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<document id="D443F1B9467720F3B33D2F66986A45F8" ID-DOI="10.5281/zenodo.275907" ID-GBIF-Dataset="d5d27825-82a5-4aaf-b47d-35d24b33aaaa" ID-ISSN="1175-5326" ID-Zenodo-Dep="275907" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460189977646" checkinUser="plazi" docAuthor="Lott, Derek A." docDate="2010" docId="4D54A843FFB51A6F0694FC5DEB5EFEA9" docLanguage="en" docName="zt02402p051.pdf" docOrigin="Zootaxa 2402" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Acylophorus tshuapensis Lott, new species" docType="treatment" docVersion="5" lastPageNumber="15" masterDocId="B16DD03BFFB81A610603FFAFEE40FFEE" masterDocTitle="The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa" masterLastPageNumber="51" masterPageNumber="1" pageNumber="14" updateTime="1698236417163" updateUser="plazi">
<mods:mods id="55DE4D9B6969A2EE46C4B2493CE0D84B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="35B096982DC29A09A62149BA1671E729">The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub-Saharan Africa</mods:title>
</mods:titleInfo>
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<mods:namePart id="16BBC48A960F1F1CB4602AB3B65AEDDD">Lott, Derek A.</mods:namePart>
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<mods:date id="D4E6FDD9A7F418E034EA8385F1746111">2010</mods:date>
<mods:detail id="68AEB10BEEEF7C050D4F8A2E89033AD7" type="volume">
<mods:number id="D904CF8855C33508E463A298F33B6998">2402</mods:number>
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<mods:classification id="7E0F4DA3AA2087D8BE24777C69AA7538">journal article</mods:classification>
<mods:identifier id="766565D9D967A700C97360118EDE757F" type="DOI">10.5281/zenodo.275907</mods:identifier>
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<mods:identifier id="19114879DED3CCDBB782B68214E77B8B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="911247507C9A94C9BAF831037FC42C47" type="Zenodo-Dep">275907</mods:identifier>
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<treatment id="4D54A843FFB51A6F0694FC5DEB5EFEA9" ID-DOI="http://doi.org/10.5281/zenodo.6196017" ID-GBIF-Taxon="127655316" ID-Zenodo-Dep="6196017" LSID="urn:lsid:plazi:treatment:4D54A843FFB51A6F0694FC5DEB5EFEA9" httpUri="http://treatment.plazi.org/id/4D54A843FFB51A6F0694FC5DEB5EFEA9" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="8DE74ADEFFB51A6C0694FC5DEFC5FBC1" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="C5421955FFB51A6C0694FC5DECE0FBE2" blockId="13.[151,672,1010,1071]" box="[151,672,1010,1036]" pageId="13" pageNumber="14">
<heading id="9E0AAE39FFB51A6C0694FC5DECE0FBE2" bold="true" box="[151,672,1010,1036]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="F789C547FFB51A6C0694FC5DECE0FBE2" bold="true" box="[151,672,1010,1036]" pageId="13" pageNumber="14">
<taxonomicName id="02FD62D6FFB51A6C0694FC5DEC43FBE2" ID-CoL="652X9" authority="Lott" authorityName="Lott" box="[151,515,1010,1036]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tshuapensis" status="sp. nov.">
<emphasis id="F789C547FFB51A6C0694FC5DEF84FBE2" bold="true" box="[151,452,1010,1036]" italics="true" pageId="13" pageNumber="14">Acylophorus tshuapensis</emphasis>
Lott
</taxonomicName>
,
<taxonomicNameLabel id="ECBA783CFFB51A6C0413FC5DECE0FBE2" box="[528,672,1010,1036]" pageId="13" pageNumber="14" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C5421955FFB51A6C0694FBBAEFC5FBC1" blockId="13.[151,672,1010,1071]" box="[151,389,1045,1071]" pageId="13" pageNumber="14">
(
<figureCitation id="5DC605D0FFB51A6C069CFBBAEEABFBC1" box="[159,235,1045,1071]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,272,1933,1957]" captionTargetBox="[171,1408,165,1908]" captionTargetId="figure@5.[156,1441,151,1941]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Forebodies of species in the Acylophorus orientalis group. 1, A. orientalis Fauvel; 2, A. nitens new species; 3, A. rossii Bordoni, 4. A. bumbunae Bordoni; 5, A. capensis Cameron; 6, A. dankalensis Bordoni; 7, A. tshuapensis new species; 8, A. makhoreae new species." httpUri="https://zenodo.org/record/275908/files/figure.png" pageId="13" pageNumber="14">Figs 7</figureCitation>
,
<figureCitation id="5DC605D0FFB51A6C06FBFBBAEF58FBC1" box="[248,280,1045,1071]" captionStart="FIGURES 51 – 76" captionStartId="36.[151,271,1818,1842]" captionTargetBox="[176,1425,305,1773]" captionTargetId="figure@36.[155,1440,293,1797]" captionTargetPageId="36" captionText="FIGURES 51 – 76. Terminal segments of the maxilliary palpi of Acylophorus species. 51, A. orientalis Fauvel; 52, A. nitens new species; 53, A. rossii Bordoni, 54, A. bumbunae Bordoni; 55, A. capensis Cameron; 56, A. dankalensis Bordoni; 57, A. tshuapensis new species; 58, A. makhoreae new species; 59, A. densipennis Bernhauer; 60, A. antennalis Cameron; 61, A. collarti Cameron; 62. A. micans new species; 63, A. allardi Levasseur; 64, A. congoensis Cameron; 65, A. ziloensis Levasseur; 66, A. mareei Bernhauer; 67, A. salifi new species; 68, A. lomaensis Levasseur; 69. A. minor new species; 70, A. trigonocephalus Cameron; 71, A. setiger new species; 72, A. usambarae Bernhauer; 73, A. grandis Bernhauer; 74, A. bafutensis Levasseur; 75, A. kambuiensis Bordoni, 76. A. lualabaensis Levasseur." httpUri="https://zenodo.org/record/275914/files/figure.png" pageId="13" pageNumber="14">57</figureCitation>
,
<figureCitation id="5DC605D0FFB51A6C0720FBBAEF03FBC1" box="[291,323,1045,1071]" captionStart="FIGURES 77 – 96" captionStartId="37.[151,271,1853,1877]" captionTargetBox="[209,1389,189,1828]" captionTargetId="figure@37.[168,1428,165,1848]" captionTargetPageId="37" captionText="FIGURES 77 – 96. Antennae of Acylophorus species. 77, A. orientalis Fauvel; 78, A. nitens new species; 79, A. rossii Bordoni, 80, A. bumbunae Bordoni; 81, A. capensis Cameron; 82, A. dankalensis Bordoni; 83, A. tshuapensis new species; 84, A. makhoreae new species; 85, A. densipennis Bernhauer; 86, A. antennalis Cameron; 87, A. collarti Cameron; 88. A. micans new species; 89, A. allardi Levasseur; 90, A. congoensis Cameron; 91, A. ziloensis Levasseur; 92, A. mareei Bernhauer; 93, A. methneri Bernhauer; 94, A. salifi new species; 95, A. lomaensis Levasseur; 96. A. minor new species." httpUri="https://zenodo.org/record/275915/files/figure.png" pageId="13" pageNumber="14">83</figureCitation>
,
<figureCitation id="5DC605D0FFB51A6C0753FBBAEF3CFBC1" box="[336,380,1045,1071]" captionStart="FIGURES 109 – 116" captionStartId="39.[151,271,1945,1969]" captionTargetBox="[226,1362,196,1910]" captionTargetId="figure@39.[207,1395,159,1942]" captionTargetPageId="39" captionText="FIGURES 109 – 116. Aedeagi of species in the Acylophorus orientalis group. 109, A. orientalis Fauvel; 110, A. nitens new species; 111, A. rossii Bordoni, 112. A. bumbunae Bordoni; 113, A. capensis Cameron; 114, A. dankalensis Bordoni; 115, A. tshuapensis new species; 116, A. makhoreae new species." httpUri="https://zenodo.org/record/275917/files/figure.png" pageId="13" pageNumber="14">115</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFB51A6C0694FBF0EA90F8F4" pageId="13" pageNumber="14" type="description">
<paragraph id="C5421955FFB51A6C0694FBF0EF0CFB2A" blockId="13.[151,1437,1119,2004]" pageId="13" pageNumber="14">
<emphasis id="F789C547FFB51A6C0694FBF0EF7BFB97" bold="true" box="[151,315,1119,1145]" pageId="13" pageNumber="14">Description.</emphasis>
Length
<quantity id="0205B4B0FFB51A6C07A4FBCFEFAAFB94" box="[423,490,1120,1146]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="14" unit="mm" value="8.0">8mm</quantity>
. Head black. Pronotum chestnut brown. Elytra brown. Abdomen dark and iridescent. Legs pale. Antennae pale with segments II to VIII slightly infuscated. The maxillary palpi are uniformly pale.
</paragraph>
<paragraph id="C5421955FFB51A6C06C7FB7FEF45FA01" blockId="13.[151,1437,1119,2004]" pageId="13" pageNumber="14">
Head somewhat small (pronotum 1.8x wider than head), as long as wide with rounded, not very pronounced temples and antennal insertion close to front margin, but separated by narrow pigmented strip (
<figureCitation id="5DC605D0FFB51A6C069CFAB5EEA8FADA" box="[159,232,1306,1332]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,272,1933,1957]" captionTargetBox="[171,1408,165,1908]" captionTargetId="figure@5.[156,1441,151,1941]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Forebodies of species in the Acylophorus orientalis group. 1, A. orientalis Fauvel; 2, A. nitens new species; 3, A. rossii Bordoni, 4. A. bumbunae Bordoni; 5, A. capensis Cameron; 6, A. dankalensis Bordoni; 7, A. tshuapensis new species; 8, A. makhoreae new species." httpUri="https://zenodo.org/record/275908/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
). Eyes relatively small. Micro-punctures very sparse. Dense short pubescence behind eyes. Two pairs of interocular setae arising from foveate punctures much closer to eyes than each other. A line of five postocular setae plus an additional short seta on hind margin of eye. Maxillary palpi with terminal segment very elongate, densely pubescent, and symmetric, longer than glabrous penultimate segment which is also elongate (
<figureCitation id="5DC605D0FFB51A6C070DFA1FEF29FA24" box="[270,361,1456,1482]" captionStart="FIGURES 51 – 76" captionStartId="36.[151,271,1818,1842]" captionTargetBox="[176,1425,305,1773]" captionTargetId="figure@36.[155,1440,293,1797]" captionTargetPageId="36" captionText="FIGURES 51 – 76. Terminal segments of the maxilliary palpi of Acylophorus species. 51, A. orientalis Fauvel; 52, A. nitens new species; 53, A. rossii Bordoni, 54, A. bumbunae Bordoni; 55, A. capensis Cameron; 56, A. dankalensis Bordoni; 57, A. tshuapensis new species; 58, A. makhoreae new species; 59, A. densipennis Bernhauer; 60, A. antennalis Cameron; 61, A. collarti Cameron; 62. A. micans new species; 63, A. allardi Levasseur; 64, A. congoensis Cameron; 65, A. ziloensis Levasseur; 66, A. mareei Bernhauer; 67, A. salifi new species; 68, A. lomaensis Levasseur; 69. A. minor new species; 70, A. trigonocephalus Cameron; 71, A. setiger new species; 72, A. usambarae Bernhauer; 73, A. grandis Bernhauer; 74, A. bafutensis Levasseur; 75, A. kambuiensis Bordoni, 76. A. lualabaensis Levasseur." httpUri="https://zenodo.org/record/275914/files/figure.png" pageId="13" pageNumber="14">Fig. 57</figureCitation>
). First segment of antenna as long as next three. Segments I to IX elongate, X transverse (
<figureCitation id="5DC605D0FFB51A6C069CFA7AEEB7FA01" box="[159,247,1493,1519]" captionStart="FIGURES 77 – 96" captionStartId="37.[151,271,1853,1877]" captionTargetBox="[209,1389,189,1828]" captionTargetId="figure@37.[168,1428,165,1848]" captionTargetPageId="37" captionText="FIGURES 77 – 96. Antennae of Acylophorus species. 77, A. orientalis Fauvel; 78, A. nitens new species; 79, A. rossii Bordoni, 80, A. bumbunae Bordoni; 81, A. capensis Cameron; 82, A. dankalensis Bordoni; 83, A. tshuapensis new species; 84, A. makhoreae new species; 85, A. densipennis Bernhauer; 86, A. antennalis Cameron; 87, A. collarti Cameron; 88. A. micans new species; 89, A. allardi Levasseur; 90, A. congoensis Cameron; 91, A. ziloensis Levasseur; 92, A. mareei Bernhauer; 93, A. methneri Bernhauer; 94, A. salifi new species; 95, A. lomaensis Levasseur; 96. A. minor new species." httpUri="https://zenodo.org/record/275915/files/figure.png" pageId="13" pageNumber="14">Fig. 83</figureCitation>
).
</paragraph>
<paragraph id="C5421955FFB51A6C06C7FA55EDD3F921" blockId="13.[151,1437,1119,2004]" pageId="13" pageNumber="14">Pronotum only slightly transverse (1.1x wider than long) with rounded sides and widest in basal half, front margins somewhat produced at sides. Shining with no micro-punctures. One pair of dorsal setae. One pair of lateral setae. Marginal setae long. Elytra transverse (1.6x wider than long) with pubescence arising from relatively fine asperate punctures. Apical fringe of thick bristles longer than the hairs on the rest of the elytra. Abdominal tergites with long semi-erect pubescence arising from asperate punctures that are denser at the base of each tergite. Hairs longer than marginal fringe of bristles.</paragraph>
<paragraph id="C5421955FFB51A6C06C7F975EA90F8F4" blockId="13.[151,1437,1119,2004]" pageId="13" pageNumber="14">
Paramere entire with wide central groove at apex, suggesting two parallel lobes that have become fused together (
<figureCitation id="5DC605D0FFB51A6C0704F8AFEF2DF8F4" box="[263,365,1792,1818]" captionStart="FIGURES 109 – 116" captionStartId="39.[151,271,1945,1969]" captionTargetBox="[226,1362,196,1910]" captionTargetId="figure@39.[207,1395,159,1942]" captionTargetPageId="39" captionText="FIGURES 109 – 116. Aedeagi of species in the Acylophorus orientalis group. 109, A. orientalis Fauvel; 110, A. nitens new species; 111, A. rossii Bordoni, 112. A. bumbunae Bordoni; 113, A. capensis Cameron; 114, A. dankalensis Bordoni; 115, A. tshuapensis new species; 116, A. makhoreae new species." httpUri="https://zenodo.org/record/275917/files/figure.png" pageId="13" pageNumber="14">Fig. 115</figureCitation>
). Median lobe much longer than paramere and strongly expanded at apex.
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFB51A6C06C7F88AEF48F864" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="C5421955FFB51A6C06C7F88AEF48F864" blockId="13.[151,1437,1119,2004]" pageId="13" pageNumber="14">
<emphasis id="F789C547FFB51A6C06C7F88AEFB4F8D1" bold="true" box="[196,500,1829,1855]" pageId="13" pageNumber="14">
<typeStatus id="1A46A7F7FFB51A6C06C7F88AEF41F8D1" box="[196,257,1829,1855]" pageId="13" pageNumber="14">Type</typeStatus>
material.
<typeStatus id="1A46A7F7FFB51A6C0781F88AEFB4F8D1" box="[386,500,1829,1855]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
3: “
<collectionCode id="A3EC8190FFB51A6C042FF88AEC3FF8D1" box="[556,639,1829,1855]" pageId="13" pageNumber="14">COLL</collectionCode>
. MUS.
<collectingCountry id="BDEA59C5FFB51A6C04DAF88AED02F8D1" box="[729,834,1829,1855]" name="Republic of the Congo" pageId="13" pageNumber="14">CONGO</collectingCountry>
Tshuapa: Flandria
<date id="B1433F95FFB51A6C0227F88AEADEF8D1" box="[1060,1182,1829,1855]" pageId="13" pageNumber="14" value="1946-10-18">18-x-1946</date>
Rév. P. Hulstaert / R. DET. UU 5694 /
<taxonomicName id="02FD62D6FFB51A6C076BF8E4EC41F88A" box="[360,513,1867,1892]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="F789C547FFB51A6C076BF8E4EC41F88A" box="[360,513,1867,1892]" italics="true" pageId="13" pageNumber="14">Acylophorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ECBA783CFFB51A6C0409F8E5EC03F88A" box="[522,579,1866,1892]" pageId="13" pageNumber="14" rank="species">sp. n</taxonomicNameLabel>
/
<typeStatus id="1A46A7F7FFB51A6C045EF8E5ECBBF88A" box="[605,763,1866,1892]" pageId="13" pageNumber="14" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="02FD62D6FFB51A6C0507F8E4EA72F88A" box="[772,1074,1867,1892]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tshuapensis" status="sp. nov.">
<emphasis id="F789C547FFB51A6C0507F8E4EA72F88A" box="[772,1074,1867,1892]" italics="true" pageId="13" pageNumber="14">Acylophorus tshuapensis</emphasis>
</taxonomicName>
<emphasis id="F789C547FFB51A6C0238F8E5EAC0F88A" bold="true" box="[1083,1152,1866,1892]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="ECBA783CFFB51A6C0238F8E5EAC0F88A" box="[1083,1152,1866,1892]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
3 det. DA Lott, 2009” (
<collectionCode id="A3EC8190FFB51A6C069CF8DFEEBCF864" box="[159,252,1904,1930]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="14">IRSNB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFB51A6F06C7F83AEF4EFECC" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="distribution">
<paragraph id="C5421955FFB51A6C06C7F83AEF53F83A" blockId="13.[151,1437,1119,2004]" pageId="13" pageNumber="14">
<emphasis id="F789C547FFB51A6C06C7F83AEC61F841" bold="true" box="[196,545,1941,1967]" pageId="13" pageNumber="14">Distribution and bionomics.</emphasis>
Known only from the
<typeStatus id="1A46A7F7FFB51A6C052DF83AEDD5F841" box="[814,917,1941,1967]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
taken in the
<collectingCountry id="BDEA59C5FFB51A6C022FF83AEBDBF841" box="[1068,1435,1941,1967]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">Democratic Republic of Congo</collectingCountry>
(
<figureCitation id="5DC605D0FFB51A6C069CF815EF46F83A" box="[159,262,1978,2004]" captionStart="FIGURE 142" captionStartId="45.[151,260,891,915]" captionTargetBox="[153,1436,186,864]" captionTargetId="figure@45.[151,1436,186,864]" captionTargetPageId="45" captionText="FIGURE 142. Distribution of species in the Acylophorus orientalis group = A. bumbunae Bordoni, = A. tshuapensis new species, = A. makhoreae new species)." httpUri="https://zenodo.org/record/275923/files/figure.png" pageId="13" pageNumber="14">Fig. 142</figureCitation>
).
</paragraph>
<paragraph id="C5421955FFB61A6F06C7FF38EF4EFECC" blockId="14.[151,1436,151,327]" pageId="14" pageNumber="15">
<emphasis id="F789C547FFB61A6F06C7FF38EFF8FF5F" bold="true" box="[196,440,151,177]" pageId="14" pageNumber="15">Comparative notes.</emphasis>
Fairly distinct within the
<taxonomicName id="02FD62D6FFB61A6F04E5FF37ED36FF5F" box="[742,886,152,177]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="F789C547FFB61A6F04E5FF37ED36FF5F" box="[742,886,152,177]" italics="true" pageId="14" pageNumber="15">A. orientalis</emphasis>
</taxonomicName>
species group by virtue of its small eyes, more sinuate front margin to the pronotum and the fused lobes of the paramere. The medial segments of the antennae and the terminal segment of the maxillary palpi are also more elongate than in the other species of the group.
</paragraph>
</subSubSection>
<subSubSection id="8DE74ADEFFB61A6F06C7FE82EB5EFEA9" box="[196,1310,301,327]" pageId="14" pageNumber="15" type="etymology">
<paragraph id="C5421955FFB61A6F06C7FE82EB5EFEA9" blockId="14.[151,1436,151,327]" box="[196,1310,301,327]" pageId="14" pageNumber="15">
<emphasis id="F789C547FFB61A6F06C7FE82EF11FEA9" bold="true" box="[196,337,301,327]" pageId="14" pageNumber="15">Etymology.</emphasis>
The specific name is the masculine form of an adjective meaning “from Tshuapa”.
</paragraph>
</subSubSection>
</treatment>
</document>
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