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<document id="FAFD8EA8D03BB52D7807F910E6C2C9E6" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D679227FDCBFD6AC9F12C81" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius vicinus Heller 1914" docType="treatment" docVersion="10" lastPageNumber="125" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="124" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="5D85938E61FD196D9C419E6536EC0A0B">
<mods:title id="E41FB3562DE1C90138C1B229022A1BB7">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="369EF0AFFCE8270FF5EB1B537CC21894" type="personal">
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<mods:namePart id="4857DCC78C3D6831FCDEDDFFBC2A2D6D">Telnov, Dmitry</mods:namePart>
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<mods:title id="59821BFE1E93DEB2B99E31359FA28F57">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="7EF45285E1A132C33F55C0FC08469815">
<mods:date id="F32D50A83FD03E559107875A4DA984EF">2020</mods:date>
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<mods:number id="D4819CCAF91B99F646BD177E3CDDE407">2020-10-19</mods:number>
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<mods:detail id="404F98BFEEBFC36888DB15CD452A26AE" type="volume">
<mods:number id="846485670B8879968119CEA5F3C1EDC7">721</mods:number>
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<mods:classification id="18DF41E512A3A7FE77B36B90430C2B32">journal article</mods:classification>
<mods:identifier id="25C0CEDCEEE27605B614B0DFE9811519" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="8B2D0D37A7782D6C30ABFD1654EA4453" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D679227FDCBFD6AC9F12C81" ID-DOI="http://doi.org/10.5281/zenodo.4344543" ID-GBIF-Taxon="168648178" ID-Zenodo-Dep="4344543" LSID="urn:lsid:plazi:treatment:1A3787F69D679227FDCBFD6AC9F12C81" httpUri="http://treatment.plazi.org/id/1A3787F69D679227FDCBFD6AC9F12C81" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124">
<subSubSection id="DA84656B9D679220FDCBFD6ACB8A2A8D" box="[600,988,724,751]" pageId="123" pageNumber="124" type="nomenclature">
<paragraph id="922136E09D679220FDCBFD6ACB8A2A8D" blockId="123.[457,1130,724,787]" box="[600,988,724,751]" pageId="123" pageNumber="124">
<heading id="C969818C9D679220FDCBFD6ACB8A2A8D" box="[600,988,724,751]" centered="true" fontSize="11" level="2" pageId="123" pageNumber="124" reason="2">
<taxonomicName id="559E4D639D679220FDCBFD6ACB8A2A8D" ID-CoL="6QTFM" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[600,988,724,751]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="A0EAEAF29D679220FDCBFD6ACB152A8D" bold="true" box="[600,835,724,751]" italics="true" pageId="123" pageNumber="124">Maechidius vicinus</emphasis>
<bibRefCitation id="F60F4B119D679220FCD9FD6BCB8A2A8D" author="Heller K. M." box="[842,988,725,751]" pageId="123" pageNumber="124" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D679220FE5AFD46CC3C2B70" box="[457,1130,760,787]" pageId="123" pageNumber="124" type="description">
<paragraph id="922136E09D679220FE5AFD46CC3C2B70" blockId="123.[457,1130,724,787]" box="[457,1130,760,787]" pageId="123" pageNumber="124">
<figureCitation id="0AA52A659D679220FE5AFD46CA752B70" box="[457,547,760,786]" captionStart="Figs 90–93" captionStartId="119.[189,241,1928,1954]" captionTargetBox="[329,1264,264,1883]" captionTargetId="figure-60@119.[321,1264,256,1905]" captionTargetPageId="119" captionText="Figs 90–93. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 90. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 91. M. trivialis sp. nov., holotype, ♂ (BMNH). 92. M. ursus sp. nov., holotype, ♀ (NME). 93. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122169" httpUri="https://zenodo.org/record/4122169/files/figure.png" pageId="123" pageNumber="124">Figs 93</figureCitation>
,
<figureCitation id="0AA52A659D679220FDBCFD46CACE2B70" box="[559,664,760,786]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="123" pageNumber="124">189–190</figureCitation>
,
<figureCitation id="0AA52A659D679220FD37FD46CA852B70" box="[676,723,760,786]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="123" pageNumber="124">262</figureCitation>
,
<figureCitation id="0AA52A659D679220FD4CFD46CB582B70" box="[735,782,760,786]" captionStart="Figs 278–285" captionStartId="151.[189,241,1697,1723]" captionTargetBox="[211,1375,265,1597]" captionTargetId="figure-104@151.[211,1376,265,1614]" captionTargetPageId="151" captionText="Figs 278–285. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventro-lateral view. 278. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 279. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 280. M. seriepunctatus Moser, 1920, lectotype, ♂ (SNSD). 281. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 282. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 283. M. trivialis sp. nov., holotype, ♂ (BMNH). 284. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 285. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122202" httpUri="https://zenodo.org/record/4122202/files/figure.png" pageId="123" pageNumber="124">284</figureCitation>
,
<figureCitation id="0AA52A659D679220FC8AFD46CB1E2B71" box="[793,840,760,787]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="123" pageNumber="124">353</figureCitation>
,
<figureCitation id="0AA52A659D679220FCC7FD46CBD52B71" box="[852,899,760,787]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="123" pageNumber="124">425</figureCitation>
, 529–530, 736–738
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D679220FF2EFC84CA302EE5" pageId="123" pageNumber="124" type="materials_examined">
<paragraph id="922136E09D679220FF2EFC84C93A2B31" blockId="123.[189,364,825,852]" box="[189,364,825,852]" pageId="123" pageNumber="124">
<heading id="C969818C9D679220FF2EFC84C93A2B31" bold="true" box="[189,364,825,852]" fontSize="11" level="3" pageId="123" pageNumber="124" reason="3">
<emphasis id="A0EAEAF29D679220FF2EFC84C93A2B31" bold="true" box="[189,364,825,852]" pageId="123" pageNumber="124">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D679220FF79FCD6CA3A2BE1" blockId="123.[189,1399,872,1004]" box="[234,620,872,899]" pageId="123" pageNumber="124">
<emphasis id="A0EAEAF29D679220FF79FCD6C9302BE0" bold="true" box="[234,358,872,898]" pageId="123" pageNumber="124">
<typeStatus id="4D2588429D679220FF79FCD6C9302BE0" box="[234,358,872,898]" pageId="123" pageNumber="124" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
</paragraph>
<paragraph id="922136E09D679220FF2EFC32CA602B8E" blockId="123.[189,1399,872,1004]" pageId="123" pageNumber="124">
<materialsCitation id="22F63CBD9D679220FF2EFC32CA602B8E" ID-GBIF-Occurrence="2898878582" collectionCode="SNSD" collectorName="Gehr. W. Muller Vermacht." country="Guinea-Bissau" location="Island of New" pageId="123" pageNumber="124" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
COUNTRY UNKNOWN (
<location id="9741603B9D679220FD90FC35CAE22BC4" LSID="urn:lsid:plazi:treatment:1A3787F69D679227FDCBFD6AC9F12C81:9741603B9D679220FD90FC35CAE22BC4" box="[515,692,907,934]" country="Guinea-Bissau" name="Island of New" pageId="123" pageNumber="124">Island of New</location>
<collectingCountry id="EA8976709D679220FD52FC35CB4E2BC4" box="[705,792,907,934]" name="Guinea-Bissau" pageId="123" pageNumber="124">Guinea</collectingCountry>
) •
<specimenCount id="8498FD699D679220FCD1FC32CB0F2BC4" box="[834,857,908,934]" pageId="123" pageNumber="124" type="female">♀</specimenCount>
; “
<collectorName id="3F6B53369D679220FCE4FC35CC902BC7" box="[887,1222,907,934]" pageId="123" pageNumber="124">Gehr. W. Müller Vermächt.</collectorName>
1909 [p, blue label]//
<typeStatus id="4D2588429D679220FE84FC11C9332BAB" box="[279,357,943,969]" pageId="123" pageNumber="124">Typus</typeStatus>
! [p]
<taxonomicName id="559E4D639D679220FE3EFC11CA0C2BAB" authority="Heller" authorityName="Heller" authorityYear="1914" box="[429,602,943,969]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="A0EAEAF29D679220FE3EFC11CA562BAA" box="[429,512,943,968]" italics="true" pageId="123" pageNumber="124">vicinus</emphasis>
Heller
</taxonomicName>
[h, red label] // N.
<collectingCountry id="EA8976709D679220FCDBFC11CBF62BAB" box="[840,928,943,969]" name="Guinea-Bissau" pageId="123" pageNumber="124">Guinea</collectingCountry>
[p, blue label] // Staatl. Museum für Tierkunde,Dresden [p]”;
<collectionCode id="F48FAE259D679220FE77FC6CCA652B8E" box="[484,563,978,1004]" collectionName="SNSD" pageId="123" pageNumber="124">SNSD</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D679220FF79FBAAC9C12C4C" blockId="123.[189,1399,1044,1141]" box="[234,407,1044,1070]" pageId="123" pageNumber="124">
<emphasis id="A0EAEAF29D679220FF79FBAAC9C12C4C" bold="true" box="[234,407,1044,1070]" pageId="123" pageNumber="124">
<typeStatus id="4D2588429D679220FF79FBAAC9C12C4C" box="[234,407,1044,1070]" pageId="123" pageNumber="124" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D679220FF2EFB89CCCD2C16" blockId="123.[189,1399,1044,1141]" pageId="123" pageNumber="124">
<materialsCitation id="22F63CBD9D679220FF2EFB89CB472C30" ID-GBIF-Occurrence="2898878547" box="[189,785,1079,1106]" country="Guinea-Bissau" location="Island of New" pageId="123" pageNumber="124" specimenCount="1" typeStatus="paralectotype">
COUNTRY UNKNOWN (
<location id="9741603B9D679220FE6EFB89CAFF2C33" LSID="urn:lsid:plazi:treatment:1A3787F69D679227FDCBFD6AC9F12C81:9741603B9D679220FE6EFB89CAFF2C33" box="[509,681,1079,1105]" country="Guinea-Bissau" name="Island of New" pageId="123" pageNumber="124">
<collectingCountry id="EA8976709D679220FE6EFB89CA102C33" box="[509,582,1079,1105]" name="Iceland" pageId="123" pageNumber="124">Island</collectingCountry>
of New
</location>
<collectingCountry id="EA8976709D679220FD21FB89CB5F2C30" box="[690,777,1079,1106]" name="Guinea-Bissau" pageId="123" pageNumber="124">Guinea</collectingCountry>
)
</materialsCitation>
•
<materialsCitation id="22F63CBD9D679220FCBDFB89CCCE2C16" ID-GBIF-Occurrence="2898878321" collectionCode="SNSD" collectorName="Gehr. W. Muller Vermacht." country="Guinea-Bissau" location="Guinea" pageId="123" pageNumber="124" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="8498FD699D679220FCBDFB89CB0B2C33" box="[814,861,1079,1105]" pageId="123" pageNumber="124" type="female">1 ♀</specimenCount>
; “N.
<collectingCountry id="EA8976709D679220FC02FB89CBBD2C33" box="[913,1003,1079,1105]" name="Guinea-Bissau" pageId="123" pageNumber="124">Guinea</collectingCountry>
[p, blue label]//
<collectorName id="3F6B53369D679220FB27FB89C9632C16" pageId="123" pageNumber="124">Gehr. W. Müller Vermächt.</collectorName>
1909 [p, blue label]// Staatl. Museum für Tierkunde Dresden [p]”;
<collectionCode id="F48FAE259D679220FBDAFBE4CCCE2C16" box="[1097,1176,1114,1140]" collectionName="SNSD" pageId="123" pageNumber="124">SNSD</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="922136E09D679220FF2EFB22C9302CD7" blockId="123.[189,358,1179,1206]" box="[189,358,1179,1206]" pageId="123" pageNumber="124">
<heading id="C969818C9D679220FF2EFB22C9302CD7" bold="true" box="[189,358,1179,1206]" fontSize="11" level="3" pageId="123" pageNumber="124" reason="3">
<emphasis id="A0EAEAF29D679220FF2EFB22C9302CD7" bold="true" box="[189,358,1179,1206]" pageId="123" pageNumber="124">New material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D679220FF2EFB74CC4A2D6A" blockId="123.[189,1399,1225,1288]" pageId="123" pageNumber="124">
<materialsCitation id="22F63CBD9D679220FF2EFB74CC4E2D6A" ID-GBIF-Occurrence="2898878626" collectionCode="SNSD" collectorName="Deutsch N. Guinea &amp; Gehr. W. Muller Vermacht." country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="123" pageNumber="124" specimenCount="3" specimenCount-female="3">
<collectingCountry id="EA8976709D679220FF2EFB74C98D2C86" box="[189,475,1226,1253]" name="Papua New Guinea" pageId="123" pageNumber="124">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D679220FE64FB74CA732C87" box="[503,549,1226,1253]" pageId="123" pageNumber="124" type="male">1 ♂</specimenCount>
,
<specimenCount id="8498FD699D679220FDA7FB74CA2E2C87" box="[564,632,1226,1253]" pageId="123" pageNumber="124" type="female">3 ♀♀</specimenCount>
; “
<collectorName id="3F6B53369D679220FD01FB74CB392C86" box="[658,879,1226,1252]" pageId="123" pageNumber="124">
Deutsch N.
<collectingCountry id="EA8976709D679220FC84FB74CB392C86" box="[791,879,1226,1252]" name="Guinea-Bissau" pageId="123" pageNumber="124">Guinea</collectingCountry>
</collectorName>
[p, blue label]//
<collectorName id="3F6B53369D679220FBA6FB74CD212C86" box="[1077,1399,1226,1253]" pageId="123" pageNumber="124">Gehr. W. Müller Vermächt.</collectorName>
1909 [p, blue label]// Staatl. Museum für Tierkunde Dresden [p]”;
<collectionCode id="F48FAE259D679220FC5AFB50CC4E2D6A" box="[969,1048,1262,1288]" collectionName="SNSD" pageId="123" pageNumber="124">SNSD</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="922136E09D679220FF2EFA91CA302EE5" blockId="123.[189,1399,1327,1671]" pageId="123" pageNumber="124">
<bibRefCitation id="F60F4B119D679220FF2EFA91C9CA2D28" author="Heller K. M." box="[189,412,1327,1354]" pageId="123" pageNumber="124" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller (1914: 628)</bibRefCitation>
based his description of
<taxonomicName id="559E4D639D679220FD5FFA8ECB1F2D2B" authorityName="Heller" authorityYear="1914" box="[716,841,1328,1353]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="A0EAEAF29D679220FD5FFA8ECB1F2D2B" box="[716,841,1328,1353]" italics="true" pageId="123" pageNumber="124">M. vicinus</emphasis>
</taxonomicName>
on an unstated number of specimens. At least two specimens with nearly identical labels are allocated in
<collectionCode id="F48FAE259D679220FC02FAECCBB62D0E" box="[913,992,1362,1388]" pageId="123" pageNumber="124">SNSD</collectionCode>
. The
<typeStatus id="4D2588429D679220FBB4FAECCCC52D0E" box="[1063,1171,1362,1388]" pageId="123" pageNumber="124" type="lectotype">lectotype</typeStatus>
here designated is the only
<typeStatus id="4D2588429D679220FEBBFAC8C9D52DF2" box="[296,387,1398,1424]" pageId="123" pageNumber="124" type="syntype">syntype</typeStatus>
with Heller’s original handwriting on a red label, which with some confidence may indicate this specimen was originally selected by the author as
<typeStatus id="4D2588429D679220FC2BFA27CC752DD1" box="[952,1059,1433,1459]" pageId="123" pageNumber="124" type="holotype">holotype</typeStatus>
. The
<typeStatus id="4D2588429D679220FBF5FA27CC842DD1" box="[1126,1234,1433,1459]" pageId="123" pageNumber="124" type="lectotype">lectotype</typeStatus>
is designated in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other (non-conspecific)
<typeStatus id="4D2588429D679220FD84FA5ECA282D98" box="[535,638,1504,1530]" pageId="123" pageNumber="124" type="syntype">syntypes</typeStatus>
are discovered in the future. Additional
<collectionCode id="F48FAE259D679220FBFAFA61CCE22D9B" box="[1129,1204,1503,1529]" pageId="123" pageNumber="124">SNSD</collectionCode>
specimens with slightly different labels (see Checklist) are not considered
<typeStatus id="4D2588429D679220FCE0F9BDCB8C2E7F" box="[883,986,1539,1565]" pageId="123" pageNumber="124" type="syntype">syntypes</typeStatus>
since
<bibRefCitation id="F60F4B119D679220FBB4F9BCCC902E7F" author="Heller K. M." box="[1063,1222,1538,1565]" pageId="123" pageNumber="124" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller (1914)</bibRefCitation>
in the original description questioned the origin of the available specimens: “Hab. Nova
<collectingCountry id="EA8976709D679220FB8FF998CC272E22" box="[1052,1137,1574,1600]" name="Guinea-Bissau" pageId="123" pageNumber="124">Guinea</collectingCountry>
(Germanica? …)” but additional specimens provided with labels exactly specifying that they coming from the former German New
<collectingCountry id="EA8976709D679220FF6FF9D2C9072EE4" box="[252,337,1644,1670]" name="Guinea-Bissau" pageId="123" pageNumber="124">Guinea</collectingCountry>
(“Deutsch N.
<collectingCountry id="EA8976709D679220FE62F9D2CA192EE5" box="[497,591,1644,1671]" name="Guinea-Bissau" pageId="123" pageNumber="124">Guinea</collectingCountry>
”).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D679227FF2EF910C9F12C81" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124" type="description">
<paragraph id="922136E09D679220FF2EF910C9182EAA" blockId="123.[189,334,1710,1736]" box="[189,334,1710,1736]" pageId="123" pageNumber="124">
<heading id="C969818C9D679220FF2EF910C9182EAA" bold="true" box="[189,334,1710,1736]" fontSize="11" level="3" pageId="123" pageNumber="124" reason="3">
<emphasis id="A0EAEAF29D679220FF2EF910C9182EAA" bold="true" box="[189,334,1710,1736]" pageId="123" pageNumber="124">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D679220FF2EF963CBF62E95" blockId="123.[189,928,1756,1783]" box="[189,928,1756,1783]" pageId="123" pageNumber="124">
MEASUREMENTS. Total body length 6.20 (
<typeStatus id="4D2588429D679220FD00F962CB542E95" box="[659,770,1756,1783]" pageId="123" pageNumber="124" type="lectotype">lectotype</typeStatus>
) to
<quantity id="55669B059D679220FCA3F962CBCD2E95" box="[816,923,1756,1783]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="123" pageNumber="124" unit="mm" value="6.9">6.90 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D679227FF2EF8A0CB502C2D" blockId="123.[189,1399,1822,2025]" lastBlockId="124.[189,1399,265,1104]" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124">
Dorsum uniformly brown, forebody dorsally with very vague green lustre, venter and appendages paler brown. Head slightly convex dorsally between compound eyes, glossy dorsally and ventrally. Male labroclypeus (
<figureCitation id="0AA52A659D679220FE3DF8DACA432F1D" box="[430,533,1892,1919]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="123" pageNumber="124">Fig. 190</figureCitation>
) broadly V-shaped emarginate on anterior margin, female labroclypeus (
<figureCitation id="0AA52A659D679220FF55F836C97F2FC0" box="[198,297,1928,1954]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="123" pageNumber="124">Fig. 189</figureCitation>
) subtruncate to very broadly emarginate, its lateral margins slightly sinuous (in male) to nearly straight (in female) in both dorsal and lateral views. Anterolateral angles of labroclypeus obtuse, nearly right-angled and moderately strongly protruding anteriad in male, rounded and not projecting in female, raised up at angle of nearly 60-70° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Upper- and underside of labroclypeus with sparse inconspicuous setae along anterior and lateral margins. Canthus obtuse angulate in dorsal view. Punctures of frons moderately deep and large, annular. Intervening spaces glossy, generally smaller than punctures. Head setae inconspicuous, suberect; each seta rises from anterior margin or centre of corresponding puncture, surpassing its length. Antenna 9-segmented, club 3-lamellate. Pronotum transverse, flattened dorsally, glossy dorsally, subopaque laterally. Anterior margin of pronotum deeply emarginate with anterolateral angles protruding anteriad. Basal margin of pronotum broadly rounded. Lateral margin of pronotum broadly rounded, delicately crenulate all along (intervals between crenulae narrow, crenulae nearly merged) (
<figureCitation id="0AA52A659D609227FBE7FD9ACC8A2A5C" box="[1140,1244,547,574]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="124" pageNumber="125">Fig. 262</figureCitation>
). Moderately long suberect curved seta present between every two crenulae. Lateral margin of pronotum slightly sinuous in lateral view. Hypomeron separated from prosternum by moderately low slightly curved carina, with long setae on its anterolateral margin opposite to compound eye (
<figureCitation id="0AA52A659D609227FB14FD30CCA22ACA" box="[1159,1268,654,680]" captionStart="Figs 278–285" captionStartId="151.[189,241,1697,1723]" captionTargetBox="[211,1375,265,1597]" captionTargetId="figure-104@151.[211,1376,265,1614]" captionTargetPageId="151" captionText="Figs 278–285. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventro-lateral view. 278. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 279. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 280. M. seriepunctatus Moser, 1920, lectotype, ♂ (SNSD). 281. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 282. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 283. M. trivialis sp. nov., holotype, ♂ (BMNH). 284. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 285. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122202" httpUri="https://zenodo.org/record/4122202/files/figure.png" pageId="124" pageNumber="125">Fig. 284</figureCitation>
). Antennal pocket moderately deep. Pronotal punctures ovoid, annular, shallow. Intervening spaces variably large, generally smaller than punctures. Setation as on head but setae longer and curved. Scutellar shield pointed apically. Elytron opaque, with tracks of three-four flat almost entirely glabrous longitudinal carinae, including sutural one. Elytral punctures linear (elongate and narrow), incision-shaped, shallow (
<figureCitation id="0AA52A659D609227FF55FC80C9792B3B" box="[198,303,830,857]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="124" pageNumber="125">Fig. 353</figureCitation>
). Intervening spaces densely microreticulate. Appressed to suberect seta rises from anterior margin of each puncture, not surpassing length of corresponding incision. Disc and lateral margins of elytra in part covered with microscopical velvety pubescence. Male pygidium flattened dorsally, with large and shallow annular punctures (Fig. 530). Female pygidium flattened dorsally, with small median hump (Fig. 529). Intervening spaces generally much smaller than punctures, subopaque. Setae of pygidium suberect, sparse. Protibia widened distally, with three acute external teeth (
<figureCitation id="0AA52A659D609227FB2CFC51CD712C68" box="[1215,1319,1007,1034]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="124" pageNumber="125">Fig. 425</figureCitation>
). Male protibial terminal spur straight, narrow and pointed, female one short and somewhat thickened. Tarsal claws with pulvilli. Aedeagus as in
<figureCitation id="0AA52A659D609227FDCEFB8BCB542C2D" box="[605,770,1077,1103]" captionStart="Figs 726–738" captionStartId="180.[189,241,1891,1917]" captionTargetBox="[343,1246,278,1849]" captionTargetId="figure-69@180.[330,1258,265,1871]" captionTargetPageId="180" captionText="Figs 726–738. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 726– 729. M. suwawa sp. nov., holotype, ♂ (BMNH). 730–732. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 733–735. M. trivialis sp. nov., holotype, ♂ (BMNH). 736–738. M. vicinus Heller, 1914, ♂ (apex of parameres broken) (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122254" httpUri="https://zenodo.org/record/4122254/files/figure.png" pageId="124" pageNumber="125">Figs 736–738</figureCitation>
.
</paragraph>
<paragraph id="922136E09D609227FF2EFBC9C9E62CF3" blockId="124.[189,432,1143,1169]" box="[189,432,1143,1169]" pageId="124" pageNumber="125">
<heading id="C969818C9D609227FF2EFBC9C9E62CF3" bold="true" box="[189,432,1143,1169]" fontSize="11" level="3" pageId="124" pageNumber="125" reason="3">
<emphasis id="A0EAEAF29D609227FF2EFBC9C9E62CF3" bold="true" box="[189,432,1143,1169]" pageId="124" pageNumber="125">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D609227FF2EFB18C9F12C81" blockId="124.[189,1398,1190,1251]" pageId="124" pageNumber="125">Female lamellae of antennal club shorter; female anterolateral angles of labroclypeus less strongly protruding anteriad.</paragraph>
</subSubSection>
</treatment>
</document>
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