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<document id="BD07C7260B67389A305FC10B6CF79D9A" ID-CLB-Dataset="22806" ID-DOI="10.11646/zootaxa.4407.1.2" ID-GBIF-Dataset="8e0aac3a-6556-4b96-a67e-fce9f0db0ba9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3766290" ID-ZooBank="B45E9ABD-CB52-4EE7-91D6-783E8E958AC5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587827908598" checkinUser="plazi" docAuthor="Short, Andrew Edward Z. &amp; Girón, Jennifer C." docDate="2018" docId="676A87E0FFB7B15E2392FB2D47900F17" docLanguage="en" docName="zootaxa.4407.1.2.pdf.imf" docOrigin="Zootaxa 4407 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helochares" docType="key" docVersion="6" lastPageNumber="49" masterDocId="9B53FF98FFA4B14A2305FFE3420C0E4A" masterDocTitle="Review of the Helochares (Hydrobaticus) MacLeay of the New World (Coleoptera: Hydrophilidae: Acidocerinae)" masterLastPageNumber="50" masterPageNumber="29" pageNumber="48" updateTime="1698835862916" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="61FDF5726100819E7D7F2321272CD4F1">Review of the Helochares (Hydrobaticus) MacLeay of the New World (Coleoptera: Hydrophilidae: Acidocerinae)</mods:title>
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<mods:namePart id="A0B9AD1A535AE9F70349A8560537318F">Short, Andrew Edward Z.</mods:namePart>
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<mods:namePart id="349B3E85201BDC1CA2CD1801A49B7365">Girón, Jennifer C.</mods:namePart>
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<mods:title id="7158328BFDC56403FB57102BA9D20879">Zootaxa</mods:title>
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<mods:date id="1AEB6ADDB65D0FA2B938EF3D4854C442">2018</mods:date>
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<mods:number id="ADB15B647AD10A9FE8A2AC753D0E4FB8">2018-04-09</mods:number>
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<mods:number id="E97B3CBB39228AF6461CE7FDC2C6747E">4407</mods:number>
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<treatment id="676A87E0FFB7B15E2392FB2D47900F17" ID-DOI="http://doi.org/10.5281/zenodo.3803384" ID-GBIF-Taxon="163743967" ID-Zenodo-Dep="3803384" LSID="urn:lsid:plazi:treatment:676A87E0FFB7B15E2392FB2D47900F17" httpUri="http://treatment.plazi.org/id/676A87E0FFB7B15E2392FB2D47900F17" lastPageId="20" lastPageNumber="49" pageId="19" pageNumber="48">
<subSubSection id="A7D9657DFFB7B1592392FB2D461C0AA2" box="[151,1040,1230,1256]" pageId="19" pageNumber="48" type="nomenclature">
<paragraph id="EF7C36F6FFB7B1592392FB2D461C0AA2" blockId="19.[151,1040,1230,1256]" box="[151,1040,1230,1256]" pageId="19" pageNumber="48">
<heading id="B434819AFFB7B1592392FB2D461C0AA2" bold="true" box="[151,1040,1230,1256]" fontSize="11" level="1" pageId="19" pageNumber="48" reason="1">
<emphasis id="DDB7EAE4FFB7B1592392FB2D461C0AA2" bold="true" box="[151,1040,1230,1256]" pageId="19" pageNumber="48">
Key to the New World species of the
<taxonomicName id="28C34D75FFB7B1592165FB2D40E70AA2" ID-CoL="8VY5S" box="[608,747,1230,1256]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="DDB7EAE4FFB7B1592165FB2D40E70AA2" bold="true" box="[608,747,1230,1256]" italics="true" pageId="19" pageNumber="48">Helochares</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="28C34D75FFB7B1592068FB2D461C0AA2" authorityName="MacLeay" authorityYear="1871" box="[877,1040,1230,1256]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="subGenus" subGenus="Hydrobaticus">
<emphasis id="DDB7EAE4FFB7B1592068FB2D461C0AA2" bold="true" box="[877,1040,1230,1256]" italics="true" pageId="19" pageNumber="48">Hydrobaticus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A7D9657DFFB7B15E2392FAF747900F17" lastPageId="20" lastPageNumber="49" pageId="19" pageNumber="48" type="key">
<keyStep id="54372E53FFB7B1592392FAF747900B2F" pageId="19" pageNumber="48">
<paragraph id="EF7C36F6FFB7B1592392FAF747970B0D" blockId="19.[151,1436,1300,2026]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392FAF747970B0D" pageId="19" pageNumber="48">
1. Elytra with distinct rows of serial punctures, sometimes deeply impressed into grooves (e.g.,
<figureCitation id="77F82A73FFB7B1592753FAF746AD0B60" box="[1110,1185,1300,1322]" captionStart="FIGURE 2" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[151,1436,265,1900]" captionTargetId="figure@6.[151,1436,265,1900]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. championi Sharp, (C) H. nexus n. sp., (D) H. zamora n. sp., (E) H. sp. A." figureDoi="http://doi.org/10.5281/zenodo.3766294" httpUri="https://zenodo.org/record/3766294/files/figure.png" pageId="19" pageNumber="48">Fig. 2A</figureCitation>
, C–E). Size variable (
<quantity id="283B9B13FFB7B1592677FAF743220B0D" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="6.6" metricValueMin="4.2" pageId="19" pageNumber="48" unit="mm" value="5.4" valueMax="6.6" valueMin="4.2">4.2– 6.6 mm</quantity>
). Widely distributed............................................................................. 2
</keyLead>
</paragraph>
<paragraph id="EF7C36F6FFB7B1592392FAAC47900B2F" blockId="19.[151,1436,1300,2026]" box="[151,1436,1359,1381]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392FAAC47900B2F" box="[151,1436,1359,1381]" pageId="19" pageNumber="48">
- Elytra without rows of serial punctures (
<figureCitation id="77F82A73FFB7B1592166FAAC40BB0B2F" box="[611,695,1359,1381]" captionStart="FIGURE 2" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[151,1436,265,1900]" captionTargetId="figure@6.[151,1436,265,1900]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. championi Sharp, (C) H. nexus n. sp., (D) H. zamora n. sp., (E) H. sp. A." figureDoi="http://doi.org/10.5281/zenodo.3766294" httpUri="https://zenodo.org/record/3766294/files/figure.png" pageId="19" pageNumber="48">Figs. 2B</figureCitation>
,
<figureCitation id="77F82A73FFB7B15921C4FAAC40DC0B2F" box="[705,720,1359,1381]" captionStart="FIGURE 4" captionStartId="10.[151,250,1284,1306]" captionTargetBox="[151,1436,193,1262]" captionTargetId="figure@10.[151,1436,193,1262]" captionTargetPageId="10" captionText="FIGURE 4. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. politus n. sp., (B) H. laevis n. sp." figureDoi="http://doi.org/10.5281/zenodo.3766298" httpUri="https://zenodo.org/record/3766298/files/figure.png" pageId="19" pageNumber="48">4</figureCitation>
). Size larger than
<quantity id="283B9B13FFB7B1592078FAAC41C50B2E" box="[893,969,1359,1381]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="19" pageNumber="48" unit="mm" value="5.5">5.5 mm</quantity>
.
<collectingCountry id="97D47666FFB7B15920D7FAAC46170B2F" box="[978,1051,1359,1381]" name="Mexico" pageId="19" pageNumber="48">Mexico</collectingCountry>
and Central America.............. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="54372E53FFB7B1592392FA8F47900BBD" pageId="19" pageNumber="48">
<paragraph id="EF7C36F6FFB7B1592392FA8F47900BF7" blockId="19.[151,1436,1300,2026]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392FA8F47900BF7" pageId="19" pageNumber="48">
2. Abdominal pubescence sparse and not uniformly distributed on each ventrite (
<figureCitation id="77F82A73FFB7B15920D9FA8F46270BC8" box="[988,1067,1388,1410]" captionStart="FIGURE 6" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[152,1436,581,1933]" captionTargetId="figure@12.[151,1436,581,1966]" captionTargetPageId="12" captionText="FIGURE 6. Abdomens of Helochares (Hydrobaticus) spp. (A) H. normatus, (B) H. maculicollis, (C) H. nexus n. sp. (Panama, ventrites paler on lateral thirds), (D) H. nexus n. sp. (Panama, uniformly pale ventrites)." figureDoi="http://doi.org/10.5281/zenodo.3766302" httpUri="https://zenodo.org/record/3766302/files/figure.png" pageId="19" pageNumber="48">Fig. 6A</figureCitation>
). Abdominal ventrites uniformly very dark brown to almost black in coloration. Elytra with rows of serial punctures not impressed into striae............................................................................................
<taxonomicName id="28C34D75FFB7B159275FFA4447490BF7" ID-CoL="8WSR6" authority="(LeConte)" baseAuthorityName="LeConte" baseAuthorityYear="1861" box="[1114,1349,1447,1469]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="normatus">
<emphasis id="DDB7EAE4FFB7B159275FFA4447490BF7" bold="true" box="[1114,1349,1447,1469]" pageId="19" pageNumber="48">
<emphasis id="DDB7EAE4FFB7B159275FFA4446DB0BF7" bold="true" box="[1114,1239,1447,1469]" italics="true" pageId="19" pageNumber="48">H. normatus</emphasis>
(LeConte)
</emphasis>
</taxonomicName>
<emphasis id="DDB7EAE4FFB7B159264EFA4447900BF7" box="[1355,1436,1447,1469]" italics="true" pageId="19" pageNumber="48">(in part)</emphasis>
</keyLead>
</paragraph>
<paragraph id="EF7C36F6FFB7B1592392FA2647900BBD" blockId="19.[151,1436,1300,2026]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392FA2647900BBD" pageId="19" pageNumber="48">
- Abdominal pubescence dense and uniform, not appearing sparse or absent on anterior half of each ventrite (e.g.
<figureCitation id="77F82A73FFB7B1592622FA27476C0B90" box="[1319,1376,1476,1498]" captionStart="FIGURE 6" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[152,1436,581,1933]" captionTargetId="figure@12.[151,1436,581,1966]" captionTargetPageId="12" captionText="FIGURE 6. Abdomens of Helochares (Hydrobaticus) spp. (A) H. normatus, (B) H. maculicollis, (C) H. nexus n. sp. (Panama, ventrites paler on lateral thirds), (D) H. nexus n. sp. (Panama, uniformly pale ventrites)." figureDoi="http://doi.org/10.5281/zenodo.3766302" httpUri="https://zenodo.org/record/3766302/files/figure.png" pageId="19" pageNumber="48">Fig. 6</figureCitation>
B–D). Abdominal ventrites variously colored. Elytra with rows of serial punctures impressed into striae or not................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="54372E53FFB7B1592392FA1C47970827" pageId="19" pageNumber="48">
<paragraph id="EF7C36F6FFB7B1592392FA1C47900878" blockId="19.[151,1436,1300,2026]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392FA1C47900878" pageId="19" pageNumber="48">
3 Size larger (&gt;
<quantity id="283B9B13FFB7B1592260FA1C43B8085E" box="[357,436,1535,1557]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="19" pageNumber="48" unit="mm" value="5.8">5.8 mm</quantity>
). Elytral serial punctures not impressed into striae or only very weakly so; absent or almost appearing absent anteromedially (
<figureCitation id="77F82A73FFB7B15922C4F9FF40010878" box="[449,525,1564,1586]" captionStart="FIGURE 2" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[151,1436,265,1900]" captionTargetId="figure@6.[151,1436,265,1900]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. championi Sharp, (C) H. nexus n. sp., (D) H. zamora n. sp., (E) H. sp. A." figureDoi="http://doi.org/10.5281/zenodo.3766294" httpUri="https://zenodo.org/record/3766294/files/figure.png" pageId="19" pageNumber="48">Fig. 2A</figureCitation>
)..............................................................
<emphasis id="DDB7EAE4FFB7B15927F8F9FF47900878" bold="true" box="[1277,1436,1564,1586]" pageId="19" pageNumber="48">
<taxonomicName id="28C34D75FFB7B15927F8F9FF476C0878" ID-CoL="8XQKL" authority="Short &amp; Girón, 2018" authorityName="Short &amp; Girón" authorityYear="2018" box="[1277,1376,1564,1586]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="trujillo" status="sp. nov.">
<emphasis id="DDB7EAE4FFB7B15927F8F9FF476C0878" bold="true" box="[1277,1376,1564,1586]" italics="true" pageId="19" pageNumber="48">H. trujillo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C684579FFFB7B1592663F9FF47900878" box="[1382,1436,1564,1586]" pageId="19" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="EF7C36F6FFB7B1592392F9D947970827" blockId="19.[151,1436,1300,2026]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392F9D947970827" pageId="19" pageNumber="48">- Size variable (4.2–6.0 mm). Elytra with rows of serial punctures impressed into striae or not; rows distinctly present anteromedially............................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="54372E53FFB7B1592392F9974790088F" pageId="19" pageNumber="48">
<paragraph id="EF7C36F6FFB7B1592392F997479708ED" blockId="19.[151,1436,1300,2026]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392F997479708ED" pageId="19" pageNumber="48">4 Abdominal ventrites pale in color, either uniformly yellowish brown or yellowish brown on lateral thirds with median third darker brown......................................................................................... 5</keyLead>
</paragraph>
<paragraph id="EF7C36F6FFB7B1592392F94C4790088F" blockId="19.[151,1436,1300,2026]" box="[151,1436,1711,1733]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392F94C4790088F" box="[151,1436,1711,1733]" pageId="19" pageNumber="48">- Abdominal ventrites uniformly dark brown to almost black in color..............................................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="54372E53FFB7B1592392F92F47900970" pageId="19" pageNumber="48">
<paragraph id="EF7C36F6FFB7B1592392F92F4790094A" blockId="19.[151,1436,1300,2026]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392F92F4790094A" pageId="19" pageNumber="48">
5 Parameres of aedeagus truncate at apex, with outer corners widely rounded (
<figureCitation id="77F82A73FFB7B15920C4F92F41F708A8" box="[961,1019,1740,1762]" captionStart="FIGURE 7" captionStartId="13.[151,250,1200,1222]" captionTargetBox="[347,1229,206,1166]" captionTargetId="figure@13.[340,1247,193,1179]" captionTargetPageId="13" captionText="FIGURE 7. Aedeagi of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. zamora n. sp., (C) H. maculicollis (Florida), (D) H. maculicollis (Kansas), (E) H. maculicollis (Pennsylvania), (F) H. nexus n. sp. (Venezuela), (G) H. nexus n. sp. (Panama), (H) H. nexus n. sp. (Ecuador), (I) H. politus n. sp., (J) H. laevis n. sp., (K) H. championi." figureDoi="http://doi.org/10.5281/zenodo.3766304" httpUri="https://zenodo.org/record/3766304/files/figure.png" pageId="19" pageNumber="48">Fig. 7</figureCitation>
F–H). More southern distribution (
<collectingCountry id="97D47666FFB7B159264DF92F479408A8" box="[1352,1432,1740,1762]" name="Panama" pageId="19" pageNumber="48">Panama</collectingCountry>
,
<collectingCountry id="97D47666FFB7B15923E4F90A433F08B5" box="[225,307,1769,1791]" name="Ecuador" pageId="19" pageNumber="48">Ecuador</collectingCountry>
,
<collectingCountry id="97D47666FFB7B1592239F90A43AC08B5" box="[316,416,1769,1791]" name="Venezuela" pageId="19" pageNumber="48">Venezuela</collectingCountry>
).......................................................................
<emphasis id="DDB7EAE4FFB7B1592603F9094790094A" bold="true" box="[1286,1436,1770,1792]" pageId="19" pageNumber="48">
<taxonomicName id="28C34D75FFB7B1592603F909476C094A" ID-CoL="8XQKD" authority="Short &amp; Girón, 2018" authorityName="Short &amp; Girón" authorityYear="2018" box="[1286,1376,1770,1792]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="nexus" status="sp. nov.">
<emphasis id="DDB7EAE4FFB7B1592603F909476C094A" bold="true" box="[1286,1376,1770,1792]" italics="true" pageId="19" pageNumber="48">H. nexus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C684579FFFB7B1592663F9094790094A" box="[1382,1436,1770,1792]" pageId="19" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="EF7C36F6FFB7B1592392F8E447900970" blockId="19.[151,1436,1300,2026]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392F8E447900970" pageId="19" pageNumber="48">
<emphasis id="DDB7EAE4FFB7B1592392F8E442930957" bold="true" box="[151,159,1799,1821]" pageId="19" pageNumber="48">-</emphasis>
Parameres of aedeagus with apex oblique to truncate, with outer corners widely rounded (
<figureCitation id="77F82A73FFB7B159274EF8E4468A0957" box="[1099,1158,1799,1821]" captionStart="FIGURE 8" captionStartId="15.[151,250,1934,1956]" captionTargetBox="[360,1216,1351,1885]" captionTargetId="figure@15.[340,1247,1345,1913]" captionTargetPageId="15" captionText="FIGURE 8. Aedeagi of Helochares (Hydrobaticus) normatus. (A) Oregon, (B) California, (C) Arizona, (D) Mexico, (E) Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.3766306" httpUri="https://zenodo.org/record/3766306/files/figure.png" pageId="19" pageNumber="48">Fig. 8</figureCitation>
). More northern distribution (this rarer form known from
<collectingCountry id="97D47666FFB7B15922F0F8C7404C0970" box="[501,576,1828,1850]" name="Mexico" pageId="19" pageNumber="48">Mexico</collectingCountry>
)............................................
<taxonomicName id="28C34D75FFB7B159275FF8C747490970" ID-CoL="8WSR6" authority="(LeConte)" baseAuthorityName="LeConte" baseAuthorityYear="1861" box="[1114,1349,1828,1850]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="normatus">
<emphasis id="DDB7EAE4FFB7B159275FF8C747490970" bold="true" box="[1114,1349,1828,1850]" pageId="19" pageNumber="48">
<emphasis id="DDB7EAE4FFB7B159275FF8C746DB0970" bold="true" box="[1114,1239,1828,1850]" italics="true" pageId="19" pageNumber="48">H. normatus</emphasis>
(LeConte)
</emphasis>
</taxonomicName>
<emphasis id="DDB7EAE4FFB7B159264EF8C747900970" box="[1355,1436,1828,1850]" italics="true" pageId="19" pageNumber="48">(in part)</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="54372E53FFB7B1592392F8A24797093E" pageId="19" pageNumber="48">
<paragraph id="EF7C36F6FFB7B1592392F8A247970912" blockId="19.[151,1436,1300,2026]" box="[151,1435,1857,1880]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392F8A247970912" box="[151,1435,1857,1880]" pageId="19" pageNumber="48">
6 Clypeus pale in coloration, not darkened medially (
<figureCitation id="77F82A73FFB7B15921C9F8A24115091D" box="[716,793,1857,1879]" captionStart="FIGURE 5" captionStartId="11.[151,250,1936,1958]" captionTargetBox="[151,1436,493,1914]" captionTargetId="figure@11.[151,1436,493,1914]" captionTargetPageId="11" captionText="FIGURE 5. Heads of Helochares (Hydrobaticus) spp. (A) H. maculicollis (Pennsylvania), (B) H. normatus (California), (C) H. normatus (Arizona), (D) H. nexus n. sp., (E) H. zamora n. sp., (F) H. trujillo n. sp., (G) H. championi, (H) H. sp. A., (I) H. politus n. sp." figureDoi="http://doi.org/10.5281/zenodo.3766300" httpUri="https://zenodo.org/record/3766300/files/figure.png" pageId="19" pageNumber="48">Fig. 5A</figureCitation>
). Eastern North America.............
<taxonomicName id="28C34D75FFB7B15927AFF8A147970912" ID-CoL="3KGWQ" authority="Mulsant" authorityName="Mulsant" authorityYear="1844" box="[1194,1435,1858,1880]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="maculicollis">
<emphasis id="DDB7EAE4FFB7B15927AFF8A147970912" bold="true" box="[1194,1435,1858,1880]" pageId="19" pageNumber="48">
<emphasis id="DDB7EAE4FFB7B15927AFF8A1474D0912" bold="true" box="[1194,1345,1858,1880]" italics="true" pageId="19" pageNumber="48">H. maculicollis</emphasis>
Mulsant
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EF7C36F6FFB7B1592392F8BC4797093E" blockId="19.[151,1436,1300,2026]" box="[151,1435,1887,1909]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392F8BC4797093E" box="[151,1435,1887,1909]" pageId="19" pageNumber="48">
- Clypeus distinctly darkened medially (
<figureCitation id="77F82A73FFB7B1592156F8BC40B7093E" box="[595,699,1887,1909]" captionStart="FIGURE 5" captionStartId="11.[151,250,1936,1958]" captionTargetBox="[151,1436,493,1914]" captionTargetId="figure@11.[151,1436,493,1914]" captionTargetPageId="11" captionText="FIGURE 5. Heads of Helochares (Hydrobaticus) spp. (A) H. maculicollis (Pennsylvania), (B) H. normatus (California), (C) H. normatus (Arizona), (D) H. nexus n. sp., (E) H. zamora n. sp., (F) H. trujillo n. sp., (G) H. championi, (H) H. sp. A., (I) H. politus n. sp." figureDoi="http://doi.org/10.5281/zenodo.3766300" httpUri="https://zenodo.org/record/3766300/files/figure.png" pageId="19" pageNumber="48">Fig. 5E, H</figureCitation>
). South America............................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="54372E53FFB7B1592392F89E479709FA" pageId="19" pageNumber="48">
<paragraph id="EF7C36F6FFB7B1592392F89E479009D8" blockId="19.[151,1436,1300,2026]" box="[151,1436,1916,1938]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392F89E479009D8" box="[151,1436,1916,1938]" pageId="19" pageNumber="48">
7 Pronotum almost entirely darkly colored (
<figureCitation id="77F82A73FFB7B1592177F89F40CC09D8" box="[626,704,1916,1938]" captionStart="FIGURE 2" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[151,1436,265,1900]" captionTargetId="figure@6.[151,1436,265,1900]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. championi Sharp, (C) H. nexus n. sp., (D) H. zamora n. sp., (E) H. sp. A." figureDoi="http://doi.org/10.5281/zenodo.3766294" httpUri="https://zenodo.org/record/3766294/files/figure.png" pageId="19" pageNumber="48">Fig. 2D</figureCitation>
). Size smaller (
<quantity id="283B9B13FFB7B1592056F89F41C709D8" box="[851,971,1916,1938]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="4.8" metricValueMin="4.7" pageId="19" pageNumber="48" unit="mm" value="4.75" valueMax="4.8" valueMin="4.7">4.7–4.8 mm</quantity>
).
<collectingCountry id="97D47666FFB7B15920DEF89F462709D8" box="[987,1067,1916,1938]" name="Ecuador" pageId="19" pageNumber="48">Ecuador</collectingCountry>
................
<emphasis id="DDB7EAE4FFB7B15927FDF89F479009D8" bold="true" box="[1272,1436,1916,1938]" pageId="19" pageNumber="48">
<taxonomicName id="28C34D75FFB7B15927FDF89F476C09D8" ID-CoL="8Y4FN" authority="Short &amp; Girón, 2018" authorityName="Short &amp; Girón" authorityYear="2018" box="[1272,1376,1916,1938]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="zamora" status="sp. nov.">
<emphasis id="DDB7EAE4FFB7B15927FDF89F476C09D8" bold="true" box="[1272,1376,1916,1938]" italics="true" pageId="19" pageNumber="48">H. zamora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C684579FFFB7B1592663F89F479009D8" box="[1382,1436,1916,1938]" pageId="19" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="EF7C36F6FFB7B1592392F879479709FA" blockId="19.[151,1436,1300,2026]" box="[151,1435,1946,1968]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392F879479709FA" box="[151,1435,1946,1968]" pageId="19" pageNumber="48">
- Pronotum only darkened on central disc (
<figureCitation id="77F82A73FFB7B1592169F87940BB09FA" box="[620,695,1946,1968]" captionStart="FIGURE 2" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[151,1436,265,1900]" captionTargetId="figure@6.[151,1436,265,1900]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus images of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. championi Sharp, (C) H. nexus n. sp., (D) H. zamora n. sp., (E) H. sp. A." figureDoi="http://doi.org/10.5281/zenodo.3766294" httpUri="https://zenodo.org/record/3766294/files/figure.png" pageId="19" pageNumber="48">Fig. 2E</figureCitation>
). Size larger (
<quantity id="283B9B13FFB7B159203EF87941B809E5" box="[827,948,1946,1968]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.5" metricValueMin="5.2" pageId="19" pageNumber="48" unit="mm" value="5.35" valueMax="5.5" valueMin="5.2">5.2–5.5 mm</quantity>
).
<collectingCountry id="97D47666FFB7B15920C1F87941FC09E5" box="[964,1008,1946,1967]" name="Peru" pageId="19" pageNumber="48">Peru</collectingCountry>
............................
<emphasis id="DDB7EAE4FFB7B159264CF879479709FA" bold="true" box="[1353,1435,1946,1968]" pageId="19" pageNumber="48">
<emphasis id="DDB7EAE4FFB7B159264CF879475009FA" bold="true" box="[1353,1372,1946,1968]" italics="true" pageId="19" pageNumber="48">H</emphasis>
. sp. A
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="54372E53FFB7B15E2392F85447900EE7" lastPageId="20" lastPageNumber="49" pageId="19" pageNumber="48">
<paragraph id="EF7C36F6FFB7B1592392F854479709A0" blockId="19.[151,1436,1300,2026]" pageId="19" pageNumber="48">
<keyLead id="543295C3FFB7B1592392F854479709A0" pageId="19" pageNumber="48">
8 Clypeus pale in coloration, not darkened medially (
<figureCitation id="77F82A73FFB7B15921D6F854412D0987" box="[723,801,1975,1997]" captionStart="FIGURE 5" captionStartId="11.[151,250,1936,1958]" captionTargetBox="[151,1436,493,1914]" captionTargetId="figure@11.[151,1436,493,1914]" captionTargetPageId="11" captionText="FIGURE 5. Heads of Helochares (Hydrobaticus) spp. (A) H. maculicollis (Pennsylvania), (B) H. normatus (California), (C) H. normatus (Arizona), (D) H. nexus n. sp., (E) H. zamora n. sp., (F) H. trujillo n. sp., (G) H. championi, (H) H. sp. A., (I) H. politus n. sp." figureDoi="http://doi.org/10.5281/zenodo.3766300" httpUri="https://zenodo.org/record/3766300/files/figure.png" pageId="19" pageNumber="48">Fig. 5G</figureCitation>
). Aedeagus with median lobe broadly rounded, with only a small “pinched” apex (
<figureCitation id="77F82A73FFB7B1592287F83743C209A0" box="[386,462,2004,2026]" captionStart="FIGURE 7" captionStartId="13.[151,250,1200,1222]" captionTargetBox="[347,1229,206,1166]" captionTargetId="figure@13.[340,1247,193,1179]" captionTargetPageId="13" captionText="FIGURE 7. Aedeagi of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. zamora n. sp., (C) H. maculicollis (Florida), (D) H. maculicollis (Kansas), (E) H. maculicollis (Pennsylvania), (F) H. nexus n. sp. (Venezuela), (G) H. nexus n. sp. (Panama), (H) H. nexus n. sp. (Ecuador), (I) H. politus n. sp., (J) H. laevis n. sp., (K) H. championi." figureDoi="http://doi.org/10.5281/zenodo.3766304" httpUri="https://zenodo.org/record/3766304/files/figure.png" pageId="19" pageNumber="48">Fig. 7K</figureCitation>
)...............................................................
<taxonomicName id="28C34D75FFB7B15927CAF837479709A0" ID-CoL="3KGWC" authority="Sharp" authorityName="Sharp" authorityYear="1882" box="[1231,1435,2004,2026]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="DDB7EAE4FFB7B15927CAF837479709A0" bold="true" box="[1231,1435,2004,2026]" pageId="19" pageNumber="48">
<emphasis id="DDB7EAE4FFB7B15927CAF837475A09A0" bold="true" box="[1231,1366,2004,2026]" italics="true" pageId="19" pageNumber="48">H. championi</emphasis>
Sharp
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="EF7C36F6FFB0B15E2392FF7447900EE7" blockId="20.[151,1436,151,349]" box="[151,1436,151,173]" pageId="20" pageNumber="49">
<keyLead id="543295C3FFB0B15E2392FF7447900EE7" box="[151,1436,151,173]" pageId="20" pageNumber="49">
- Clypeus distinctly darkened medially (
<figureCitation id="77F82A73FFB0B15E2156FF74409A0EE7" box="[595,662,151,173]" captionStart="FIGURE 5" captionStartId="11.[151,250,1936,1958]" captionTargetBox="[151,1436,493,1914]" captionTargetId="figure@11.[151,1436,493,1914]" captionTargetPageId="11" captionText="FIGURE 5. Heads of Helochares (Hydrobaticus) spp. (A) H. maculicollis (Pennsylvania), (B) H. normatus (California), (C) H. normatus (Arizona), (D) H. nexus n. sp., (E) H. zamora n. sp., (F) H. trujillo n. sp., (G) H. championi, (H) H. sp. A., (I) H. politus n. sp." figureDoi="http://doi.org/10.5281/zenodo.3766300" httpUri="https://zenodo.org/record/3766300/files/figure.png" pageId="20" pageNumber="49">Fig. 5I</figureCitation>
). Aedeagus with median lobe tapered to a point at apex (
<figureCitation id="77F82A73FFB0B15E27B2FF7446E30EE6" box="[1207,1263,151,173]" captionStart="FIGURE 7" captionStartId="13.[151,250,1200,1222]" captionTargetBox="[347,1229,206,1166]" captionTargetId="figure@13.[340,1247,193,1179]" captionTargetPageId="13" captionText="FIGURE 7. Aedeagi of Helochares (Hydrobaticus) spp. (A) H. trujillo n. sp., (B) H. zamora n. sp., (C) H. maculicollis (Florida), (D) H. maculicollis (Kansas), (E) H. maculicollis (Pennsylvania), (F) H. nexus n. sp. (Venezuela), (G) H. nexus n. sp. (Panama), (H) H. nexus n. sp. (Ecuador), (I) H. politus n. sp., (J) H. laevis n. sp., (K) H. championi." figureDoi="http://doi.org/10.5281/zenodo.3766304" httpUri="https://zenodo.org/record/3766304/files/figure.png" pageId="20" pageNumber="49">Fig. 7</figureCitation>
I– J)......... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="54372E53FFB0B15E2392FF5747900F17" pageId="20" pageNumber="49">
<paragraph id="EF7C36F6FFB0B15E2392FF57479A0F4F" blockId="20.[151,1436,151,349]" pageId="20" pageNumber="49">
<keyLead id="543295C3FFB0B15E2392FF57479A0F4F" pageId="20" pageNumber="49">
9 Aedeagus with apex of parameres rounded (
<figureCitation id="77F82A73FFB0B15E218DFF5740C00E80" box="[648,716,180,202]" captionStart="FIGURE 6" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[152,1436,581,1933]" captionTargetId="figure@12.[151,1436,581,1966]" captionTargetPageId="12" captionText="FIGURE 6. Abdomens of Helochares (Hydrobaticus) spp. (A) H. normatus, (B) H. maculicollis, (C) H. nexus n. sp. (Panama, ventrites paler on lateral thirds), (D) H. nexus n. sp. (Panama, uniformly pale ventrites)." figureDoi="http://doi.org/10.5281/zenodo.3766302" httpUri="https://zenodo.org/record/3766302/files/figure.png" pageId="20" pageNumber="49">Fig. 6I</figureCitation>
); lateral outer margins of parameres parallel, slightly constricted near midlength; maximum width of aedeagus at basal half, nearly as wide as maximum width of aedeagus at apical half...................................................................................................
<emphasis id="DDB7EAE4FFB0B15E27FDFF0C479A0F4F" bold="true" box="[1272,1430,239,261]" pageId="20" pageNumber="49">
<taxonomicName id="28C34D75FFB0B15E27FDFF0C47560F4F" ID-CoL="8Y4DP" authority="Short &amp; Girón, 2018" authorityName="Short &amp; Girón" authorityYear="2018" box="[1272,1370,239,261]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="politus" status="sp. nov.">
<emphasis id="DDB7EAE4FFB0B15E27FDFF0C47560F4F" bold="true" box="[1272,1370,239,261]" italics="true" pageId="20" pageNumber="49">H. politus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C684579FFFB0B15E2665FF0C479A0F4F" box="[1376,1430,239,261]" pageId="20" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="EF7C36F6FFB0B15E2392FEEE47900F17" blockId="20.[151,1436,151,349]" pageId="20" pageNumber="49">
<keyLead id="543295C3FFB0B15E2392FEEE47900F17" pageId="20" pageNumber="49">
- Aedeagus with apex of parameres truncate (
<figureCitation id="77F82A73FFB0B15E2181FEEF40C50F68" box="[644,713,268,290]" captionStart="FIGURE 6" captionStartId="12.[151,250,1965,1987]" captionTargetBox="[152,1436,581,1933]" captionTargetId="figure@12.[151,1436,581,1966]" captionTargetPageId="12" captionText="FIGURE 6. Abdomens of Helochares (Hydrobaticus) spp. (A) H. normatus, (B) H. maculicollis, (C) H. nexus n. sp. (Panama, ventrites paler on lateral thirds), (D) H. nexus n. sp. (Panama, uniformly pale ventrites)." figureDoi="http://doi.org/10.5281/zenodo.3766302" httpUri="https://zenodo.org/record/3766302/files/figure.png" pageId="20" pageNumber="49">Fig. 6J</figureCitation>
); lateral outer margins of parameres parallel along basal two thirds, slightly constricted at two thirds, diverging along apical third; maximum width of aedeagus at basal half, clearly narrower than maximum width of aedeagus at apical half...........................................................
<emphasis id="DDB7EAE4FFB0B15E260CFEA447900F17" bold="true" box="[1289,1436,327,349]" pageId="20" pageNumber="49">
<taxonomicName id="28C34D75FFB0B15E260CFEA4476C0F17" ID-CoL="8Y4CJ" authority="Short &amp; Girón, 2018" authorityName="Short &amp; Girón" authorityYear="2018" box="[1289,1376,327,349]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="49" phylum="Arthropoda" rank="species" species="laevis" status="sp. nov.">
<emphasis id="DDB7EAE4FFB0B15E260CFEA4476C0F17" bold="true" box="[1289,1376,327,349]" italics="true" pageId="20" pageNumber="49">H. laevis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C684579FFFB0B15E2663FEA447900F17" box="[1382,1436,327,349]" pageId="20" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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