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<document id="00033F721692787FF6649C0BD7D1B7D9" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D709235FDE8F993CB5C2A49" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius skalei Telnov 2020, sp. nov." docType="treatment" docUuid="C9FAA274-0AB0-4E8A-B2E3-2CF7C564CCC1" docUuidSource="ZooBank" docVersion="10" lastPageNumber="111" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="109" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="770BA7EF101E1A0BE280CF74B325EC4E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="627F901CE88380BA316ECBD193E579EE">
<mods:title id="1A51AB54B6E2264C15F1AD9DDADD8B28">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="882F1D2BB844DB5D8C898D163E40EFFD" type="personal">
<mods:role id="A1C8ECA00BFB35A8B6070ABEC7AC61A9">
<mods:roleTerm id="64C24A7C899F2F8A2C1AAF149EDFAEB8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6BA611542216182351A329F98E1151F2">Telnov, Dmitry</mods:namePart>
</mods:name>
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<mods:title id="622172D0C40FD0FD9B61EBE349F243A6">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="6E2BCD3925F548EE93CA0F4B411B002B">
<mods:date id="7E57F0C2C37995B5C6AFD5CBC303E034">2020</mods:date>
<mods:detail id="2425876943E9F06503CFDBEB3D3C90D2" type="pubDate">
<mods:number id="9BEBDC970A871D95B78C7B0B4758A89F">2020-10-19</mods:number>
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<mods:detail id="4EDA683E9597E562E443D271283AF37A" type="volume">
<mods:number id="159BCC7B12B272FE955A22259F8FBB49">721</mods:number>
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<mods:classification id="DC485D2CF454B221F4ED2EE42586BDD6">journal article</mods:classification>
<mods:identifier id="B06E650986D07EBBC7A0A5C3213B775D" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="F34F51B67BD66F2AA4D343BF35D33F4B" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="00ED0E56BD9DD23737165B152682A870" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="8F553E5F5060D2566B26469456666B18" type="Zenodo-Dep">4122118</mods:identifier>
<mods:identifier id="699DAA43ABA3E42B0775BD4F641FC6F5" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
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<treatment id="1A3787F69D709235FDE8F993CB5C2A49" ID-DOI="http://doi.org/10.5281/zenodo.4344281" ID-GBIF-Taxon="168648122" ID-Zenodo-Dep="4344281" LSID="urn:lsid:zoobank.org:act:C9FAA274-0AB0-4E8A-B2E3-2CF7C564CCC1" httpUri="http://treatment.plazi.org/id/1A3787F69D709235FDE8F993CB5C2A49" lastPageId="110" lastPageNumber="111" pageId="108" pageNumber="109">
<subSubSection id="DA84656B9D709237FDE8F993CBEF2E2A" box="[635,953,1581,1608]" pageId="108" pageNumber="109" type="nomenclature">
<paragraph id="922136E09D709237FDE8F993CBEF2E2A" blockId="108.[356,1231,1581,1682]" box="[635,953,1581,1608]" pageId="108" pageNumber="109">
<heading id="C969818C9D709237FDE8F993CBEF2E2A" box="[635,953,1581,1608]" centered="true" fontSize="11" level="2" pageId="108" pageNumber="109" reason="2">
<taxonomicName id="559E4D639D709237FDE8F993CB002E2A" ID-CoL="7X3XQ" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[635,854,1581,1608]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="skalei" status="sp. nov.">
<emphasis id="A0EAEAF29D709237FDE8F993CB002E2A" bold="true" box="[635,854,1581,1608]" italics="true" pageId="108" pageNumber="109">Maechidius skalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D709237FCCDF990CBEF2E2A" box="[862,953,1582,1608]" pageId="108" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D709237FEF7F9ECCC662EF0" pageId="108" pageNumber="109" type="description">
<paragraph id="922136E09D709237FEF7F9ECCC992E0F" blockId="108.[356,1231,1581,1682]" box="[356,1231,1618,1645]" pageId="108" pageNumber="109">
<uri id="E60F3AE29D709237FEF7F9ECCC992E0F" box="[356,1231,1618,1645]" pageId="108" pageNumber="109">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D709237FD16F9ECCC992E0F" box="[645,1231,1618,1645]" pageId="108" pageNumber="109">C9FAA274-0AB0-4E8A-B2E3-2CF7C564CCC1</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D709237FD90F9C6CC662EF0" blockId="108.[356,1231,1581,1682]" box="[515,1072,1655,1682]" pageId="108" pageNumber="109">
<figureCitation id="0AA52A659D709237FD90F9C6CA0B2EF0" box="[515,605,1656,1682]" captionStart="Figs 82–85" captionStartId="106.[189,241,1894,1920]" captionTargetBox="[330,1255,264,1870]" captionTargetId="figure-67@106.[330,1258,252,1883]" captionTargetPageId="106" captionText="Figs 82–85. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 82. M. similis sp. nov., paratype, ♂ (BMNH). 83. M. skalei sp. nov., paratype, ♂ (NME). 84. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). 85. M. sougb sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122165" httpUri="https://zenodo.org/record/4122165/files/figure.png" pageId="108" pageNumber="109">Figs 83</figureCitation>
,
<figureCitation id="0AA52A659D709237FDFAF9C6CAC12EF0" box="[617,663,1656,1682]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="108" pageNumber="109">177</figureCitation>
,
<figureCitation id="0AA52A659D709237FD37F9C9CA852EF0" box="[676,723,1655,1682]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="108" pageNumber="109">253</figureCitation>
,
<figureCitation id="0AA52A659D709237FD4CF9C6CB582EF0" box="[735,782,1656,1682]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="108" pageNumber="109">344</figureCitation>
, 416, 446, 517, 712–714
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D709237FF2EF905CA8B2F32" pageId="108" pageNumber="109" type="diagnosis">
<paragraph id="922136E09D709237FF2EF905C99F2EB7" blockId="108.[189,457,1723,1749]" box="[189,457,1723,1749]" pageId="108" pageNumber="109">
<heading id="C969818C9D709237FF2EF905C99F2EB7" bold="true" box="[189,457,1723,1749]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="A0EAEAF29D709237FF2EF905C99F2EB7" bold="true" box="[189,457,1723,1749]" pageId="108" pageNumber="109">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D709237FF2EF955CA8B2F32" blockId="108.[189,1399,1771,1872]" pageId="108" pageNumber="109">
<taxonomicName id="559E4D639D709237FF2EF955C9C72F67" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[189,401,1771,1797]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="skalei" status="sp. nov.">
<emphasis id="A0EAEAF29D709237FF2EF955C9C72F67" box="[189,401,1771,1797]" italics="true" pageId="108" pageNumber="109">Maechidius skalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D709237FE0AF952C9A52F64" box="[409,499,1772,1798]" pageId="108" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
is peculiar among its congeners from
<collectingRegion id="505AF8029D709237FC20F955CC4B2F67" box="[947,1053,1771,1797]" country="Indonesia" name="Sulawesi Utara" pageId="108" pageNumber="109">Sulawesi</collectingRegion>
primarily due to the shape of the male aedeagus in combination with a widened and flattened male basal metatarsomere (also occurs in
<taxonomicName id="559E4D639D709237FF48F888C9372F2D" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[219,353,1845,1871]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="legalovi" status="sp. nov.">
<emphasis id="A0EAEAF29D709237FF48F888C9372F2D" box="[219,353,1845,1871]" italics="true" pageId="108" pageNumber="109">M. legalovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D709237FEFBF888C9952F32" box="[360,451,1846,1872]" pageId="108" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="559E4D639D709237FE68F888CA3C2F2D" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[507,618,1845,1871]" class="Aves" family="Megapodiidae" genus="Macrocephalon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galliformes" pageId="108" pageNumber="109" phylum="Chordata" rank="species" species="maleo" status="sp. nov.">
<emphasis id="A0EAEAF29D709237FE68F888CA3C2F2D" box="[507,618,1845,1871]" italics="true" pageId="108" pageNumber="109">M. maleo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D709237FDE2F888CA862F32" box="[625,720,1846,1872]" pageId="108" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D709237FF2EF8C7CB102F8A" pageId="108" pageNumber="109" type="etymology">
<paragraph id="922136E09D709237FF2EF8C7C9122FF1" blockId="108.[189,324,1913,1939]" box="[189,324,1913,1939]" pageId="108" pageNumber="109">
<heading id="C969818C9D709237FF2EF8C7C9122FF1" bold="true" box="[189,324,1913,1939]" fontSize="11" level="3" pageId="108" pageNumber="109" reason="3">
<emphasis id="A0EAEAF29D709237FF2EF8C7C9122FF1" bold="true" box="[189,324,1913,1939]" pageId="108" pageNumber="109">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D709237FF2EF817CB102F8A" blockId="108.[189,1398,1961,2025]" pageId="108" pageNumber="109">
Patronymic. The new species is named after its first collector, André Skale (Gera,
<collectingCountry id="EA8976709D709237FB0FF817CD582FA1" box="[1180,1294,1961,1987]" name="Germany" pageId="108" pageNumber="109">Germany</collectingCountry>
), a wellknown coleopterist, respected colleague and my friend.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D719236FF2EFEB7C9942A21" pageId="109" pageNumber="110" type="materials_examined">
<paragraph id="922136E09D719236FF2EFEB7C93A2940" blockId="109.[189,364,264,291]" box="[189,364,264,291]" pageId="109" pageNumber="110">
<heading id="C969818C9D719236FF2EFEB7C93A2940" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="109" pageNumber="110" reason="3">
<emphasis id="A0EAEAF29D719236FF2EFEB7C93A2940" bold="true" box="[189,364,264,291]" pageId="109" pageNumber="110">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D719236FF79FE89C95029D9" ID-GBIF-Occurrence="2898878450" collectingDate="2009-03-03" collectionCode="NME" collectorName="A. Skale &amp; Coll. A. Skale Hof" country="Indonesia" elevation="3" location="Skale river" pageId="109" pageNumber="110" specimenCount="1" typeStatus="holotype">
<paragraph id="922136E09D719236FF79FE89C90A2933" blockId="109.[189,1398,311,443]" box="[234,348,311,337]" pageId="109" pageNumber="110">
<emphasis id="A0EAEAF29D719236FF79FE89C90A2933" bold="true" box="[234,348,311,337]" pageId="109" pageNumber="110">
<typeStatus id="4D2588429D719236FF79FE89C90A2933" box="[234,348,311,337]" pageId="109" pageNumber="110" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D719236FF2EFEE4C95029D9" blockId="109.[189,1398,311,443]" pageId="109" pageNumber="110">
<collectingCountry id="EA8976709D719236FF2EFEE4C90D2916" box="[189,347,346,372]" name="Indonesia" pageId="109" pageNumber="110">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D719236FEE8FEE5C9C52917" box="[379,403,347,373]" pageId="109" pageNumber="110">♂</specimenCount>
; “
<collectingCountry id="EA8976709D719236FE3DFEE4CA192916" box="[430,591,346,372]" name="Indonesia" pageId="109" pageNumber="110">INDONESIA</collectingCountry>
C-Sulawesi ca.
<locationDeviation id="83932FC29D719236FC8AFEE4CB862916" box="[793,976,346,373]" location="Palu" pageId="109" pageNumber="110">
<quantity id="55669B059D719236FC8AFEE4CB0D2916" box="[793,859,346,372]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="109" pageNumber="110" unit="km" value="20.0">20km</quantity>
NE Palu
</locationDeviation>
, ca.
<quantity id="55669B059D719236FB98FEE4CC672916" box="[1035,1073,346,372]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="109" pageNumber="110" unit="m" value="3.0">
<elevation id="19B3D1D39D719236FB98FEE4CC672916" box="[1035,1073,346,372]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="109" pageNumber="110" unit="m" value="3.0">3m</elevation>
</quantity>
[sic
<specimenCount id="8498FD699D719236FBFEFEE4CC272917" box="[1133,1137,346,373]" pageId="109" pageNumber="110">♂</specimenCount>
] W Tawaeli,
<quantity id="55669B059D719236FA8BFEE4CD0D2916" box="[1304,1371,346,372]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="109" pageNumber="110" unit="m" value="170.0">
<elevation id="19B3D1D39D719236FA8BFEE4CD0D2916" box="[1304,1371,346,372]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="109" pageNumber="110" unit="m" value="170.0">170m</elevation>
</quantity>
<geoCoordinate id="F7AA50279D719236FAF5FEE4C97A29FA" degrees="0" direction="south" minutes="43" orientation="latitude" pageId="109" pageNumber="110" precision="15" seconds="56" value="-0.7322222">S 0°43’56”</geoCoordinate>
,
<geoCoordinate id="F7AA50279D719236FEAAFEC0C98829FA" box="[313,478,381,408]" degrees="119" direction="east" minutes="55" orientation="longitude" pageId="109" pageNumber="110" precision="15" seconds="30" value="119.924995">E 119°55’30”</geoCoordinate>
,
<date id="E62010209D719236FE7EFEC0CA2629FA" box="[493,624,382,408]" pageId="109" pageNumber="110" value="2009-03-03">
<collectingDate id="F664E9C89D719236FE7EFEC0CA2629FA" box="[493,624,382,408]" pageId="109" pageNumber="110" value="2009-03-03">03.III.2009</collectingDate>
</date>
leg.
<collectorName id="3F6B53369D719236FD3EFEC0CB4529F5" box="[685,787,381,408]" pageId="109" pageNumber="110">
A.
<location id="9741603B9D719236FD41FEC3CB4529F5" LSID="urn:lsid:plazi:treatment:1A3787F69D709235FDE8F993CB5C2A49:9741603B9D719236FD41FEC3CB4529F5" box="[722,787,381,407]" country="Indonesia" name="Skale river" pageId="109" pageNumber="110">Skale</location>
</collectorName>
river valley (020) //
<collectorName id="3F6B53369D719236FB96FEC3CCB129F5" box="[1029,1255,381,408]" pageId="109" pageNumber="110">Coll. A. Skale Hof</collectorName>
,
<collectingCountry id="EA8976709D719236FB67FEC0CD3E29FA" box="[1268,1384,382,408]" name="Germany" pageId="109" pageNumber="110">Germany</collectingCountry>
”;
<collectionCode id="F48FAE259D719236FF2EFE1FC95429D9" box="[189,258,417,443]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="109" pageNumber="110">NME</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="922136E09D719236FF79FE5CCA45299F" blockId="109.[189,1398,482,579]" box="[234,531,482,509]" pageId="109" pageNumber="110">
<emphasis id="A0EAEAF29D719236FF79FE5CC931299E" bold="true" box="[234,359,482,508]" pageId="109" pageNumber="110">
<typeStatus id="4D2588429D719236FF79FE5CC931299E" box="[234,359,482,508]" pageId="109" pageNumber="110" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="8498FD699D719236FEEBFE5CCA5D299F" box="[376,523,482,509]" pageId="109" pageNumber="110" type="generic">2 specimens</specimenCount>
)
</paragraph>
<paragraph id="922136E09D719236FF2EFDBBC9942A21" blockId="109.[189,1398,482,579]" pageId="109" pageNumber="110">
<materialsCitation id="22F63CBD9D719236FF2EFDBBCB672A7D" ID-GBIF-Occurrence="2898878524" box="[189,817,517,544]" pageId="109" pageNumber="110" typeStatus="holotype">
<collectingCountry id="EA8976709D719236FF2EFDBBC90D2A7D" box="[189,347,517,543]" name="Indonesia" pageId="109" pageNumber="110">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D719236FEE0FDBBC9F62A42" box="[371,416,517,544]" pageId="109" pageNumber="110" type="male">1 ♂</specimenCount>
; same labels as for
<typeStatus id="4D2588429D719236FD17FDBBCAB82A7D" box="[644,750,517,543]" pageId="109" pageNumber="110" type="holotype">holotype</typeStatus>
; ASC
</materialsCitation>
•
<materialsCitation id="22F63CBD9D719236FCDAFDBBC9E82A21" ID-GBIF-Occurrence="2898878388" collectionCode="NME" country="Indonesia" location="Sulawesi" pageId="109" pageNumber="110" specimenCount="1" stateProvince="Sulawesi Utara">
<specimenCount id="8498FD699D719236FCDAFDBBCB212A42" box="[841,887,517,544]" pageId="109" pageNumber="110" type="male">1 ♂</specimenCount>
; “
<collectingCountry id="EA8976709D719236FC1EFDBBCC782A7D" box="[909,1070,517,543]" name="Indonesia" pageId="109" pageNumber="110">INDONESIA</collectingCountry>
<collectingRegion id="505AF8029D719236FBA5FDBBCCF62A7D" box="[1078,1184,517,543]" country="Indonesia" name="Sulawesi Utara" pageId="109" pageNumber="110">Sulawesi</collectingRegion>
Puncak W Palopo 6.2000 Beneš”;
<collectionCode id="F48FAE259D719236FEEAFD97C9E82A21" box="[377,446,553,579]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="109" pageNumber="110">NME</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D719235FF2EFDD4C9FD2932" lastPageId="110" lastPageNumber="111" pageId="109" pageNumber="110" type="description">
<paragraph id="922136E09D719236FF2EFDD4C9182AE6" blockId="109.[189,334,618,644]" box="[189,334,618,644]" pageId="109" pageNumber="110">
<heading id="C969818C9D719236FF2EFDD4C9182AE6" bold="true" box="[189,334,618,644]" fontSize="11" level="3" pageId="109" pageNumber="110" reason="3">
<emphasis id="A0EAEAF29D719236FF2EFDD4C9182AE6" bold="true" box="[189,334,618,644]" pageId="109" pageNumber="110">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D719236FF2EFD27C8AF2B7E" blockId="109.[189,1399,664,796]" pageId="109" pageNumber="110">
MEASUREMENTS. Male
<typeStatus id="4D2588429D719236FE52FD26CA7D2AD0" box="[449,555,664,690]" pageId="109" pageNumber="110" type="holotype">holotype</typeStatus>
, total body length
<quantity id="55669B059D719236FC9AFD26CB202AD1" box="[777,886,664,691]" metricMagnitude="-3" metricUnit="m" metricValue="7.45" pageId="109" pageNumber="110" unit="mm" value="7.45">7.45 mm</quantity>
. Head
<quantity id="55669B059D719236FC5AFD27CC652AD1" box="[969,1075,665,691]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="109" pageNumber="110" unit="mm" value="1.3">1.30 mm</quantity>
long, across eyes
<quantity id="55669B059D719236FA9EFD27CD212AD1" box="[1293,1399,665,691]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="109" pageNumber="110" unit="mm" value="1.7">1.70 mm</quantity>
wide. Pronotum
<quantity id="55669B059D719236FEEDFD02C9B02AB4" box="[382,486,700,726]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="109" pageNumber="110" unit="mm" value="1.7">1.70 mm</quantity>
long, maximum width
<quantity id="55669B059D719236FD66FD02CB092AB4" box="[757,863,700,726]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="109" pageNumber="110" unit="mm" value="2.8">2.80 mm</quantity>
. Elytral length
<quantity id="55669B059D719236FB83FD05CC2D2AB4" box="[1040,1147,699,726]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" pageId="109" pageNumber="110" unit="mm" value="4.45">4.45 mm</quantity>
, maximum combined width
<quantity id="55669B059D719236FE95FD61C9272A9B" box="[262,369,735,762]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" pageId="109" pageNumber="110" unit="mm" value="3.45">3.45 mm</quantity>
.
<typeStatus id="4D2588429D719236FEE8FD61C9B42A9B" box="[379,482,735,761]" pageId="109" pageNumber="110" type="paratype">Paratype</typeStatus>
from locus typicus is
<quantity id="55669B059D719236FD4CFD61CB1C2A9B" box="[735,842,735,761]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="109" pageNumber="110" unit="mm" value="7.3">7.30 mm</quantity>
,
<typeStatus id="4D2588429D719236FCC7FD61CBEF2A9B" box="[852,953,735,761]" pageId="109" pageNumber="110" type="paratype">paratype</typeStatus>
from Palopo surroundings is
<quantity id="55669B059D719236FA9CFD61CD212A9B" box="[1295,1399,735,761]" metricMagnitude="-3" metricUnit="m" metricValue="8.67" pageId="109" pageNumber="110" unit="mm" value="8.67">8.67 mm</quantity>
long.
</paragraph>
<paragraph id="922136E09D719236FF2EFCFACC8D2F8A" blockId="109.[189,1399,835,2025]" pageId="109" pageNumber="110">
Dorsum and venter uniformly black-brown with brown labroclypeus, antennae and legs. Head transverse, convex dorsally between eyes, subopaque dorsally and ventrally, with large not prominent eyes occupying less than half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D719236FC0BFC34CBAD2BC6" box="[920,1019,906,932]" captionStart="Figs 169–180" captionStartId="143.[189,241,1787,1813]" captionTargetBox="[359,1217,271,1750]" captionTargetId="figure-123@143.[359,1228,265,1755]" captionTargetPageId="143" captionText="Figs 169–180. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 169. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 170. Same, paratype, ♂ (BMNH). 171. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 172. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 173. Same, ♂ (ZMHB). 174. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 175. M. similis sp. nov., paratype, ♂ (BMNH). 176. Same, paratype, ♀ (BMNH). 177. M. skalei sp. nov., paratype, ♂ (NME). 178. M. sougb sp. nov., holotype, ♂ (NME). 179. M. speciosus (Frey, 1969) comb. nov., ♂ (NHMB). 180. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122185" httpUri="https://zenodo.org/record/4122185/files/figure.png" pageId="109" pageNumber="110">Fig. 177</figureCitation>
) subtruncate on anterior margin, with lateral margins smooth, slightly sinuous in dorsal and nearly straight in lateral view. Anterolateral angles of labroclypeus not protruding anteriad, strongly rounded. Head punctures irregularly ovoid, deep and dense, filled with dense microscopical velvety pubescence. Intervening spaces glossy, generally smaller than punctures, in part wrinkled. Head setae dirty yellowish, moderately long and suberect; each seta rises from anterior margin of corresponding puncture, surpassing its length. Labroclypeus laterally and frontally with more delicate long sparse setae. Three longest setae present near eyes. Male antenna 9-segmented, club 3-lamellate. Scape moderately large, bulbous dorsally in distal half, provided with few long erect setae on posterior and distal margins. Antennomere 2 short, slightly transverse. Pronotum transverse, subopaque dorsally and laterally. Anterior margin of pronotum slightly sinuous with anterolateral angles (stronger) and mesal portion (slightly) protruding anteriad. Basal margin of pronotum broadly rounded. Lateral margins of pronotum broadly rounded with maximum width posterior to midlength, crenulate all along (
<figureCitation id="0AA52A659D719236FD24FA8ECB4B2D29" box="[695,797,1328,1355]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="109" pageNumber="110">Fig. 253</figureCitation>
). Long curved erect scale-like seta present between every two crenulae. Lateral margin of pronotum nearly straight in lateral view. Pronotal punctures ovoid, deep and dense. Intervening spaces glossy and glabrous, much smaller than punctures. Setae similar to those on head, stronger curved and in part clavate. Lateral and basal margins, antero- and posterolateral angles partly covered with microscopical velvety pubescence. Hypomeron smooth, sparsely covered by circular punctures, flange-like, with large emargination opposite to eye (to receive canthus separating compound eye) provided with long setae. Antennal pocket deep. Scutellar shield rounded apically. Elytra cylindrical, maximum width in median third, subopaque, with rounded humeri and slightly elevated omoplates. Indistinct tracks of three flat longitudinal carinae on each elytron, including sutural one. Disc and lateral sides of elytra irregularly densely punctured, punctures deep, of irregular size and shape (
<figureCitation id="0AA52A659D719236FE9CF92EC9202EC9" box="[271,374,1680,1707]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="109" pageNumber="110">Fig. 344</figureCitation>
). Perimeter of each puncture covered with microscopical velvety pubescence (
<figureCitation id="0AA52A659D719236FA97F92ECD3C2EC9" box="[1284,1386,1680,1707]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="109" pageNumber="110">Fig. 344</figureCitation>
). Intervening spaces glossy, generally smaller than punctures. Setae of two kinds on elytral disc. Long erect, in part clavate setae arranged in irregular longitudinal rows on each elytron. Inconspicuous short setae rising from anterior margin of corresponding punctures and not surpassing their length, positioned between longitudinal rows of longer setae. Male pygidium flattened dorsally, with moderately deep punctures and microreticulate intervening spaces of variable size (
<figureCitation id="0AA52A659D719236FC29F8FECC772F39" box="[954,1057,1856,1883]" captionStart="Figs 504–521" captionStartId="166.[189,241,1681,1707]" captionTargetBox="[433,1155,265,1647]" captionTargetId="figure-177@166.[433,1155,265,1658]" captionTargetPageId="166" captionText="Figs 504–521. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 504. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 505. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 506. Same, ♀ (BMNH). 507. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 508. Same, syntype, ♂ (BMNH). 509. M. peregrinus Lansberge, 1886, ♂ (DTC). 510. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 511. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 512. Same, paratype, ♂ (BMNH). 513. M. riedeli sp. nov., holotype, ♂ (SMNS). 514. M. rugicollis Moser, 1920, ♂ (BMNH). 515. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 516. M. similis sp. nov., paratype, ♂ (BMNH). 517. M. skalei sp. nov., paratype, ♂ (NME). 518. M. sougb sp. nov., holotype, ♂ (NME). 519. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 520. Same, different ♂ (BMNH). 521. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122228" httpUri="https://zenodo.org/record/4122228/files/figure.png" pageId="109" pageNumber="110">Fig. 517</figureCitation>
). Setae of pygidium suberect, moderately long, sparse. Male protibia with two strong distal teeth on external margin (Fig. 416). Male lower meso- and metatibial terminal spur strongly curved. Distal margin of male metatibia with projection over basal metatarsomere. Male basal metatarsomere widened and dorso-ventrally flattened (
<figureCitation id="0AA52A659D719236FF55F870C9782F8B" box="[198,302,1998,2025]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="109" pageNumber="110">Fig. 446</figureCitation>
). Tarsal claws with pulvilli (in males only?). Male aedeagus as in
<figureCitation id="0AA52A659D719236FBA1F870CC802F8A" box="[1074,1238,1998,2024]" captionStart="Figs 712–725" captionStartId="179.[189,241,1892,1918]" captionTargetBox="[433,1144,277,1858]" captionTargetId="figure-72@179.[427,1160,265,1870]" captionTargetPageId="179" captionText="Figs 712–725. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 712– 714. M. skalei sp. nov., holotype, ♂ (NME). 715–717. M. sougb sp. nov., holotype, ♂ (NME). 718–720. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 721–723. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 724–725. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122252" httpUri="https://zenodo.org/record/4122252/files/figure.png" pageId="109" pageNumber="110">Figs 712–714</figureCitation>
.
</paragraph>
<paragraph id="922136E09D729235FF2EFEB6C9E62940" blockId="110.[189,432,264,290]" box="[189,432,264,290]" pageId="110" pageNumber="111">
<heading id="C969818C9D729235FF2EFEB6C9E62940" bold="true" box="[189,432,264,290]" fontSize="11" level="3" pageId="110" pageNumber="111" reason="3">
<emphasis id="A0EAEAF29D729235FF2EFEB6C9E62940" bold="true" box="[189,432,264,290]" pageId="110" pageNumber="111">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D729235FF2EFE88C9FD2932" blockId="110.[189,427,310,336]" box="[189,427,310,336]" pageId="110" pageNumber="111">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D729235FF2EFEC8CA4929DF" box="[189,543,374,445]" pageId="110" pageNumber="111" type="biology_ecology">
<paragraph id="922136E09D729235FF2EFEC8C97629F2" blockId="110.[189,288,374,400]" box="[189,288,374,400]" pageId="110" pageNumber="111">
<heading id="C969818C9D729235FF2EFEC8C97629F2" bold="true" box="[189,288,374,400]" fontSize="11" level="3" pageId="110" pageNumber="111" reason="3">
<emphasis id="A0EAEAF29D729235FF2EFEC8C97629F2" bold="true" box="[189,288,374,400]" pageId="110" pageNumber="111">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D729235FF2EFE1DCA4929DF" blockId="110.[189,543,419,445]" box="[189,543,419,445]" pageId="110" pageNumber="111">Occurs in lowland rainforests.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D729235FF2EFE5DCB5C2A49" box="[189,778,483,555]" pageId="110" pageNumber="111" type="distribution">
<paragraph id="922136E09D729235FF2EFE5DC900299F" blockId="110.[189,342,483,509]" box="[189,342,483,509]" pageId="110" pageNumber="111">
<heading id="C969818C9D729235FF2EFE5DC900299F" bold="true" box="[189,342,483,509]" fontSize="11" level="3" pageId="110" pageNumber="111" reason="3">
<emphasis id="A0EAEAF29D729235FF2EFE5DC900299F" bold="true" box="[189,342,483,509]" pageId="110" pageNumber="111">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D729235FF2EFDAFCB5C2A49" blockId="110.[189,778,529,555]" box="[189,778,529,555]" pageId="110" pageNumber="111">
Hitherto known from Central and
<collectingRegion id="505AF8029D729235FDDEFDAFCB512A49" box="[589,775,529,555]" country="Indonesia" name="Sulawesi Selatan" pageId="110" pageNumber="111">South Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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