FF9FA406FF3FD4AEFEDEA9A6" bold="true" box="[151,277,1934,1957]" pageId="4" pageNumber="61">FIGURE 3.</emphasis>
<taxonomicName id="30A026D1FF9FA406FEB6D4AFFE05A9A7" box="[286,462,1935,1957]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9FA406FEB6D4AFFE05A9A7" box="[286,462,1935,1957]" italics="true" pageId="4" pageNumber="61">Calappa karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9FA406FE7FD4AFFDE1A9A6" bold="true" box="[471,554,1935,1956]" pageId="4" pageNumber="61">
<taxonomicNameLabel id="DEE73C3BFF9FA406FE7FD4AFFDE1A9A6" box="[471,554,1935,1956]" pageId="4" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites.
</paragraph>
</caption>
<caption id="A3DF0DDAFF9EA407FF3FD7AAFCEEAADC" httpUri="https://zenodo.org/record/211201/files/figure.png" pageId="5" pageNumber="62" targetBox="[151,1435,193,1140]" targetPageId="5">
<paragraph id="F71F5D52FF9EA407FF3FD7AAFCEEAADC" blockId="5.[151,1436,1162,1246]" pageId="5" pageNumber="62">
<emphasis id="C5D48140FF9EA407FF3FD7AAFED8AAA2" bold="true" box="[151,275,1162,1185]" pageId="5" pageNumber="62">FIGURE 4.</emphasis>
<taxonomicName id="30A026D1FF9EA407FEB2D7ABFE03AAA3" box="[282,456,1163,1185]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9EA407FEB2D7ABFE03AAA3" box="[282,456,1163,1185]" italics="true" pageId="5" pageNumber="62">Calappa karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9EA407FE67D7ABFDEBAAA2" bold="true" box="[463,544,1163,1184]" pageId="5" pageNumber="62">
<taxonomicNameLabel id="DEE73C3BFF9EA407FE67D7ABFDEBAAA2" box="[463,544,1163,1184]" pageId="5" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, outer view of right chela, carpus and merus; B, outer view of left chela; C, dorsal view of merus and carpus of left cheliped; D, frontal view showing third maxillipeds, front, orbit, antennules and antennae.
</paragraph>
</caption>
<subSubSection id="BFBA0ED9FF9EA405FF6FD60FFA4DAA9D" lastPageId="7" lastPageNumber="64" pageId="5" pageNumber="62" type="reference_group">
<paragraph id="F71F5D52FF9EA407FF6FD60FFECBABB1" blockId="5.[151,1437,1327,2036]" pageId="5" pageNumber="62">
<taxonomicName id="30A026D1FF9EA407FF6FD60FFE40AB45" ID-CoL="7T362" box="[199,395,1327,1351]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9EA407FF6FD60FFE40AB45" box="[199,395,1327,1351]" italics="true" pageId="5" pageNumber="62">Calappa karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9EA407FE3ED60FFE38AB45" bold="true" box="[406,499,1327,1351]" pageId="5" pageNumber="62">
<taxonomicNameLabel id="DEE73C3BFF9EA407FE3ED60FFE38AB45" box="[406,499,1327,1351]" pageId="5" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was originally identified as 
<taxonomicName id="30A026D1FF9EA407FCECD610FBAEAB45" ID-CoL="PFSH" authority="Galil, 1997" authorityName="Galil" authorityYear="1997" box="[836,1125,1327,1351]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="sebastieni">
<emphasis id="C5D48140FF9EA407FCECD610FC12AB45" box="[836,985,1327,1351]" italics="true" pageId="5" pageNumber="62">C. sebastieni</emphasis>
<bibRefCitation id="933120A3FF9EA407FC4BD60FFBAEAB45" author="Galil" box="[995,1125,1327,1351]" pageId="5" pageNumber="62" refString="Galil, B. S. (1997). Crustacea Decapoda: A revision of the Indo-Pacific species of the genus Calappa Weber, 1795 (Calappidae) In A. Crosnier (ed.) Resultats des Campagnes MUSORTORM, Volume 18. Memoires du Museum national d'Histoire naturelle, Paris, 176, 271 - 335." type="journal article" year="1997">Galil, 1997</bibRefCitation>
</taxonomicName>
(labelled as “ 
<taxonomicName id="30A026D1FF9EA407FAA3D60FFECCAB69" ID-CoL="PFSH" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="sebastieni">
<emphasis id="C5D48140FF9EA407FAA3D60FFAA0AB45" box="[1291,1387,1327,1351]" italics="true" pageId="5" pageNumber="62">Calappa</emphasis>
aff. 
<emphasis id="C5D48140FF9EA407FF3FD673FECCAB69" box="[151,263,1363,1387]" italics="true" pageId="5" pageNumber="62">sebastieni</emphasis>
</taxonomicName>
” in 
<bibRefCitation id="933120A3FF9EA407FE9FD673FDC8AB69" author="Paulay" box="[311,515,1363,1387]" pageId="5" pageNumber="62" refString="Paulay, G., Kropp, R., Ng, P. K. L. &amp; Eldredge, L. G. (2003) The crustaceans and pycnogonids of the Mariana Islands. Micronesica, 35 - 36, 456 - 513." type="book chapter" year="2003">
Paulay 
<emphasis id="C5D48140FF9EA407FE22D674FE70AB69" box="[394,443,1363,1387]" italics="true" pageId="5" pageNumber="62">et al</emphasis>
. 2003
</bibRefCitation>
) on the basis of the photograph taken by Roy Kropp, but they are clearly different species, with 
<taxonomicName id="30A026D1FF9EA407FE85D658FE75AB8D" ID-CoL="PFSH" box="[301,446,1399,1423]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="sebastieni">
<emphasis id="C5D48140FF9EA407FE85D658FE75AB8D" box="[301,446,1399,1423]" italics="true" pageId="5" pageNumber="62">C. sebastieni</emphasis>
</taxonomicName>
possessing a very different rostrum (
<bibRefCitation id="933120A3FF9EA407FCF2D657FBFFAB8D" author="Galil" box="[858,1076,1399,1423]" pageId="5" pageNumber="62" refString="Galil, B. S. (1997). Crustacea Decapoda: A revision of the Indo-Pacific species of the genus Calappa Weber, 1795 (Calappidae) In A. Crosnier (ed.) Resultats des Campagnes MUSORTORM, Volume 18. Memoires du Museum national d'Histoire naturelle, Paris, 176, 271 - 335." type="journal article" year="1997" yearSuffix="f">Galil 1997: fig. 20f</bibRefCitation>
) and a proportionately narrower carapace.
</paragraph>
<paragraph id="F71F5D52FF9EA407FF6FD69FFC2EA9F1" blockId="5.[151,1437,1327,2036]" pageId="5" pageNumber="62">
The species perhaps closest to 
<taxonomicName id="30A026D1FF9EA407FD80D6E0FD68ABD5" ID-CoL="7T362" box="[552,675,1471,1495]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9EA407FD80D6E0FD68ABD5" box="[552,675,1471,1495]" italics="true" pageId="5" pageNumber="62">C. karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9EA407FD04D69FFCC3ABD5" bold="true" box="[684,776,1471,1495]" pageId="5" pageNumber="62">
<taxonomicNameLabel id="DEE73C3BFF9EA407FD04D69FFCC3ABD5" box="[684,776,1471,1495]" pageId="5" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the poorly known 
<taxonomicName id="30A026D1FF9EA407FC56D6E0FAE8ABD5" ID-CoL="PFT4" authority="Sakai, 1980" authorityName="Sakai" authorityYear="1980" box="[1022,1315,1471,1495]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9EA407FC56D6E0FB59ABD5" box="[1022,1170,1472,1495]" italics="true" pageId="5" pageNumber="62">C. yamasitae</emphasis>
<bibRefCitation id="933120A3FF9EA407FB33D69FFAE8ABD5" author="Sakai" box="[1179,1315,1471,1495]" pageId="5" pageNumber="62" refString="Sakai, T. (1980) New species of crabs of the families Lithodidae and Calappidae. Researches on Crustacea, 10, 1 - 11, frontispiece, text-figs. 1 - 3, pl. 1." type="journal article" year="1980">Sakai, 1980</bibRefCitation>
</taxonomicName>
, described from two specimens (
<typeStatus id="281BE3F0FF9EA407FE22D6C3FE25ABF9" box="[394,494,1507,1531]" pageId="5" pageNumber="62" type="holotype">holotype</typeStatus>
male 110.0 × 66.0 mm and 
<typeStatus id="281BE3F0FF9EA407FC82D6C4FC41ABFE" box="[810,906,1508,1532]" pageId="5" pageNumber="62" type="paratype">paratype</typeStatus>
female 100.0 × 62.0 mm) from Mie Prefecture in Japan. Unfortunately, comparisons between the two species can only be made based on Sakai’s publication, as the whereabouts of the only two known specimens of 
<taxonomicName id="30A026D1FF9EA407FCA0D50CFC56A841" ID-CoL="PFT4" box="[776,925,1580,1603]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9EA407FCA0D50CFC56A841" box="[776,925,1580,1603]" italics="true" pageId="5" pageNumber="62">C. yamasitae</emphasis>
</taxonomicName>
is unknown. They are neither present in the various Japanese museums where Sakai’s material are known to be deposited, nor in Senckenberg Museum in Frankfurt where a good part of his collection was posthumously donated. The specimens may be lost, mislabeled, or misplaced in one of these museums. Sakai provided a colour photograph of 
<taxonomicName id="30A026D1FF9EA407FBC8D5B8FB32A8AD" ID-CoL="PFT4" box="[1120,1273,1688,1711]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9EA407FBC8D5B8FB32A8AD" box="[1120,1273,1688,1711]" italics="true" pageId="5" pageNumber="62">C. yamasitae</emphasis>
</taxonomicName>
(
<bibRefCitation id="933120A3FF9EA407FAA6D5B7FEAAA8D1" author="Sakai" pageId="5" pageNumber="62" refString="Sakai, T. (1980) New species of crabs of the families Lithodidae and Calappidae. Researches on Crustacea, 10, 1 - 11, frontispiece, text-figs. 1 - 3, pl. 1." type="journal article" year="1980">Sakai 1980: frontispiece fig. 2</bibRefCitation>
) although we do not know if it is an accurate representation of its live colour. The 
<typeStatus id="281BE3F0FF9EA407FAB2D59BFAB7A8D1" box="[1306,1404,1723,1747]" pageId="5" pageNumber="62" type="holotype">holotype</typeStatus>
as figured by 
<bibRefCitation id="933120A3FF9EA407FEB3D5FFFD51A8F5" author="Sakai" box="[283,666,1759,1783]" pageId="5" pageNumber="62" refString="Sakai, T. (1980) New species of crabs of the families Lithodidae and Calappidae. Researches on Crustacea, 10, 1 - 11, frontispiece, text-figs. 1 - 3, pl. 1." type="journal article" year="1980">
Sakai (1980: frontispiece 
<figureCitation id="6F9B41D7FF9EA407FDE2D5FFFD59A8F5" box="[586,658,1759,1783]" captionStart="FIGURE 2" captionStartId="3.[151,250,1959,1982]" captionTargetBox="[203,1384,193,1936]" captionTargetId="figure@3.[203,1386,193,1937]" captionTargetPageId="3" captionText="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="5" pageNumber="62">Fig. 2</figureCitation>
)
</bibRefCitation>
had a carapace that was reddish-orange, with the large tubercles relatively darker in colour. The colouration, presence of mammary-type tubercles on carapace and general form of the chela are shared characters in both species. Their carapace proportions, however, are different; the width to length ratio is 1.6 for 
<taxonomicName id="30A026D1FF9EA407FE23D46CFDD5A961" ID-CoL="PFT4" box="[395,542,1868,1891]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9EA407FE23D46CFDD5A961" box="[395,542,1868,1891]" italics="true" pageId="5" pageNumber="62">C. yamasitae</emphasis>
</taxonomicName>
whereas it is 
<quantity id="3058F0B7FF9EA407FD11D46BFCC3A961" box="[697,776,1867,1891]" metricMagnitude="-2" metricUnit="m" metricValue="4.7752" pageId="5" pageNumber="62" unit="in" value="1.88">1.88 in</quantity>
the 
<typeStatus id="281BE3F0FF9EA407FC90D46BFC51A961" box="[824,922,1867,1891]" pageId="5" pageNumber="62" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="30A026D1FF9EA407FC68D46CFBF1A961" ID-CoL="7T362" box="[960,1082,1867,1891]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9EA407FC68D46CFBF1A961" box="[960,1082,1867,1891]" italics="true" pageId="5" pageNumber="62">C. karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9EA407FBEAD46BFB57A961" bold="true" box="[1090,1180,1867,1891]" pageId="5" pageNumber="62">
<taxonomicNameLabel id="DEE73C3BFF9EA407FBEAD46BFB57A961" box="[1090,1180,1867,1891]" pageId="5" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<quantity id="3058F0B7FF9EA407FB03D46BFB31A961" box="[1195,1274,1867,1891]" metricMagnitude="-2" metricUnit="m" metricValue="5.2578" pageId="5" pageNumber="62" unit="in" value="2.07">2.07 in</quantity>
the 
<typeStatus id="281BE3F0FF9EA407FA83D46BFA5AA961" box="[1323,1425,1867,1891]" pageId="5" pageNumber="62" type="paratype">paratype</typeStatus>
). While this disparity may be due to the much larger sizes of our specimens (CW 110.0 mm, 
<taxonomicName id="30A026D1FF9EA407FB34D450FAE4A985" ID-CoL="PFT4" box="[1180,1327,1904,1927]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9EA407FB34D450FAE4A985" box="[1180,1327,1904,1927]" italics="true" pageId="5" pageNumber="62">C. yamasitae</emphasis>
</taxonomicName>
vs. 
<quantity id="3058F0B7FF9EA407FAF4D44FFF0EA9A9" metricMagnitude="-1" metricUnit="m" metricValue="1.8039999999999998" pageId="5" pageNumber="62" unit="mm" value="180.4">180.4 mm</quantity>
, 
<taxonomicName id="30A026D1FF9EA407FF7BD4B4FE9BA9A9" ID-CoL="7T362" box="[211,336,1939,1963]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="karenae">
<emphasis id="C5D48140FF9EA407FF7BD4B4FE9BA9A9" box="[211,336,1939,1963]" italics="true" pageId="5" pageNumber="62">C. karenae</emphasis>
</taxonomicName>
), it seems rather unlikely. The carapace width to length ratios of adult 
<taxonomicName id="30A026D1FF9EA407FB25D4B3FB25A9A9" ID-CoL="7NRJ9" box="[1165,1262,1939,1963]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="C5D48140FF9EA407FB25D4B3FB25A9A9" box="[1165,1262,1939,1963]" italics="true" pageId="5" pageNumber="62">Calappa</emphasis>
</taxonomicName>
species do not usually vary by more than 0.2 even when considering changes associated with size are considered (unpublished data; based on 10 
<taxonomicName id="30A026D1FF9EA407FEC9D4FCFE17A9F1" ID-CoL="PFQL" box="[353,476,2011,2035]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="calappa">
<emphasis id="C5D48140FF9EA407FEC9D4FCFE17A9F1" box="[353,476,2011,2035]" italics="true" pageId="5" pageNumber="62">C. calappa</emphasis>
</taxonomicName>
adult males with CW greater than 100.0 mm).
</paragraph>
<caption id="A3DF0DDAFF9DA404FF3FD458FB63A9B3" httpUri="https://zenodo.org/record/211202/files/figure.png" pageId="6" pageNumber="63" targetBox="[213,1370,204,1858]" targetPageId="6">
<paragraph id="F71F5D52FF9DA404FF3FD458FB63A9B3" blockId="6.[151,1436,1912,1969]" pageId="6" pageNumber="63">
<emphasis id="C5D48140FF9DA404FF3FD458FED8A98C" bold="true" box="[151,275,1912,1935]" pageId="6" pageNumber="63">FIGURE 5.</emphasis>
<taxonomicName id="30A026D1FF9DA404FEB2D458FE03A98C" ID-CoL="7T362" box="[282,456,1912,1934]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9DA404FEB2D458FE03A98C" box="[282,456,1912,1934]" italics="true" pageId="6" pageNumber="63">Calappa karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9DA404FE78D459FDEAA98C" bold="true" box="[464,545,1913,1934]" pageId="6" pageNumber="63">
<taxonomicNameLabel id="DEE73C3BFF9DA404FE78D459FDEAA98C" box="[464,545,1913,1934]" pageId="6" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, left basal antennal segment; B, C, left G1; D, E, distal part of left G1; F, left G2; G, distal part of left G2. Scale = 1.0 mm.
</paragraph>
</caption>
<paragraph id="F71F5D52FF9CA405FF6FD3B8FDE6AF66" blockId="7.[151,1436,151,1184]" pageId="7" pageNumber="64">
From the colour photograph of 
<taxonomicName id="30A026D1FF9CA405FD92D3B8FD1BAEB2" ID-CoL="PFT4" box="[570,720,152,176]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9CA405FD92D3B8FD1BAEB2" box="[570,720,152,176]" italics="true" pageId="7" pageNumber="64">C. yamasitae</emphasis>
</taxonomicName>
(
<bibRefCitation id="933120A3FF9CA405FD4AD3B7FB8DAEAD" author="Sakai" box="[738,1094,151,175]" pageId="7" pageNumber="64" refString="Sakai, T. (1980) New species of crabs of the families Lithodidae and Calappidae. Researches on Crustacea, 10, 1 - 11, frontispiece, text-figs. 1 - 3, pl. 1." type="journal article" year="1980">Sakai 1980: frontispiece fig. 2</bibRefCitation>
), this species also appears to possess a more depressed hepatic (similar to 
<taxonomicName id="30A026D1FF9CA405FD00D39DFCDBAED6" ID-CoL="696D5" box="[680,784,188,212]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="gallus">
<emphasis id="C5D48140FF9CA405FD00D39DFCDBAED6" box="[680,784,188,212]" italics="true" pageId="7" pageNumber="64">C. gallus</emphasis>
</taxonomicName>
) region than 
<taxonomicName id="30A026D1FF9CA405FC1AD39DFBFBAED6" ID-CoL="7T362" box="[946,1072,188,212]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9CA405FC1AD39DFBFBAED6" box="[946,1072,188,212]" italics="true" pageId="7" pageNumber="64">C. karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9CA405FB92D39CFB5CAED6" bold="true" box="[1082,1175,188,212]" pageId="7" pageNumber="64">
<taxonomicNameLabel id="DEE73C3BFF9CA405FB92D39CFB5CAED6" box="[1082,1175,188,212]" pageId="7" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The curvatures of the antero-lateral margins of both species are different. It is more when compared with convex in 
<taxonomicName id="30A026D1FF9CA405FB1AD3C0FA8EAEFA" ID-CoL="PFT4" box="[1202,1349,224,248]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9CA405FB1AD3C0FA8EAEFA" box="[1202,1349,224,248]" italics="true" pageId="7" pageNumber="64">C. yamasitae</emphasis>
</taxonomicName>
than 
<taxonomicName id="30A026D1FF9CA405FA2AD3C0FF3BAF1E" ID-CoL="7T362" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9CA405FA2AD3C0FF3BAF1E" italics="true" pageId="7" pageNumber="64">C. karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9CA405FF50D224FE98AF1E" bold="true" box="[248,339,260,284]" pageId="7" pageNumber="64">
<taxonomicNameLabel id="DEE73C3BFF9CA405FF50D224FE98AF1E" box="[248,339,260,284]" pageId="7" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, although this is likely due to the relative width differences between the two species. The lateral spines along the clypeiform expansions on the posterolateral margin of the carapace points laterally in 
<taxonomicName id="30A026D1FF9CA405FAB8D208FA50AF42" box="[1296,1435,296,320]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="yamisitae">
<emphasis id="C5D48140FF9CA405FAB8D208FA50AF42" box="[1296,1435,296,320]" italics="true" pageId="7" pageNumber="64">C. yamisitae</emphasis>
</taxonomicName>
but anteriorly in 
<taxonomicName id="30A026D1FF9CA405FEFAD26DFE07AF66" ID-CoL="7T362" box="[338,460,332,356]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9CA405FEFAD26DFE07AF66" box="[338,460,332,356]" italics="true" pageId="7" pageNumber="64">C. karenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEE73C3BFF9CA405FE7BD26CFDE6AF66" box="[467,557,332,356]" pageId="7" pageNumber="64" rank="species">
<emphasis id="C5D48140FF9CA405FE7BD26CFDE6AF66" bold="true" box="[467,557,332,356]" pageId="7" pageNumber="64">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="F71F5D52FF9CA405FF6FD24FFC55AC86" blockId="7.[151,1436,151,1184]" pageId="7" pageNumber="64">
The dorsal crest of the chela was also described as having 9 or 10 long teeth in 
<taxonomicName id="30A026D1FF9CA405FBEFD250FB10AF8A" ID-CoL="PFT4" box="[1095,1243,368,392]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9CA405FBEFD250FB10AF8A" box="[1095,1243,368,392]" italics="true" pageId="7" pageNumber="64">C. yamasitae</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="933120A3FF9CA405FABED24FFA53AF85" author="Sakai" box="[1302,1432,367,391]" pageId="7" pageNumber="64" refString="Sakai, T. (1980) New species of crabs of the families Lithodidae and Calappidae. Researches on Crustacea, 10, 1 - 11, frontispiece, text-figs. 1 - 3, pl. 1." type="journal article" year="1980">Sakai 1980</bibRefCitation>
: 7), but there are only six such teeth in 
<taxonomicName id="30A026D1FF9CA405FDFAD2B5FD06AFAE" ID-CoL="7T362" box="[594,717,404,428]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9CA405FDFAD2B5FD06AFAE" box="[594,717,404,428]" italics="true" pageId="7" pageNumber="64">C. karenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEE73C3BFF9CA405FD7DD2B4FCFBAFAE" box="[725,816,404,428]" pageId="7" pageNumber="64" rank="species">
<emphasis id="C5D48140FF9CA405FD7DD2B4FCFBAFAE" bold="true" box="[725,816,404,428]" pageId="7" pageNumber="64">sp. nov.</emphasis>
</taxonomicNameLabel>
, and even counting the two low teeth on the proximal part of the palm, there are a total of eight (
<figureCitation id="6F9B41D7FF9CA405FDC4D297FD65AFCD" box="[620,686,439,463]" captionStart="FIGURE 4" captionStartId="5.[151,250,1162,1185]" captionTargetBox="[151,1435,193,1140]" captionTargetId="figure@5.[151,1436,193,1141]" captionTargetPageId="5" captionText="FIGURE 4. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, outer view of right chela, carpus and merus; B, outer view of left chela; C, dorsal view of merus and carpus of left cheliped; D, frontal view showing third maxillipeds, front, orbit, antennules and antennae." httpUri="https://zenodo.org/record/211201/files/figure.png" pageId="7" pageNumber="64">Fig. 4</figureCitation>
A, B). The teeth of the dorsal crest are more acute in 
<taxonomicName id="30A026D1FF9CA405FB53D298FABFAFD2" ID-CoL="7T362" box="[1275,1396,440,464]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9CA405FB53D298FABFAFD2" box="[1275,1396,440,464]" italics="true" pageId="7" pageNumber="64">C. karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9CA405FAD3D298FF03AFF6" bold="true" pageId="7" pageNumber="64">
<taxonomicNameLabel id="DEE73C3BFF9CA405FAD3D298FF03AFF6" pageId="7" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; they are higher and have narrower bases than those seen in 
<taxonomicName id="30A026D1FF9CA405FC2ED2FDFBD0AFF6" ID-CoL="PFT4" box="[902,1051,477,500]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9CA405FC2ED2FDFBD0AFF6" box="[902,1051,477,500]" italics="true" pageId="7" pageNumber="64">C. yamasitae</emphasis>
</taxonomicName>
. The front also does not protrude very much in 
<taxonomicName id="30A026D1FF9CA405FE92D120FE04AC1A" ID-CoL="PFT4" box="[314,463,512,536]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9CA405FE92D120FE04AC1A" box="[314,463,512,536]" italics="true" pageId="7" pageNumber="64">C. yamasitae</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="933120A3FF9CA405FDA7D2DFFD58AC15" author="Sakai" box="[527,659,511,535]" pageId="7" pageNumber="64" refString="Sakai, T. (1980) New species of crabs of the families Lithodidae and Calappidae. Researches on Crustacea, 10, 1 - 11, frontispiece, text-figs. 1 - 3, pl. 1." type="journal article" year="1980">Sakai 1980</bibRefCitation>
: frontispiece fig. 2, text fig. 2a) while it does so prominently in 
<taxonomicName id="30A026D1FF9CA405FA2AD120FF3BAC3E" ID-CoL="7T362" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9CA405FA2AD120FF3BAC3E" italics="true" pageId="7" pageNumber="64">C. karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9CA405FF5FD104FE9BAC3E" bold="true" box="[247,336,548,572]" pageId="7" pageNumber="64">
<taxonomicNameLabel id="DEE73C3BFF9CA405FF5FD104FE9BAC3E" box="[247,336,548,572]" pageId="7" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="6F9B41D7FF9CA405FEF7D104FE67AC3E" box="[351,428,548,572]" captionStart="FIGURE 1" captionStartId="2.[151,250,1979,2002]" captionTargetBox="[151,1435,1065,1958]" captionTargetId="figure@2.[151,1436,1055,1958]" captionTargetPageId="2" captionText="FIGURE 1. Calappa karenae sp. nov. Paratype male (89.61 mm) (USNM 1150291), Guam. Colour in life, photograph by Roy Kropp." httpUri="https://zenodo.org/record/211198/files/figure.png" pageId="7" pageNumber="64">Figs. 1</figureCitation>
, 
<figureCitation id="6F9B41D7FF9CA405FE12D104FE03AC3E" box="[442,456,548,572]" captionStart="FIGURE 2" captionStartId="3.[151,250,1959,1982]" captionTargetBox="[203,1384,193,1936]" captionTargetId="figure@3.[203,1386,193,1937]" captionTargetPageId="3" captionText="FIGURE 2. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. Differently angled dorsal views to show features and convexity of the carapace." httpUri="https://zenodo.org/record/211199/files/figure.png" pageId="7" pageNumber="64">2</figureCitation>
A, B). 
<bibRefCitation id="933120A3FF9CA405FDBCD104FD0AAC3E" author="Sakai" box="[532,705,548,572]" pageId="7" pageNumber="64" refString="Sakai, T. (1980) New species of crabs of the families Lithodidae and Calappidae. Researches on Crustacea, 10, 1 - 11, frontispiece, text-figs. 1 - 3, pl. 1." type="journal article" year="1980">Sakai (1980: 6)</bibRefCitation>
described the front of 
<taxonomicName id="30A026D1FF9CA405FC16D105FB9AAC3E" ID-CoL="PFT4" box="[958,1105,549,572]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9CA405FC16D105FB9AAC3E" box="[958,1105,549,572]" italics="true" pageId="7" pageNumber="64">C. yamasitae</emphasis>
</taxonomicName>
as “consisting of two median obtuse teeth, which are separated medially by a U-shaped sinus”. The frontal margin of 
<taxonomicName id="30A026D1FF9CA405FBDFD168FB3AAC62" ID-CoL="7T362" box="[1143,1265,584,608]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="karenae">
<emphasis id="C5D48140FF9CA405FBDFD168FB3AAC62" box="[1143,1265,584,608]" italics="true" pageId="7" pageNumber="64">C. karenae</emphasis>
</taxonomicName>
comprises two relatively sharp teeth, separated by a deeper U-shaped sulcus instead.
</paragraph>
<paragraph id="F71F5D52FF9CA405FF61D1B0FAE7AD35" blockId="7.[151,1436,151,1184]" pageId="7" pageNumber="64">
More importantly, the male abdominal somite 6 of 
<taxonomicName id="30A026D1FF9CA405FCB6D1B0FC7FACAA" ID-CoL="PFT4" box="[798,948,656,680]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9CA405FCB6D1B0FC7FACAA" box="[798,948,656,680]" italics="true" pageId="7" pageNumber="64">C. yamasitae</emphasis>
</taxonomicName>
is more transversely rectangular, with the telson evenly triangular, and the lateral margins of both structures are convex (
<bibRefCitation id="933120A3FF9CA405FBA9D194FAC1ACCE" author="Sakai" box="[1025,1290,692,716]" pageId="7" pageNumber="64" refString="Sakai, T. (1980) New species of crabs of the families Lithodidae and Calappidae. Researches on Crustacea, 10, 1 - 11, frontispiece, text-figs. 1 - 3, pl. 1." type="journal article" year="1980" yearSuffix="b">Sakai 1980: text fig. 3b</bibRefCitation>
). In contrast, the male somite 6 is quadrate, with curved, concave lateral margins and the telson more elongated, with straight lateral margins in 
<taxonomicName id="30A026D1FF9CA405FEC9D1DDFE10AD16" ID-CoL="7T362" box="[353,475,764,788]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9CA405FEC9D1DDFE10AD16" box="[353,475,764,788]" italics="true" pageId="7" pageNumber="64">C. karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9CA405FE4AD1DCFDF0AD16" bold="true" box="[482,571,764,788]" pageId="7" pageNumber="64">
<taxonomicNameLabel id="DEE73C3BFF9CA405FE4AD1DCFDF0AD16" box="[482,571,764,788]" pageId="7" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="6F9B41D7FF9CA405FDE2D1DCFD46AD16" box="[586,653,764,788]" captionStart="FIGURE 3" captionStartId="4.[151,250,1934,1957]" captionTargetBox="[224,1364,193,1899]" captionTargetId="figure@4.[224,1366,193,1900]" captionTargetPageId="4" captionText="FIGURE 3. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, frontal view of carapace; B, ventral view showing anterior thoracic sternites, distal abdominal somites and telson; C, ventral view showing posterior thoracic sternites and proximal abdominal somites." httpUri="https://zenodo.org/record/211200/files/figure.png" pageId="7" pageNumber="64">Fig. 3</figureCitation>
B). The G1 is also relatively more slender in 
<taxonomicName id="30A026D1FF9CA405FB2DD1DDFB34AD16" ID-CoL="7T362" box="[1157,1279,764,788]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9CA405FB2DD1DDFB34AD16" box="[1157,1279,764,788]" italics="true" pageId="7" pageNumber="64">C. karenae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEE73C3BFF9CA405FAAED1DCFA94AD16" box="[1286,1375,764,788]" pageId="7" pageNumber="64" rank="species">
<emphasis id="C5D48140FF9CA405FAAED1DCFA94AD16" bold="true" box="[1286,1375,764,788]" pageId="7" pageNumber="64">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="6F9B41D7FF9CA405FAC6D1DCFF6EAD35" captionStart="FIGURE 5" captionStartId="6.[151,250,1912,1935]" captionTargetBox="[213,1370,204,1858]" captionTargetId="figure@6.[200,1387,193,1891]" captionTargetPageId="6" captionText="FIGURE 5. Calappa karenae sp. nov. Holotype male (180.44 by 95.92 mm) (USNM 1150290), Guam. A, left basal antennal segment; B, C, left G 1; D, E, distal part of left G 1; F, left G 2; G, distal part of left G 2. Scale = 1.0 mm." httpUri="https://zenodo.org/record/211202/files/figure.png" pageId="7" pageNumber="64">Fig. 5</figureCitation>
B, C), with the median and basal parts proportionately stouter in 
<taxonomicName id="30A026D1FF9CA405FCD4D000FBC5AD3A" ID-CoL="PFT4" box="[892,1038,800,824]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="yamasitae">
<emphasis id="C5D48140FF9CA405FCD4D000FBC5AD3A" box="[892,1038,800,824]" italics="true" pageId="7" pageNumber="64">C. yamasitae</emphasis>
</taxonomicName>
(
<bibRefCitation id="933120A3FF9CA405FBB5D03FFAEFAD35" author="Sakai" box="[1053,1316,799,824]" pageId="7" pageNumber="64" refString="Sakai, T. (1980) New species of crabs of the families Lithodidae and Calappidae. Researches on Crustacea, 10, 1 - 11, frontispiece, text-figs. 1 - 3, pl. 1." type="journal article" year="1980" yearSuffix="c">Sakai 1980: text fig. 3c</bibRefCitation>
)
</paragraph>
<paragraph id="F71F5D52FF9CA405FF61D064FA59AA0D" blockId="7.[151,1436,151,1184]" pageId="7" pageNumber="64">
The new species also resembles species in the 
<taxonomicName id="30A026D1FF9CA405FCA5D065FB92AD5E" ID-CoL="PFRG" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[781,1113,836,860]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="C5D48140FF9CA405FCA5D065FC53AD5E" box="[781,920,837,860]" italics="true" pageId="7" pageNumber="64">C. japonica</emphasis>
<bibRefCitation id="933120A3FF9CA405FC0ED064FB92AD5E" author="Ortmann" box="[934,1113,836,860]" pageId="7" pageNumber="64" refString="Ortmann, A. E. (1892) Die Decapoda-Krebe des Strassburger Museum. Theil 5, Die Abtheilungen Hippidea, Dromiidea und Oxystomata, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum sufbewahrten Formen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588." type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
</taxonomicName>
, group, which includes 
<taxonomicName id="30A026D1FF9CA405FA2AD065FDBFAD7D" ID-CoL="PFR6" authority="Alcock &amp; Anderson, 1894" authorityName="Alcock &amp; Anderson" authorityYear="1894" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="exanthematosa">
<emphasis id="C5D48140FF9CA405FA2AD065FEF4AD82" italics="true" pageId="7" pageNumber="64">C. exanthematosa</emphasis>
<bibRefCitation id="933120A3FF9CA405FEEFD047FDBFAD7D" author="Alcock" box="[327,628,871,895]" pageId="7" pageNumber="64" refString="Alcock, A. &amp; Anderson, A. R. (1894) Natural History notes from H. M. Indian marine survey steamer &quot; Investigator &quot; Ser. II. No. 17. List of the shore and shallow water Brachyura collected during the season 1893 - 1894. Journal of the Asiatic Society of Bengal, 63 (2), 197 - 209." type="journal article" year="1894">Alcock &amp; Anderson, 1894</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="30A026D1FF9CA405FD18D048FC26AD7D" ID-CoL="PFQ9" authority="Lai &amp; Ng, 2006" authorityName="Lai &amp; Ng" authorityYear="2006" box="[688,1005,871,897]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="C5D48140FF9CA405FD18D048FCFAAD82" box="[688,817,872,896]" italics="true" pageId="7" pageNumber="64">C. africana</emphasis>
<bibRefCitation id="933120A3FF9CA405FC91D047FC26AD7D" author="Lai" box="[825,1005,871,897]" pageId="7" pageNumber="64" refString="Lai, J. C. Y. &amp; Ng, P. K. L. (2006) A new species of Calappa Weber, 1795 (Crustacea: Decapoda: Calappidae) from East and South Africa. Zootaxa, 1358, 39 - 48." type="journal article" year="2006">Lai &amp; Ng, 2006</bibRefCitation>
</taxonomicName>
(see Ng 
<emphasis id="C5D48140FF9CA405FBFCD049FB46AD82" box="[1108,1165,872,896]" italics="true" pageId="7" pageNumber="64">et al.</emphasis>
2011), especially in the general form of the mammary-like granules on the carapace. Other than the different colour patterns, 
<taxonomicName id="30A026D1FF9CA405FA88D0ADFA57ADA6" ID-CoL="7T362" box="[1312,1436,908,932]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9CA405FA88D0ADFA57ADA6" box="[1312,1436,908,932]" italics="true" pageId="7" pageNumber="64">C. karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9CA405FF3FD090FF3BADCA" bold="true" box="[151,240,944,968]" pageId="7" pageNumber="64">
<taxonomicNameLabel id="DEE73C3BFF9CA405FF3FD090FF3BADCA" box="[151,240,944,968]" pageId="7" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from 
<taxonomicName id="30A026D1FF9CA405FE5DD090FDB3ADCA" ID-CoL="PFRG" box="[501,632,944,968]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="C5D48140FF9CA405FE5DD090FDB3ADCA" box="[501,632,944,968]" italics="true" pageId="7" pageNumber="64">C. japonica</emphasis>
</taxonomicName>
, 
<taxonomicName id="30A026D1FF9CA405FD2ED090FC85ADCA" ID-CoL="PFR6" box="[646,846,944,968]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="exanthematosa">
<emphasis id="C5D48140FF9CA405FD2ED090FC85ADCA" box="[646,846,944,968]" italics="true" pageId="7" pageNumber="64">C. exanthematosa</emphasis>
</taxonomicName>
and 
<taxonomicName id="30A026D1FF9CA405FC2DD090FBCFADCA" ID-CoL="PFQ9" box="[901,1028,944,968]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="C5D48140FF9CA405FC2DD090FBCFADCA" box="[901,1028,944,968]" italics="true" pageId="7" pageNumber="64">C. africana</emphasis>
</taxonomicName>
by its much broader carapace, more deeply clefted distal margin of the first maxilliped endopod, the rounded rather than triangular and compressed proepistome, and proportionately broader groove on the pterygostomian lobe (cf. 
<bibRefCitation id="933120A3FF9CA405FB81D0D7FB1DAA0D" author="Lai" box="[1065,1238,1015,1041]" pageId="7" pageNumber="64" refString="Lai, J. C. Y. &amp; Ng, P. K. L. (2006) A new species of Calappa Weber, 1795 (Crustacea: Decapoda: Calappidae) from East and South Africa. Zootaxa, 1358, 39 - 48." type="journal article" year="2006">Lai &amp; Ng 2006</bibRefCitation>
; Ng 
<emphasis id="C5D48140FF9CA405FAA4D0D9FA8FAA12" box="[1292,1348,1016,1040]" italics="true" pageId="7" pageNumber="64">et al.</emphasis>
2011).
</paragraph>
<paragraph id="F71F5D52FF9CA405FF61D73CFA4DAA9D" blockId="7.[151,1436,151,1184]" pageId="7" pageNumber="64">
The use of deep-water traps in Guam and the rest of the Mariana Is have already uncovered many new and/or rare species of homolids, poupiniids, inachids and mathildellids (
<bibRefCitation id="933120A3FF9CA405FCD0D71FFBE9AA55" author="Eldredge" box="[888,1058,1087,1111]" pageId="7" pageNumber="64" refString="Eldredge, L. G. (1980) Two species of Homola (Dromiacea, Homolidae) from Guam. Micronesica, 16 (2), 271 - 277." type="journal article" year="1980">Eldredge 1980</bibRefCitation>
; 
<bibRefCitation id="933120A3FF9CA405FB87D71FFA84AA55" author="Williams" box="[1071,1359,1087,1111]" pageId="7" pageNumber="64" refString="Williams, A. B. &amp; Moffitt, R. B. (1991) Crabs from the Mariana Archipelago: Bothromaia griffini new genus and species (Brachyura: Majidae), and remarks on Poupinia hirsuta Guinot (Homoloidea, Poupiniidae). Proceedings of the Biological Society of Washington, 104 (3), 569 - 582, figs. 1 - 5, Table 1." type="journal article" year="1991">Williams &amp; Moffitt 1991</bibRefCitation>
; 
<bibRefCitation id="933120A3FF9CA405FAF4D761FED1AA7E" author="Ng" pageId="7" pageNumber="64" refString="Ng, P. K. L. &amp; Wang, C. - H. (2002) Notes on two species of deep-water porter crabs of the genus Lamoha (Crustacea: Decapoda: Brachyura: Homolidae) from the Pacific. Journal of the National Taiwan Museum, 2001, 54 (2), 13 - 24." type="journal article" year="2002">Ng &amp; Wang 2002</bibRefCitation>
; 
<bibRefCitation id="933120A3FF9CA405FE8DD744FDDAAA7E" author="Crosnier" box="[293,529,1124,1149]" pageId="7" pageNumber="64" refString="Crosnier, A. &amp; Ng, P. K. L. (2004) Remarques sur le genre Intesius (Crustacea, Decapoda, Brachyura, Goneplacidae) et description de deux especes nouvelles. Zoosystema, 26 (2), 263 - 277." type="journal article" year="2004">Crosnier &amp; Ng 2004</bibRefCitation>
). The present discovery of 
<taxonomicName id="30A026D1FF9CA405FCE8D744FBCAAA7E" ID-CoL="7T362" box="[832,1025,1124,1148]" class="Malacostraca" family="Calappidae" genus="Calappa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="64" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="C5D48140FF9CA405FCE8D744FBCAAA7E" box="[832,1025,1124,1148]" italics="true" pageId="7" pageNumber="64">Calappa karenae</emphasis>
</taxonomicName>
<emphasis id="C5D48140FF9CA405FBA0D744FBA9AA7E" bold="true" box="[1032,1122,1124,1148]" pageId="7" pageNumber="64">
<taxonomicNameLabel id="DEE73C3BFF9CA405FBA0D744FBA9AA7E" box="[1032,1122,1124,1148]" pageId="7" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
attests to the value of using trapping in areas with deep steep cliffs which cannot be sampled by trawls or dredges (see 
<bibRefCitation id="933120A3FF9CA405FB27D7A7FAB1AA9D" author="Mendoza" box="[1167,1402,1159,1184]" pageId="7" pageNumber="64" refString="Mendoza, J. C. E., Naruse, T., Tan, S. - H., Chan, T. - Y., Richer De Forges, B. &amp; Ng, P. K. L. (2010) Case studies on decapod crustaceans from the Philippines reveal deep, steep underwater slopes as prime habitats for ' rare' species. Biological Conservation, 19, 575 - 586." type="journal article" year="2010">
Mendoza 
<emphasis id="C5D48140FF9CA405FB56D7A9FAFCAAA2" box="[1278,1335,1160,1184]" italics="true" pageId="7" pageNumber="64">et al.</emphasis>
2010
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>