370 lines
54 KiB
XML
370 lines
54 KiB
XML
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<mods:title id="B24A09013EDB48D298B67D10F745D87E">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
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<mods:namePart id="C6A28FB3C9B7E392808E9320205CEA79">Telnov, Dmitry</mods:namePart>
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<mods:title id="E0AD5ABCEEC705A2A397660B92562F35">European Journal of Taxonomy</mods:title>
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<mods:part id="CF6A2427FAA21ADECD5B3351BBFB48A8">
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<mods:date id="F4156D46315E9B954603ED5D9E96DDF1">2020</mods:date>
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<mods:number id="F7E8833738E84062464E282E5D2943FB">2020-10-19</mods:number>
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<mods:number id="B696150DCC12959A60BC086A0E27D9BF">721</mods:number>
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<treatment id="1A3787F69D759230FDE4FC26C90E2F8A" ID-DOI="http://doi.org/10.5281/zenodo.4344331" ID-GBIF-Taxon="168648125" ID-Zenodo-Dep="4344331" LSID="urn:lsid:zoobank.org:act:3727E57E-1211-4A7A-BC4A-804DBF844DAF" httpUri="http://treatment.plazi.org/id/1A3787F69D759230FDE4FC26C90E2F8A" lastPageId="107" lastPageNumber="108" pageId="105" pageNumber="106">
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<subSubSection id="DA84656B9D759232FDE4FC26CBEB2BD1" box="[631,957,920,947]" pageId="105" pageNumber="106" type="nomenclature">
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<paragraph id="922136E09D759232FDE4FC26CBEB2BD1" blockId="105.[361,1225,920,1028]" box="[631,957,920,947]" pageId="105" pageNumber="106">
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<heading id="C969818C9D759232FDE4FC26CBEB2BD1" box="[631,957,920,947]" centered="true" fontSize="11" level="2" pageId="105" pageNumber="106" reason="2">
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<taxonomicName id="559E4D639D759232FDE4FC26CB0C2BD1" ID-CoL="7X3XP" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[631,858,920,947]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
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<emphasis id="A0EAEAF29D759232FDE4FC26CB0C2BD1" bold="true" box="[631,858,920,947]" italics="true" pageId="105" pageNumber="106">Maechidius similis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBD957899D759232FCF1FC27CBEB2BD1" box="[866,957,921,947]" pageId="105" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D759232FEFAFC7FCC792C61" pageId="105" pageNumber="106" type="description">
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<paragraph id="922136E09D759232FEFAFC7FCC9F2BBE" blockId="105.[361,1225,920,1028]" box="[361,1225,961,988]" pageId="105" pageNumber="106">
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<uri id="E60F3AE29D759232FEFAFC7FCC9F2BBE" box="[361,1225,961,988]" pageId="105" pageNumber="106">
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urn:lsid:zoobank.org:act:
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<uuid id="E6380C359D759232FD16FC7FCC9F2BBE" box="[645,1225,961,988]" pageId="105" pageNumber="106">3727E57E-1211-4A7A-BC4A-804DBF844DAF</uuid>
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</uri>
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</paragraph>
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<paragraph id="922136E09D759232FD97FC57CC792C61" blockId="105.[361,1225,920,1028]" box="[516,1071,1001,1028]" pageId="105" pageNumber="106">
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<figureCitation id="0AA52A659D759232FD97FC57CA082C61" box="[516,606,1001,1027]" captionStart="Figs 82–85" captionStartId="106.[189,241,1894,1920]" captionTargetBox="[330,1255,264,1870]" captionTargetId="figure-67@106.[330,1258,252,1883]" captionTargetPageId="106" captionText="Figs 82–85. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 82. M. similis sp. nov., paratype, ♂ (BMNH). 83. M. skalei sp. nov., paratype, ♂ (NME). 84. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). 85. M. sougb sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122165" httpUri="https://zenodo.org/record/4122165/files/figure.png" pageId="105" pageNumber="106">Figs 82</figureCitation>
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,
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<figureCitation id="0AA52A659D759232FDF9FC57CA852C66" box="[618,723,1001,1028]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="105" pageNumber="106">175–176</figureCitation>
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,
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<figureCitation id="0AA52A659D759232FD4CFC57CB582C66" box="[735,782,1001,1028]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="105" pageNumber="106">252</figureCitation>
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,
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<figureCitation id="0AA52A659D759232FC89FC57CB1F2C61" box="[794,841,1001,1027]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="105" pageNumber="106">343</figureCitation>
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, 415, 516, 708–711
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</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D759232FF2EFB91C91F2D7F" pageId="105" pageNumber="106" type="diagnosis">
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<paragraph id="922136E09D759232FF2EFB91C99F2C2B" blockId="105.[189,457,1071,1097]" box="[189,457,1071,1097]" pageId="105" pageNumber="106">
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||
<heading id="C969818C9D759232FF2EFB91C99F2C2B" bold="true" box="[189,457,1071,1097]" fontSize="11" level="3" pageId="105" pageNumber="106" reason="3">
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<emphasis id="A0EAEAF29D759232FF2EFB91C99F2C2B" bold="true" box="[189,457,1071,1097]" pageId="105" pageNumber="106">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="922136E09D759232FF2EFBDDC91F2D7F" blockId="105.[189,1399,1123,1309]" pageId="105" pageNumber="106">
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This species is generally close to
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<taxonomicName id="559E4D639D759232FDA8FBDDCBB02C1C" authority="Heller, 1895" authorityName=", Hell. Determined" authorityYear="1895" box="[571,998,1123,1150]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="subcostatus">
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<emphasis id="A0EAEAF29D759232FDA8FBDDCB182C1F" box="[571,846,1123,1149]" italics="true" pageId="105" pageNumber="106">Maechidius subcostatus</emphasis>
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<bibRefCitation id="F60F4B119D759232FCC6FBDDCBB02C1C" author="Heller K. M." box="[853,998,1123,1150]" pageId="105" pageNumber="106" pagination="1 - 17" refId="ref98131" refString="Heller K. M. 1895. Erster Beitrag zur Papuaniscen [sic!] Kaferfauna. Abhandlungen und Berichte des Koniglichen Zoologischen und Anthropologisch-Ethnographischen Museums zu Dresden 1894 / 1895 (16): 1 - 17." type="journal article" year="1895">Heller, 1895</bibRefCitation>
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</taxonomicName>
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, but readily differs primarily in the shape of the male aedeagus. The aedeagus of
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<taxonomicName id="559E4D639D759232FD44FB32CB1C2CC7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[727,842,1163,1189]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
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<emphasis id="A0EAEAF29D759232FD44FB32CB1C2CC7" box="[727,842,1163,1189]" italics="true" pageId="105" pageNumber="106">M. similis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBD957899D759232FCC0FB32CBFB2CC4" box="[851,941,1164,1190]" pageId="105" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
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is similar to that in
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<taxonomicName id="559E4D639D759232FB0EFB32C8A02CAF" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="hirtipes">
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<emphasis id="A0EAEAF29D759232FB0EFB32CD4A2CC7" box="[1181,1308,1163,1189]" italics="true" pageId="105" pageNumber="106">M. hirtipes</emphasis>
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<bibRefCitation id="F60F4B119D759232FAB5FB35C8A02CAF" author="Arrow G. E." pageId="105" pageNumber="106" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
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</taxonomicName>
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(cf.
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<figureCitation id="0AA52A659D759232FEBAFB0DC99E2CAF" box="[297,456,1203,1229]" captionStart="Figs 617–631" captionStartId="173.[189,241,1892,1918]" captionTargetBox="[439,1134,274,1859]" captionTargetId="figure-71@173.[434,1154,265,1868]" captionTargetPageId="173" captionText="Figs 617–631. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 617– 619. M. hamatus sp. nov., holotype, ♂ (IECA). 620–622. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 623–625. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 626–628. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 629–631. M. konjo sp. nov., holotype, ♂ (NMPC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122240" httpUri="https://zenodo.org/record/4122240/files/figure.png" pageId="105" pageNumber="106">Figs 620–622</figureCitation>
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,
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<figureCitation id="0AA52A659D759232FE41FB0DCA6F2CAF" box="[466,569,1203,1229]" captionStart="Figs 697–711" captionStartId="178.[189,241,1892,1918]" captionTargetBox="[435,1146,268,1853]" captionTargetId="figure-75@178.[425,1163,265,1860]" captionTargetPageId="178" captionText="Figs 697–711. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 697– 699. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH?). 700–702. M. riedeli sp. nov., holotype, ♂ (SMNS). 703–705. M. rugicollis Moser, 1920, ♂ (NHMB). 706–708. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 709–711. M. similis sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122250" httpUri="https://zenodo.org/record/4122250/files/figure.png" pageId="105" pageNumber="106">708–711</figureCitation>
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), but specifically different in the dorsal and lateral aspects, with a broader dorsal opening of the parameres with their lateral and posterior margins somewhat raised and visible in lateral view.
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</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D759232FF2EFAF4C98B2DA2" pageId="105" pageNumber="106" type="etymology">
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<paragraph id="922136E09D759232FF2EFAF4C9122D06" blockId="105.[189,324,1354,1380]" box="[189,324,1354,1380]" pageId="105" pageNumber="106">
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||
<heading id="C969818C9D759232FF2EFAF4C9122D06" bold="true" box="[189,324,1354,1380]" fontSize="11" level="3" pageId="105" pageNumber="106" reason="3">
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||
<emphasis id="A0EAEAF29D759232FF2EFAF4C9122D06" bold="true" box="[189,324,1354,1380]" pageId="105" pageNumber="106">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="922136E09D759232FF2EFAC3C98B2DA2" blockId="105.[189,1399,1405,1472]" pageId="105" pageNumber="106">
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Named from the Latin ‘
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<taxonomicName id="559E4D639D759232FE41FAC3CA492DF5" authorityName="Telnov" authorityYear="2020" box="[466,543,1405,1431]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="similis">
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||
<emphasis id="A0EAEAF29D759232FE41FAC3CA492DF5" box="[466,543,1405,1431]" italics="true" pageId="105" pageNumber="106">similis</emphasis>
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</taxonomicName>
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’ (meaning ‘similar’), because of the similarity in external morphology to
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<taxonomicName id="559E4D639D759232FF2EFA1BC9802DDD" authorityName=", Hell. Determined" authorityYear="1895" box="[189,470,1445,1471]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="subcostatus">
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<emphasis id="A0EAEAF29D759232FF2EFA1BC9802DDD" box="[189,470,1445,1471]" italics="true" pageId="105" pageNumber="106">Maechidius subcostatus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="DA84656B9D759232FF2EFA52CB412F8B" pageId="105" pageNumber="106" type="materials_examined">
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<paragraph id="922136E09D759232FF2EFA52C93A2E67" blockId="105.[189,364,1515,1542]" box="[189,364,1515,1542]" pageId="105" pageNumber="106">
|
||
<heading id="C969818C9D759232FF2EFA52C93A2E67" bold="true" box="[189,364,1515,1542]" fontSize="11" level="3" pageId="105" pageNumber="106" reason="3">
|
||
<emphasis id="A0EAEAF29D759232FF2EFA52C93A2E67" bold="true" box="[189,364,1515,1542]" pageId="105" pageNumber="106">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="22F63CBD9D759232FF79F9A1CC782EE8" ID-GBIF-Occurrence="2898878574" collectingDate="1964-10-15" collectingDateMax="1964-10-24" collectingDateMin="1964-10-15" collectionCode="NEW, BMNH" collectorName="M. E. Bacchus." country="Papua New Guinea" county="Mts. Budemu" elevation="1219" location="Madang Dist." municipality="Finisterre Mts." pageId="105" pageNumber="106" specimenCount="1" typeStatus="holotype">
|
||
<paragraph id="922136E09D759232FF79F9A1C90A2E5B" blockId="105.[189,1398,1567,1674]" box="[234,348,1567,1593]" pageId="105" pageNumber="106">
|
||
<emphasis id="A0EAEAF29D759232FF79F9A1C90A2E5B" bold="true" box="[234,348,1567,1593]" pageId="105" pageNumber="106">
|
||
<typeStatus id="4D2588429D759232FF79F9A1C90A2E5B" box="[234,348,1567,1593]" pageId="105" pageNumber="106" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="922136E09D759232FF2EF9F9CC782EE8" blockId="105.[189,1398,1567,1674]" pageId="105" pageNumber="106">
|
||
<collectingCountry id="EA8976709D759232FF2EF9F9C9812E03" box="[189,471,1607,1634]" name="Papua New Guinea" pageId="105" pageNumber="106">PAPUA NEW GUINEA</collectingCountry>
|
||
•
|
||
<specimenCount id="8498FD699D759232FE7DF9F6CA502E00" box="[494,518,1608,1634]" pageId="105" pageNumber="106">♂</specimenCount>
|
||
; “Stn. No. 51. [p] //
|
||
<collectionCode id="F48FAE259D759232FD7FF9F6CB782E00" box="[748,814,1608,1634]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="105" pageNumber="106">NEW</collectionCode>
|
||
<collectingCountry id="EA8976709D759232FCA6F9F9CBFC2E03" box="[821,938,1607,1633]" name="Guinea-Bissau" pageId="105" pageNumber="106">GUINEA</collectingCountry>
|
||
:
|
||
<location id="9741603B9D759232FC27F9F9CC002E03" LSID="urn:lsid:plazi:treatment:1A3787F69D759230FDE4FC26C90E2F8A:9741603B9D759232FC27F9F9CC002E03" box="[948,1110,1607,1633]" country="Papua New Guinea" county="Mts. Budemu" municipality="Finisterre Mts." name="Madang Dist." pageId="105" pageNumber="106">Madang Dist.</location>
|
||
,
|
||
<collectingMunicipality id="7245AC9A9D759232FBF2F9F9CD5B2E00" box="[1121,1293,1607,1634]" pageId="105" pageNumber="106">
|
||
Finisterre
|
||
<collectingCounty id="7B404E6C9D759232FB44F9F6CD5B2E00" box="[1239,1293,1608,1634]" pageId="105" pageNumber="106">Mts.</collectingCounty>
|
||
</collectingMunicipality>
|
||
Budemu c.
|
||
<quantity id="55669B059D759232FF4AF9D1C9622EEB" box="[217,308,1647,1673]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="105" pageNumber="106" unit="ft" value="4000.0">
|
||
<elevation id="19B3D1D39D759232FF4AF9D1C9622EEB" box="[217,308,1647,1673]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="105" pageNumber="106" unit="ft" value="4000.0">4000 ft.</elevation>
|
||
</quantity>
|
||
<date id="E62010209D759232FEA8F9D1C98E2EE8" box="[315,472,1647,1674]" pageId="105" pageNumber="106" value="1964-10-15" valueMax="1964-10-24" valueMin="1964-10-15">
|
||
<collectingDate id="F664E9C89D759232FEA8F9D1C98E2EE8" box="[315,472,1647,1674]" pageId="105" pageNumber="106" value="1964-10-15" valueMax="1964-10-24" valueMin="1964-10-15">15-24.x.1964</collectingDate>
|
||
</date>
|
||
[p] //
|
||
<collectorName id="3F6B53369D759232FD8AF9CECA902EEB" box="[537,710,1647,1674]" pageId="105" pageNumber="106">M.E. Bacchus.</collectorName>
|
||
B.M. 1965-120. [p]”;
|
||
<collectionCode id="F48FAE259D759232FC5DF9CECC7C2EE8" box="[974,1066,1648,1674]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="105" pageNumber="106">BMNH</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="922136E09D759232FF79F908CA742EB2" blockId="105.[189,1399,1717,2025]" box="[234,546,1717,1744]" pageId="105" pageNumber="106">
|
||
<emphasis id="A0EAEAF29D759232FF79F908C9312EB2" bold="true" box="[234,359,1718,1744]" pageId="105" pageNumber="106">
|
||
<typeStatus id="4D2588429D759232FF79F908C9312EB2" box="[234,359,1718,1744]" pageId="105" pageNumber="106" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="8498FD699D759232FEEBF908CA4C2EB2" box="[376,538,1717,1744]" pageId="105" pageNumber="106" type="generic">12 specimens</specimenCount>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="922136E09D759232FF2EF960CB412F8B" blockId="105.[189,1399,1717,2025]" pageId="105" pageNumber="106">
|
||
<materialsCitation id="22F63CBD9D759232FF2EF960CBBB2E9A" ID-GBIF-Occurrence="2898878572" box="[189,1005,1758,1784]" collectionCode="BMNH" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="105" pageNumber="106" specimenCount="6" specimenCount-female="6" typeStatus="holotype">
|
||
<collectingCountry id="EA8976709D759232FF2EF960C98F2E9A" box="[189,473,1758,1784]" name="Papua New Guinea" pageId="105" pageNumber="106">PAPUA NEW GUINEA</collectingCountry>
|
||
•
|
||
<specimenCount id="8498FD699D759232FE60F960CA602E9A" box="[499,566,1758,1784]" pageId="105" pageNumber="106" type="female">6 ♀♀</specimenCount>
|
||
; same labels as for
|
||
<typeStatus id="4D2588429D759232FC8DF960CBDE2E9A" box="[798,904,1758,1784]" pageId="105" pageNumber="106" type="holotype">holotype</typeStatus>
|
||
;
|
||
<collectionCode id="F48FAE259D759232FC06F960CBBB2E9A" box="[917,1005,1758,1784]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="105" pageNumber="106">BMNH</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="22F63CBD9D759232FB95F960C9832F2A" ID-GBIF-Occurrence="2898878550" collectingDate="1964-10-02" collectingDateMax="1964-10-11" collectingDateMin="1964-10-02" collectionCode="NEW, BMNH" country="Guinea-Bissau" elevation="1082" location="Madang Dist." municipality="Finisterre Mts." pageId="105" pageNumber="106" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
|
||
<specimenCount id="8498FD699D759232FB95F960CC1F2E9A" box="[1030,1097,1758,1784]" pageId="105" pageNumber="106" type="female">2 ♀♀</specimenCount>
|
||
; “Stn. No. 46. [p] //
|
||
<collectionCode id="F48FAE259D759232FAA7F960CD202E9A" box="[1332,1398,1758,1784]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="105" pageNumber="106">NEW</collectionCode>
|
||
<collectingCountry id="EA8976709D759232FF2EF8B8C9642F42" box="[189,306,1798,1824]" name="Guinea-Bissau" pageId="105" pageNumber="106">GUINEA</collectingCountry>
|
||
:
|
||
<location id="9741603B9D759232FED0F8B8C9BA2F42" LSID="urn:lsid:plazi:treatment:1A3787F69D759230FDE4FC26C90E2F8A:9741603B9D759232FED0F8B8C9BA2F42" box="[323,492,1798,1824]" country="Guinea-Bissau" municipality="Finisterre Mts." name="Madang Dist." pageId="105" pageNumber="106">Madang Dist.</location>
|
||
,
|
||
<collectingMunicipality id="7245AC9A9D759232FE6EF8B8CAE72F42" box="[509,689,1798,1824]" pageId="105" pageNumber="106">Finisterre Mts.</collectingMunicipality>
|
||
Damanti
|
||
<quantity id="55669B059D759232FCA3F8B8CBCF2F42" box="[816,921,1798,1825]" metricMagnitude="3" metricUnit="m" metricValue="1.08204" pageId="105" pageNumber="106" unit="ft" value="3550.0">
|
||
<elevation id="19B3D1D39D759232FCA3F8B8CBCF2F42" box="[816,921,1798,1825]" metricMagnitude="3" metricUnit="m" metricValue="1.08204" pageId="105" pageNumber="106" unit="ft" value="3550.0">3,550 ft.</elevation>
|
||
</quantity>
|
||
<date id="E62010209D759232FC35F8B8CC652F42" box="[934,1075,1798,1824]" pageId="105" pageNumber="106" value="1964-10-02" valueMax="1964-10-11" valueMin="1964-10-02">
|
||
<collectingDate id="F664E9C89D759232FC35F8B8CC652F42" box="[934,1075,1798,1824]" pageId="105" pageNumber="106" value="1964-10-02" valueMax="1964-10-11" valueMin="1964-10-02">2-11.x.1964</collectingDate>
|
||
</date>
|
||
[p] // M.E. Bacchus. B.M. 1965-120. [p]”;
|
||
<collectionCode id="F48FAE259D759232FEEEF890C9832F2A" box="[381,469,1838,1864]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="105" pageNumber="106">BMNH</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="22F63CBD9D759232FE62F890CA142FFB" ID-GBIF-Occurrence="2898878318" collectingDate="1964-10-02" collectingDateMax="1964-10-11" collectingDateMin="1964-10-02" collectionCode="NEW, BMNH" country="Guinea-Bissau" determinerName="G. Frey" elevation="1082" location="Madang Dist." municipality="Finisterre Mts." pageId="105" pageNumber="106" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<specimenCount id="8498FD699D759232FE62F890CA492F2A" box="[497,543,1838,1864]" pageId="105" pageNumber="106" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="8498FD699D759232FDBDF890CA0B2F2A" box="[558,605,1838,1864]" pageId="105" pageNumber="106" type="female">1 ♀</specimenCount>
|
||
; “Stn. No. 46. [p]//
|
||
<collectionCode id="F48FAE259D759232FCD8F890CBDB2F2A" box="[843,909,1838,1864]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="105" pageNumber="106">NEW</collectionCode>
|
||
<collectingCountry id="EA8976709D759232FC05F890CC5D2F2A" box="[918,1035,1838,1864]" name="Guinea-Bissau" pageId="105" pageNumber="106">GUINEA</collectingCountry>
|
||
:
|
||
<location id="9741603B9D759232FB84F890CCED2F2A" LSID="urn:lsid:plazi:treatment:1A3787F69D759230FDE4FC26C90E2F8A:9741603B9D759232FB84F890CCED2F2A" box="[1047,1211,1838,1864]" country="Guinea-Bissau" municipality="Finisterre Mts." name="Madang Dist." pageId="105" pageNumber="106">Madang Dist.</location>
|
||
,
|
||
<collectingMunicipality id="7245AC9A9D759232FB5BF890CD212F2A" box="[1224,1399,1838,1864]" pageId="105" pageNumber="106">Finisterre Mts.</collectingMunicipality>
|
||
Damanti
|
||
<quantity id="55669B059D759232FEBEF8E8C9C52F12" box="[301,403,1878,1905]" metricMagnitude="3" metricUnit="m" metricValue="1.08204" pageId="105" pageNumber="106" unit="ft" value="3550.0">
|
||
<elevation id="19B3D1D39D759232FEBEF8E8C9C52F12" box="[301,403,1878,1905]" metricMagnitude="3" metricUnit="m" metricValue="1.08204" pageId="105" pageNumber="106" unit="ft" value="3550.0">3,550 ft.</elevation>
|
||
</quantity>
|
||
<date id="E62010209D759232FE0DF8E8CA7C2F12" box="[414,554,1878,1904]" pageId="105" pageNumber="106" value="1964-10-02" valueMax="1964-10-11" valueMin="1964-10-02">
|
||
<collectingDate id="F664E9C89D759232FE0DF8E8CA7C2F12" box="[414,554,1878,1904]" pageId="105" pageNumber="106" value="1964-10-02" valueMax="1964-10-11" valueMin="1964-10-02">2-11.x.1964</collectingDate>
|
||
</date>
|
||
[p]// M.E. Bacchus. B.M. 1965-120. [p] //
|
||
<taxonomicName id="559E4D639D759232FBBDF8E8CD1C2F12" authority="Ar." authorityName="Arrow" authorityYear="1941" box="[1070,1354,1878,1904]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="hirtipes">
|
||
<emphasis id="A0EAEAF29D759232FBBDF8E8CD4A2F12" box="[1070,1308,1878,1904]" italics="true" pageId="105" pageNumber="106">Maechidius hirtipes</emphasis>
|
||
Ar.
|
||
</taxonomicName>
|
||
[h] det.
|
||
<determinerName id="67FDDA249D759232FF74F8C0C96C2FFA" box="[231,314,1918,1944]" pageId="105" pageNumber="106">G.Frey</determinerName>
|
||
,1967/68 [p]”;
|
||
<collectionCode id="F48FAE259D759232FE79F8C1CA142FFB" box="[490,578,1919,1945]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="105" pageNumber="106">BMNH</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="22F63CBD9D759232FDCCF8C0CB452F8B" ID-GBIF-Occurrence="2898878371" collectingDate="1965-02-05" collectionCode="NEW, BMNH" collectorName="M. E. Bacchus." country="Guinea-Bissau" county="Nr. Okapa" determinerName="G. Frey" elevation="1524" location="Highlands Dist." municipality="Wanatabe valley" pageId="105" pageNumber="106" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
|
||
<specimenCount id="8498FD699D759232FDCCF8C0CAD82FFB" box="[607,654,1918,1945]" pageId="105" pageNumber="106" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="8498FD699D759232FD0DF8C0CA9B2FFB" box="[670,717,1918,1945]" pageId="105" pageNumber="106" type="female">1 ♀</specimenCount>
|
||
; “Stn. No. 174. [p] //
|
||
<collectionCode id="F48FAE259D759232FC5DF8C1CC462FFB" box="[974,1040,1919,1945]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="105" pageNumber="106">NEW</collectionCode>
|
||
<collectingCountry id="EA8976709D759232FB89F8C0CCD92FFA" box="[1050,1167,1918,1944]" name="Guinea-Bissau" pageId="105" pageNumber="106">GUINEA</collectingCountry>
|
||
: E.
|
||
<location id="9741603B9D759232FB22F8C0CD242FFA" LSID="urn:lsid:plazi:treatment:1A3787F69D759230FDE4FC26C90E2F8A:9741603B9D759232FB22F8C0CD242FFA" box="[1201,1394,1918,1944]" country="Guinea-Bissau" county="Nr. Okapa" municipality="Wanatabe valley" name="Highlands Dist." pageId="105" pageNumber="106">Highlands Dist.</location>
|
||
,
|
||
<collectingMunicipality id="7245AC9A9D759232FF2EF818C9D72FA2" box="[189,385,1958,1984]" pageId="105" pageNumber="106">Wanatabe valley</collectingMunicipality>
|
||
.
|
||
<collectingCounty id="7B404E6C9D759232FE18F819CA502FA2" box="[395,518,1958,1985]" pageId="105" pageNumber="106">Nr. Okapa</collectingCounty>
|
||
, c.
|
||
<quantity id="55669B059D759232FDBFF818CAD92FA2" box="[556,655,1958,1985]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="105" pageNumber="106" unit="ft" value="5000.0">
|
||
<elevation id="19B3D1D39D759232FDBFF818CAD92FA2" box="[556,655,1958,1985]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="105" pageNumber="106" unit="ft" value="5000.0">5,000 ft.</elevation>
|
||
</quantity>
|
||
<date id="E62010209D759232FD05F818CAA82FA3" box="[662,766,1958,1985]" pageId="105" pageNumber="106" value="1965-02-05">
|
||
<collectingDate id="F664E9C89D759232FD05F818CAA82FA3" box="[662,766,1958,1985]" pageId="105" pageNumber="106" value="1965-02-05">5.ii.1965</collectingDate>
|
||
</date>
|
||
[p] //
|
||
<collectorName id="3F6B53369D759232FCACF819CBBD2FA2" box="[831,1003,1958,1985]" pageId="105" pageNumber="106">M.E. Bacchus.</collectorName>
|
||
B.M. 1965-120. [p] //
|
||
<taxonomicName id="559E4D639D759232FB7DF818C90B2F8B" authority="♀ Ar." authorityName="Ar." class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="tarsalis">
|
||
<emphasis id="A0EAEAF29D759232FB7DF818C9432F8A" italics="true" pageId="105" pageNumber="106">Maechidius tarsalis</emphasis>
|
||
<specimenCount id="8498FD699D759232FE8FF871C9642F8B" box="[284,306,1999,2025]" pageId="105" pageNumber="106" type="female">♀</specimenCount>
|
||
Ar.
|
||
</taxonomicName>
|
||
[h] det.
|
||
<determinerName id="67FDDA249D759232FE2BF870CA5D2F8A" box="[440,523,1998,2024]" pageId="105" pageNumber="106">G.Frey</determinerName>
|
||
,1967/68 [p]”;
|
||
<collectionCode id="F48FAE259D759232FD24F871CB452F8B" box="[695,787,1999,2025]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="105" pageNumber="106">BMNH</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="C6E166689D769231FF2EF8D8C8A92F88" ID-DOI="http://doi.org/10.5281/zenodo.4122165" ID-Zenodo-Dep="4122165" httpUri="https://zenodo.org/record/4122165/files/figure.png" pageId="106" pageNumber="107" startId="106.[189,241,1894,1920]" targetBox="[330,1255,264,1870]" targetPageId="106">
|
||
<paragraph id="922136E09D769231FF2EF8D8C8A92F88" blockId="106.[189,1399,1894,2026]" pageId="106" pageNumber="107">
|
||
<emphasis id="A0EAEAF29D769231FF2EF8D8C91B2FE2" bold="true" box="[189,333,1894,1920]" pageId="106" pageNumber="107">Figs 82–85.</emphasis>
|
||
Papuan and Wallacean
|
||
<taxonomicName id="559E4D639D769231FDE7F8D8CBED2FE3" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[628,955,1894,1921]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0EAEAF29D769231FDE7F8D8CAAB2FE2" box="[628,765,1894,1920]" italics="true" pageId="106" pageNumber="107">Maechidius</emphasis>
|
||
<bibRefCitation id="F60F4B119D769231FC9BF8D8CBED2FE3" author="Macleay W. S." box="[776,955,1894,1921]" pageId="106" pageNumber="107" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
|
||
</taxonomicName>
|
||
, habitus, dorsal view.
|
||
<emphasis id="A0EAEAF29D769231FB5CF8D8CCBA2FE2" bold="true" box="[1231,1260,1894,1920]" pageId="106" pageNumber="107">82</emphasis>
|
||
.
|
||
<taxonomicName id="559E4D639D769231FB6CF8D9CD212FE2" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1279,1399,1894,1920]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
|
||
<emphasis id="A0EAEAF29D769231FB6CF8D9CD212FE2" box="[1279,1399,1894,1920]" italics="true" pageId="106" pageNumber="107">M. similis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BBD957899D769231FF2EF834C94D2FC6" box="[189,283,1930,1956]" pageId="106" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂ (BMNH).
|
||
<emphasis id="A0EAEAF29D769231FDA5F837CA052FC1" bold="true" box="[566,595,1929,1955]" pageId="106" pageNumber="107">83</emphasis>
|
||
.
|
||
<taxonomicName id="559E4D639D769231FDF0F834CA992FC6" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[611,719,1930,1956]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="skalei" status="sp. nov.">
|
||
<emphasis id="A0EAEAF29D769231FDF0F834CA992FC6" box="[611,719,1930,1956]" italics="true" pageId="106" pageNumber="107">M. skalei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BBD957899D769231FD4BF834CB602FC6" box="[728,822,1930,1956]" pageId="106" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype, ♂ (NME).
|
||
<emphasis id="A0EAEAF29D769231FBA9F837CC012FC1" bold="true" box="[1082,1111,1929,1955]" pageId="106" pageNumber="107">84</emphasis>
|
||
.
|
||
<taxonomicName id="559E4D639D769231FBF4F834CD272FC6" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[1127,1393,1930,1956]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="simplex">
|
||
<emphasis id="A0EAEAF29D769231FBF4F834CCBF2FC6" box="[1127,1257,1930,1956]" italics="true" pageId="106" pageNumber="107">M. simplex</emphasis>
|
||
<bibRefCitation id="F60F4B119D769231FB62F834CD272FC6" author="Frey G." box="[1265,1393,1930,1956]" pageId="106" pageNumber="107" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype, ♂ (photograph courtesy BPBM) (BPBM).
|
||
<emphasis id="A0EAEAF29D769231FCB9F813CB112FA5" bold="true" box="[810,839,1965,1991]" pageId="106" pageNumber="107">85</emphasis>
|
||
.
|
||
<taxonomicName id="559E4D639D769231FCC5F810CB952FA5" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[854,963,1965,1991]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="sougb" status="sp. nov.">
|
||
<emphasis id="A0EAEAF29D769231FCC5F810CB952FA5" box="[854,963,1965,1991]" italics="true" pageId="106" pageNumber="107">M. sougb</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BBD957899D769231FC58F813CC7F2FA5" box="[971,1065,1965,1991]" pageId="106" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (NME). Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="DA84656B9D779230FF2EFEB6CBE82E86" pageId="107" pageNumber="108" type="description">
|
||
<paragraph id="922136E09D779230FF2EFEB6C9182940" blockId="107.[189,334,264,290]" box="[189,334,264,290]" pageId="107" pageNumber="108">
|
||
<heading id="C969818C9D779230FF2EFEB6C9182940" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="107" pageNumber="108" reason="3">
|
||
<emphasis id="A0EAEAF29D779230FF2EFEB6C9182940" bold="true" box="[189,334,264,290]" pageId="107" pageNumber="108">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D779230FF2EFE86CB4529FB" blockId="107.[189,1399,311,409]" pageId="107" pageNumber="108">
|
||
MEASUREMENTS.
|
||
<typeStatus id="4D2588429D779230FEEFFE89C9BA2933" box="[380,492,311,337]" pageId="107" pageNumber="108" type="holotype">Holotype</typeStatus>
|
||
, total body length
|
||
<quantity id="55669B059D779230FD54FE89CB652930" box="[711,819,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="9.45" pageId="107" pageNumber="108" unit="mm" value="9.45">9.45 mm</quantity>
|
||
. Head
|
||
<quantity id="55669B059D779230FC16FE89CBB82930" box="[901,1006,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="107" pageNumber="108" unit="mm" value="1.65">1.65 mm</quantity>
|
||
long, across eyes
|
||
<quantity id="55669B059D779230FB55FE89CD792930" box="[1222,1327,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="107" pageNumber="108" unit="mm" value="2.15">2.15 mm</quantity>
|
||
wide. Pronotum
|
||
<quantity id="55669B059D779230FEA4FEE5C9C92914" box="[311,415,347,374]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="107" pageNumber="108" unit="mm" value="1.9">1.90 mm</quantity>
|
||
long, maximum width 3.00 mm. Elytral length
|
||
<quantity id="55669B059D779230FC54FEE5CC642914" box="[967,1074,347,374]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="107" pageNumber="108" unit="mm" value="5.9">5.90 mm</quantity>
|
||
, maximum combined width
|
||
<quantity id="55669B059D779230FF2EFEC1C97F29FB" box="[189,297,383,409]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="107" pageNumber="108" unit="mm" value="4.8">4.80 mm</quantity>
|
||
. Selected
|
||
<typeStatus id="4D2588429D779230FE0CFEC1CA5929FB" box="[415,527,383,409]" pageId="107" pageNumber="108" type="paratype">paratypes</typeStatus>
|
||
<quantity id="55669B059D779230FD85FEC1CA8629FB" box="[534,720,383,409]" metricMagnitude="-3" metricUnit="m" metricValue="9.55" metricValueMax="10.3" metricValueMin="8.8" pageId="107" pageNumber="108" unit="mm" value="9.55" valueMax="10.3" valueMin="8.8">8.80–10.30 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="922136E09D779230FF2EFE7FC9D42E2D" blockId="107.[189,1399,448,1615]" pageId="107" pageNumber="108">
|
||
Dorsum uniformly black-brown, venter and appendages brown. Head flattened dorsally between compound eyes, glossy dorsally and ventrally. Male labroclypeus (
|
||
<figureCitation id="0AA52A659D779230FB96FE5ACC26299D" box="[1029,1136,484,511]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="107" pageNumber="108">Fig. 175</figureCitation>
|
||
) deeply and broadly V-shaped emarginate on anterior margin, its lateral margins strongly sinuous in both dorsal and lateral views. Anterolateral angles acute, apically rounded, strongly protruding, raised up at angle of nearly 90° to frons in lateral view. Female labroclypeus (
|
||
<figureCitation id="0AA52A659D779230FD4AFDF1CB682A08" box="[729,830,591,618]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="107" pageNumber="108">Fig. 176</figureCitation>
|
||
) with much less protruding anterolateral angles. Anterior and lateral margins of labroclypeus smooth. Upper- and underside of labroclypeus with sparse setae along anterior and lateral margins. Canthus broadly rounded in dorsal view. Punctures of frons irregularly hexagonal, variably large, deep and dense. Intervening spaces glossy, much smaller than punctures, in part wrinkled. Head setae inconspicuous, appressed to suberect, not or hardly surpassing length of corresponding punctures. Antenna 9-segmented, club 3-lamellate. Pronotum transverse, flattened dorsally, glossy to subopaque dorsally and laterally. Anterior margin broadly emarginate with slightly protruding anterolateral angles, basal margin broadly rounded. Lateral margin in dorsal view slightly gradually widened towards middle, deeply emarginate in basal third, crenulate in anterior half (crenulae acute angulate at area of emargination), irregularly rugulose in emargination area. Lateral margin of pronotum arched in lateral view. Inconspicuous erect seta present between every two crenulae (
|
||
<figureCitation id="0AA52A659D779230FEF8FC69C9812B90" box="[363,471,983,1010]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="107" pageNumber="108">Fig. 252</figureCitation>
|
||
). Hypomeron separated from prosternum by moderately high nearly straight carina which is obtusely angulate medially, with long setae on its anterolateral margin opposite to compound eye. Antennal pocket moderately deep. Punctures of pronotal disc irregularly hexagonal, large and coarse, deep and dense. Intervening spaces glossy, distinctly smaller than punctures, in part wrinkled. Pronotal setae inconspicuous; seta rises from anterior margin of each puncture, not surpassing length of corresponding puncture. Scutellar shield narrowly rounded apically. Elytron opaque, with tracks of 3–4 variously broadly interrupted glabrous longitudinal carinae, including sutural one. Elytral punctures linear (elongate and narrow), incision-shaped, moderately deep. Intervening spaces densely microreticulate. Appressed to suberect seta rises from anterior margin of each incision, not surpassing length of corresponding incision (
|
||
<figureCitation id="0AA52A659D779230FD5CFAA9CB602D50" box="[719,822,1303,1330]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="107" pageNumber="108">Fig. 343</figureCitation>
|
||
). Slightly longer setae present along remnants of longitudinal carinae. Male and female pygidium flattened dorsally, with large shallow dense irregularly circular punctures. Intervening spaces microreticulate, much smaller than punctures. Male and female pygidium flattened dorsally, opaque, with dense large and shallow annular punctures (
|
||
<figureCitation id="0AA52A659D779230FB2AFA3CCD742DFF" box="[1209,1314,1410,1437]" captionStart="Figs 504–521" captionStartId="166.[189,241,1681,1707]" captionTargetBox="[433,1155,265,1647]" captionTargetId="figure-177@166.[433,1155,265,1658]" captionTargetPageId="166" captionText="Figs 504–521. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 504. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 505. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 506. Same, ♀ (BMNH). 507. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 508. Same, syntype, ♂ (BMNH). 509. M. peregrinus Lansberge, 1886, ♂ (DTC). 510. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 511. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 512. Same, paratype, ♂ (BMNH). 513. M. riedeli sp. nov., holotype, ♂ (SMNS). 514. M. rugicollis Moser, 1920, ♂ (BMNH). 515. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 516. M. similis sp. nov., paratype, ♂ (BMNH). 517. M. skalei sp. nov., paratype, ♂ (NME). 518. M. sougb sp. nov., holotype, ♂ (NME). 519. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 520. Same, different ♂ (BMNH). 521. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122228" httpUri="https://zenodo.org/record/4122228/files/figure.png" pageId="107" pageNumber="108">Fig. 516</figureCitation>
|
||
). Setae of pygidium inconspicuous, short and suberect, in anterior slightly, in distal part distinctly surpassing length of corresponding punctures. Protibia slender, with three external teeth: two distal acute and rather large, basal one situated close to midlength of protibia very inconspicuous (Fig. 415). Male protibial terminal spur long and straight, female one longer and stouter. Tarsal claws with pulvilli. Aedeagus as in
|
||
<figureCitation id="0AA52A659D779230FF48F98AC9292E2D" box="[219,383,1588,1615]" captionStart="Figs 697–711" captionStartId="178.[189,241,1892,1918]" captionTargetBox="[435,1146,268,1853]" captionTargetId="figure-75@178.[425,1163,265,1860]" captionTargetPageId="178" captionText="Figs 697–711. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 697– 699. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH?). 700–702. M. riedeli sp. nov., holotype, ♂ (SMNS). 703–705. M. rugicollis Moser, 1920, ♂ (NHMB). 706–708. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 709–711. M. similis sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122250" httpUri="https://zenodo.org/record/4122250/files/figure.png" pageId="107" pageNumber="108">Figs 709–711</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="922136E09D779230FF2EF9C8C9E62EF2" blockId="107.[189,432,1654,1680]" box="[189,432,1654,1680]" pageId="107" pageNumber="108">
|
||
<heading id="C969818C9D779230FF2EF9C8C9E62EF2" bold="true" box="[189,432,1654,1680]" fontSize="11" level="3" pageId="107" pageNumber="108" reason="3">
|
||
<emphasis id="A0EAEAF29D779230FF2EF9C8C9E62EF2" bold="true" box="[189,432,1654,1680]" pageId="107" pageNumber="108">Sexual dimorphism</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D779230FF2EF91BCBE82E86" blockId="107.[189,1399,1701,1764]" pageId="107" pageNumber="108">Female generally more robust, with comparatively shorter antennal lamellae, longer protibial terminal spur and apically rounded (not pointed) metatibial terminal spurs.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA84656B9D779230FF2EF8B5CA3E2F36" box="[189,616,1803,1877]" pageId="107" pageNumber="108" type="biology_ecology">
|
||
<paragraph id="922136E09D779230FF2EF8B5C9762F47" blockId="107.[189,288,1803,1829]" box="[189,288,1803,1829]" pageId="107" pageNumber="108">
|
||
<heading id="C969818C9D779230FF2EF8B5C9762F47" bold="true" box="[189,288,1803,1829]" fontSize="11" level="3" pageId="107" pageNumber="108" reason="3">
|
||
<emphasis id="A0EAEAF29D779230FF2EF8B5C9762F47" bold="true" box="[189,288,1803,1829]" pageId="107" pageNumber="108">Ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D779230FF2EF884CA3E2F36" blockId="107.[189,616,1850,1877]" box="[189,616,1850,1877]" pageId="107" pageNumber="108">
|
||
Occurs at altitudes of
|
||
<quantity id="55669B059D779230FE2DF884CA322F36" box="[446,612,1850,1877]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.52" metricValueMin="1.08" pageId="107" pageNumber="108" unit="m" value="1300.0" valueMax="1520.0" valueMin="1080.0">1080–1520 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA84656B9D779230FF2EF8C5C90E2F8A" pageId="107" pageNumber="108" type="distribution">
|
||
<paragraph id="922136E09D779230FF2EF8C5C9002FF7" blockId="107.[189,342,1915,1941]" box="[189,342,1915,1941]" pageId="107" pageNumber="108">
|
||
<heading id="C969818C9D779230FF2EF8C5C9002FF7" bold="true" box="[189,342,1915,1941]" fontSize="11" level="3" pageId="107" pageNumber="108" reason="3">
|
||
<emphasis id="A0EAEAF29D779230FF2EF8C5C9002FF7" bold="true" box="[189,342,1915,1941]" pageId="107" pageNumber="108">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="922136E09D779230FF2EF815C90E2F8A" blockId="107.[189,1399,1963,2025]" pageId="107" pageNumber="108">
|
||
Hitherto known from Finisterre Mountains and Central Cordillera (Eastern Highlands Province), East New
|
||
<collectingCountry id="EA8976709D779230FF6FF870C9032F8A" box="[252,341,1998,2024]" name="Guinea-Bissau" pageId="107" pageNumber="108">Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |