treatments-xml/data/1A/37/87/1A3787F69D9D92D3FF2EF9DBCD212F8B.xml

<document id="FA9B20943831D608BB325A53E6B43B16" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D9D92D3FF2EF9DBCD212F8B" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidiini Telnov 2020" docType="key" docVersion="7" lastPageNumber="137" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="130" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4793E58B0045FCB40CEF556A582160FD">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
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<mods:namePart id="6121553A5C98A46FFD8BF8E0A0405A75">Telnov, Dmitry</mods:namePart>
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<subSubSection id="DA84656B9D9D92DAFF2EF9DBCCD22EE3" box="[189,1156,1636,1665]" pageId="129" pageNumber="130" type="nomenclature">
<paragraph id="922136E09D9D92DAFF2EF9DBCCD22EE3" blockId="129.[189,1156,1636,1665]" box="[189,1156,1636,1665]" pageId="129" pageNumber="130">
<emphasis id="A0EAEAF29D9D92DAFF2EF9DBCCD22EE3" bold="true" box="[189,1156,1636,1665]" italics="true" pageId="129" pageNumber="130">
Key to the 
<taxonomicName id="559E4D639D9D92DAFEDAF9DAC9BF2EE3" ID-CoL="628FF" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[329,489,1636,1665]" class="Insecta" family="Scarabaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="129" pageNumber="5" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Maechidiini">Maechidiini</taxonomicName>
from the Indo-Australian transition zone and 
<collectingRegion id="505AF8029D9D92DAFBDEF9DACCD22EE3" box="[1101,1156,1636,1665]" country="Indonesia" name="Bali" pageId="129" pageNumber="130">Bali</collectingRegion>
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<subSubSection id="DA84656B9D9D92DAFF2EF927CD222ED6" box="[189,1396,1689,1716]" pageId="129" pageNumber="130" type="description">
<paragraph id="922136E09D9D92DAFF2EF927CD262ED6" blockId="129.[189,1396,1689,1716]" box="[189,1392,1689,1716]" pageId="129" pageNumber="130">This key is generally adapted for males and is mainly based on male features, including genital organs</paragraph>
<paragraph id="922136E09D9D92DAFAE3F924CD222ED6" blockId="129.[189,1396,1689,1716]" box="[1392,1396,1690,1716]" pageId="129" pageNumber="130">.</paragraph>
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<keyLead id="296F95D59D9D92DAFF2EF965CD202F7B" pageId="129" pageNumber="130">1. Punctures of elytral disc ordinary: circular, variably ovoid or hexagonal, annular or not but not elongate linear or sinusoid incision-shaped..................................................................................... 31</keyLead>
</paragraph>
<paragraph id="922136E09D9D92DAFF52F89DCD212FE6" blockId="129.[189,1399,1755,1924]" pageId="129" pageNumber="130">
<keyLead id="296F95D59D9D92DAFF52F89DCD212FE6" pageId="129" pageNumber="130">
– Punctures of elytral disc at least in part conspicuous: elongate linear or sinusoid, incision-shaped (cf. 
<figureCitation id="0AA52A659D9D92DAFE89F8F8CA632F02" box="[282,565,1862,1888]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="129" pageNumber="130">Figs 288, 290–291, 296</figureCitation>
, 
<figureCitation id="0AA52A659D9D92DAFDD2F8F8CAFA2F02" box="[577,684,1862,1888]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="129" pageNumber="130">303, 314</figureCitation>
, 
<figureCitation id="0AA52A659D9D92DAFD2BF8F8CB822F02" box="[696,980,1862,1888]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="129" pageNumber="130">317–320, 323, 328, 331</figureCitation>
, 
<figureCitation id="0AA52A659D9D92DAFC73F8F8CC962F02" box="[992,1216,1862,1888]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="129" pageNumber="130">339–340, 343, 348</figureCitation>
, 
<figureCitation id="0AA52A659D9D92DAFB5FF8F8CD242F03" box="[1228,1394,1862,1889]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="129" pageNumber="130">351, 353, 355</figureCitation>
, etc.) .................................................................................................................................................... 2
</keyLead>
</paragraph>
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<keyStep id="296A2E459D9D92D9FF2EF812CD21290E" lastPageId="130" lastPageNumber="131" pageId="129" pageNumber="130">
<paragraph id="922136E09D9D92DAFF2EF812CD212F8A" blockId="129.[189,1399,1963,2025]" pageId="129" pageNumber="130">
<keyLead id="296F95D59D9D92DAFF2EF812CD212F8A" pageId="129" pageNumber="130">2. Lateral margins of pronotum in dorsal view distinctly rounded (more or less distinctly constricted anteriad and posteriad), without or with prebasal emargination or constriction ............................... 6</keyLead>
</paragraph>
<paragraph id="922136E09D9E92D9FF52FEB7CD21290E" blockId="130.[193,1399,265,364]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF52FEB7CD21290E" pageId="130" pageNumber="131">
– Lateral margins of pronotum in dorsal view slightly sinuous, somewhat widened from anterolateral angles towards base or nearly straight in basal half, not distinctly constricted both anteriad and posteriad (
<figureCitation id="0AA52A659D9E92D9FEF6FEECC986290E" box="[357,464,338,364]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="130" pageNumber="131">Figs 218</figureCitation>
, 
<figureCitation id="0AA52A659D9E92D9FE4FFEECCA15290E" box="[476,579,338,364]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="130" pageNumber="131">226, 241</figureCitation>
) ........................................................................................................... 3
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<keyStep id="296A2E459D9E92D9FF2EFE2ACD21299A" pageId="130" pageNumber="131">
<paragraph id="922136E09D9E92D9FF2EFE2ACD2129B1" blockId="130.[189,1399,404,504]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF2EFE2ACD2129B1" pageId="130" pageNumber="131">
3. Dorsal punctures of forebody comparatively large, distinctly annular; aedeagus not as in 
<figureCitation id="0AA52A659D9E92D9FA92FE2AC94029B0" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="130" pageNumber="131">Figs 646– 648</figureCitation>
..................................................................................................................................................... 4
</keyLead>
</paragraph>
<paragraph id="922136E09D9E92D9FF52FE60CD21299A" blockId="130.[189,1399,404,504]" box="[193,1399,477,504]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF52FE60CD21299A" box="[193,1399,477,504]" pageId="130" pageNumber="131">– Dorsal punctures of forebody comparatively less large, not annular ................................................ 5</keyLead>
</paragraph>
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<keyStep id="296A2E459D9E92D9FF2EFD9ECD212B3F" pageId="130" pageNumber="131">
<paragraph id="922136E09D9E92D9FF2EFD9ECD212AE0" blockId="130.[189,1399,543,861]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF2EFD9ECD212AE0" pageId="130" pageNumber="131">
4. Lateral margin of pronotum with about 20 flat crenulae; elytron without tracks of longitudinal carinae; intervening spaces of elytra not transversely wrinkled; aedeagus unknown ............................................................................................. 
<taxonomicName id="559E4D639D9E92D9FB9CFDD7CD212AE0" ID-CoL="3XFKL" authority="Moser, 1920" authorityName="Moser" authorityYear="1920" box="[1039,1399,616,643]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="parallelicollis">
<emphasis id="A0EAEAF29D9E92D9FB9CFDD7CC8A2AE1" bold="true" box="[1039,1244,616,643]" italics="true" pageId="130" pageNumber="131">M. parallelicollis</emphasis>
<bibRefCitation id="F60F4B119D9E92D9FB70FDD7CD212AE0" author="Moser J." box="[1251,1399,616,643]" pageId="130" pageNumber="131" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922136E09D9E92D9FF2EFD33CD212A8D" blockId="130.[189,1399,543,861]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF2EFD33CD212A8D" pageId="130" pageNumber="131">
– Lateral margin of pronotum with about 19–20 flat crenulae; elytron with vague tracks of longitudinal carinae; intervening spaces of elytra transversely wrinkled; aedeagus as in 
<figureCitation id="0AA52A659D9E92D9FA93FD0FC9402A8D" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="130" pageNumber="131">Figs 646– 648</figureCitation>
............................................................................................................ 
<taxonomicName id="559E4D639D9E92D9FBA8FD68CD212A8D" ID-CoL="3XFJK" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[1083,1399,725,752]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="A0EAEAF29D9E92D9FBA8FD68CC8B2A92" bold="true" box="[1083,1245,725,752]" italics="true" pageId="130" pageNumber="131">M. humeralis</emphasis>
<bibRefCitation id="F60F4B119D9E92D9FB76FD6BCD212A8D" author="Heller K. M." box="[1253,1399,725,751]" pageId="130" pageNumber="131" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922136E09D9E92D9FF52FD44CD212B3F" blockId="130.[189,1399,543,861]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF52FD44CD212B3F" pageId="130" pageNumber="131">
– Lateral margin of pronotum with about 24–25 crenulae; elytron with vague tracks of three longitudinal carinae; intervening spaces of elytra transversely wrinkled; aedeagus as in 
<bibRefCitation id="F60F4B119D9E92D9FBE2FCA0C9652B3F" author="Narakusumo R. P. &amp; Balke M." pageId="130" pageNumber="131" pagination="35 - 50" refId="ref98467" refString="Narakusumo R. P. &amp; Balke M. 2019. Four new species of Epholcis Waterhouse, 1875 (Coleoptera: Scarabaeidae: Melolonthinae: Maechidiini) from the Moluccas, Indonesia. Treubia 46: 35 - 50. https: // doi. org / 10.14203 / treubia. v 46 i 0.3791" type="journal article" year="2019">Narakusumo &amp; Balke (2019</bibRefCitation>
: fig. 1e–f) ................................................ 
<taxonomicName id="559E4D639D9E92D9FC82FCFDCCB82B3F" ID-CoL="7X3VZ" authority="(Narakusumo &amp; Balke, 2019)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Narakusumo &amp; Balke" baseAuthorityYear="2019" box="[785,1262,834,861]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="acutus" status="comb. nov.">
<emphasis id="A0EAEAF29D9E92D9FC82FCFDCBDC2B3F" bold="true" box="[785,906,835,861]" italics="true" pageId="130" pageNumber="131">M. acutus</emphasis>
(
<bibRefCitation id="F60F4B119D9E92D9FC0AFCFCCCB32B3F" author="Narakusumo R. P. &amp; Balke M." box="[921,1253,834,861]" pageId="130" pageNumber="131" pagination="35 - 50" refId="ref98467" refString="Narakusumo R. P. &amp; Balke M. 2019. Four new species of Epholcis Waterhouse, 1875 (Coleoptera: Scarabaeidae: Melolonthinae: Maechidiini) from the Moluccas, Indonesia. Treubia 46: 35 - 50. https: // doi. org / 10.14203 / treubia. v 46 i 0.3791" type="journal article" year="2019">Narakusumo &amp; Balke, 2019</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9E92D9FB66FCFCCD212B3F" box="[1269,1399,834,861]" pageId="130" pageNumber="131" rank="species">comb. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9E92D9FF2EFC3BCD212C37" pageId="130" pageNumber="131">
<paragraph id="922136E09D9E92D9FF2EFC3BCD212B8A" blockId="130.[189,1399,901,1109]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF2EFC3BCD212B8A" pageId="130" pageNumber="131">
5. Anterolateral angles of labroclypeus broadly obtuse, not or slightly produced anteriad (
<figureCitation id="0AA52A659D9E92D9FB6CFC3BCD3E2BC2" box="[1279,1384,901,928]" captionStart="Figs 133–144" captionStartId="140.[189,241,1822,1848]" captionTargetBox="[346,1223,273,1784]" captionTargetId="figure-120@140.[345,1234,265,1799]" captionTargetPageId="140" captionText="Figs 133–144. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 133. M. lapsus sp. nov., holotype, ♂ (BMNH). 134. Same, paratype, ♀ (BMNH). 135. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 136. M. legalovi sp. nov., holotype, ♂ (BMNH). 137. Same, paratype, ♀ (BMNH). 138. M. longipes sp. nov., holotype, ♂ (BMNH). 139. M. lobaticeps Frey, 1969, ♂ (BMNH). 140. Same, ♀. 141. M. mailu sp. nov., holotype, ♂ (NHMB). 142. M. luniceps Fairmaire, 1883, ♂ (BMNH). 143. Same, ♀ (BMNH). 144. M. maleo sp. nov., holotype, ♂ (IRSN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122179" httpUri="https://zenodo.org/record/4122179/files/figure.png" pageId="130" pageNumber="131">Fig. 135</figureCitation>
); elytral setae very inconspicuous, strongly appressed; elytra without metallic shine; aedeagus as in 
<figureCitation id="0AA52A659D9E92D9FF79FC70C9DD2B8A" box="[234,395,974,1000]" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="130" pageNumber="131">Figs 646–648</figureCitation>
.................................................................................... 
<taxonomicName id="559E4D639D9E92D9FC6CFC70CD212B8A" ID-CoL="3XFJX" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[1023,1399,973,1000]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="lineatopunctatus">
<emphasis id="A0EAEAF29D9E92D9FC6CFC70CCA42B8A" bold="true" box="[1023,1266,973,1000]" italics="true" pageId="130" pageNumber="131">M. lineatopunctatus</emphasis>
<bibRefCitation id="F60F4B119D9E92D9FB6AFC70CD212B8A" author="Frey G." box="[1273,1399,974,1000]" pageId="130" pageNumber="131" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922136E09D9E92D9FF52FC4DCD212C37" blockId="130.[189,1399,901,1109]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF52FC4DCD212C37" pageId="130" pageNumber="131">
– Anterolateral angles of labroclypeus subacute, strongly produced anteriad (
<figureCitation id="0AA52A659D9E92D9FBF5FC4CCC982C6E" box="[1126,1230,1010,1036]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="130" pageNumber="131">Fig. 120</figureCitation>
); some elytral setae are longer and stouter than others, suberect, arranged in longidutinal rows; elytra with vague metallic shine; aedeagus not studied................................................................ 
<taxonomicName id="559E4D639D9E92D9FBF8FB85CD212C37" ID-CoL="3XFJD" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[1131,1399,1083,1109]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="gressitti">
<emphasis id="A0EAEAF29D9E92D9FBF8FB85CCA42C37" bold="true" box="[1131,1266,1083,1109]" italics="true" pageId="130" pageNumber="131">M. gressitti</emphasis>
<bibRefCitation id="F60F4B119D9E92D9FB6AFB85CD212C37" author="Frey G." box="[1273,1399,1083,1109]" pageId="130" pageNumber="131" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9E92D9FF2EFBC3CD212D4B" pageId="130" pageNumber="131">
<paragraph id="922136E09D9E92D9FF2EFBC3CD212CDE" blockId="130.[189,1399,1149,1321]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF2EFBC3CD212CDE" pageId="130" pageNumber="131">6. Lateral margins of pronotum crenulate (rarely crenulate-denticulate); male labroclypeus broadly emarginate to subtruncate on anterior margin ................................................................................... 7</keyLead>
</paragraph>
<paragraph id="922136E09D9E92D9FF52FB79CD212D4B" blockId="130.[189,1399,1149,1321]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF52FB79CD212D4B" pageId="130" pageNumber="131">
– Lateral margins of pronotum distinctly denticulate (
<figureCitation id="0AA52A659D9E92D9FCDBFB78CBF92C82" box="[840,943,1222,1248]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="130" pageNumber="131">Fig. 199</figureCitation>
); anterior margin of male labroclypeus deeply U-shaped emarginate on anterior margin, its anterolateral angles strongly protruding anteriad (
<figureCitation id="0AA52A659D9E92D9FF60FAB1C91F2D4B" box="[243,329,1295,1321]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="130" pageNumber="131">Fig. 99</figureCitation>
) .......................................................................................................... 
<taxonomicName id="559E4D639D9E92D9FBF0FAB1CD432D4B" ID-CoL="7X3W5" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1123,1301,1294,1321]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="alesbezdeki" status="sp. nov.">
<emphasis id="A0EAEAF29D9E92D9FBF0FAB1CD432D4B" bold="true" box="[1123,1301,1294,1321]" italics="true" pageId="130" pageNumber="131">M. alesbezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9E92D9FA8FFAB1CD212D4B" box="[1308,1399,1295,1321]" pageId="130" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9E92D9FF2EFAECCD212DBB" pageId="130" pageNumber="131">
<paragraph id="922136E09D9E92D9FF2EFAECCD212DF2" blockId="130.[189,1399,1361,1497]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF2EFAECCD212DF2" pageId="130" pageNumber="131">
7. At least male metatibia distinctly thickened distally, not long and slender; all tarsi with short to moderately long tarsomeres; aedeagus not as in 
<figureCitation id="0AA52A659D9E92D9FC8EFAC8CBE82DF3" box="[797,958,1398,1425]" captionStart="Figs 652–666" captionStartId="175.[189,241,1892,1918]" captionTargetBox="[436,1156,267,1863]" captionTargetId="figure-70@175.[426,1161,265,1868]" captionTargetPageId="175" captionText="Figs 652–666. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 652– 654. M. longipes sp. nov., holotype, ♂ (BMNH). 655–657. M. luniceps Fairmaire, 1883, ♂ (BMNH). 658–660. M. mailu sp. nov., holotype, ♂ (NHMB). 661–663. M. maleo sp. nov., holotype, ♂ (IRSN). 664–666. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122244" httpUri="https://zenodo.org/record/4122244/files/figure.png" pageId="130" pageNumber="131">Figs 652–654</figureCitation>
......................................................... 8
</keyLead>
</paragraph>
<paragraph id="922136E09D9E92D9FF52FA25CD212DBB" blockId="130.[189,1399,1361,1497]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF52FA25CD212DBB" pageId="130" pageNumber="131">
– Male metatibia slender and hardly widened distally (
<figureCitation id="0AA52A659D9E92D9FCC2FA24CBFF2DD7" box="[849,937,1434,1461]" captionStart="Figs 54–57" captionStartId="69.[189,241,1925,1951]" captionTargetBox="[359,1240,274,1913]" captionTargetId="figure-59@69.[346,1240,265,1925]" captionTargetPageId="69" captionText="Figs 54–57. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 54. M. lobaticeps Frey, 1969, ♂ (DTC). 55. M. longipes sp. nov., holotype, ♂ (BMNH). 56. M. mailu sp. nov., holotype, ♂ (NHMB). 57. M. luniceps Fairmaire, 1883, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122150" httpUri="https://zenodo.org/record/4122150/files/figure.png" pageId="130" pageNumber="131">Fig. 55</figureCitation>
); all tarsomeres slender and lengthened (
<figureCitation id="0AA52A659D9E92D9FF60FA00C91D2DBB" box="[243,331,1470,1497]" captionStart="Figs 54–57" captionStartId="69.[189,241,1925,1951]" captionTargetBox="[359,1240,274,1913]" captionTargetId="figure-59@69.[346,1240,265,1925]" captionTargetPageId="69" captionText="Figs 54–57. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 54. M. lobaticeps Frey, 1969, ♂ (DTC). 55. M. longipes sp. nov., holotype, ♂ (BMNH). 56. M. mailu sp. nov., holotype, ♂ (NHMB). 57. M. luniceps Fairmaire, 1883, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122150" httpUri="https://zenodo.org/record/4122150/files/figure.png" pageId="130" pageNumber="131">Fig. 55</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9E92D9FD82FA00CAE42DBB" box="[529,690,1470,1497]" captionStart="Figs 652–666" captionStartId="175.[189,241,1892,1918]" captionTargetBox="[436,1156,267,1863]" captionTargetId="figure-70@175.[426,1161,265,1868]" captionTargetPageId="175" captionText="Figs 652–666. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 652– 654. M. longipes sp. nov., holotype, ♂ (BMNH). 655–657. M. luniceps Fairmaire, 1883, ♂ (BMNH). 658–660. M. mailu sp. nov., holotype, ♂ (NHMB). 661–663. M. maleo sp. nov., holotype, ♂ (IRSN). 664–666. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122244" httpUri="https://zenodo.org/record/4122244/files/figure.png" pageId="130" pageNumber="131">Figs 652–654</figureCitation>
............................................................... 
<taxonomicName id="559E4D639D9E92D9FB14FA01CD432DBB" ID-CoL="7X3X3" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1159,1301,1470,1497]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="longipes" status="sp. nov.">
<emphasis id="A0EAEAF29D9E92D9FB14FA01CD432DBB" bold="true" box="[1159,1301,1470,1497]" italics="true" pageId="130" pageNumber="131">M. longipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9E92D9FA8FFA01CD212DBB" box="[1308,1399,1471,1497]" pageId="130" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9E92D9FF2EF9BFCD212ECF" pageId="130" pageNumber="131">
<paragraph id="922136E09D9E92D9FF2EF9BFCD202E06" blockId="130.[189,1399,1537,1709]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF2EF9BFCD202E06" pageId="130" pageNumber="131">
8. Setae on lateral margins of pronotum and elytra shorter than length of compound eye (in older specimens setae often become broken and look shorter); aedeagus not as in 
<figureCitation id="0AA52A659D9E92D9FB27F99BCD002E5D" box="[1204,1366,1573,1599]" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="130" pageNumber="131">Figs 614–616</figureCitation>
or 703–705 ........................................................................................................................................... 10
</keyLead>
</paragraph>
<paragraph id="922136E09D9E92D9FF52F9D1CD212ECF" blockId="130.[189,1399,1537,1709]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF52F9D1CD212ECF" pageId="130" pageNumber="131">
– Setae on lateral margins of pronotum and elytra in fresh specimens as long or longer than length of compound eye; aedeagus as in 
<figureCitation id="0AA52A659D9E92D9FDC5F92CCAA12ECE" box="[598,759,1682,1708]" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="130" pageNumber="131">Figs 614–616</figureCitation>
or 703–705 ................................................................ 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9E92D9FF2EF96BCD212F3E" pageId="130" pageNumber="131">
<paragraph id="922136E09D9E92D9FF2EF96BCD212F76" blockId="130.[189,1399,1749,1885]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF2EF96BCD212F76" pageId="130" pageNumber="131">
9. Hypomeron separated from prosternum by low straight carina; aedeagus as in 
<figureCitation id="0AA52A659D9E92D9FB13F96BCD772E8D" box="[1152,1313,1749,1775]" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="130" pageNumber="131">Figs 614–616</figureCitation>
.......... ............................................................................................................................ 
<taxonomicName id="559E4D639D9E92D9FBEEF944CD212F76" ID-CoL="3XFJ3" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[1149,1399,1785,1812]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="esau">
<emphasis id="A0EAEAF29D9E92D9FBEEF944CC8B2F76" bold="true" box="[1149,1245,1786,1812]" italics="true" pageId="130" pageNumber="131">M. esau</emphasis>
<bibRefCitation id="F60F4B119D9E92D9FB76F947CD212F76" author="Heller K. M." box="[1253,1399,1785,1812]" pageId="130" pageNumber="131" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922136E09D9E92D9FF52F8A0CD212F3E" blockId="130.[189,1399,1749,1885]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF52F8A0CD212F3E" pageId="130" pageNumber="131">
– Hypomeron almost fused to prosternum, carina at place of fusion not indicated or inconspicuous; aedeagus as in 
<figureCitation id="0AA52A659D9E92D9FE08F8FCCA6A2F3F" box="[411,572,1858,1885]" captionStart="Figs 697–711" captionStartId="178.[189,241,1892,1918]" captionTargetBox="[435,1146,268,1853]" captionTargetId="figure-75@178.[425,1163,265,1860]" captionTargetPageId="178" captionText="Figs 697–711. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 697– 699. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH?). 700–702. M. riedeli sp. nov., holotype, ♂ (SMNS). 703–705. M. rugicollis Moser, 1920, ♂ (NHMB). 706–708. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 709–711. M. similis sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122250" httpUri="https://zenodo.org/record/4122250/files/figure.png" pageId="130" pageNumber="131">Figs 703–705</figureCitation>
..................................................................... 
<taxonomicName id="559E4D639D9E92D9FBD2F8FDCD212F3E" ID-CoL="3XFL4" authority="Moser, 1920" authorityName=", Moser Determined" authorityYear="1920" box="[1089,1399,1858,1885]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="A0EAEAF29D9E92D9FBD2F8FDCC8A2F3F" bold="true" box="[1089,1244,1858,1885]" italics="true" pageId="130" pageNumber="131">M. rugicollis</emphasis>
<bibRefCitation id="F60F4B119D9E92D9FB70F8FDCD212F3E" author="Moser J." box="[1251,1399,1858,1885]" pageId="130" pageNumber="131" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="296A2E459D9E92D9FF2EF83BCD202F8A" pageId="130" pageNumber="131">
<paragraph id="922136E09D9E92D9FF2EF83BCD232FFD" blockId="130.[189,1398,1925,2025]" box="[189,1397,1925,1951]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF2EF83BCD232FFD" box="[189,1397,1925,1951]" pageId="130" pageNumber="131">10. Lateral margin of pronotum distinctly emarginate or strongly constricted prebasally.................... 11</keyLead>
</paragraph>
<paragraph id="922136E09D9E92D9FF52F814CD202F8A" blockId="130.[189,1398,1925,2025]" pageId="130" pageNumber="131">
<keyLead id="296F95D59D9E92D9FF52F814CD202F8A" pageId="130" pageNumber="131">– Lateral margin of pronotum not or hardly emarginate prebasally, evenly rounded all along or slightly constricted prebasally ..................................................................................................................... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9F92D8FF2EFEB7CD2129BB" pageId="131" pageNumber="132">
<paragraph id="922136E09D9F92D8FF2EFEB7CD202925" blockId="131.[189,1399,265,473]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF2EFEB7CD202925" pageId="131" pageNumber="132">
11. Lateral margin of pronotum moderately strongly emarginate or constricted, upper angle of constriction not produced posteriad, either obtuse or completely rounded; aedeagus not as in 
<figureCitation id="0AA52A659D9F92D8FB08FE93CD6D2925" box="[1179,1339,301,327]" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="131" pageNumber="132">Figs 601–603</figureCitation>
... 12
</keyLead>
</paragraph>
<paragraph id="922136E09D9F92D8FF52FEECCD2129BB" blockId="131.[189,1399,265,473]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF52FEECCD2129BB" pageId="131" pageNumber="132">
– Lateral margin of pronotum very deeply emarginate prebasally, upper angle of emargination produced posteriad and acute (
<figureCitation id="0AA52A659D9F92D8FDC8FEC8CA9329F3" box="[603,709,374,401]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="131" pageNumber="132">Fig. 206</figureCitation>
); lateral margin of pronotum in emargination area almost smooth, markedly less strongly crenulate than anterior part of it; aedeagus as in 
<figureCitation id="0AA52A659D9F92D8FB69FE24C94029BB" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="131" pageNumber="132">Figs 601– 603</figureCitation>
.......................................................................................................................... 
<taxonomicName id="559E4D639D9F92D8FB33FE01CD4329BB" ID-CoL="7X3WF" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1184,1301,446,473]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="brocki" status="sp. nov.">
<emphasis id="A0EAEAF29D9F92D8FB33FE01CD4329BB" bold="true" box="[1184,1301,446,473]" italics="true" pageId="131" pageNumber="132">M. brocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9F92D8FA8FFE01CD2129BB" box="[1308,1399,447,473]" pageId="131" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9F92D8FF2EFDBCCD202AEB" pageId="131" pageNumber="132">
<paragraph id="922136E09D9F92D8FF2EFDBCCD202A22" blockId="131.[189,1399,513,649]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF2EFDBCCD202A22" pageId="131" pageNumber="132">12. Underside /lateral margins of male basal metatarsomere with ordinary short or long setae, which are not arranged brush-like .................................................................................................................... 14</keyLead>
</paragraph>
<paragraph id="922136E09D9F92D8FF52FDF5CD202AEB" blockId="131.[189,1399,513,649]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF52FDF5CD202AEB" pageId="131" pageNumber="132">
– Ventrolateral margins of male basal metatarsomere with a brush of long (sub)erect setae (cf. 
<figureCitation id="0AA52A659D9F92D8FF79FDD0C9DA2AEB" box="[234,396,622,649]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="131" pageNumber="132">Figs 436–437</figureCitation>
) .................................................................................................................................. 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9F92D8FF2EFD0FCD212B5A" pageId="131" pageNumber="132">
<paragraph id="922136E09D9F92D8FF2EFD0FCD212A92" blockId="131.[189,1399,689,825]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF2EFD0FCD212A92" pageId="131" pageNumber="132">
13. Abdominal ventrites medially with golden (sub)erect setae (
<figureCitation id="0AA52A659D9F92D8FC52FD0FCC7D2AAE" box="[961,1067,689,716]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="131" pageNumber="132">Fig. 435</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9F92D8FB6CFD0FC9402A92" captionStart="Figs 726–738" captionStartId="180.[189,241,1891,1917]" captionTargetBox="[343,1246,278,1849]" captionTargetId="figure-69@180.[330,1258,265,1871]" captionTargetPageId="180" captionText="Figs 726–738. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 726– 729. M. suwawa sp. nov., holotype, ♂ (BMNH). 730–732. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 733–735. M. trivialis sp. nov., holotype, ♂ (BMNH). 736–738. M. vicinus Heller, 1914, ♂ (apex of parameres broken) (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122254" httpUri="https://zenodo.org/record/4122254/files/figure.png" pageId="131" pageNumber="132">Figs 730– 732</figureCitation>
................................................................................................................. 
<taxonomicName id="559E4D639D9F92D8FBC9FD68CD212A92" ID-CoL="3XFLP" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[1114,1399,725,752]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis id="A0EAEAF29D9F92D8FBC9FD68CC8D2A92" bold="true" box="[1114,1243,725,752]" italics="true" pageId="131" pageNumber="132">M. tarsalis</emphasis>
<bibRefCitation id="F60F4B119D9F92D8FB71FD68CD212A92" author="Arrow G. E." box="[1250,1399,726,752]" pageId="131" pageNumber="132" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922136E09D9F92D8FF52FD44CD212B5A" blockId="131.[189,1399,689,825]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF52FD44CD212B5A" pageId="131" pageNumber="132">
– Abdominal ventrites without golden (sub)erect setae; aedeagus as in 
<figureCitation id="0AA52A659D9F92D8FB81FD44CCE42B76" box="[1042,1202,762,788]" captionStart="Figs 617–631" captionStartId="173.[189,241,1892,1918]" captionTargetBox="[439,1134,274,1859]" captionTargetId="figure-71@173.[434,1154,265,1868]" captionTargetPageId="173" captionText="Figs 617–631. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 617– 619. M. hamatus sp. nov., holotype, ♂ (IECA). 620–622. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 623–625. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 626–628. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 629–631. M. konjo sp. nov., holotype, ♂ (NMPC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122240" httpUri="https://zenodo.org/record/4122240/files/figure.png" pageId="131" pageNumber="132">Figs 620–622</figureCitation>
........................ ...................................................................................................................... 
<taxonomicName id="559E4D639D9F92D8FBC4FCA1CD212B5A" ID-CoL="3XFJG" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[1111,1399,798,825]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="hirtipes">
<emphasis id="A0EAEAF29D9F92D8FBC4FCA1CC8D2B5B" bold="true" box="[1111,1243,798,825]" italics="true" pageId="131" pageNumber="132">M. hirtipes</emphasis>
<bibRefCitation id="F60F4B119D9F92D8FB71FCA1CD212B5A" author="Arrow G. E." box="[1250,1399,798,825]" pageId="131" pageNumber="132" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9F92D8FF2EFCDFCD212C6F" pageId="131" pageNumber="132">
<paragraph id="922136E09D9F92D8FF2EFCDFCD202BC2" blockId="131.[189,1399,865,1037]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF2EFCDFCD202BC2" pageId="131" pageNumber="132">
14. Lateral margin of pronotum markedly emarginate prebasally; intervening spaces of elytra densely microreticulate, opaque; basal protarsomere without tooth; aedeagus not as in 
<figureCitation id="0AA52A659D9F92D8FBECFC3BCD762BC2" box="[1151,1312,901,928]" captionStart="Figs 739–750" captionStartId="181.[189,241,1892,1918]" captionTargetBox="[360,1207,271,1867]" captionTargetId="figure-66@181.[347,1240,265,1871]" captionTargetPageId="181" captionText="Figs 739–750. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 739– 741. M. weigeli sp. nov., paratype, ♂ (NME). 742–744. M. woodlarkianus Heller, 1914, ♂ (SNSD). 745–747. M. yamdena sp. nov., paratype, ♂ (BMNH). 748–750. M. paupianus Heller, 1910 (paratype ♂ of M. arrowi Frey, 1969) (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122256" httpUri="https://zenodo.org/record/4122256/files/figure.png" pageId="131" pageNumber="132">Figs 745–747</figureCitation>
...... 15
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</paragraph>
<paragraph id="922136E09D9F92D8FF52FC14CD212C6F" blockId="131.[189,1399,865,1037]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF52FC14CD212C6F" pageId="131" pageNumber="132">
– Lateral margin of pronotum rather constricted prebasally, not markedly emarginate; intervening spaces of elytra smooth or microscopically wrinkled, glossy; basal protarsomere in both sexes with central tooth (
<figureCitation id="0AA52A659D9F92D8FE1DFC4CC9A02C6F" box="[398,502,1010,1037]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="131" pageNumber="132">Fig. 447</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9F92D8FD28FC4CCB0D2C6F" box="[699,859,1010,1037]" captionStart="Figs 739–750" captionStartId="181.[189,241,1892,1918]" captionTargetBox="[360,1207,271,1867]" captionTargetId="figure-66@181.[347,1240,265,1871]" captionTargetPageId="181" captionText="Figs 739–750. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 739– 741. M. weigeli sp. nov., paratype, ♂ (NME). 742–744. M. woodlarkianus Heller, 1914, ♂ (SNSD). 745–747. M. yamdena sp. nov., paratype, ♂ (BMNH). 748–750. M. paupianus Heller, 1910 (paratype ♂ of M. arrowi Frey, 1969) (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122256" httpUri="https://zenodo.org/record/4122256/files/figure.png" pageId="131" pageNumber="132">Figs 745–747</figureCitation>
....................................... 
<taxonomicName id="559E4D639D9F92D8FBECFC4DCD432C6F" ID-CoL="7X3XZ" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1151,1301,1010,1037]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="yamdena" status="sp. nov.">
<emphasis id="A0EAEAF29D9F92D8FBECFC4DCD432C6F" bold="true" box="[1151,1301,1010,1037]" italics="true" pageId="131" pageNumber="132">M. yamdena</emphasis>
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<taxonomicNameLabel id="BBD957899D9F92D8FA8FFC4DCD212C6F" box="[1308,1399,1011,1037]" pageId="131" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
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<keyStep id="296A2E459D9F92D8FF2EFB8BCD212CDF" pageId="131" pageNumber="132">
<paragraph id="922136E09D9F92D8FF2EFB8BCD212C16" blockId="131.[189,1399,1077,1213]" pageId="131" pageNumber="132">
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15. Underside of male basal metatarsomere with brush of long erect setae; aedeagus as in 
<figureCitation id="0AA52A659D9F92D8FB6CFB8BC9402C16" captionStart="Figs 712–725" captionStartId="179.[189,241,1892,1918]" captionTargetBox="[433,1144,277,1858]" captionTargetId="figure-72@179.[427,1160,265,1870]" captionTargetPageId="179" captionText="Figs 712–725. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 712– 714. M. skalei sp. nov., holotype, ♂ (NME). 715–717. M. sougb sp. nov., holotype, ♂ (NME). 718–720. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 721–723. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 724–725. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122252" httpUri="https://zenodo.org/record/4122252/files/figure.png" pageId="131" pageNumber="132">Figs 724– 725</figureCitation>
.......................................................................................................... 
<taxonomicName id="559E4D639D9F92D8FBBAFBE4CD212C16" ID-CoL="3XFLN" authority="Heller, 1895" authorityName=", Hell. Determined" authorityYear="1895" box="[1065,1399,1113,1140]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="subcostatus">
<emphasis id="A0EAEAF29D9F92D8FBBAFBE4CC8B2C16" bold="true" box="[1065,1245,1113,1140]" italics="true" pageId="131" pageNumber="132">M. subcostatus</emphasis>
<bibRefCitation id="F60F4B119D9F92D8FB76FBE7CD212C16" author="Heller K. M." box="[1253,1399,1113,1140]" pageId="131" pageNumber="132" pagination="1 - 17" refId="ref98131" refString="Heller K. M. 1895. Erster Beitrag zur Papuaniscen [sic!] Kaferfauna. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden 1894 / 1895 (16): 1 - 17." type="journal article" year="1895">Heller, 1895</bibRefCitation>
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</paragraph>
<paragraph id="922136E09D9F92D8FF52FBC0CD212CDF" blockId="131.[189,1399,1077,1213]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF52FBC0CD212CDF" pageId="131" pageNumber="132">
– Underside of male basal metatarsomere with a long setae which are not brush-like arranged and rather suberect; aedeagus as in 
<figureCitation id="0AA52A659D9F92D8FDCBFB1CCAA12CDE" box="[600,759,1186,1212]" captionStart="Figs 697–711" captionStartId="178.[189,241,1892,1918]" captionTargetBox="[435,1146,268,1853]" captionTargetId="figure-75@178.[425,1163,265,1860]" captionTargetPageId="178" captionText="Figs 697–711. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 697– 699. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH?). 700–702. M. riedeli sp. nov., holotype, ♂ (SMNS). 703–705. M. rugicollis Moser, 1920, ♂ (NHMB). 706–708. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 709–711. M. similis sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122250" httpUri="https://zenodo.org/record/4122250/files/figure.png" pageId="131" pageNumber="132">Figs 709–711</figureCitation>
......................................................... 
<taxonomicName id="559E4D639D9F92D8FB0DFB1DCD432CDF" ID-CoL="7X3XP" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1182,1301,1186,1213]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="A0EAEAF29D9F92D8FB0DFB1DCD432CDF" bold="true" box="[1182,1301,1186,1213]" italics="true" pageId="131" pageNumber="132">M. similis</emphasis>
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<taxonomicNameLabel id="BBD957899D9F92D8FA8FFB1DCD212CDF" box="[1308,1399,1187,1213]" pageId="131" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<keyStep id="296A2E459D9F92D8FF2EFB5BCD212D0E" pageId="131" pageNumber="132">
<paragraph id="922136E09D9F92D8FF2EFB5BCD202D41" blockId="131.[189,1399,1253,1388]" pageId="131" pageNumber="132">
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16. Lateral margin of pronotum with 20 or more delicate crenulae; lateral margin of male labroclypeus variably strongly sinuous; aedeagus not as in 
<figureCitation id="0AA52A659D9F92D8FC93FAB7CBF62D41" box="[768,928,1289,1315]" captionStart="Figs 697–711" captionStartId="178.[189,241,1892,1918]" captionTargetBox="[435,1146,268,1853]" captionTargetId="figure-75@178.[425,1163,265,1860]" captionTargetPageId="178" captionText="Figs 697–711. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 697– 699. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH?). 700–702. M. riedeli sp. nov., holotype, ♂ (SMNS). 703–705. M. rugicollis Moser, 1920, ♂ (NHMB). 706–708. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 709–711. M. similis sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122250" httpUri="https://zenodo.org/record/4122250/files/figure.png" pageId="131" pageNumber="132">Figs 706–708</figureCitation>
.......................................................... 18
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</paragraph>
<paragraph id="922136E09D9F92D8FF52FA90CD212D0E" blockId="131.[189,1399,1253,1388]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF52FA90CD212D0E" pageId="131" pageNumber="132">
– Lateral margin of pronotum with 13–16 rough crenulae; lateral margin of male labroclypeus strongly sinuous (
<figureCitation id="0AA52A659D9F92D8FEC7FAECC9EB2D0E" box="[340,445,1362,1388]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="131" pageNumber="132">Figs 117</figureCitation>
, 
<figureCitation id="0AA52A659D9F92D8FE5AFAECC9A02D0E" box="[457,502,1362,1388]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="131" pageNumber="132">171</figureCitation>
)..................................................................17 [two possibly conspecific taxa]
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<keyStep id="296A2E459D9F92D8FF2EFA2BCD212EB0" pageId="131" pageNumber="132">
<paragraph id="922136E09D9F92D8FF2EFA2BCD212E7E" blockId="131.[189,1399,1428,1746]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF2EFA2BCD212E7E" pageId="131" pageNumber="132">
17. Intervening spaces on pronotal dorsum completely flat; pronotal disc without impressions; dorsal punctures of pronotum comparatively stronger ovoid (
<figureCitation id="0AA52A659D9F92D8FC44FA07CC132DB1" box="[983,1093,1465,1491]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="131" pageNumber="132">Figs 171</figureCitation>
, 
<figureCitation id="0AA52A659D9F92D8FBC4FA07CCDE2DB6" box="[1111,1160,1465,1492]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="131" pageNumber="132">250</figureCitation>
); each elytron with vague tracks of three longitudinal carinae (including sutural one); aedeagus as in 
<figureCitation id="0AA52A659D9F92D8FB68FA63C9402E7E" captionStart="Figs 697–711" captionStartId="178.[189,241,1892,1918]" captionTargetBox="[435,1146,268,1853]" captionTargetId="figure-75@178.[425,1163,265,1860]" captionTargetPageId="178" captionText="Figs 697–711. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 697– 699. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH?). 700–702. M. riedeli sp. nov., holotype, ♂ (SMNS). 703–705. M. rugicollis Moser, 1920, ♂ (NHMB). 706–708. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 709–711. M. similis sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122250" httpUri="https://zenodo.org/record/4122250/files/figure.png" pageId="131" pageNumber="132">Figs 706– 708</figureCitation>
...................................................................................................... 
<taxonomicName id="559E4D639D9F92D8FB9EF9BCCD212E7E" ID-CoL="7X3XL" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[1037,1399,1538,1564]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="seriegranosus">
<emphasis id="A0EAEAF29D9F92D8FB9EF9BCCC882E7E" bold="true" box="[1037,1246,1538,1564]" italics="true" pageId="131" pageNumber="132">M. seriegranosus</emphasis>
<bibRefCitation id="F60F4B119D9F92D8FB76F9BCCD212E7E" author="Heller K. M." box="[1253,1399,1538,1564]" pageId="131" pageNumber="132" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="922136E09D9F92D8FF52F999CD212EB0" blockId="131.[189,1399,1428,1746]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF52F999CD212EB0" pageId="131" pageNumber="132">
– Intervening spaces on pronotal dorsum in part slightly globose, not completely flat; pronotal disc with vague circular impression on either side of hardly longitudinally impressed midline; dorsal punctures of pronotum rather circular to irregularly hexagonal, distinctly ovoid along lateral sides only (
<figureCitation id="0AA52A659D9F92D8FEA3F92DC9DB2ECF" box="[304,397,1683,1709]" captionStart="Figs 38–41" captionStartId="47.[189,241,1893,1919]" captionTargetBox="[360,1224,277,1874]" captionTargetId="figure-68@47.[360,1227,265,1875]" captionTargetPageId="47" captionText="Figs 38–41. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 38. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 39. M. hamatus sp. nov., holotype, ♂ (IECA). 40. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (BPBM) (photograph courtesy BPBM). 41. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122142" httpUri="https://zenodo.org/record/4122142/files/figure.png" pageId="131" pageNumber="132">Figs 38</figureCitation>
, 
<figureCitation id="0AA52A659D9F92D8FE09F92DC99F2ECF" box="[410,457,1683,1709]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="131" pageNumber="132">117</figureCitation>
); each elytron with vague tracks of four longitudinal carinae (including sutural one); aedeagus unknown.......................................................................... 
<taxonomicName id="559E4D639D9F92D8FBBDF906CD212EB0" ID-CoL="3XFJ8" authority="Moser, 1920" authorityName="Moser Determined" authorityYear="1920" box="[1070,1399,1719,1746]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="fraterculus">
<emphasis id="A0EAEAF29D9F92D8FBBDF906CC8A2EB0" bold="true" box="[1070,1244,1719,1746]" italics="true" pageId="131" pageNumber="132">M. fraterculus</emphasis>
<bibRefCitation id="F60F4B119D9F92D8FB70F906CD212EB0" author="Moser J." box="[1251,1399,1720,1746]" pageId="131" pageNumber="132" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="296A2E459D9F92D8FF2EF944CD202F5B" pageId="131" pageNumber="132">
<paragraph id="922136E09D9F92D8FF2EF944CD202F76" blockId="131.[189,1398,1786,1849]" box="[189,1398,1786,1813]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF2EF944CD202F76" box="[189,1398,1786,1813]" pageId="131" pageNumber="132">18. Intervening spaces of elytra opaque, microreticulate ...................................................................... 19</keyLead>
</paragraph>
<paragraph id="922136E09D9F92D8FF52F8A1CD202F5B" blockId="131.[189,1398,1786,1849]" box="[193,1398,1822,1849]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF52F8A1CD202F5B" box="[193,1398,1822,1849]" pageId="131" pageNumber="132">– Intervening spaces of elytra glossy, smooth or wrinkled, not microreticulate ................................ 23</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9F92DFFF2EF8DFCD2129C9" lastPageId="132" lastPageNumber="133" pageId="131" pageNumber="132">
<paragraph id="922136E09D9F92D8FF2EF8DFCD202F8A" blockId="131.[189,1398,1889,2025]" pageId="131" pageNumber="132">
<keyLead id="296F95D59D9F92D8FF2EF8DFCD202F8A" pageId="131" pageNumber="132">19. Male and female labroclypeus emarginate or subtruncate (not truncate) on anterior margin, bent up to with regard to axis of frons in lateral view; head dorsally flattened or convex on frons, impressed dorsally at anterolateral angles of labroclypeus; pronotal punctures annular or not; crenulae lateral pronotal margin separated by delicate shallow or broad and distinct spaces ................................. 20</keyLead>
</paragraph>
<paragraph id="922136E09D9892DFFF52FEB7CD2129C9" blockId="132.[193,1399,265,428]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF52FEB7CD2129C9" pageId="132" pageNumber="133">
– Female labroclypeus gradually constricted anteriad on lateral margins, its anterior margin truncate (
<figureCitation id="0AA52A659D9892DFFF60FE95C9092924" box="[243,351,299,326]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="132" pageNumber="133">Fig. 156</figureCitation>
), not or hardly bent up with regard to axis of frons in lateral view; head dorsally flattened, hardly impressed dorsally at anterolateral angles of labroclypeus; pronotal punctures not annular; crenulae of lateral pronotal margin separated by delicate shallow spaces; male is unknown .................................................................................................... 
<taxonomicName id="559E4D639D9892DFFBAFFE2FCD2129C9" ID-CoL="3XFKK" authority="Moser, 1926" authorityName="Moser" authorityYear="1926" box="[1084,1399,401,428]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="A0EAEAF29D9892DFFBAFFE2FCC8A29C9" bold="true" box="[1084,1244,401,427]" italics="true" pageId="132" pageNumber="133">M. papuanus</emphasis>
<bibRefCitation id="F60F4B119D9892DFFB70FE2CCD2129C9" author="Moser J." box="[1251,1399,401,428]" pageId="132" pageNumber="133" pagination="196 - 203" refId="ref98419" refString="Moser J. 1926. Neue Melolonthiden und Cetoniden aus dem Suden von Hollandisch Neu-Guinea. Nova Guinea: a Journal of Botany, Zoology, Anthropology, Ethnography, Geology and Palaeontology of the Papuan Region. Zoology 15 (2): 196 - 203." type="journal article" year="1926">Moser, 1926</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9892DFFF2EFE6FCD212A6C" pageId="132" pageNumber="133">
<paragraph id="922136E09D9892DFFF2EFE6FCD202989" blockId="132.[189,1399,465,526]" box="[189,1398,465,492]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF2EFE6FCD202989" box="[189,1398,465,492]" pageId="132" pageNumber="133">20. Parameres long and slender, constricted towards apices ................................................................. 21</keyLead>
</paragraph>
<paragraph id="922136E09D9892DFFF52FE4ACD212A6C" blockId="132.[189,1399,465,526]" box="[193,1399,499,526]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF52FE4ACD212A6C" box="[193,1399,499,526]" pageId="132" pageNumber="133">
– Parameres short and broad, widened apically (
<figureCitation id="0AA52A659D9892DFFC9BFE4DCBFA2A6C" box="[776,940,499,526]" captionStart="Figs 667–681" captionStartId="176.[189,241,1892,1918]" captionTargetBox="[444,1142,267,1860]" captionTargetId="figure-72@176.[419,1168,265,1868]" captionTargetPageId="176" captionText="Figs 667–681. Papuan Maechidius Macleay, 1819, aedeagus, different aspects. 667– 669. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 670–672. M. milneanus Heller, 1914, ♂ (SNSD). 673–675. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 676–678. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 679-681. M. nepenthephilus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122246" httpUri="https://zenodo.org/record/4122246/files/figure.png" pageId="132" pageNumber="133">Figs 679–680</figureCitation>
) ............... 
<taxonomicName id="559E4D639D9892DFFBA1FE4ACD432A6C" ID-CoL="7X3X9" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1074,1301,499,526]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="nepenthephilus" status="sp. nov.">
<emphasis id="A0EAEAF29D9892DFFBA1FE4ACD432A6C" bold="true" box="[1074,1301,499,526]" italics="true" pageId="132" pageNumber="133">M. nepenthephilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9892DFFA8FFE4ACD212A6C" box="[1308,1399,500,526]" pageId="132" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9892DFFF2EFD8ACD212AD6" pageId="132" pageNumber="133">
<paragraph id="922136E09D9892DFFF2EFD8ACD202A12" blockId="132.[189,1399,563,692]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF2EFD8ACD202A12" pageId="132" pageNumber="133">
21. Parameres narrowly separated at base (
<figureCitation id="0AA52A659D9892DFFD21FD8ACB002A2C" box="[690,854,563,590]" captionStart="Figs 652–666" captionStartId="175.[189,241,1892,1918]" captionTargetBox="[436,1156,267,1863]" captionTargetId="figure-70@175.[426,1161,265,1868]" captionTargetPageId="175" captionText="Figs 652–666. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 652– 654. M. longipes sp. nov., holotype, ♂ (BMNH). 655–657. M. luniceps Fairmaire, 1883, ♂ (BMNH). 658–660. M. mailu sp. nov., holotype, ♂ (NHMB). 661–663. M. maleo sp. nov., holotype, ♂ (IRSN). 664–666. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122244" httpUri="https://zenodo.org/record/4122244/files/figure.png" pageId="132" pageNumber="133">Figs 655–657</figureCitation>
, 
<figureCitation id="0AA52A659D9892DFFCF1FD8DCB9B2A2C" box="[866,973,563,590]" captionStart="Figs 739–750" captionStartId="181.[189,241,1892,1918]" captionTargetBox="[360,1207,271,1867]" captionTargetId="figure-66@181.[347,1240,265,1871]" captionTargetPageId="181" captionText="Figs 739–750. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 739– 741. M. weigeli sp. nov., paratype, ♂ (NME). 742–744. M. woodlarkianus Heller, 1914, ♂ (SNSD). 745–747. M. yamdena sp. nov., paratype, ♂ (BMNH). 748–750. M. paupianus Heller, 1910 (paratype ♂ of M. arrowi Frey, 1969) (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122256" httpUri="https://zenodo.org/record/4122256/files/figure.png" pageId="132" pageNumber="133">748–750</figureCitation>
); crenulae of lateral pronotal margin separated by distinct moderately deep and broad intervening spaces ............................................. 22
</keyLead>
</paragraph>
<paragraph id="922136E09D9892DFFF52FDC6CD212AD6" blockId="132.[189,1399,563,692]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF52FDC6CD212AD6" pageId="132" pageNumber="133">
– Parameres moderately broadly separated at base (
<figureCitation id="0AA52A659D9892DFFCBAFDC6CB862AF1" box="[809,976,632,659]" captionStart="Figs 726–738" captionStartId="180.[189,241,1891,1917]" captionTargetBox="[343,1246,278,1849]" captionTargetId="figure-69@180.[330,1258,265,1871]" captionTargetPageId="180" captionText="Figs 726–738. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 726– 729. M. suwawa sp. nov., holotype, ♂ (BMNH). 730–732. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 733–735. M. trivialis sp. nov., holotype, ♂ (BMNH). 736–738. M. vicinus Heller, 1914, ♂ (apex of parameres broken) (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122254" httpUri="https://zenodo.org/record/4122254/files/figure.png" pageId="132" pageNumber="133">Figs 736–738</figureCitation>
); crenulae of lateral pronotal margin narrowly separated in anterior half, nearly merged ....................................... 
<taxonomicName id="559E4D639D9892DFFBCCFD24CD212AD6" ID-CoL="6QTFM" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[1119,1399,666,692]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="A0EAEAF29D9892DFFBCCFD24CC8B2AD6" bold="true" box="[1119,1245,666,692]" italics="true" pageId="132" pageNumber="133">M. vicinus</emphasis>
<bibRefCitation id="F60F4B119D9892DFFB76FD24CD212AD6" author="Heller K. M." box="[1253,1399,666,692]" pageId="132" pageNumber="133" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9892DFFF2EFD64CD212B5A" pageId="132" pageNumber="133">
<paragraph id="922136E09D9892DFFF2EFD64CD212A96" blockId="132.[189,1399,729,825]" box="[189,1399,729,756]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF2EFD64CD212A96" box="[189,1399,729,756]" pageId="132" pageNumber="133">
22. Lateral margin of pronotum evenly broadly rounded (
<figureCitation id="0AA52A659D9892DFFCCDFD64CB952A96" box="[862,963,730,756]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="132" pageNumber="133">Fig. 229</figureCitation>
) ........... 
<taxonomicName id="559E4D639D9892DFFBB5FD64CD212A96" ID-CoL="3XFK2" authority="Fairmaire, 1883" authorityName="Fairmaire" authorityYear="1883" box="[1062,1399,729,756]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="luniceps">
<emphasis id="A0EAEAF29D9892DFFBB5FD64CCE22A96" bold="true" box="[1062,1204,729,756]" italics="true" pageId="132" pageNumber="133">M. luniceps</emphasis>
<bibRefCitation id="F60F4B119D9892DFFB28FD64CD212A96" author="Fairmaire M. L." box="[1211,1399,730,756]" pageId="132" pageNumber="133" pagination="1 - 58" refId="ref97970" refString="Fairmaire M. L. 1883. Essai sur les coleopteres de l'Archipel de la Nouvelle Bretagne. Annales de la Societe entomologique de Belgique 27: 1 - 58." type="journal article" year="1883">Fairmaire, 1883</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922136E09D9892DFFF2EFD43CD212B5A" blockId="132.[189,1399,729,825]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF2EFD43CD212B5A" pageId="132" pageNumber="133">
– Lateral margin of pronotum somewhat constricted towards base in basal third (
<figureCitation id="0AA52A659D9892DFFB14FD42CCBA2B74" box="[1159,1260,764,790]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="132" pageNumber="133">Fig. 242</figureCitation>
) ................ .................................................................................................................. 
<taxonomicName id="559E4D639D9892DFFBA5FCA0CD212B5A" ID-CoL="3XFKP" authority="Heller, 1910" authorityName=", Hell. Determined" authorityYear="1910" box="[1078,1399,798,824]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="paupianus">
<emphasis id="A0EAEAF29D9892DFFBA5FCA0CC8B2B5A" bold="true" box="[1078,1245,798,824]" italics="true" pageId="132" pageNumber="133">M. paupianus</emphasis>
<bibRefCitation id="F60F4B119D9892DFFB76FCA0CD212B5A" author="Heller K. M." box="[1253,1399,798,824]" pageId="132" pageNumber="133" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller, 1910</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9892DFFF2EFCE0CD202BF8" pageId="132" pageNumber="133">
<paragraph id="922136E09D9892DFFF2EFCE0CD202B1A" blockId="132.[189,1398,862,923]" box="[189,1398,862,889]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF2EFCE0CD202B1A" box="[189,1398,862,889]" pageId="132" pageNumber="133">23. Elytral incision-shaped punctures distinctly sinuous....................................................................... 27</keyLead>
</paragraph>
<paragraph id="922136E09D9892DFFF52FC3FCD202BF8" blockId="132.[189,1398,862,923]" box="[193,1398,896,923]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF52FC3FCD202BF8" box="[193,1398,896,923]" pageId="132" pageNumber="133">– Elytral incision-shaped punctures more or less straight .................................................................. 24</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9892DFFF2EFC7FCD212C01" pageId="132" pageNumber="133">
<paragraph id="922136E09D9892DFFF2EFC7FCD202B9F" blockId="132.[189,1399,960,1123]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF2EFC7FCD202B9F" pageId="132" pageNumber="133">24. Dorsal forebody with greenish metallic lustre; dorsal punctures of pronotum regularly circular to ovoid ................................................................................................................................................ 25</keyLead>
</paragraph>
<paragraph id="922136E09D9892DFFF52FBBBCD212C01" blockId="132.[189,1399,960,1123]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF52FBBBCD212C01" pageId="132" pageNumber="133">
– Forebody without metallic lustre; dorsal punctures of pronotum, especially lateral and laterobasal ones, irregularly ovoid, double annular and large (
<figureCitation id="0AA52A659D9892DFFC03FB99CBB82C23" box="[912,1006,1062,1089]" captionStart="Figs 94–96" captionStartId="126.[189,241,1928,1954]" captionTargetBox="[346,1191,267,1904]" captionTargetId="figure-55@126.[339,1249,265,1909]" captionTargetPageId="126" captionText="Figs 94–96. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 94. M. weigeli sp. nov., holotype, ♂ (NME). 95. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 96. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122171" httpUri="https://zenodo.org/record/4122171/files/figure.png" pageId="132" pageNumber="133">Figs 95</figureCitation>
, 
<figureCitation id="0AA52A659D9892DFFC6EFB99CC782C20" box="[1021,1070,1063,1090]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="132" pageNumber="133">264</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9892DFFB6CFB99C9402C01" captionStart="Figs 739–750" captionStartId="181.[189,241,1892,1918]" captionTargetBox="[360,1207,271,1867]" captionTargetId="figure-66@181.[347,1240,265,1871]" captionTargetPageId="181" captionText="Figs 739–750. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 739– 741. M. weigeli sp. nov., paratype, ♂ (NME). 742–744. M. woodlarkianus Heller, 1914, ♂ (SNSD). 745–747. M. yamdena sp. nov., paratype, ♂ (BMNH). 748–750. M. paupianus Heller, 1910 (paratype ♂ of M. arrowi Frey, 1969) (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122256" httpUri="https://zenodo.org/record/4122256/files/figure.png" pageId="132" pageNumber="133">Figs 742– 744</figureCitation>
..................................................................................................... 
<taxonomicName id="559E4D639D9892DFFB91FBF7CD212C01" ID-CoL="3XFLW" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[1026,1399,1096,1123]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="woodlarkianus">
<emphasis id="A0EAEAF29D9892DFFB91FBF7CC8B2C01" bold="true" box="[1026,1245,1096,1123]" italics="true" pageId="132" pageNumber="133">M. woodlarkianus</emphasis>
<bibRefCitation id="F60F4B119D9892DFFB76FBF7CD212C01" author="Heller K. M." box="[1253,1399,1097,1123]" pageId="132" pageNumber="133" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
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<keyStep id="296A2E459D9892DFFF2EFB37CD212D6B" pageId="132" pageNumber="133">
<paragraph id="922136E09D9892DFFF2EFB37CD202CA7" blockId="132.[189,1399,1161,1290]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF2EFB37CD202CA7" pageId="132" pageNumber="133">25. Intervening spaces of elytra smooth or in part delicately wrinkled; elytral longitudinal carinae vague and inconspicuous; aedeagus is unknown ....................................................................................... 26</keyLead>
</paragraph>
<paragraph id="922136E09D9892DFFF52FB70CD212D6B" blockId="132.[189,1399,1161,1290]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF52FB70CD212D6B" pageId="132" pageNumber="133">
– Intervening spaces of elytra irregularly roughly wrinkled; elytral longitudinal carinae rather broad and elevated; aedeagus as in 
<figureCitation id="0AA52A659D9892DFFDADFB51CA892D6B" box="[574,735,1263,1289]" captionStart="Figs 667–681" captionStartId="176.[189,241,1892,1918]" captionTargetBox="[444,1142,267,1860]" captionTargetId="figure-72@176.[419,1168,265,1868]" captionTargetPageId="176" captionText="Figs 667–681. Papuan Maechidius Macleay, 1819, aedeagus, different aspects. 667– 669. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 670–672. M. milneanus Heller, 1914, ♂ (SNSD). 673–675. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 676–678. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 679-681. M. nepenthephilus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122246" httpUri="https://zenodo.org/record/4122246/files/figure.png" pageId="132" pageNumber="133">Figs 670–672</figureCitation>
.............................................. 
<taxonomicName id="559E4D639D9892DFFBA5FB51CD212D6B" ID-CoL="3XFK8" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[1078,1399,1262,1289]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="milneanus">
<emphasis id="A0EAEAF29D9892DFFBA5FB51CC8B2D6B" bold="true" box="[1078,1245,1262,1289]" italics="true" pageId="132" pageNumber="133">M. milneanus</emphasis>
<bibRefCitation id="F60F4B119D9892DFFB76FB51CD212D6B" author="Heller K. M." box="[1253,1399,1263,1289]" pageId="132" pageNumber="133" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
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</paragraph>
</keyStep>
<keyStep id="296A2E459D9892DFFF2EFA91CD212D96" pageId="132" pageNumber="133">
<paragraph id="922136E09D9892DFFF2EFA91CD212DEC" blockId="132.[189,1399,1327,1524]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF2EFA91CD212DEC" pageId="132" pageNumber="133">
26. Male labroclypeus narrowed anteriad, truncate on anterior margin (female unknown); anterolateral angles of labroclypeus obtuse, not marked (
<figureCitation id="0AA52A659D9892DFFC3CFAEFCC4B2D09" box="[943,1053,1361,1387]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="132" pageNumber="133">Fig. 187</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9892DFFB6FFAEFC9402DEC" captionStart="Figs 726–738" captionStartId="180.[189,241,1891,1917]" captionTargetBox="[343,1246,278,1849]" captionTargetId="figure-69@180.[330,1258,265,1871]" captionTargetPageId="180" captionText="Figs 726–738. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 726– 729. M. suwawa sp. nov., holotype, ♂ (BMNH). 730–732. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 733–735. M. trivialis sp. nov., holotype, ♂ (BMNH). 736–738. M. vicinus Heller, 1914, ♂ (apex of parameres broken) (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122254" httpUri="https://zenodo.org/record/4122254/files/figure.png" pageId="132" pageNumber="133">Figs 733– 735</figureCitation>
........................................................................................................................ 
<taxonomicName id="559E4D639D9892DFFB02FACACD432DEC" ID-CoL="7X3XV" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1169,1301,1395,1422]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="trivialis" status="sp. nov.">
<emphasis id="A0EAEAF29D9892DFFB02FACACD432DEC" bold="true" box="[1169,1301,1395,1422]" italics="true" pageId="132" pageNumber="133">M. trivialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9892DFFA8FFACACD212DEC" box="[1308,1399,1396,1422]" pageId="132" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9892DFFF52FA28CD212D96" blockId="132.[189,1399,1327,1524]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF52FA28CD212D96" pageId="132" pageNumber="133">
– Female labroclypeus not markedly narrowed anteriad, subtruncate on anterior margin (male unknown); anterolateral angles of labroclypeus obtuse but marked (
<figureCitation id="0AA52A659D9892DFFB90FA09CC0F2DB0" box="[1027,1113,1463,1490]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="132" pageNumber="133">Fig. 97</figureCitation>
) .................................... ................................................................................................................. 
<taxonomicName id="559E4D639D9892DFFBA8FA64CD212D96" ID-CoL="3XFH6" authority="Heller, 1910" authorityName="Heller" authorityYear="1910" box="[1083,1399,1497,1524]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="aenescens">
<emphasis id="A0EAEAF29D9892DFFBA8FA64CC8B2D96" bold="true" box="[1083,1245,1498,1524]" italics="true" pageId="132" pageNumber="133">M. aenescens</emphasis>
<bibRefCitation id="F60F4B119D9892DFFB76FA67CD212D96" author="Heller K. M." box="[1253,1399,1497,1524]" pageId="132" pageNumber="133" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller, 1910</bibRefCitation>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="296A2E459D9892DFFF2EF9A4CD202E34" pageId="132" pageNumber="133">
<paragraph id="922136E09D9892DFFF2EF9A4CD202E56" blockId="132.[189,1398,1562,1622]" box="[189,1398,1562,1588]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF2EF9A4CD202E56" box="[189,1398,1562,1588]" pageId="132" pageNumber="133">
27. Total body length less than 
<quantity id="55669B059D9892DFFDBEF9A4CA222E56" box="[557,628,1562,1588]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="132" pageNumber="133" unit="mm" value="8.0">8 mm</quantity>
; body brown to castaneous brown ............................................. 29
</keyLead>
</paragraph>
<paragraph id="922136E09D9892DFFF52F982CD202E34" blockId="132.[189,1398,1562,1622]" box="[193,1398,1596,1622]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF52F982CD202E34" box="[193,1398,1596,1622]" pageId="132" pageNumber="133">
– Total body length about 
<quantity id="55669B059D9892DFFD9AF982CA2A2E34" box="[521,636,1596,1622]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="132" pageNumber="133" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
; body black to dark brown........................................................ 28
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9892DFFF2EF9C2CD212F22" pageId="132" pageNumber="133">
<paragraph id="922136E09D9892DFFF2EF9C2CD212EB8" blockId="132.[189,1399,1660,1857]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF2EF9C2CD212EB8" pageId="132" pageNumber="133">
28. Dorsal punctures of pronotum oblong-ovoid, horseshoe-shaped, moderately dense; regular transverse row of punctures present along anterior and basal margin of pronotum; male labroclypeus broadly V-shape emarginate on anterior margin (
<figureCitation id="0AA52A659D9892DFFD57F97ECB7C2EB8" box="[708,810,1728,1754]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="132" pageNumber="133">Fig. 130</figureCitation>
); aedeagus unknown .... 
<taxonomicName id="559E4D639D9892DFFBDAF97ECD212EB8" ID-CoL="3XFJP" authority="Moser, 1920" authorityName="Moser" authorityYear="1920" box="[1097,1399,1727,1755]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="jobiensis">
<emphasis id="A0EAEAF29D9892DFFBDAF97ECC8A2EB8" bold="true" box="[1097,1244,1727,1754]" italics="true" pageId="132" pageNumber="133">M. jobiensis</emphasis>
<bibRefCitation id="F60F4B119D9892DFFB70F97FCD212EB8" author="Moser J." box="[1251,1399,1728,1755]" pageId="132" pageNumber="133" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922136E09D9892DFFF52F95DCD212F22" blockId="132.[189,1399,1660,1857]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF52F95DCD212F22" pageId="132" pageNumber="133">
– Dorsal punctures of pronotum circular to ovoid, horseshoe-shaped along lateral margins only, sparse; no row of punctures along either anterior and basal margins of pronotum, male labroclypeus shallowly emarginate on anterior margin (
<figureCitation id="0AA52A659D9892DFFD44F898CB6B2F22" box="[727,829,1830,1856]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="132" pageNumber="133">Fig. 101</figureCitation>
); aedeagus unknown........ 
<taxonomicName id="559E4D639D9892DFFBE3F898CD212F22" ID-CoL="3XFHC" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[1136,1399,1830,1856]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="aroae">
<emphasis id="A0EAEAF29D9892DFFBE3F898CC8B2F22" bold="true" box="[1136,1245,1830,1856]" italics="true" pageId="132" pageNumber="133">M. aroae</emphasis>
<bibRefCitation id="F60F4B119D9892DFFB76F898CD212F22" author="Heller K. M." box="[1253,1399,1830,1856]" pageId="132" pageNumber="133" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="296A2E459D9892DFFF2EF8D8CD212F8B" pageId="132" pageNumber="133">
<paragraph id="922136E09D9892DFFF2EF8D8CD202FC1" blockId="132.[189,1399,1894,2025]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF2EF8D8CD202FC1" pageId="132" pageNumber="133">29. Lateral margin of pronotum stronger rounded, constricted anteriad and posteriad; posterolateral angle of pronotum obtuse ................................................................................................................ 30</keyLead>
</paragraph>
<paragraph id="922136E09D9892DFFF52F815CD212F8B" blockId="132.[189,1399,1894,2025]" pageId="132" pageNumber="133">
<keyLead id="296F95D59D9892DFFF52F815CD212F8B" pageId="132" pageNumber="133">
– Lateral margin of pronotum less rounded, stronger constricted anteriad than posteriad; posterolateral angle of pronotum nearly right-angled ............................... 
<taxonomicName id="559E4D639D9892DFFC0BF871CCB82F8A" ID-CoL="7X3X8" authority="Moser, 1920" authorityName="Moser" authorityYear="1920" box="[920,1262,1998,2025]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="132" pageNumber="133" phylum="Arthropoda" rank="species" species="moluccanus">
<emphasis id="A0EAEAF29D9892DFFC0BF871CC052F8B" bold="true" box="[920,1107,1998,2025]" italics="true" pageId="132" pageNumber="133">M. moluccanus</emphasis>
<bibRefCitation id="F60F4B119D9892DFFBC9F871CCB82F8A" author="Moser J." box="[1114,1262,1998,2025]" pageId="132" pageNumber="133" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
comb. rest.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9992DEFF2EFEB7CD21292B" pageId="133" pageNumber="134">
<paragraph id="922136E09D9992DEFF2EFEB7CD212941" blockId="133.[189,1399,264,329]" box="[189,1399,264,292]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF2EFEB7CD212941" box="[189,1399,264,292]" pageId="133" pageNumber="134">
30. Aedeagus as in 
<figureCitation id="0AA52A659D9992DEFE30FEB7CA152946" box="[419,579,265,292]" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="133" pageNumber="134">Figs 649–651</figureCitation>
...................................................................... 
<taxonomicName id="559E4D639D9992DEFBC2FEB7CD212941" ID-CoL="3XFJY" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[1105,1399,264,291]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="lobaticeps">
<emphasis id="A0EAEAF29D9992DEFBC2FEB7CCA42941" bold="true" box="[1105,1266,264,291]" italics="true" pageId="133" pageNumber="134">M. lobaticeps</emphasis>
<bibRefCitation id="F60F4B119D9992DEFB6AFEB7CD212941" author="Frey G." box="[1273,1399,265,291]" pageId="133" pageNumber="134" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922136E09D9992DEFF52FE91CD21292B" blockId="133.[189,1399,264,329]" box="[193,1399,302,329]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF52FE91CD21292B" box="[193,1399,302,329]" pageId="133" pageNumber="134">
– Aedeagus as in 
<figureCitation id="0AA52A659D9992DEFE30FE91CA12292B" box="[419,580,303,329]" captionStart="Figs 697–711" captionStartId="178.[189,241,1892,1918]" captionTargetBox="[435,1146,268,1853]" captionTargetId="figure-75@178.[425,1163,265,1860]" captionTargetPageId="178" captionText="Figs 697–711. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 697– 699. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH?). 700–702. M. riedeli sp. nov., holotype, ♂ (SMNS). 703–705. M. rugicollis Moser, 1920, ♂ (NHMB). 706–708. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 709–711. M. similis sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122250" httpUri="https://zenodo.org/record/4122250/files/figure.png" pageId="133" pageNumber="134">Figs 700–702</figureCitation>
................................................................................. 
<taxonomicName id="559E4D639D9992DEFB33FE91CD43292B" ID-CoL="7X3XK" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1184,1301,302,329]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="A0EAEAF29D9992DEFB33FE91CD43292B" bold="true" box="[1184,1301,302,329]" italics="true" pageId="133" pageNumber="134">M. riedeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9992DEFA8FFE91CD21292B" box="[1308,1399,303,329]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
</keyStep>
<keyStep id="296A2E459D9992DEFF2EFECDCD2029BA" pageId="133" pageNumber="134">
<paragraph id="922136E09D9992DEFF2EFECDCD2029D0" blockId="133.[189,1398,370,472]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF2EFECDCD2029D0" pageId="133" pageNumber="134">
31. With conspicuous dense brush-like cluster of setae at least along lateral margins of pronotum or on its postero-lateral angles (cf. 
<figureCitation id="0AA52A659D9992DEFDD3FE26CAFF29D0" box="[576,681,408,434]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="133" pageNumber="134">Figs 122</figureCitation>
, 
<figureCitation id="0AA52A659D9992DEFD27FE26CAB529D0" box="[692,739,408,434]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="133" pageNumber="134">221</figureCitation>
, 
<figureCitation id="0AA52A659D9992DEFD7CFE26CB4829D0" box="[751,798,408,434]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="133" pageNumber="134">238</figureCitation>
, 
<figureCitation id="0AA52A659D9992DEFCB9FE26CB0F29D0" box="[810,857,408,434]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="133" pageNumber="134">246</figureCitation>
, 
<figureCitation id="0AA52A659D9992DEFCF7FE26CBC529D0" box="[868,915,408,434]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="133" pageNumber="134">261</figureCitation>
, 
<figureCitation id="0AA52A659D9992DEFC0CFE26CB9A29D1" box="[927,972,408,435]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="133" pageNumber="134">358</figureCitation>
).................................................... 32
</keyLead>
</paragraph>
<paragraph id="922136E09D9992DEFF52FE00CD2029BA" blockId="133.[189,1398,370,472]" box="[193,1398,446,472]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF52FE00CD2029BA" box="[193,1398,446,472]" pageId="133" pageNumber="134">– Dorsal pubescence ordinary or scale-like, not brush-like clustered ............................................... 36</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9992DEFF2EFDBCCD212A20" pageId="133" pageNumber="134">
<paragraph id="922136E09D9992DEFF2EFDBCCD202A7E" blockId="133.[189,1399,514,578]" box="[189,1398,514,540]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF2EFDBCCD202A7E" box="[189,1398,514,540]" pageId="133" pageNumber="134">32. Pronotum slightly narrower than combined width of elytra............................................................ 33</keyLead>
</paragraph>
<paragraph id="922136E09D9992DEFF52FD96CD212A20" blockId="133.[189,1399,514,578]" box="[193,1399,551,578]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF52FD96CD212A20" box="[193,1399,551,578]" pageId="133" pageNumber="134">
– Pronotum distinctly narrower than combined width of elytra...... 
<taxonomicName id="559E4D639D9992DEFC4CFD96CCB82A20" ID-CoL="7XWMM" authority="(Frey, 1969)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" box="[991,1262,551,578]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="helleri" status="comb. nov.">
<emphasis id="A0EAEAF29D9992DEFC4CFD96CC002A20" bold="true" box="[991,1110,551,578]" italics="true" pageId="133" pageNumber="134">M. helleri</emphasis>
(
<bibRefCitation id="F60F4B119D9992DEFBF6FD99CCB32A20" author="Frey G." box="[1125,1253,551,578]" pageId="133" pageNumber="134" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9992DEFB66FD99CD212A20" box="[1269,1399,551,578]" pageId="133" pageNumber="134" rank="species">comb. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9992DEFF2EFDD5CD212A95" pageId="133" pageNumber="134">
<paragraph id="922136E09D9992DEFF2EFDD5CD212AC9" blockId="133.[189,1399,619,759]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF2EFDD5CD212AC9" pageId="133" pageNumber="134">
33. Lateral margin of pronotum flattened and laterally expanded in anterior half, strongly emarginate postmedially (
<figureCitation id="0AA52A659D9992DEFE02FD2FC9AA2AC9" box="[401,508,657,683]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="133" pageNumber="134">Figs 246</figureCitation>
, 
<figureCitation id="0AA52A659D9992DEFD9BFD2FCA632ACE" box="[520,565,657,684]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="133" pageNumber="134">261</figureCitation>
) ..............................................................................34 [
<emphasis id="A0EAEAF29D9992DEFB3CFD2FCD3A2AC9" bold="true" box="[1199,1388,657,683]" pageId="133" pageNumber="134">
<taxonomicName id="559E4D639D9992DEFB3CFD2FCD4D2AC9" ID-CoL="7X3XX" authorityName="Telnov" authorityYear="2020" box="[1199,1307,657,683]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="A0EAEAF29D9992DEFB3CFD2FCD4D2AC9" bold="true" box="[1199,1307,657,683]" italics="true" pageId="133" pageNumber="134">M. ursus</emphasis>
</taxonomicName>
group
</emphasis>
]
</keyLead>
</paragraph>
<paragraph id="922136E09D9992DEFF52FD09CD212A95" blockId="133.[189,1399,619,759]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF52FD09CD212A95" pageId="133" pageNumber="134">
– Lateral margin of pronotum not expanded laterally in anterior half, postmedian emargination short, inconspicuous (
<figureCitation id="0AA52A659D9992DEFE31FD62CA502A94" box="[418,518,732,758]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="133" pageNumber="134">Fig. 221</figureCitation>
) ................................................................................. 
<taxonomicName id="559E4D639D9992DEFBFCFD63CD422A95" ID-CoL="7X3WV" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1135,1300,732,759]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="kazantsevi" status="sp. nov.">
<emphasis id="A0EAEAF29D9992DEFBFCFD63CD422A95" bold="true" box="[1135,1300,732,759]" italics="true" pageId="133" pageNumber="134">M. kazantsevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9992DEFA8FFD63CD212A95" box="[1308,1399,733,759]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9992DEFF2EFC9ECD212BE4" pageId="133" pageNumber="134">
<paragraph id="922136E09D9992DEFF2EFC9ECD202B59" blockId="133.[189,1399,800,902]" box="[189,1398,800,827]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF2EFC9ECD202B59" box="[189,1398,800,827]" pageId="133" pageNumber="134">34. Elytra without longitudinal rows of glossy glabrous humps ........................................................... 35</keyLead>
</paragraph>
<paragraph id="922136E09D9992DEFF52FCF8CD212BE4" blockId="133.[189,1399,800,902]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF52FCF8CD212BE4" pageId="133" pageNumber="134">
– Longitudinal carinae of elytra formed by large irregular glossy glabrous humps (
<figureCitation id="0AA52A659D9992DEFB58FCF8CD7D2B02" box="[1227,1323,838,864]" captionStart="Figs 74–77" captionStartId="97.[189,241,1928,1954]" captionTargetBox="[328,1255,263,1887]" captionTargetId="figure-56@97.[325,1262,256,1897]" captionTargetPageId="97" captionText="Figs 74–77. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 74. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 75. Same, ♂ (SMNS). 76. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 77. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122160" httpUri="https://zenodo.org/record/4122160/files/figure.png" pageId="133" pageNumber="134">Figs 74</figureCitation>
, 
<figureCitation id="0AA52A659D9992DEFAAFFCF8C9412BE4" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="133" pageNumber="134">337– 338</figureCitation>
) ............................................................................................ 
<taxonomicName id="559E4D639D9992DEFC58FCD2CCB82BE4" ID-CoL="7XWMN" authority="(Frey, 1969)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" box="[971,1262,875,902]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="perlatus" status="comb. nov.">
<emphasis id="A0EAEAF29D9992DEFC58FCD2CC002BE4" bold="true" box="[971,1110,875,902]" italics="true" pageId="133" pageNumber="134">M. perlatus</emphasis>
(
<bibRefCitation id="F60F4B119D9992DEFBF6FCD2CCB32BE4" author="Frey G." box="[1125,1253,875,902]" pageId="133" pageNumber="134" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9992DEFB66FCD5CD212BE4" box="[1269,1399,875,902]" pageId="133" pageNumber="134" rank="species">comb. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9992DEFF2EFC11CD212C03" pageId="133" pageNumber="134">
<paragraph id="922136E09D9992DEFF2EFC11CD212B92" blockId="133.[189,1399,943,1121]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF2EFC11CD212B92" pageId="133" pageNumber="134">
35. Metatibia widened, flattened and glabrous on inner margin (
<figureCitation id="0AA52A659D9992DEFC5FFC11CC6E2BAB" box="[972,1080,943,969]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="133" pageNumber="134">Fig. 444</figureCitation>
); sutural carina of elytron present; lateral postmedian emargination of pronotum deep (
<figureCitation id="0AA52A659D9992DEFC23FC6BCC422B92" box="[944,1044,981,1008]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="133" pageNumber="134">Fig. 261</figureCitation>
) ................. 
<taxonomicName id="559E4D639D9992DEFB3AFC6BCD432B8D" ID-CoL="7X3XX" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1193,1301,981,1007]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="A0EAEAF29D9992DEFB3AFC6BCD432B8D" bold="true" box="[1193,1301,981,1007]" italics="true" pageId="133" pageNumber="134">M. ursus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9992DEFA8FFC68CD212B92" box="[1308,1399,982,1008]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9992DEFF52FC45CD212C03" blockId="133.[189,1399,943,1121]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF52FC45CD212C03" pageId="133" pageNumber="134">
– Metatibia slender, not flattened and glabrous on inner margin; sutural carina of elytron absent; lateral postmedian emargination of pronotum moderately strong (
<figureCitation id="0AA52A659D9992DEFC0BFB9ECBAB2C58" box="[920,1021,1056,1082]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="133" pageNumber="134">Fig. 238</figureCitation>
)................................................. .............................................................................................. 
<taxonomicName id="559E4D639D9992DEFC3FFBF8CCB82C02" ID-CoL="7X3XC" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[940,1262,1093,1120]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="opatroides">
<emphasis id="A0EAEAF29D9992DEFC3FFBF8CC042C02" bold="true" box="[940,1106,1093,1120]" italics="true" pageId="133" pageNumber="134">M. opatroides</emphasis>
<bibRefCitation id="F60F4B119D9992DEFBC9FBF8CCB82C02" author="Arrow G. E." box="[1114,1262,1094,1120]" pageId="133" pageNumber="134" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
comb. rest.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9992DEFF2EFB34CD202C8D" pageId="133" pageNumber="134">
<paragraph id="922136E09D9992DEFF2EFB34CD212CA8" blockId="133.[189,1399,1162,1264]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF2EFB34CD212CA8" pageId="133" pageNumber="134">
36. Dorsal sculpture of pronotum and elytra of strongly irregularly arranged punctures forming conspicuous fingerprint-like pattern (
<figureCitation id="0AA52A659D9992DEFD0EFB11CB512CA8" box="[669,775,1199,1226]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="133" pageNumber="134">Figs 297</figureCitation>
, 
<figureCitation id="0AA52A659D9992DEFC80FB0ECB162CA8" box="[787,832,1200,1226]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="133" pageNumber="134">347</figureCitation>
).......................................37 [
<emphasis id="A0EAEAF29D9992DEFB04FB0ECD3A2CA8" bold="true" box="[1175,1388,1200,1226]" pageId="133" pageNumber="134">
<taxonomicName id="559E4D639D9992DEFB04FB0ECD4D2CA8" ID-CoL="3XFLM" authorityName="Arrow" authorityYear="1941" box="[1175,1307,1200,1226]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="sturnus">
<emphasis id="A0EAEAF29D9992DEFB04FB0ECD4D2CA8" bold="true" box="[1175,1307,1200,1226]" italics="true" pageId="133" pageNumber="134">M. sturnus</emphasis>
</taxonomicName>
group
</emphasis>
]
</keyLead>
</paragraph>
<paragraph id="922136E09D9992DEFF52FB68CD202C8D" blockId="133.[189,1399,1162,1264]" box="[193,1398,1237,1264]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF52FB68CD202C8D" box="[193,1398,1237,1264]" pageId="133" pageNumber="134">
– Punctures of pronotum and elytra not fingerprint-like arranged, not like in 
<figureCitation id="0AA52A659D9992DEFBC7FB6BCCEB2C8D" box="[1108,1213,1237,1263]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="133" pageNumber="134">Figs 297</figureCitation>
, 
<figureCitation id="0AA52A659D9992DEFB5BFB6BCCA22C8D" box="[1224,1268,1237,1263]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="133" pageNumber="134">347</figureCitation>
............ 38
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9992DEFF2EFAA7CD212DC7" pageId="133" pageNumber="134">
<paragraph id="922136E09D9992DEFF2EFAA7CD212D3B" blockId="133.[189,1399,1305,1445]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF2EFAA7CD212D3B" pageId="133" pageNumber="134">
37. Middle of posterior margin of male abdominal sternites 2–3 each with a paired bunch of three long setae on each side (
<figureCitation id="0AA52A659D9992DEFE5EFA81CA622D3B" box="[461,564,1343,1369]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="133" pageNumber="134">Fig. 433</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9992DEFD69FA81CBCD2D3B" box="[762,923,1343,1369]" captionStart="Figs 712–725" captionStartId="179.[189,241,1892,1918]" captionTargetBox="[433,1144,277,1858]" captionTargetId="figure-72@179.[427,1160,265,1870]" captionTargetPageId="179" captionText="Figs 712–725. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 712– 714. M. skalei sp. nov., holotype, ♂ (NME). 715–717. M. sougb sp. nov., holotype, ♂ (NME). 718–720. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 721–723. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 724–725. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122252" httpUri="https://zenodo.org/record/4122252/files/figure.png" pageId="133" pageNumber="134">Figs 721–723</figureCitation>
........................ 
<taxonomicName id="559E4D639D9992DEFBC4FA81CD212D3B" ID-CoL="3XFLM" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[1111,1399,1343,1369]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="sturnus">
<emphasis id="A0EAEAF29D9992DEFBC4FA81CC8D2D3B" bold="true" box="[1111,1243,1343,1369]" italics="true" pageId="133" pageNumber="134">M. sturnus</emphasis>
<bibRefCitation id="F60F4B119D9992DEFB71FA81CD212D3B" author="Arrow G. E." box="[1250,1399,1343,1369]" pageId="133" pageNumber="134" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922136E09D9992DEFF52FADBCD212DC7" blockId="133.[189,1399,1305,1445]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF52FADBCD212DC7" pageId="133" pageNumber="134">
– Male abdominal sternites medially with somewhat stronger appressed curved setae but without paired bunches of long setae; aedeagus as in 
<figureCitation id="0AA52A659D9992DEFD6AFA34CBCF2DC7" box="[761,921,1418,1445]" captionStart="Figs 586–600" captionStartId="171.[189,241,1892,1918]" captionTargetBox="[400,1175,268,1865]" captionTargetId="figure-72@171.[391,1196,265,1868]" captionTargetPageId="171" captionText="Figs 586–600. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 586– 588. M. bintang sp. nov., holotype, ♂ (NME). 589–591. M. boessnecki sp. nov., holotype, ♂ (SNSD). 592–594. M. caperatus sp. nov., holotype, ♂ (NME). 595–597. M. ciliatus sp. nov., holotype, ♂ (IECA). 598–600. M. crypticus sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122234" httpUri="https://zenodo.org/record/4122234/files/figure.png" pageId="133" pageNumber="134">Figs 592–594</figureCitation>
.............................. 
<taxonomicName id="559E4D639D9992DEFBE4FA34CD432DC6" ID-CoL="7X3WH" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1143,1301,1418,1444]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="caperatus" status="sp. nov.">
<emphasis id="A0EAEAF29D9992DEFBE4FA34CD432DC6" bold="true" box="[1143,1301,1418,1444]" italics="true" pageId="133" pageNumber="134">M. caperatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9992DEFA8FFA35CD212DC7" box="[1308,1399,1419,1445]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="922136E09D9992DEFF2EFA70CD212E6C" blockId="133.[189,1399,1485,1588]" pageId="133" pageNumber="134">
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38. Male 3 
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visible abdominal sternite with obtuse ventral median triangular hump (
<figureCitation id="0AA52A659D9992DEFB6BFA70C94E2E6C" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="133" pageNumber="134">Figs 429– 432</figureCitation>
) .......................................................................................................................39 [
<emphasis id="A0EAEAF29D9992DEFB2FFA4ACD3A2E6C" bold="true" box="[1212,1388,1523,1550]" pageId="133" pageNumber="134">
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<emphasis id="A0EAEAF29D9992DEFB2FFA4ACD4D2E6C" bold="true" box="[1212,1307,1523,1550]" italics="true" pageId="133" pageNumber="134">M. dani</emphasis>
</taxonomicName>
group
</emphasis>
]
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</paragraph>
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– Male 3 
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visible abdominal sternite not humped, evenly rounded or flattened ventrally ................ 40
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</paragraph>
</keyStep>
<keyStep id="296A2E459D9992DEFF2EF9E3CD212F56" pageId="133" pageNumber="134">
<paragraph id="922136E09D9992DEFF2EF9E3CD212EA1" blockId="133.[189,1399,1629,1844]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF2EF9E3CD212EA1" pageId="133" pageNumber="134">
39. Male labroclypeus comparatively shallower emarginate on anterior margin (
<figureCitation id="0AA52A659D9992DEFBF2F9E3CC902E15" box="[1121,1222,1629,1655]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="133" pageNumber="134">Fig. 121</figureCitation>
); ventral hump of visible abdominal ventrite 3 acute in ventral view (
<figureCitation id="0AA52A659D9992DEFC13F93DCC7C2EFF" box="[896,1066,1667,1693]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="133" pageNumber="134">Figs 431–432</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9992DEFB6DF93DC9402EA0" captionStart="Figs 617–631" captionStartId="173.[189,241,1892,1918]" captionTargetBox="[439,1134,274,1859]" captionTargetId="figure-71@173.[434,1154,265,1868]" captionTargetPageId="173" captionText="Figs 617–631. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 617– 619. M. hamatus sp. nov., holotype, ♂ (IECA). 620–622. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 623–625. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 626–628. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 629–631. M. konjo sp. nov., holotype, ♂ (NMPC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122240" httpUri="https://zenodo.org/record/4122240/files/figure.png" pageId="133" pageNumber="134">Figs 617– 619</figureCitation>
...................................................................................................................... 
<taxonomicName id="559E4D639D9992DEFB17F917CD432EA1" ID-CoL="7X3WR" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1156,1301,1704,1731]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="A0EAEAF29D9992DEFB17F917CD432EA1" bold="true" box="[1156,1301,1704,1731]" italics="true" pageId="133" pageNumber="134">M. hamatus</emphasis>
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<taxonomicNameLabel id="BBD957899D9992DEFA8FF917CD212EA1" box="[1308,1399,1705,1731]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="922136E09D9992DEFF52F971CD212F56" blockId="133.[189,1399,1629,1844]" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF52F971CD212F56" pageId="133" pageNumber="134">
– Male labroclypeus comparatively deeper emarginate on anterior margin (
<figureCitation id="0AA52A659D9992DEFBD2F970CCF12E8A" box="[1089,1191,1742,1768]" captionStart="Figs 109–120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109–120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109–119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="133" pageNumber="134">Fig. 114</figureCitation>
); ventral hump of visible abdominal ventrite 3 rather obtuse in ventral view (
<figureCitation id="0AA52A659D9992DEFC04F94ACC682F6C" box="[919,1086,1780,1806]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="133" pageNumber="134">Figs 429–430</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9992DEFA91F94AC9402F51" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="133" pageNumber="134">Figs 604– 606</figureCitation>
............................................................................................................................. 
<taxonomicName id="559E4D639D9992DEFB25F8A4CD432F56" ID-CoL="7X3WL" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1206,1301,1817,1844]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="dani" status="sp. nov.">
<emphasis id="A0EAEAF29D9992DEFB25F8A4CD432F56" bold="true" box="[1206,1301,1817,1844]" italics="true" pageId="133" pageNumber="134">M. dani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9992DEFA8FF8A4CD212F56" box="[1308,1399,1818,1844]" pageId="133" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
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</keyStep>
<paragraph id="922136E09D9992DEFF2EF8E3CD202FFF" blockId="133.[189,1399,1885,2025]" pageId="133" pageNumber="134">
<keyStep id="296A2E459D9992DEFF2EF8E3CD202FFF" pageId="133" pageNumber="134">
<keyLead id="296F95D59D9992DEFF2EF8E3CD202FFF" pageId="133" pageNumber="134">
40. Punctures of elytra circular to ovoid, arranged in more or less regular paired longitudinal rows (cf. 
<figureCitation id="0AA52A659D9992DEFF79F83DC9052FFF" box="[234,339,1923,1949]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="133" pageNumber="134">Figs 327</figureCitation>
, 
<figureCitation id="0AA52A659D9992DEFECCF83DC9DA2FFC" box="[351,396,1923,1950]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="133" pageNumber="134">345</figureCitation>
) .................................................................................................................................. 41
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</paragraph>
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<keyLead id="296F95D59D9992DEFF52F817CD202F8A" pageId="133" pageNumber="134">– Punctures of elytra circular, ovoid or of different shape, if arranged in longitudinal rows then those not paired ......................................................................................................................................... 46</keyLead>
<keyStep id="296A2E459D9A92DDFF2EFEB7CD20292B" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF2EFEB7CD20292B" pageId="134" pageNumber="135">
41. Lateral margin of pronotum distinctly emarginate prebasally (
<figureCitation id="0AA52A659D9A92DDFC5CFEB7CC622946" box="[975,1076,265,292]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="134" pageNumber="135">Fig. 255</figureCitation>
and 
<bibRefCitation id="F60F4B119D9A92DDFBE1FEB7C97F292B" author="Narakusumo R. P. &amp; Balke M." pageId="134" pageNumber="135" pagination="35 - 50" refId="ref98467" refString="Narakusumo R. P. &amp; Balke M. 2019. Four new species of Epholcis Waterhouse, 1875 (Coleoptera: Scarabaeidae: Melolonthinae: Maechidiini) from the Moluccas, Indonesia. Treubia 46: 35 - 50. https: // doi. org / 10.14203 / treubia. v 46 i 0.3791" type="journal article" year="2019">Narakusumo &amp; Balke 2019</bibRefCitation>
: fig. 5a).................................................................................................................................... 42
</keyLead>
</keyStep>
</paragraph>
<paragraph id="922136E09D9A92DDFF52FEEBCD20290C" blockId="134.[189,1398,265,367]" box="[193,1398,340,367]" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF52FEEBCD20290C" box="[193,1398,340,367]" pageId="134" pageNumber="135">– Lateral margin of pronotum evenly broadly rounded, not or indistinctly emarginate prebasally ... 43</keyLead>
</paragraph>
<keyStep id="296A2E459D9A92DDFF2EFE26CD212A2B" pageId="134" pageNumber="135">
<paragraph id="922136E09D9A92DDFF2EFE26CD2129BA" blockId="134.[189,1399,408,586]" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF2EFE26CD2129BA" pageId="134" pageNumber="135">
42. Generally larger species, total body length over 
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; male metatibia slender, inner margin straight; aedeagus as in 
<figureCitation id="0AA52A659D9A92DDFE08FE00CA6A29BB" box="[411,572,446,473]" captionStart="Figs 712–725" captionStartId="179.[189,241,1892,1918]" captionTargetBox="[433,1144,277,1858]" captionTargetId="figure-72@179.[427,1160,265,1870]" captionTargetPageId="179" captionText="Figs 712–725. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 712– 714. M. skalei sp. nov., holotype, ♂ (NME). 715–717. M. sougb sp. nov., holotype, ♂ (NME). 718–720. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 721–723. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 724–725. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122252" httpUri="https://zenodo.org/record/4122252/files/figure.png" pageId="134" pageNumber="135">Figs 715–717</figureCitation>
................................................................................... 
<taxonomicName id="559E4D639D9A92DDFB37FE00CD4329BA" ID-CoL="7X3XR" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1188,1301,445,472]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="sougb" status="sp. nov.">
<emphasis id="A0EAEAF29D9A92DDFB37FE00CD4329BA" bold="true" box="[1188,1301,445,472]" italics="true" pageId="134" pageNumber="135">M. sougb</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9A92DDFA8FFE00CD2129BA" box="[1308,1399,446,472]" pageId="134" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9A92DDFF52FE5ACD212A2B" blockId="134.[189,1399,408,586]" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF52FE5ACD212A2B" pageId="134" pageNumber="135">
– Generally smaller species, total body length under 
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; male metatibia thickened and somewhat curved on inner margin (cf. 
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: fig. 5e); aedeagus as in 
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: fig. 5f–g)................................... 
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<emphasis id="A0EAEAF29D9A92DDFD6CFD91CBDC2A2B" bold="true" box="[767,906,558,585]" italics="true" pageId="134" pageNumber="135">M. obiensis</emphasis>
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43. Generally larger species, total body length over 
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; labroclypeus deeply emarginate anteriorly; aedeagus not as in 
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..................................................................................................... 44
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– Generally smaller species, total body length under 
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; labroclypeus shallowly emarginate anteriorly; aedeagus as in 
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......................................................... 
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<emphasis id="A0EAEAF29D9A92DDFBF4FD5ACC8D2A9C" bold="true" box="[1127,1243,740,766]" italics="true" pageId="134" pageNumber="135">M. nanus</emphasis>
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<keyLead id="296F95D59D9A92DDFF2EFC96CD202B20" box="[189,1398,807,834]" pageId="134" pageNumber="135">44. Male metatibia straight, not curved ................................................................................................. 45</keyLead>
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– Male metatibia strongly curved (cf. 
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: fig. 2e); aedeagus as in 
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: fig. 2f–g).............................. 
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<emphasis id="A0EAEAF29D9A92DDFD64FCCDCBDC2BEF" bold="true" box="[759,906,883,909]" italics="true" pageId="134" pageNumber="135">M. arcuatus</emphasis>
(
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45. Crenulae of lateral margin of pronotum flat, nearly merged, intervening spaces indistinct (
<figureCitation id="0AA52A659D9A92DDFA91FC09CD3F2BB3" box="[1282,1385,951,977]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="134" pageNumber="135">Fig. 244</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9A92DDFE08FC62CA6D2B94" box="[411,571,988,1014]" captionStart="Figs 682–696" captionStartId="177.[189,241,1892,1918]" captionTargetBox="[436,1163,269,1860]" captionTargetId="figure-72@177.[423,1174,265,1866]" captionTargetPageId="177" captionText="Figs 682–696. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 682– 684. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 685–687. M. pauxillus Heller, 1910 comb. rest., ♂ (BMNH). 688–690. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 691–693. M. peregrinus Lansberge, 1886, ♂ (DTC). 694-696. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122248" httpUri="https://zenodo.org/record/4122248/files/figure.png" pageId="134" pageNumber="135">Figs 688–690</figureCitation>
........................................................................... 
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<emphasis id="A0EAEAF29D9A92DDFBFAFC63CD4B2B95" bold="true" box="[1129,1309,988,1015]" italics="true" pageId="134" pageNumber="135">M. pedarioides</emphasis>
Arrow
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– Crenulae of lateral margin of pronotum strong, intervening spaces rather deep and broad (cf. 
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: fig. 3a–b); aedeagus as in 
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: fig. 3e–f)........ ....................................................................... 
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<emphasis id="A0EAEAF29D9A92DDFD6DFBF0CBDF2C0A" bold="true" box="[766,905,1101,1128]" italics="true" pageId="134" pageNumber="135">M. cakalele</emphasis>
(
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<keyLead id="296F95D59D9A92DDFF2EFB2FCD202CCE" box="[189,1398,1169,1196]" pageId="134" pageNumber="135">46. Male basal metatarsomere ordinary, not conspicuously widened and flattened ............................. 50</keyLead>
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– Male basal metatarsomere conspicuously widened and flattened (cf. 
<figureCitation id="0AA52A659D9A92DDFB9EFB09CC262CB0" box="[1037,1136,1207,1234]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="134" pageNumber="135">Fig. 446</figureCitation>
) ............................. 47
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47. Dorsal setae of forebody long, surpassing length of corresponding punctures; aedeagus not as in 
<figureCitation id="0AA52A659D9A92DDFF79FA9EC9DD2D58" box="[234,395,1312,1338]" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="134" pageNumber="135">Figs 640–642</figureCitation>
................................................................................................................................... 48
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<keyLead id="296F95D59D9A92DDFF52FAF9CD212DE4" pageId="134" pageNumber="135">
– Dorsal setae of forebody inconspicuous, short, hardly surpassing length of corresponding punctures (
<figureCitation id="0AA52A659D9A92DDFF60FAD2C90C2DE5" box="[243,346,1388,1415]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="134" pageNumber="135">Fig. 315</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9A92DDFDB3FAD2CA972DE4" box="[544,705,1388,1414]" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="134" pageNumber="135">Figs 640–642</figureCitation>
.............................................................. 
<taxonomicName id="559E4D639D9A92DDFB1DFAD2CD432DE4" ID-CoL="7X3WY" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1166,1301,1387,1414]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="legalovi" status="sp. nov.">
<emphasis id="A0EAEAF29D9A92DDFB1DFAD2CD432DE4" bold="true" box="[1166,1301,1387,1414]" italics="true" pageId="134" pageNumber="135">M. legalovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9A92DDFA8FFAD2CD212DE4" box="[1308,1399,1388,1414]" pageId="134" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9A92DDFF2EFA0ECD212E03" pageId="134" pageNumber="135">
<paragraph id="922136E09D9A92DDFF2EFA0ECD202D8D" blockId="134.[189,1399,1456,1633]" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF2EFA0ECD202D8D" pageId="134" pageNumber="135">48. Dorsal setae at least on forebody long, pointed or hardly widened apically; male and female labroclypeus subtruncate on anterior margin................................................................................... 49</keyLead>
</paragraph>
<paragraph id="922136E09D9A92DDFF52FA42CD212E03" blockId="134.[189,1399,1456,1633]" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF52FA42CD212E03" pageId="134" pageNumber="135">
– Dorsal setae slightly clavate (widened apically), comparatively shorter (
<figureCitation id="0AA52A659D9A92DDFBD8FA45CCFF2E74" box="[1099,1193,1531,1558]" captionStart="Figs 58–61" captionStartId="75.[189,241,1928,1954]" captionTargetBox="[336,1249,266,1914]" captionTargetId="figure-62@75.[333,1249,257,1923]" captionTargetPageId="75" captionText="Figs 58–61. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 58. M. maleo sp. nov., holotype, ♂ (IRSN). 59. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 60. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 61. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122152" httpUri="https://zenodo.org/record/4122152/files/figure.png" pageId="134" pageNumber="135">Figs 58</figureCitation>
, 
<figureCitation id="0AA52A659D9A92DDFB2BFA45CCB12E77" box="[1208,1255,1531,1557]" captionStart="Figs 133–144" captionStartId="140.[189,241,1822,1848]" captionTargetBox="[346,1223,273,1784]" captionTargetId="figure-120@140.[345,1234,265,1799]" captionTargetPageId="140" captionText="Figs 133–144. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 133. M. lapsus sp. nov., holotype, ♂ (BMNH). 134. Same, paratype, ♀ (BMNH). 135. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 136. M. legalovi sp. nov., holotype, ♂ (BMNH). 137. Same, paratype, ♀ (BMNH). 138. M. longipes sp. nov., holotype, ♂ (BMNH). 139. M. lobaticeps Frey, 1969, ♂ (BMNH). 140. Same, ♀. 141. M. mailu sp. nov., holotype, ♂ (NHMB). 142. M. luniceps Fairmaire, 1883, ♂ (BMNH). 143. Same, ♀ (BMNH). 144. M. maleo sp. nov., holotype, ♂ (IRSN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122179" httpUri="https://zenodo.org/record/4122179/files/figure.png" pageId="134" pageNumber="135">144</figureCitation>
, 
<figureCitation id="0AA52A659D9A92DDFB65FA45CD702E77" box="[1270,1318,1531,1557]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="134" pageNumber="135">231</figureCitation>
); male labroclypeus comparatively deeper emarginate on anterior margin (
<figureCitation id="0AA52A659D9A92DDFBA0F99FCCF72E59" box="[1075,1185,1569,1595]" captionStart="Figs 133–144" captionStartId="140.[189,241,1822,1848]" captionTargetBox="[346,1223,273,1784]" captionTargetId="figure-120@140.[345,1234,265,1799]" captionTargetPageId="140" captionText="Figs 133–144. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 133. M. lapsus sp. nov., holotype, ♂ (BMNH). 134. Same, paratype, ♀ (BMNH). 135. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 136. M. legalovi sp. nov., holotype, ♂ (BMNH). 137. Same, paratype, ♀ (BMNH). 138. M. longipes sp. nov., holotype, ♂ (BMNH). 139. M. lobaticeps Frey, 1969, ♂ (BMNH). 140. Same, ♀. 141. M. mailu sp. nov., holotype, ♂ (NHMB). 142. M. luniceps Fairmaire, 1883, ♂ (BMNH). 143. Same, ♀ (BMNH). 144. M. maleo sp. nov., holotype, ♂ (IRSN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122179" httpUri="https://zenodo.org/record/4122179/files/figure.png" pageId="134" pageNumber="135">Fig. 144</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9A92DDFF79F9F8C9DD2E02" box="[234,395,1606,1632]" captionStart="Figs 652–666" captionStartId="175.[189,241,1892,1918]" captionTargetBox="[436,1156,267,1863]" captionTargetId="figure-70@175.[426,1161,265,1868]" captionTargetPageId="175" captionText="Figs 652–666. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 652– 654. M. longipes sp. nov., holotype, ♂ (BMNH). 655–657. M. luniceps Fairmaire, 1883, ♂ (BMNH). 658–660. M. mailu sp. nov., holotype, ♂ (NHMB). 661–663. M. maleo sp. nov., holotype, ♂ (IRSN). 664–666. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122244" httpUri="https://zenodo.org/record/4122244/files/figure.png" pageId="134" pageNumber="135">Figs 661–663</figureCitation>
........................................................................................................... 
<taxonomicName id="559E4D639D9A92DDFB30F9F9CD432E03" ID-CoL="3X3FH" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1187,1301,1606,1633]" class="Aves" family="Megapodiidae" genus="Macrocephalon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galliformes" pageId="134" pageNumber="135" phylum="Chordata" rank="species" species="maleo" status="sp. nov.">
<emphasis id="A0EAEAF29D9A92DDFB30F9F9CD432E03" bold="true" box="[1187,1301,1606,1633]" italics="true" pageId="134" pageNumber="135">M. maleo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9A92DDFA8FF9F9CD212E03" box="[1308,1399,1607,1633]" pageId="134" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9A92DDFF2EF934CD212EA8" pageId="134" pageNumber="135">
<paragraph id="922136E09D9A92DDFF2EF934CD212EC7" blockId="134.[189,1399,1674,1738]" box="[189,1399,1674,1701]" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF2EF934CD212EC7" box="[189,1399,1674,1701]" pageId="134" pageNumber="135">
49. Aedeagus as in 
<figureCitation id="0AA52A659D9A92DDFE30F934CA122EC6" box="[419,580,1674,1700]" captionStart="Figs 712–725" captionStartId="179.[189,241,1892,1918]" captionTargetBox="[433,1144,277,1858]" captionTargetId="figure-72@179.[427,1160,265,1870]" captionTargetPageId="179" captionText="Figs 712–725. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 712– 714. M. skalei sp. nov., holotype, ♂ (NME). 715–717. M. sougb sp. nov., holotype, ♂ (NME). 718–720. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 721–723. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 724–725. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122252" httpUri="https://zenodo.org/record/4122252/files/figure.png" pageId="134" pageNumber="135">Figs 712–714</figureCitation>
.................................................................................. 
<taxonomicName id="559E4D639D9A92DDFB35F935CD432EC7" ID-CoL="7X3XQ" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1190,1301,1674,1701]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="skalei" status="sp. nov.">
<emphasis id="A0EAEAF29D9A92DDFB35F935CD432EC7" bold="true" box="[1190,1301,1674,1701]" italics="true" pageId="134" pageNumber="135">M. skalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9A92DDFA8FF935CD212EC7" box="[1308,1399,1675,1701]" pageId="134" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9A92DDFF52F90ECD212EA8" blockId="134.[189,1399,1674,1738]" box="[193,1399,1712,1738]" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF52F90ECD212EA8" box="[193,1399,1712,1738]" pageId="134" pageNumber="135">
– Aedeagus as in 
<figureCitation id="0AA52A659D9A92DDFE30F90ECA122EA8" box="[419,580,1712,1738]" captionStart="Figs 682–696" captionStartId="177.[189,241,1892,1918]" captionTargetBox="[436,1163,269,1860]" captionTargetId="figure-72@177.[423,1174,265,1866]" captionTargetPageId="177" captionText="Figs 682–696. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 682– 684. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 685–687. M. pauxillus Heller, 1910 comb. rest., ♂ (BMNH). 688–690. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 691–693. M. peregrinus Lansberge, 1886, ♂ (DTC). 694-696. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122248" httpUri="https://zenodo.org/record/4122248/files/figure.png" pageId="134" pageNumber="135">Figs 691–693</figureCitation>
............................................................ 
<taxonomicName id="559E4D639D9A92DDFB92F90ECD212EA8" ID-CoL="7X3XG" authority="Lansberge, 1886" authorityName="Lansberge" authorityYear="1886" box="[1025,1399,1712,1738]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="A0EAEAF29D9A92DDFB92F90ECCFA2EA8" bold="true" box="[1025,1196,1712,1738]" italics="true" pageId="134" pageNumber="135">M. peregrinus</emphasis>
<bibRefCitation id="F60F4B119D9A92DDFB20F90ECD212EA8" author="Lansberge J. W. van" box="[1203,1399,1712,1738]" pageId="134" pageNumber="135" pagination="131 - 137" refId="ref98256" refString="Lansberge J. W. van 1886. Description des quelques scarabaeides des Indes neerlandaises, appartenant au Musee du Leyde. Notes from the Leyden Museum 8: 131 - 137." type="journal article" year="1886">Lansberge, 1886</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9A92DDFF2EF94ACD212F56" pageId="134" pageNumber="135">
<paragraph id="922136E09D9A92DDFF2EF94ACD202F6D" blockId="134.[189,1399,1780,1844]" box="[189,1398,1780,1807]" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF2EF94ACD202F6D" box="[189,1398,1780,1807]" pageId="134" pageNumber="135">50. Parameres entirely glabrous............................................................................................................. 51</keyLead>
</paragraph>
<paragraph id="922136E09D9A92DDFF52F8A4CD212F56" blockId="134.[189,1399,1780,1844]" box="[193,1399,1817,1844]" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF52F8A4CD212F56" box="[193,1399,1817,1844]" pageId="134" pageNumber="135">
– Parameres on apical margin with delicate cilia-like setae (
<figureCitation id="0AA52A659D9A92DDFC03F8A7CC622F56" box="[912,1076,1817,1844]" captionStart="Figs 586–600" captionStartId="171.[189,241,1892,1918]" captionTargetBox="[400,1175,268,1865]" captionTargetId="figure-72@171.[391,1196,265,1868]" captionTargetPageId="171" captionText="Figs 586–600. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 586– 588. M. bintang sp. nov., holotype, ♂ (NME). 589–591. M. boessnecki sp. nov., holotype, ♂ (SNSD). 592–594. M. caperatus sp. nov., holotype, ♂ (NME). 595–597. M. ciliatus sp. nov., holotype, ♂ (IECA). 598–600. M. crypticus sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122234" httpUri="https://zenodo.org/record/4122234/files/figure.png" pageId="134" pageNumber="135">Figs 595–596</figureCitation>
) .......... 
<taxonomicName id="559E4D639D9A92DDFB07F8A4CD432F56" ID-CoL="7X3WJ" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1172,1301,1817,1844]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="ciliatus" status="sp. nov.">
<emphasis id="A0EAEAF29D9A92DDFB07F8A4CD432F56" bold="true" box="[1172,1301,1817,1844]" italics="true" pageId="134" pageNumber="135">M. ciliatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9A92DDFA8FF8A4CD212F56" box="[1308,1399,1818,1844]" pageId="134" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9A92DDFF2EF8E3CD212F8B" pageId="134" pageNumber="135">
<paragraph id="922136E09D9A92DDFF2EF8E3CD202FFC" blockId="134.[189,1399,1885,2025]" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF2EF8E3CD202FFC" pageId="134" pageNumber="135">51. Lateral margins of pronotum in dorsal view distinctly rounded (more or less distinctly constricted anteriad and posteriad)..................................................................................................................... 52</keyLead>
</paragraph>
<paragraph id="922136E09D9A92DDFF52F817CD212F8B" blockId="134.[189,1399,1885,2025]" pageId="134" pageNumber="135">
<keyLead id="296F95D59D9A92DDFF52F817CD212F8B" pageId="134" pageNumber="135">
– Lateral margins of pronotum in dorsal view nearly straight, not distinctly constricted anteriad and posteriad (
<figureCitation id="0AA52A659D9A92DDFEF5F870C9982F8A" box="[358,462,1998,2024]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="134" pageNumber="135">Fig. 230</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9A92DDFD00F870CB652F8B" box="[659,819,1998,2025]" captionStart="Figs 652–666" captionStartId="175.[189,241,1892,1918]" captionTargetBox="[436,1156,267,1863]" captionTargetId="figure-70@175.[426,1161,265,1868]" captionTargetPageId="175" captionText="Figs 652–666. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 652– 654. M. longipes sp. nov., holotype, ♂ (BMNH). 655–657. M. luniceps Fairmaire, 1883, ♂ (BMNH). 658–660. M. mailu sp. nov., holotype, ♂ (NHMB). 661–663. M. maleo sp. nov., holotype, ♂ (IRSN). 664–666. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122244" httpUri="https://zenodo.org/record/4122244/files/figure.png" pageId="134" pageNumber="135">Figs 658–660</figureCitation>
.................................................. 
<taxonomicName id="559E4D639D9A92DDFB35F871CD432F8B" ID-CoL="7X3X4" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1190,1301,1998,2025]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="mailu" status="sp. nov.">
<emphasis id="A0EAEAF29D9A92DDFB35F871CD432F8B" bold="true" box="[1190,1301,1998,2025]" italics="true" pageId="134" pageNumber="135">M. mailu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9A92DDFA8FF871CD212F8B" box="[1308,1399,1999,2025]" pageId="134" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EFEB7CD2129F9" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EFEB7CD20292E" blockId="135.[189,1399,265,412]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EFEB7CD20292E" pageId="135" pageNumber="136">52. Hypomeron separated from prosternum by flange-like carina; preapical hump of elytron not Г- shaped........................................................................................................................................... 53</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52FEE4CD2129F9" blockId="135.[189,1399,265,412]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52FEE4CD2129F9" pageId="135" pageNumber="136">
– Hypomeron separated from prosternum by moderately high carina; preapical hump of elytron Г- shaped.......................................................................................................... 
<taxonomicName id="559E4D639D9B92DCFBFFFE3FCD2129F9" ID-CoL="3XFLF" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[1132,1399,384,412]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="A0EAEAF29D9B92DCFBFFFE3FCCA429F9" bold="true" box="[1132,1266,384,411]" italics="true" pageId="135" pageNumber="136">M. simplex</emphasis>
<bibRefCitation id="F60F4B119D9B92DCFB6AFE3CCD2129F9" author="Frey G." box="[1273,1399,385,412]" pageId="135" pageNumber="136" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EFE79CD202A50" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EFE79CD212A68" blockId="135.[189,1399,455,562]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EFE79CD212A68" pageId="135" pageNumber="136">
53. Dorsal setae at least in part distinctly clavate (distinctly widened apically) ...................................... ..................................................................................................................... 66 [
<emphasis id="A0EAEAF29D9B92DCFB11FE4ECD3A2A6B" bold="true" box="[1154,1388,495,522]" pageId="135" pageNumber="136">
<taxonomicName id="559E4D639D9B92DCFB11FE4ECD4C2A68" authorityName="Heller" authorityYear="1910" box="[1154,1306,495,522]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="pauxillus">
<emphasis id="A0EAEAF29D9B92DCFB11FE4ECD4C2A68" bold="true" box="[1154,1306,495,522]" italics="true" pageId="135" pageNumber="136">M. pauxillus</emphasis>
</taxonomicName>
group
</emphasis>
]
</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52FDA6CD202A50" blockId="135.[189,1399,455,562]" box="[193,1398,535,562]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52FDA6CD202A50" box="[193,1398,535,562]" pageId="135" pageNumber="136">– Dorsal setae ordinary or slightly widened apically.......................................................................... 54</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EFDE3CD212AC2" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EFDE3CD202A1A" blockId="135.[189,1399,605,673]" box="[189,1398,605,632]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EFDE3CD202A1A" box="[189,1398,605,632]" pageId="135" pageNumber="136">
54. Species from New 
<collectingCountry id="EA8976709D9B92DCFE59FDE3CA492A15" box="[458,543,605,631]" name="Guinea-Bissau" pageId="135" pageNumber="136">Guinea</collectingCountry>
or Sulawesi ........................................................................................... 55
</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52FD38CD212AC2" blockId="135.[189,1399,605,673]" box="[193,1399,645,673]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52FD38CD212AC2" box="[193,1399,645,673]" pageId="135" pageNumber="136">
– Species from Lesser Sunda Islands (Bali); aedeagus as in 
<figureCitation id="0AA52A659D9B92DCFC1CFD38CC662AC2" box="[911,1072,645,672]" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="135" pageNumber="136">Figs 643–645</figureCitation>
........ 
<taxonomicName id="559E4D639D9B92DCFBE4FD38CD432AC2" ID-CoL="7X3WZ" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1143,1301,645,672]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="leucopsar" status="sp. nov.">
<emphasis id="A0EAEAF29D9B92DCFBE4FD38CD432AC2" bold="true" box="[1143,1301,645,672]" italics="true" pageId="135" pageNumber="136">M. leucopsar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9B92DCFA8FFD38CD212AC2" box="[1308,1399,646,672]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EFD72CD202B6D" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EFD72CD202A85" blockId="135.[189,1398,716,783]" box="[189,1398,716,743]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EFD72CD202A85" box="[189,1398,716,743]" pageId="135" pageNumber="136">55. Male labroclypeus subtruncate, truncate or shallowly emarginate on anterior margin .................. 56</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52FD4ACD202B6D" blockId="135.[189,1398,716,783]" box="[193,1398,756,783]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52FD4ACD202B6D" box="[193,1398,756,783]" pageId="135" pageNumber="136">– Male labroclypeus distinctly emarginate on anterior margin .......................................................... 58</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EFC84CD212B1E" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EFC84CD202B37" blockId="135.[189,1399,826,893]" box="[189,1398,826,853]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EFC84CD202B37" box="[189,1398,826,853]" pageId="135" pageNumber="136">
56. Species from Sulawesi; aedeagus as in 
<figureCitation id="0AA52A659D9B92DCFD25FC84CB002B37" box="[694,854,826,853]" captionStart="Figs 586–600" captionStartId="171.[189,241,1892,1918]" captionTargetBox="[400,1175,268,1865]" captionTargetId="figure-72@171.[391,1196,265,1868]" captionTargetPageId="171" captionText="Figs 586–600. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 586– 588. M. bintang sp. nov., holotype, ♂ (NME). 589–591. M. boessnecki sp. nov., holotype, ♂ (SNSD). 592–594. M. caperatus sp. nov., holotype, ♂ (NME). 595–597. M. ciliatus sp. nov., holotype, ♂ (IECA). 598–600. M. crypticus sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122234" httpUri="https://zenodo.org/record/4122234/files/figure.png" pageId="135" pageNumber="136">Figs 589–591</figureCitation>
or 726–729 ................................................. 57
</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52FCDDCD212B1E" blockId="135.[189,1399,826,893]" box="[193,1399,866,893]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52FCDDCD212B1E" box="[193,1399,866,893]" pageId="135" pageNumber="136">
– Species from New 
<collectingCountry id="EA8976709D9B92DCFE59FCDCCA742B1E" box="[458,546,866,892]" name="Guinea-Bissau" pageId="135" pageNumber="136">Guinea</collectingCountry>
; aedeagus as in 
<figureCitation id="0AA52A659D9B92DCFD4CFCDCCBD62B1F" box="[735,896,866,893]" captionStart="Figs 667–681" captionStartId="176.[189,241,1892,1918]" captionTargetBox="[444,1142,267,1860]" captionTargetId="figure-72@176.[419,1168,265,1868]" captionTargetPageId="176" captionText="Figs 667–681. Papuan Maechidius Macleay, 1819, aedeagus, different aspects. 667– 669. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 670–672. M. milneanus Heller, 1914, ♂ (SNSD). 673–675. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 676–678. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 679-681. M. nepenthephilus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122246" httpUri="https://zenodo.org/record/4122246/files/figure.png" pageId="135" pageNumber="136">Figs 673–675</figureCitation>
............................ 
<taxonomicName id="559E4D639D9B92DCFBC1FCDCCD212B1E" ID-CoL="3XFKC" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[1106,1399,866,892]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="muticus">
<emphasis id="A0EAEAF29D9B92DCFBC1FCDCCC8D2B1E" bold="true" box="[1106,1243,866,892]" italics="true" pageId="135" pageNumber="136">M. muticus</emphasis>
<bibRefCitation id="F60F4B119D9B92DCFB71FCDCCD212B1E" author="Arrow G. E." box="[1250,1399,866,892]" pageId="135" pageNumber="136" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EFC16CD212C59" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EFC16CD212B89" blockId="135.[189,1399,936,1083]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EFC16CD212B89" pageId="135" pageNumber="136">
57. Aedeagus as in 
<figureCitation id="0AA52A659D9B92DCFE3BFC16CA1B2BA1" box="[424,589,936,963]" captionStart="Figs 586–600" captionStartId="171.[189,241,1892,1918]" captionTargetBox="[400,1175,268,1865]" captionTargetId="figure-72@171.[391,1196,265,1868]" captionTargetPageId="171" captionText="Figs 586–600. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 586– 588. M. bintang sp. nov., holotype, ♂ (NME). 589–591. M. boessnecki sp. nov., holotype, ♂ (SNSD). 592–594. M. caperatus sp. nov., holotype, ♂ (NME). 595–597. M. ciliatus sp. nov., holotype, ♂ (IECA). 598–600. M. crypticus sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122234" httpUri="https://zenodo.org/record/4122234/files/figure.png" pageId="135" pageNumber="136">Figs 589–591</figureCitation>
; lateral margin of each elytron with a row of short setae; elytron on disc with track of single longitudinal carina.................................................... 
<taxonomicName id="559E4D639D9B92DCFBF9FC6FCD432B89" ID-CoL="7X3WD" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1130,1301,976,1003]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="boessnecki" status="sp. nov.">
<emphasis id="A0EAEAF29D9B92DCFBF9FC6FCD432B89" bold="true" box="[1130,1301,976,1003]" italics="true" pageId="135" pageNumber="136">M. boessnecki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9B92DCFA8FFC6FCD212B89" box="[1308,1399,977,1003]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52FC47CD212C59" blockId="135.[189,1399,936,1083]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52FC47CD212C59" pageId="135" pageNumber="136">
– Aedeagus as in 
<figureCitation id="0AA52A659D9B92DCFE0CFC46CA142C70" box="[415,578,1016,1042]" captionStart="Figs 726–738" captionStartId="180.[189,241,1891,1917]" captionTargetBox="[343,1246,278,1849]" captionTargetId="figure-69@180.[330,1258,265,1871]" captionTargetPageId="180" captionText="Figs 726–738. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 726– 729. M. suwawa sp. nov., holotype, ♂ (BMNH). 730–732. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 733–735. M. trivialis sp. nov., holotype, ♂ (BMNH). 736–738. M. vicinus Heller, 1914, ♂ (apex of parameres broken) (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122254" httpUri="https://zenodo.org/record/4122254/files/figure.png" pageId="135" pageNumber="136">Figs 726–729</figureCitation>
; lateral margin of each elytron with a row of long setae; elytron on disc with tracks of two longitudinal carinae ............................................................... 
<taxonomicName id="559E4D639D9B92DCFB1FFB9FCD432C59" ID-CoL="7X3XT" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1164,1301,1057,1083]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="suwawa" status="sp. nov.">
<emphasis id="A0EAEAF29D9B92DCFB1FFB9FCD432C59" bold="true" box="[1164,1301,1057,1083]" italics="true" pageId="135" pageNumber="136">M. suwawa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9B92DCFA8FFB9FCD212C59" box="[1308,1399,1057,1083]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EFBD9CD202D40" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EFBD9CD202CB3" blockId="135.[189,1399,1127,1314]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EFBD9CD202CB3" pageId="135" pageNumber="136">58. Dorsal setae of forebody inconspicuous, appressed to suberect; setae short, generally not or slightly surpassing length of corresponding punctures (some setae can be longer than corresponding punctures, but most of them are shorter) ......................................................................................... 62</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52FB61CD202D40" blockId="135.[189,1399,1127,1314]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52FB61CD202D40" pageId="135" pageNumber="136">– Dorsal setae of forebody suberect to erect, distinctly longer than length of corresponding punctures.......................................................................................................................................... 59</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EFAF3CD212DF2" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EFAF3CD202D05" blockId="135.[189,1399,1357,1424]" box="[189,1398,1357,1384]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EFAF3CD202D05" box="[189,1398,1357,1384]" pageId="135" pageNumber="136">59. Lateral margin of elytron with a row of short setae......................................................................... 60</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52FAC8CD212DF2" blockId="135.[189,1399,1357,1424]" box="[193,1399,1397,1424]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52FAC8CD212DF2" box="[193,1399,1397,1424]" pageId="135" pageNumber="136">
– Lateral margin of elytron with a row of extraordinarily long setae (
<figureCitation id="0AA52A659D9B92DCFC6AFACBCC192DED" box="[1017,1103,1397,1423]" captionStart="Figs 22–24" captionStartId="22.[189,241,1928,1954]" captionTargetBox="[354,1240,272,1893]" captionTargetId="figure-55@22.[344,1244,264,1901]" captionTargetPageId="22" captionText="Figs 22–24. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 22. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 23. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 24. M. awu sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122132" httpUri="https://zenodo.org/record/4122132/files/figure.png" pageId="135" pageNumber="136">Fig. 24</figureCitation>
) ........... 
<taxonomicName id="559E4D639D9B92DCFB29FACBCD432DED" ID-CoL="7X3W8" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1210,1301,1397,1423]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="awu" status="sp. nov.">
<emphasis id="A0EAEAF29D9B92DCFB29FACBCD432DED" bold="true" box="[1210,1301,1397,1423]" italics="true" pageId="135" pageNumber="136">M. awu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9B92DCFA8FFAC8CD212DF2" box="[1308,1399,1398,1424]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EFA05CD212E2C" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EFA05CD202D9F" blockId="135.[189,1399,1467,1614]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EFA05CD202D9F" pageId="135" pageNumber="136">60. Pronotum dorsally with shallow circular impression on either side of median part; punctures of pronotum comparatively larger and more circular ......................................................................... 61</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52F9B2CD212E2C" blockId="135.[189,1399,1467,1614]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52F9B2CD212E2C" pageId="135" pageNumber="136">
– Pronotum dorsally without impressions; punctures of pronotum comparatively smaller and stronger ovoid (
<figureCitation id="0AA52A659D9B92DCFEAEF98DC9F22E2C" box="[317,420,1587,1614]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="135" pageNumber="136">Fig. 287</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9B92DCFDF9F98DCB5D2E2C" box="[618,779,1587,1614]" captionStart="Figs 570–585" captionStartId="170.[189,241,1892,1918]" captionTargetBox="[425,1155,265,1858]" captionTargetId="figure-74@170.[416,1172,265,1864]" captionTargetPageId="170" captionText="Figs 570–585. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 570– 572. M. aiyura sp. nov., holotype, ♂ (BMNH). 573–575. M. alesbezdeki sp. nov., holotype, ♂ (NME). 576–578. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 579–581. M. awu sp. nov., holotype, ♂ (IECA). 582–585. M. babyrousa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122232" httpUri="https://zenodo.org/record/4122232/files/figure.png" pageId="135" pageNumber="136">Figs 570–572</figureCitation>
...................................................... 
<taxonomicName id="559E4D639D9B92DCFB0DF98ACD432E2F" ID-CoL="7X3W3" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1182,1301,1587,1614]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="aiyura" status="sp. nov.">
<emphasis id="A0EAEAF29D9B92DCFB0DF98ACD432E2F" bold="true" box="[1182,1301,1587,1614]" italics="true" pageId="135" pageNumber="136">M. aiyura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9B92DCFA8FF98ACD212E2C" box="[1308,1399,1588,1614]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EF9C4CD212F6E" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EF9C4CD212EDE" blockId="135.[189,1399,1657,1804]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EF9C4CD212EDE" pageId="135" pageNumber="136">
61. Aedeagus as in 
<figureCitation id="0AA52A659D9B92DCFE51F9C4CA272EF6" box="[450,625,1657,1684]" captionStart="Figs 570–585" captionStartId="170.[189,241,1892,1918]" captionTargetBox="[425,1155,265,1858]" captionTargetId="figure-74@170.[416,1172,265,1864]" captionTargetPageId="170" captionText="Figs 570–585. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 570– 572. M. aiyura sp. nov., holotype, ♂ (BMNH). 573–575. M. alesbezdeki sp. nov., holotype, ♂ (NME). 576–578. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 579–581. M. awu sp. nov., holotype, ♂ (IECA). 582–585. M. babyrousa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122232" httpUri="https://zenodo.org/record/4122232/files/figure.png" pageId="135" pageNumber="136">Figs 582–585</figureCitation>
; dorsal punctures of pronotum smaller and generally ovoid (
<figureCitation id="0AA52A659D9B92DCFF60F91CC9012EDE" box="[243,343,1698,1724]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="135" pageNumber="136">Fig. 203</figureCitation>
) ......................................................................................................... 
<taxonomicName id="559E4D639D9B92DCFBFCF91CCD422EDE" ID-CoL="7X3W9" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1135,1300,1697,1724]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="babyrousa" status="sp. nov.">
<emphasis id="A0EAEAF29D9B92DCFBFCF91CCD422EDE" bold="true" box="[1135,1300,1697,1724]" italics="true" pageId="135" pageNumber="136">M. babyrousa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9B92DCFA8FF91CCD212EDE" box="[1308,1399,1698,1724]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52F974CD212F6E" blockId="135.[189,1399,1657,1804]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52F974CD212F6E" pageId="135" pageNumber="136">
– Aedeagus as in 
<figureCitation id="0AA52A659D9B92DCFE52F974CA392E86" box="[449,623,1738,1764]" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="135" pageNumber="136">Figs 607–610</figureCitation>
; dorsal punctures of pronotum larger and generally circular (
<figureCitation id="0AA52A659D9B92DCFF60F94CC9002F6E" box="[243,342,1778,1804]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="135" pageNumber="136">Fig. 211</figureCitation>
)............................................................................................................... 
<taxonomicName id="559E4D639D9B92DCFB1DF94CCD432F6E" ID-CoL="7X3WM" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1166,1301,1777,1804]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="deltouri" status="sp. nov.">
<emphasis id="A0EAEAF29D9B92DCFB1DF94CCD432F6E" bold="true" box="[1166,1301,1777,1804]" italics="true" pageId="135" pageNumber="136">M. deltouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9B92DCFA8FF94CCD212F6E" box="[1308,1399,1778,1804]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EF886CD212F19" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EF886CD202F30" blockId="135.[189,1399,1848,1915]" box="[189,1398,1848,1875]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EF886CD202F30" box="[189,1398,1848,1875]" pageId="135" pageNumber="136">
62. Species from New 
<collectingCountry id="EA8976709D9B92DCFE59F886CA742F30" box="[458,546,1848,1874]" name="Guinea-Bissau" pageId="135" pageNumber="136">Guinea</collectingCountry>
; aedeagus not as in 
<figureCitation id="0AA52A659D9B92DCFC9FF886CBFA2F30" box="[780,940,1848,1874]" captionStart="Figs 617–631" captionStartId="173.[189,241,1892,1918]" captionTargetBox="[439,1134,274,1859]" captionTargetId="figure-71@173.[434,1154,265,1868]" captionTargetPageId="173" captionText="Figs 617–631. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 617– 619. M. hamatus sp. nov., holotype, ♂ (IECA). 620–622. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 623–625. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 626–628. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 629–631. M. konjo sp. nov., holotype, ♂ (NMPC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122240" httpUri="https://zenodo.org/record/4122240/files/figure.png" pageId="135" pageNumber="136">Figs 629–631</figureCitation>
........................................................ 63
</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52F8DFCD212F19" blockId="135.[189,1399,1848,1915]" box="[193,1399,1887,1915]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52F8DFCD212F19" box="[193,1399,1887,1915]" pageId="135" pageNumber="136">
– Species from Sulawesi; aedeagus as in 
<figureCitation id="0AA52A659D9B92DCFD25F8DECB002F18" box="[694,854,1888,1914]" captionStart="Figs 617–631" captionStartId="173.[189,241,1892,1918]" captionTargetBox="[439,1134,274,1859]" captionTargetId="figure-71@173.[434,1154,265,1868]" captionTargetPageId="173" captionText="Figs 617–631. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 617– 619. M. hamatus sp. nov., holotype, ♂ (IECA). 620–622. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 623–625. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 626–628. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 629–631. M. konjo sp. nov., holotype, ♂ (NMPC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122240" httpUri="https://zenodo.org/record/4122240/files/figure.png" pageId="135" pageNumber="136">Figs 629–631</figureCitation>
............................................. 
<taxonomicName id="559E4D639D9B92DCFB3BF8DECD422F18" ID-CoL="7X3WW" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1192,1300,1887,1914]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="konjo" status="sp. nov.">
<emphasis id="A0EAEAF29D9B92DCFB3BF8DECD422F18" bold="true" box="[1192,1300,1887,1914]" italics="true" pageId="135" pageNumber="136">M. konjo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9B92DCFA8FF8DFCD212F19" box="[1308,1399,1889,1915]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9B92DCFF2EF818CD202F8B" pageId="135" pageNumber="136">
<paragraph id="922136E09D9B92DCFF2EF818CD202FA2" blockId="135.[189,1398,1958,2025]" box="[189,1398,1958,1985]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF2EF818CD202FA2" box="[189,1398,1958,1985]" pageId="135" pageNumber="136">63. Anterolateral angles of labroclypeus either acute or right-angled................................................... 64</keyLead>
</paragraph>
<paragraph id="922136E09D9B92DCFF52F871CD202F8B" blockId="135.[189,1398,1958,2025]" box="[193,1398,1998,2025]" pageId="135" pageNumber="136">
<keyLead id="296F95D59D9B92DCFF52F871CD202F8B" box="[193,1398,1998,2025]" pageId="135" pageNumber="136">– Anterolateral angles of labroclypeus strongly obtuse...................................................................... 65</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9492D3FF2EFEB7CD2129B7" pageId="136" pageNumber="137">
<paragraph id="922136E09D9492D3FF2EFEB7CD212909" blockId="136.[189,1399,265,469]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EFEB7CD212909" pageId="136" pageNumber="137">
64. Aedeagus as in 
<figureCitation id="0AA52A659D9492D3FE5FFEB7CA2B2941" box="[460,637,265,291]" captionStart="Figs 667–681" captionStartId="176.[189,241,1892,1918]" captionTargetBox="[444,1142,267,1860]" captionTargetId="figure-72@176.[419,1168,265,1868]" captionTargetPageId="176" captionText="Figs 667–681. Papuan Maechidius Macleay, 1819, aedeagus, different aspects. 667– 669. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 670–672. M. milneanus Heller, 1914, ♂ (SNSD). 673–675. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 676–678. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 679-681. M. nepenthephilus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122246" httpUri="https://zenodo.org/record/4122246/files/figure.png" pageId="136" pageNumber="137">Figs 667–669</figureCitation>
; male upper metatibial terminal spur half the length of male basal metatarsomere, not surpassing distal margin of metatibia in dorsal view (
<figureCitation id="0AA52A659D9492D3FF60FEEEC91F2909" box="[243,329,336,363]" captionStart="Figs 58–61" captionStartId="75.[189,241,1928,1954]" captionTargetBox="[336,1249,266,1914]" captionTargetId="figure-62@75.[333,1249,257,1923]" captionTargetPageId="75" captionText="Figs 58–61. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 58. M. maleo sp. nov., holotype, ♂ (IRSN). 59. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 60. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 61. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122152" httpUri="https://zenodo.org/record/4122152/files/figure.png" pageId="136" pageNumber="137">Fig. 59</figureCitation>
) ................................................................................................ 
<taxonomicName id="559E4D639D9492D3FB89FEEECD432908" ID-CoL="7X3X7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1050,1301,335,362]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="miklouhomaclayi" status="sp. nov.">
<emphasis id="A0EAEAF29D9492D3FB89FEEECD432908" bold="true" box="[1050,1301,335,362]" italics="true" pageId="136" pageNumber="137">M. miklouhomaclayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9492D3FA8FFEEFCD212909" box="[1308,1399,337,363]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF52FECACD2129B7" blockId="136.[189,1399,265,469]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF52FECACD2129B7" pageId="136" pageNumber="137">
– Aedeagus as in 
<figureCitation id="0AA52A659D9492D3FE39FECDCA0729EC" box="[426,593,371,398]" captionStart="Figs 570–585" captionStartId="170.[189,241,1892,1918]" captionTargetBox="[425,1155,265,1858]" captionTargetId="figure-74@170.[416,1172,265,1864]" captionTargetPageId="170" captionText="Figs 570–585. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 570– 572. M. aiyura sp. nov., holotype, ♂ (BMNH). 573–575. M. alesbezdeki sp. nov., holotype, ♂ (NME). 576–578. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 579–581. M. awu sp. nov., holotype, ♂ (IECA). 582–585. M. babyrousa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122232" httpUri="https://zenodo.org/record/4122232/files/figure.png" pageId="136" pageNumber="137">Figs 576–578</figureCitation>
; male upper metatibial terminal spur longer than half the length of male basal metatarsomere, surpassing distal margin of metatibia in dorsal view ............................... .............................................................................................................. 
<taxonomicName id="559E4D639D9492D3FBB6FE05CD2129B7" ID-CoL="72TG2" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[1061,1399,442,469]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="A0EAEAF29D9492D3FBB6FE05CC8D29B6" bold="true" box="[1061,1243,442,469]" italics="true" pageId="136" pageNumber="137">M. angusticeps</emphasis>
<bibRefCitation id="F60F4B119D9492D3FB71FE05CD2129B7" author="Arrow G. E." box="[1250,1399,443,469]" pageId="136" pageNumber="137" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9492D3FF2EFE42CD212AC6" pageId="136" pageNumber="137">
<paragraph id="922136E09D9492D3FF2EFE42CD212A3F" blockId="136.[189,1399,508,677]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EFE42CD212A3F" pageId="136" pageNumber="137">
65. Elytral longitudinal carinae glabrous, glossy, stronger elevated; sutural carina present; intervening spaces on pygidium opaque, microreticulate; aedeagus unknown ...................................................... ............................................................................................... 
<taxonomicName id="559E4D639D9492D3FC23FDFACD212A3F" ID-CoL="3XFJN" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[944,1399,579,606]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="interruptocarinulatus">
<emphasis id="A0EAEAF29D9492D3FC23FDFACC882A3C" bold="true" box="[944,1246,579,606]" italics="true" pageId="136" pageNumber="137">M. interruptocarinulatus</emphasis>
<bibRefCitation id="F60F4B119D9492D3FB76FDFDCD212A3F" author="Heller K. M." box="[1253,1399,579,605]" pageId="136" pageNumber="137" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF52FDD9CD212AC6" blockId="136.[189,1399,508,677]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF52FDD9CD212AC6" pageId="136" pageNumber="137">
– Elytra without elevated tracks of longitudinal carinae; sutural carina not indicated; intervening spaces on pygidium glossy; aedeagus unknown................................ 
<taxonomicName id="559E4D639D9492D3FB90FD35CD212AC6" ID-CoL="3XFL9" authority="Moser, 1920" authorityName="Moser" authorityYear="1920" box="[1027,1399,650,677]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="seriepunctatus">
<emphasis id="A0EAEAF29D9492D3FB90FD35CC8A2AC6" bold="true" box="[1027,1244,650,677]" italics="true" pageId="136" pageNumber="137">M. seriepunctatus</emphasis>
<bibRefCitation id="F60F4B119D9492D3FB70FD35CD212AC6" author="Moser J." box="[1251,1399,650,677]" pageId="136" pageNumber="137" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9492D3FF2EFD72CD202B33" pageId="136" pageNumber="137">
<paragraph id="922136E09D9492D3FF2EFD72CD202B4F" blockId="136.[189,1398,716,849]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EFD72CD202B4F" pageId="136" pageNumber="137">66. Elytron with elevated glossy and glabrous longitudinal carina(e) (these carinae are more or less widely interrupted, in some specimens strongly reduced to few humps per carina and can be limited to sutural carina only) ...................................................................................................................... 67</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF52FC89CD202B33" blockId="136.[189,1398,716,849]" box="[193,1398,822,849]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF52FC89CD202B33" box="[193,1398,822,849]" pageId="136" pageNumber="137">– Longitudinal carina(e) of elytron when present not elevated nor glossy or glabrous...................... 70</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9492D3FF2EFCC6CD212B9F" pageId="136" pageNumber="137">
<paragraph id="922136E09D9492D3FF2EFCC6CD202BD4" blockId="136.[189,1399,888,1022]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EFCC6CD202BD4" pageId="136" pageNumber="137">
67. Dorsal setae on lateral parts of pronotum scale-like but not extraordinarily enlarged and not arranged in two lateral longitudinal stripes; aedeagus not as in 
<figureCitation id="0AA52A659D9492D3FCC8FC22CBAD2BD5" box="[859,1019,924,951]" captionStart="Figs 586–600" captionStartId="171.[189,241,1892,1918]" captionTargetBox="[400,1175,268,1865]" captionTargetId="figure-72@171.[391,1196,265,1868]" captionTargetPageId="171" captionText="Figs 586–600. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 586– 588. M. bintang sp. nov., holotype, ♂ (NME). 589–591. M. boessnecki sp. nov., holotype, ♂ (SNSD). 592–594. M. caperatus sp. nov., holotype, ♂ (NME). 595–597. M. ciliatus sp. nov., holotype, ♂ (IECA). 598–600. M. crypticus sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122234" httpUri="https://zenodo.org/record/4122234/files/figure.png" pageId="136" pageNumber="137">Figs 586–588</figureCitation>
.............................................. 68
</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF52FC7ECD212B9F" blockId="136.[189,1399,888,1022]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF52FC7ECD212B9F" pageId="136" pageNumber="137">
– Dorsal setae on lateral parts of pronotum scale-like and extraordinarily enlarged, as in 
<figureCitation id="0AA52A659D9492D3FB4DFC01CD612BBB" box="[1246,1335,959,985]" captionStart="Figs 25–27" captionStartId="26.[189,241,1928,1954]" captionTargetBox="[381,1192,273,1894]" captionTargetId="figure-49@26.[361,1255,262,1894]" captionTargetPageId="26" captionText="Figs 25–27. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal and lateral views. 25. M. babyrousa sp. nov., holotype, ♂ (BMNH). 26–27. M. bintang sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122134" httpUri="https://zenodo.org/record/4122134/files/figure.png" pageId="136" pageNumber="137">Figs 26</figureCitation>
, 
<figureCitation id="0AA52A659D9492D3FAD0FC01CD242BBB" box="[1347,1394,959,985]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="136" pageNumber="137">204</figureCitation>
, arranged in two lateral longitudinal rows; aedeagus as in 
<figureCitation id="0AA52A659D9492D3FC1AFC5DCC7F2B9C" box="[905,1065,995,1022]" captionStart="Figs 586–600" captionStartId="171.[189,241,1892,1918]" captionTargetBox="[400,1175,268,1865]" captionTargetId="figure-72@171.[391,1196,265,1868]" captionTargetPageId="171" captionText="Figs 586–600. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 586– 588. M. bintang sp. nov., holotype, ♂ (NME). 589–591. M. boessnecki sp. nov., holotype, ♂ (SNSD). 592–594. M. caperatus sp. nov., holotype, ♂ (NME). 595–597. M. ciliatus sp. nov., holotype, ♂ (IECA). 598–600. M. crypticus sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122234" httpUri="https://zenodo.org/record/4122234/files/figure.png" pageId="136" pageNumber="137">Figs 586–588</figureCitation>
............. 
<taxonomicName id="559E4D639D9492D3FB1CFC5DCD432B9F" ID-CoL="7X3WC" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1167,1301,994,1021]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="bintang" status="sp. nov.">
<emphasis id="A0EAEAF29D9492D3FB1CFC5DCD432B9F" bold="true" box="[1167,1301,994,1021]" italics="true" pageId="136" pageNumber="137">M. bintang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9492D3FA8FFC5DCD212B9F" box="[1308,1399,995,1021]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9492D3FF2EFB9ACD212C93" pageId="136" pageNumber="137">
<paragraph id="922136E09D9492D3FF2EFB9ACD202CE4" blockId="136.[189,1399,1060,1265]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EFB9ACD202CE4" pageId="136" pageNumber="137">68. Lateral margin of pronotum nearly straight to gradually slightly widened towards postmedium, with shallow prebasal emargination; elytron with less than four longitudinal carinae including sutural one.................................................................................................................................................... 69</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF52FB2ECD212C93" blockId="136.[189,1399,1060,1265]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF52FB2ECD212C93" pageId="136" pageNumber="137">
– Lateral margin of pronotum sinuous – somewhat emarginate in anterior and posterior half, obtuse angulate medially (
<figureCitation id="0AA52A659D9492D3FE47FB0DCA162CAC" box="[468,576,1203,1230]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="136" pageNumber="137">Fig. 216</figureCitation>
); elytron with four in part interrupted elevated longitudinal carinae including sutural one................................................... 
<taxonomicName id="559E4D639D9492D3FCC9FB68CCB82C92" ID-CoL="7X3WT" authority="Heller, 1910" authorityName="Heller" authorityYear="1910" box="[858,1262,1238,1265]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="heterosquamosus">
<emphasis id="A0EAEAF29D9492D3FCC9FB68CC032C93" bold="true" box="[858,1109,1238,1265]" italics="true" pageId="136" pageNumber="137">M. heterosquamosus</emphasis>
<bibRefCitation id="F60F4B119D9492D3FBCFFB68CCB82C92" author="Heller K. M." box="[1116,1262,1238,1264]" pageId="136" pageNumber="137" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller, 1910</bibRefCitation>
</taxonomicName>
comb. rest.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9492D3FF2EFAA6CD212DFF" pageId="136" pageNumber="137">
<paragraph id="922136E09D9492D3FF2EFAA6CD212D50" blockId="136.[189,1399,1303,1437]" box="[189,1399,1303,1331]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EFAA6CD212D50" box="[189,1399,1303,1331]" pageId="136" pageNumber="137">
69. Aedeagus as in 
<figureCitation id="0AA52A659D9492D3FE30FAA6CA102D50" box="[419,582,1304,1330]" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="136" pageNumber="137">Figs 611–613</figureCitation>
; male pygidium as in Fig. 543 ...................... 
<taxonomicName id="559E4D639D9492D3FBC5FAA6CD432D50" ID-CoL="7X3WN" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1110,1301,1303,1330]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="dendrolagus" status="sp. nov.">
<emphasis id="A0EAEAF29D9492D3FBC5FAA6CD432D50" bold="true" box="[1110,1301,1303,1330]" italics="true" pageId="136" pageNumber="137">M. dendrolagus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9492D3FA8FFAA6CD212D50" box="[1308,1399,1304,1330]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF2EFA82CD212D34" blockId="136.[189,1399,1303,1437]" box="[189,1399,1339,1366]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EFA82CD212D34" box="[189,1399,1339,1366]" pageId="136" pageNumber="137">
– Aedeagus as in 
<figureCitation id="0AA52A659D9492D3FE30FA85CA112D37" box="[419,583,1339,1365]" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="136" pageNumber="137">Figs 632–639</figureCitation>
; male and female pygidium as in 
<figureCitation id="0AA52A659D9492D3FC41FA85CC252D34" box="[978,1139,1339,1366]" captionStart="Figs 469–486" captionStartId="164.[189,241,1681,1707]" captionTargetBox="[450,1137,265,1633]" captionTargetId="figure-184@164.[450,1138,265,1647]" captionTargetPageId="164" captionText="Figs 469–486. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pygidium, dorsal view. 469. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 470. M. esau Heller, 1914, ♂ (BMNH). 471. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 472. M. hamatus sp. nov., holotype, ♂ (IECA). 473. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 474. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 475. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 476. Same, paralectotype, ♀ (SNSD). 477. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 478. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 479. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 480. M. konjo sp. nov., holotype, ♂ (NMPC). 481. M. lapsus sp. nov., holotype, ♂ (BMNH). 482. Same, paratype, ♀ (BMNH). 483. M. legalovi sp. nov., holotype, ♂ (BMNH). 484. Same, paratype, ♀ (BMNH). 485. M. leucopsar sp. nov., holotype, ♂ (IECA). 486. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122224" httpUri="https://zenodo.org/record/4122224/files/figure.png" pageId="136" pageNumber="137">Figs 481–482</figureCitation>
... 
<taxonomicName id="559E4D639D9492D3FB33FA82CD432D34" ID-CoL="7X3WX" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1184,1301,1339,1366]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="lapsus" status="sp. nov.">
<emphasis id="A0EAEAF29D9492D3FB33FA82CD432D34" bold="true" box="[1184,1301,1339,1366]" italics="true" pageId="136" pageNumber="137">M. lapsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9492D3FA8FFA82CD212D34" box="[1308,1399,1340,1366]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF52FAE1CD212DFF" blockId="136.[189,1399,1303,1437]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF52FAE1CD212DFF" pageId="136" pageNumber="137">
– Aedeagus as in 
<figureCitation id="0AA52A659D9492D3FE30FAE1CA1E2D1B" box="[419,584,1375,1401]" captionStart="Figs 712–725" captionStartId="179.[189,241,1892,1918]" captionTargetBox="[433,1144,277,1858]" captionTargetId="figure-72@179.[427,1160,265,1870]" captionTargetPageId="179" captionText="Figs 712–725. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 712– 714. M. skalei sp. nov., holotype, ♂ (NME). 715–717. M. sougb sp. nov., holotype, ♂ (NME). 718–720. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 721–723. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 724–725. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122252" httpUri="https://zenodo.org/record/4122252/files/figure.png" pageId="136" pageNumber="137">Figs 718–720</figureCitation>
; male and female pygidium as in 
<figureCitation id="0AA52A659D9492D3FC41FAE1CC382D18" box="[978,1134,1375,1402]" captionStart="Figs 504–521" captionStartId="166.[189,241,1681,1707]" captionTargetBox="[433,1155,265,1647]" captionTargetId="figure-177@166.[433,1155,265,1658]" captionTargetPageId="166" captionText="Figs 504–521. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 504. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 505. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 506. Same, ♀ (BMNH). 507. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 508. Same, syntype, ♂ (BMNH). 509. M. peregrinus Lansberge, 1886, ♂ (DTC). 510. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 511. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 512. Same, paratype, ♂ (BMNH). 513. M. riedeli sp. nov., holotype, ♂ (SMNS). 514. M. rugicollis Moser, 1920, ♂ (BMNH). 515. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 516. M. similis sp. nov., paratype, ♂ (BMNH). 517. M. skalei sp. nov., paratype, ♂ (NME). 518. M. sougb sp. nov., holotype, ♂ (NME). 519. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 520. Same, different ♂ (BMNH). 521. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122228" httpUri="https://zenodo.org/record/4122228/files/figure.png" pageId="136" pageNumber="137">Figs 519-521</figureCitation>
.................................. ................................................................................................ 
<taxonomicName id="559E4D639D9492D3FC28FA3DCCB82DFF" ID-CoL="7XWMQ" authority="(Frey, 1969)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" box="[955,1262,1410,1437]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="speciosus" status="comb. nov.">
<emphasis id="A0EAEAF29D9492D3FC28FA3DCC002DFE" bold="true" box="[955,1110,1410,1437]" italics="true" pageId="136" pageNumber="137">M. speciosus</emphasis>
(
<bibRefCitation id="F60F4B119D9492D3FBF6FA3DCCB32DFF" author="Frey G." box="[1125,1253,1410,1437]" pageId="136" pageNumber="137" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9492D3FB66FA3CCD212DFF" box="[1269,1399,1410,1437]" pageId="136" pageNumber="137" rank="species">comb. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9492D3FF2EFA7ACD212E44" pageId="136" pageNumber="137">
<paragraph id="922136E09D9492D3FF2EFA7ACD202DBC" blockId="136.[189,1399,1476,1574]" box="[189,1398,1476,1503]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EFA7ACD202DBC" box="[189,1398,1476,1503]" pageId="136" pageNumber="137">70. Anterolateral angles of labroclypeus acute in dorsal view .............................................................. 71</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF52FA56CD212E44" blockId="136.[189,1399,1476,1574]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF52FA56CD212E44" pageId="136" pageNumber="137">
– Anterolateral angles of labroclypeus obtuse in dorsal view (
<figureCitation id="0AA52A659D9492D3FC0FFA56CC152E61" box="[924,1091,1512,1539]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="136" pageNumber="137">Figs 169–170</figureCitation>
); aedeagus as in 
<figureCitation id="0AA52A659D9492D3FA90FA56C9402E47" captionStart="Figs 697–711" captionStartId="178.[189,241,1892,1918]" captionTargetBox="[435,1146,268,1853]" captionTargetId="figure-75@178.[425,1163,265,1860]" captionTargetPageId="178" captionText="Figs 697–711. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 697– 699. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH?). 700–702. M. riedeli sp. nov., holotype, ♂ (SMNS). 703–705. M. rugicollis Moser, 1920, ♂ (NHMB). 706–708. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 709–711. M. similis sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122250" httpUri="https://zenodo.org/record/4122250/files/figure.png" pageId="136" pageNumber="137">Figs 697– 699</figureCitation>
.................................................................................................. 
<taxonomicName id="559E4D639D9492D3FC7FF9B2CCB82E44" ID-CoL="7XWMP" authority="(Frey, 1969)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1969" box="[1004,1262,1547,1574]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="popei" status="comb. nov.">
<emphasis id="A0EAEAF29D9492D3FC7FF9B2CC002E47" bold="true" box="[1004,1110,1547,1574]" italics="true" pageId="136" pageNumber="137">M. popei</emphasis>
(
<bibRefCitation id="F60F4B119D9492D3FBF6F9B5CCB32E44" author="Frey G." box="[1125,1253,1547,1574]" pageId="136" pageNumber="137" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9492D3FB66F9B5CD212E44" box="[1269,1399,1547,1574]" pageId="136" pageNumber="137" rank="species">comb. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9492D3FF2EF9F3CD212ECD" pageId="136" pageNumber="137">
<paragraph id="922136E09D9492D3FF2EF9F3CD202E05" blockId="136.[189,1399,1613,1711]" box="[189,1398,1613,1640]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EF9F3CD202E05" box="[189,1398,1613,1640]" pageId="136" pageNumber="137">
71. Species from New 
<collectingCountry id="EA8976709D9492D3FE59F9F3CA492E05" box="[458,543,1613,1639]" name="Guinea-Bissau" pageId="136" pageNumber="137">Guinea</collectingCountry>
............................................................................................................... 72
</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF52F9CFCD212ECD" blockId="136.[189,1399,1613,1711]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF52F9CFCD212ECD" pageId="136" pageNumber="137">
– Species from Central 
<collectingCountry id="EA8976709D9492D3FE7AF9CECA0D2EE8" box="[489,603,1648,1674]" name="Indonesia" pageId="136" pageNumber="137">Moluccas</collectingCountry>
(Seram); aedeagus as in 
<bibRefCitation id="F60F4B119D9492D3FC12F9CECC152EE9" author="Prokofiev A. M." box="[897,1091,1648,1675]" pageId="136" pageNumber="137" pagination="159 - 163" refId="ref98551" refString="Prokofiev A. M. 2018. First finding of the genus Paramaechidius Frey, 1969 from the Indonesian province Maluku (Coleoptera: Scarabaeidae: Melolonthinae). Baltic Journal of Coleopterology 18 (2): 159 - 163." type="journal article" year="2018">Prokofiev (2018</bibRefCitation>
: fig. 5) ............................. .......................................................................................... 
<taxonomicName id="559E4D639D9492D3FC19F92ACCB82ECC" ID-CoL="7XWMK" authority="(Prokofiev, 2018)" authorityName="Telnov" authorityYear="2020" baseAuthorityName="Prokofiev" baseAuthorityYear="2018" box="[906,1262,1683,1710]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="agnellus" status="comb. nov.">
<emphasis id="A0EAEAF29D9492D3FC19F92ACC4C2ECC" bold="true" box="[906,1050,1683,1710]" italics="true" pageId="136" pageNumber="137">M. agnellus</emphasis>
(
<bibRefCitation id="F60F4B119D9492D3FBBBF92ACCB32ECC" author="Prokofiev A. M." box="[1064,1253,1684,1710]" pageId="136" pageNumber="137" pagination="159 - 163" refId="ref98551" refString="Prokofiev A. M. 2018. First finding of the genus Paramaechidius Frey, 1969 from the Indonesian province Maluku (Coleoptera: Scarabaeidae: Melolonthinae). Baltic Journal of Coleopterology 18 (2): 159 - 163." type="journal article" year="2018">Prokofiev, 2018</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9492D3FB66F92ACD212ECD" box="[1269,1399,1684,1711]" pageId="136" pageNumber="137" rank="species">comb. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296A2E459D9492D3FF2EF968CD212F8B" pageId="136" pageNumber="137">
<paragraph id="922136E09D9492D3FF2EF968CD212F76" blockId="136.[189,1399,1750,2025]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EF968CD212F76" pageId="136" pageNumber="137">
72. Aedeagus as in 
<figureCitation id="0AA52A659D9492D3FE30F968CA1E2E93" box="[419,584,1750,1777]" captionStart="Figs 586–600" captionStartId="171.[189,241,1892,1918]" captionTargetBox="[400,1175,268,1865]" captionTargetId="figure-72@171.[391,1196,265,1868]" captionTargetPageId="171" captionText="Figs 586–600. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 586– 588. M. bintang sp. nov., holotype, ♂ (NME). 589–591. M. boessnecki sp. nov., holotype, ♂ (SNSD). 592–594. M. caperatus sp. nov., holotype, ♂ (NME). 595–597. M. ciliatus sp. nov., holotype, ♂ (IECA). 598–600. M. crypticus sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122234" httpUri="https://zenodo.org/record/4122234/files/figure.png" pageId="136" pageNumber="137">Figs 598–600</figureCitation>
; male and female pygidium as in 
<figureCitation id="0AA52A659D9492D3FC41F968CC252E93" box="[978,1139,1750,1777]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="136" pageNumber="137">Figs 465–466</figureCitation>
.................................. ........................................................................................................................... 
<taxonomicName id="559E4D639D9492D3FBECF944CD432F71" ID-CoL="7X3WK" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1151,1301,1785,1812]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="crypticus" status="sp. nov.">
<emphasis id="A0EAEAF29D9492D3FBECF944CD432F71" bold="true" box="[1151,1301,1785,1812]" italics="true" pageId="136" pageNumber="137">M. crypticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9492D3FA8FF944CD212F76" box="[1308,1399,1786,1812]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF2EF8A3CD212F55" blockId="136.[189,1399,1750,2025]" box="[189,1399,1820,1847]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EF8A3CD212F55" box="[189,1399,1820,1847]" pageId="136" pageNumber="137">
– Aedeagus as in 
<figureCitation id="0AA52A659D9492D3FE30F8A3CA112F55" box="[419,583,1821,1847]" captionStart="Figs 652–666" captionStartId="175.[189,241,1892,1918]" captionTargetBox="[436,1156,267,1863]" captionTargetId="figure-70@175.[426,1161,265,1868]" captionTargetPageId="175" captionText="Figs 652–666. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 652– 654. M. longipes sp. nov., holotype, ♂ (BMNH). 655–657. M. luniceps Fairmaire, 1883, ♂ (BMNH). 658–660. M. mailu sp. nov., holotype, ♂ (NHMB). 661–663. M. maleo sp. nov., holotype, ♂ (IRSN). 664–666. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122244" httpUri="https://zenodo.org/record/4122244/files/figure.png" pageId="136" pageNumber="137">Figs 664–666</figureCitation>
; male pygidium as in 
<figureCitation id="0AA52A659D9492D3FCDAF8A3CBFD2F55" box="[841,939,1821,1847]" captionStart="Figs487–503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487–503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="136" pageNumber="137">Fig. 492</figureCitation>
............................ 
<taxonomicName id="559E4D639D9492D3FB17F8A3CD432F55" ID-CoL="7X3X6" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1156,1301,1820,1847]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="merdeka" status="sp. nov.">
<emphasis id="A0EAEAF29D9492D3FB17F8A3CD432F55" bold="true" box="[1156,1301,1820,1847]" italics="true" pageId="136" pageNumber="137">M. merdeka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9492D3FA8FF8A3CD212F55" box="[1308,1399,1821,1847]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF2EF8FFCD212F1C" blockId="136.[189,1399,1750,2025]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EF8FFCD212F1C" pageId="136" pageNumber="137">
– Aedeagus as in 
<figureCitation id="0AA52A659D9492D3FE30F8FECA112F38" box="[419,583,1856,1882]" captionStart="Figs 682–696" captionStartId="177.[189,241,1892,1918]" captionTargetBox="[436,1163,269,1860]" captionTargetId="figure-72@177.[423,1174,265,1866]" captionTargetPageId="177" captionText="Figs 682–696. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 682– 684. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 685–687. M. pauxillus Heller, 1910 comb. rest., ♂ (BMNH). 688–690. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 691–693. M. peregrinus Lansberge, 1886, ♂ (DTC). 694-696. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122248" httpUri="https://zenodo.org/record/4122248/files/figure.png" pageId="136" pageNumber="137">Figs 682–684</figureCitation>
; male and female pygidium as in 
<figureCitation id="0AA52A659D9492D3FC41F8FECC6E2F39" box="[978,1080,1856,1883]" captionStart="Figs487–503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487–503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="136" pageNumber="137">Figs 499</figureCitation>
–5000................................ .................................................................................................................... 
<taxonomicName id="559E4D639D9492D3FBC3F8DACD422F1C" ID-CoL="7X3XD" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1104,1300,1891,1918]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="owenstanleyi" status="sp. nov.">
<emphasis id="A0EAEAF29D9492D3FBC3F8DACD422F1C" bold="true" box="[1104,1300,1891,1918]" italics="true" pageId="136" pageNumber="137">M. owenstanleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9492D3FA8FF8DACD212F1C" box="[1308,1399,1892,1918]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF2EF836CD212FA7" blockId="136.[189,1399,1750,2025]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF2EF836CD212FA7" pageId="136" pageNumber="137">
– Aedeagus as in 
<figureCitation id="0AA52A659D9492D3FE30F839CA112FC0" box="[419,583,1927,1954]" captionStart="Figs 682–696" captionStartId="177.[189,241,1892,1918]" captionTargetBox="[436,1163,269,1860]" captionTargetId="figure-72@177.[423,1174,265,1866]" captionTargetPageId="177" captionText="Figs 682–696. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 682– 684. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 685–687. M. pauxillus Heller, 1910 comb. rest., ♂ (BMNH). 688–690. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 691–693. M. peregrinus Lansberge, 1886, ♂ (DTC). 694-696. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122248" httpUri="https://zenodo.org/record/4122248/files/figure.png" pageId="136" pageNumber="137">Figs 685–687</figureCitation>
; male and female pygidium as in 
<figureCitation id="0AA52A659D9492D3FC41F839CC252FC0" box="[978,1139,1927,1954]" captionStart="Figs 504–521" captionStartId="166.[189,241,1681,1707]" captionTargetBox="[433,1155,265,1647]" captionTargetId="figure-177@166.[433,1155,265,1658]" captionTargetPageId="166" captionText="Figs 504–521. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 504. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 505. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 506. Same, ♀ (BMNH). 507. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 508. Same, syntype, ♂ (BMNH). 509. M. peregrinus Lansberge, 1886, ♂ (DTC). 510. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 511. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 512. Same, paratype, ♂ (BMNH). 513. M. riedeli sp. nov., holotype, ♂ (SMNS). 514. M. rugicollis Moser, 1920, ♂ (BMNH). 515. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 516. M. similis sp. nov., paratype, ♂ (BMNH). 517. M. skalei sp. nov., paratype, ♂ (NME). 518. M. sougb sp. nov., holotype, ♂ (NME). 519. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 520. Same, different ♂ (BMNH). 521. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122228" httpUri="https://zenodo.org/record/4122228/files/figure.png" pageId="136" pageNumber="137">Figs 505–506</figureCitation>
.................................. ................................................................................................ 
<taxonomicName id="559E4D639D9492D3FC2FF815CCB82FA7" authority="Heller, 1910" authorityName="Heller" authorityYear="1910" box="[956,1262,1962,1989]" class="Insecta" family="Scarabaeidae" genus="Paramaechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="pauxillus">
<emphasis id="A0EAEAF29D9492D3FC2FF815CC022FA7" bold="true" box="[956,1108,1962,1989]" italics="true" pageId="136" pageNumber="137">M. pauxillus</emphasis>
<bibRefCitation id="F60F4B119D9492D3FBCFF815CCB82FA7" author="Heller K. M." box="[1116,1262,1963,1989]" pageId="136" pageNumber="137" pagination="1 - 42" refId="ref98170" refString="Heller K. M. 1910. Funfter Beitrag zur Papuanischen Kaferfauna hauptsachlich auf Grund der Ausbeute von Dr. Schlaginhaufen. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch- Ethnographischen Museums zu Dresden 13 (3): 1 - 42." type="journal article" year="1910">Heller, 1910</bibRefCitation>
</taxonomicName>
comb. rest.
</keyLead>
</paragraph>
<paragraph id="922136E09D9492D3FF52F871CD212F8B" blockId="136.[189,1399,1750,2025]" box="[193,1399,1998,2025]" pageId="136" pageNumber="137">
<keyLead id="296F95D59D9492D3FF52F871CD212F8B" box="[193,1399,1998,2025]" pageId="136" pageNumber="137">
– Aedeagus as in 
<figureCitation id="0AA52A659D9492D3FE30F870CA112F8A" box="[419,583,1998,2024]" captionStart="Figs 739–750" captionStartId="181.[189,241,1892,1918]" captionTargetBox="[360,1207,271,1867]" captionTargetId="figure-66@181.[347,1240,265,1871]" captionTargetPageId="181" captionText="Figs 739–750. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 739– 741. M. weigeli sp. nov., paratype, ♂ (NME). 742–744. M. woodlarkianus Heller, 1914, ♂ (SNSD). 745–747. M. yamdena sp. nov., paratype, ♂ (BMNH). 748–750. M. paupianus Heller, 1910 (paratype ♂ of M. arrowi Frey, 1969) (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122256" httpUri="https://zenodo.org/record/4122256/files/figure.png" pageId="136" pageNumber="137">Figs 739–741</figureCitation>
; male and female pygidium as in Figs 531–532.... 
<taxonomicName id="559E4D639D9492D3FB04F871CD432F8B" ID-CoL="7X3XY" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1175,1301,1998,2025]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="weigeli" status="sp. nov.">
<emphasis id="A0EAEAF29D9492D3FB04F871CD432F8B" bold="true" box="[1175,1301,1998,2025]" italics="true" pageId="136" pageNumber="137">M. weigeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9492D3FA8FF871CD212F8B" box="[1308,1399,1999,2025]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>