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<document id="8633A3374A884996327C124682AF501C" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D3D9278FDF7F98FCC582A51" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius caperatus Telnov 2020, sp. nov." docType="treatment" docUuid="9440F4C6-085C-46F1-BBDB-469D0D04AC53" docUuidSource="ZooBank" docVersion="10" lastPageNumber="36" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="34" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="90E28405A62A555ACD20408B33812E6F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FD70A52DD88C199614F90CAC217AA7CE">
<mods:title id="A1E7CF93107E0F6A6A2A38CFDB8C5F65">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="9E92104AC55A4DF44CF54CD907CE5021" type="personal">
<mods:role id="064996E5D189148B18DA27F103AF6D0F">
<mods:roleTerm id="FFADC6A75182BA39AA6CCDE59E385C38">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="63D6654487C7E00F732C05735CA930C3">Telnov, Dmitry</mods:namePart>
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<mods:titleInfo id="77E668523B9947F5942954FA3E12B93A">
<mods:title id="827569E8F45CFC46E56CB300EF285F11">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="18C205F0FA1F97478E26FD971843F59C">
<mods:date id="E7A0A406B8D2130CDC26658BFCC385A6">2020</mods:date>
<mods:detail id="ADB83AFC1497EBECC79B0C6BA86D2CDD" type="pubDate">
<mods:number id="9F486DD1B1B6C0167E62F74BD49608A9">2020-10-19</mods:number>
</mods:detail>
<mods:detail id="B66D7051D747A76163F9CB7B5DF049D4" type="volume">
<mods:number id="450FE4AB31A50B5FE1C518B687A0680C">721</mods:number>
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<mods:classification id="B1FE0914E6851EB3927417A168E712AF">journal article</mods:classification>
<mods:identifier id="138ACA4B31FFC3E7FDCF40459A510E0C" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="FA29ABB70321537F2948F1C0C3767255" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="01301EF5104B50DE7AE55DA7F4B0467B" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
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<treatment id="1A3787F69D3D9278FDF7F98FCC582A51" ID-DOI="http://doi.org/10.5281/zenodo.4344397" ID-GBIF-Taxon="168648151" ID-Zenodo-Dep="4344397" LSID="urn:lsid:zoobank.org:act:9440F4C6-085C-46F1-BBDB-469D0D04AC53" httpUri="http://treatment.plazi.org/id/1A3787F69D3D9278FDF7F98FCC582A51" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
<subSubSection id="DA84656B9D3D927AFDF7F98FCB862E2E" box="[612,976,1585,1612]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="922136E09D3D927AFDF7F98FCB862E2E" blockId="33.[363,1224,1585,1686]" box="[612,976,1585,1612]" pageId="33" pageNumber="34">
<heading id="C969818C9D3D927AFDF7F98FCB862E2E" box="[612,976,1585,1612]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="34" reason="2">
<taxonomicName id="559E4D639D3D927AFDF7F98FCB382E2E" ID-CoL="7X3WH" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[612,878,1585,1612]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="caperatus" status="sp. nov.">
<emphasis id="A0EAEAF29D3D927AFDF7F98FCB382E2E" bold="true" box="[612,878,1585,1612]" italics="true" pageId="33" pageNumber="34">Maechidius caperatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D3D927AFCE5F98CCB862E2E" box="[886,976,1586,1612]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3D927AFEF8F9E8CC662EF4" pageId="33" pageNumber="34" type="description">
<paragraph id="922136E09D3D927AFEF8F9E8CC9E2E13" blockId="33.[363,1224,1585,1686]" box="[363,1224,1622,1649]" pageId="33" pageNumber="34">
<uri id="E60F3AE29D3D927AFEF8F9E8CC9E2E13" box="[363,1224,1622,1649]" pageId="33" pageNumber="34">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D3D927AFD18F9E8CC9E2E13" box="[651,1224,1622,1649]" pageId="33" pageNumber="34">9440F4C6-085C-46F1-BBDB-469D0D04AC53</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D3D927AFD90F9C5CC662EF4" blockId="33.[363,1224,1585,1686]" box="[515,1072,1659,1686]" pageId="33" pageNumber="34">
<figureCitation id="0AA52A659D3D927AFD90F9C5CA0B2EF7" box="[515,605,1659,1685]" captionStart="Figs 28–30" captionStartId="31.[189,241,1928,1954]" captionTargetBox="[347,1240,269,1901]" captionTargetId="figure-52@31.[345,1242,265,1911]" captionTargetPageId="31" captionText="Figs 28–30. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 28. M. boessnecki sp. nov., holotype, ♂ (SNSD). 29. M. brocki sp. nov., holotype, ♂ (BMNH). 30. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122136" httpUri="https://zenodo.org/record/4122136/files/figure.png" pageId="33" pageNumber="34">Figs 30</figureCitation>
,
<figureCitation id="0AA52A659D3D927AFDFAF9C5CAC12EF7" box="[617,663,1659,1685]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="33" pageNumber="34">108</figureCitation>
,
<figureCitation id="0AA52A659D3D927AFD37F9C5CA852EF7" box="[676,723,1659,1685]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="33" pageNumber="34">208</figureCitation>
,
<figureCitation id="0AA52A659D3D927AFD4CF9C5CB582EF7" box="[735,782,1659,1685]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="33" pageNumber="34">297</figureCitation>
, 369, 463, 540, 592–594
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3D927AFF2EF900CB912F33" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="922136E09D3D927AFF2EF900C99F2EBA" blockId="33.[189,457,1726,1752]" box="[189,457,1726,1752]" pageId="33" pageNumber="34">
<heading id="C969818C9D3D927AFF2EF900C99F2EBA" bold="true" box="[189,457,1726,1752]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="A0EAEAF29D3D927AFF2EF900C99F2EBA" bold="true" box="[189,457,1726,1752]" pageId="33" pageNumber="34">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3D927AFF2EF950CB912F33" blockId="33.[189,1399,1774,1873]" pageId="33" pageNumber="34">
This species strongly resembles
<taxonomicName id="559E4D639D3D927AFDD7F950CBD32F6A" authority="Arrow" authorityName="Arrow" authorityYear="1941" box="[580,901,1774,1800]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="sturnus">
<emphasis id="A0EAEAF29D3D927AFDD7F950CB782F6A" box="[580,814,1774,1800]" italics="true" pageId="33" pageNumber="34">Maechidius sturnus</emphasis>
Arrow
</taxonomicName>
(Yapen Island), but differs readily in the shape of the aedeagus, less regular (stronger sinuous arranged) punctures of the elytra and in the absence of three long paired setae on the male abdominal sternites 2 and 3.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3D927AFF2EF8C4CB1F2F8A" pageId="33" pageNumber="34" type="etymology">
<paragraph id="922136E09D3D927AFF2EF8C4C9122FF6" blockId="33.[189,324,1914,1940]" box="[189,324,1914,1940]" pageId="33" pageNumber="34">
<heading id="C969818C9D3D927AFF2EF8C4C9122FF6" bold="true" box="[189,324,1914,1940]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="A0EAEAF29D3D927AFF2EF8C4C9122FF6" bold="true" box="[189,324,1914,1940]" pageId="33" pageNumber="34">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3D927AFF2EF814CB1F2F8A" blockId="33.[189,1399,1962,2025]" pageId="33" pageNumber="34">
The name of this species is derived from the Latin ‘
<taxonomicName id="559E4D639D3D927AFCA5F815CBFC2FA6" authorityName="Telnov" authorityYear="2020" box="[822,938,1963,1988]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="caperatus">
<emphasis id="A0EAEAF29D3D927AFCA5F815CBFC2FA6" box="[822,938,1963,1988]" italics="true" pageId="33" pageNumber="34">caperatus</emphasis>
</taxonomicName>
’, meaning ‘wrinkled’, referring to the irregularly wrinkled pattern of the pronotum and elytra.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3E9279FF2EFEB7C95029A3" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="922136E09D3E9279FF2EFEB7C93A2940" blockId="34.[189,364,264,291]" box="[189,364,264,291]" pageId="34" pageNumber="35">
<heading id="C969818C9D3E9279FF2EFEB7C93A2940" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="A0EAEAF29D3E9279FF2EFEB7C93A2940" bold="true" box="[189,364,264,291]" pageId="34" pageNumber="35">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3E9279FF79FE87C90A2931" blockId="34.[189,1399,313,449]" box="[234,348,313,339]" pageId="34" pageNumber="35">
<emphasis id="A0EAEAF29D3E9279FF79FE87C90A2931" bold="true" box="[234,348,313,339]" pageId="34" pageNumber="35">
<typeStatus id="4D2588429D3E9279FF79FE87C90A2931" box="[234,348,313,339]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D3E9279FF2EFEE3C95029A3" blockId="34.[189,1399,313,449]" pageId="34" pageNumber="35">
<materialsCitation id="22F63CBD9D3E9279FF2EFEE3C95029A3" ID-GBIF-Occurrence="2898878494" collectingDate="1998-04" collectionCode="NME" collectorName="M. Balke" country="Indonesia" elevation="750" location="Nabire area" municipality="Nabire area" pageId="34" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Irian Jaya" typeStatus="holotype">
<collectingCountry id="EA8976709D3E9279FF2EFEE3C90D2915" box="[189,347,349,375]" name="Indonesia" pageId="34" pageNumber="35">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D3E9279FEEDFEE0C9C0291A" box="[382,406,350,376]" pageId="34" pageNumber="35" type="male">♂</specimenCount>
; “
<collectingCountry id="EA8976709D3E9279FE21FEE3CA0E2915" box="[434,600,349,375]" name="Indonesia" pageId="34" pageNumber="35">INDONESIA</collectingCountry>
,
<collectingRegion id="505AF8029D3E9279FDFAFEE3CAB6291A" box="[617,736,349,376]" country="Indonesia" name="Papua Barat" pageId="34" pageNumber="35">Irian Jaya</collectingRegion>
<collectingMunicipality id="7245AC9A9D3E9279FD7EFEE3CB2B291A" box="[749,893,349,376]" pageId="34" pageNumber="35">Nabire area</collectingMunicipality>
,
<locationDeviation id="83932FC29D3E9279FC1EFEE3CC9A291A" box="[909,1228,349,376]" pageId="34" pageNumber="35">road Nabire–Ilaga, km 54</locationDeviation>
,
<geoCoordinate id="F7AA50279D3E9279FB4EFEE3C93F29FF" pageId="34" pageNumber="35">03°29’517”S 135°43’913”E</geoCoordinate>
,
<quantity id="55669B059D3E9279FEEAFE3CC9EA29FF" box="[377,444,386,413]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="34" pageNumber="35" unit="m" value="750.0">
<elevation id="19B3D1D39D3E9279FEEAFE3CC9EA29FF" box="[377,444,386,413]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="34" pageNumber="35" unit="m" value="750.0">750m</elevation>
</quantity>
NN
<date id="E62010209D3E9279FE6FFE3CCA0A29FE" box="[508,604,386,412]" pageId="34" pageNumber="35" value="1998-04">
<collectingDate id="F664E9C89D3E9279FE6FFE3CCA0A29FE" box="[508,604,386,412]" pageId="34" pageNumber="35" value="1998-04">IV.1998</collectingDate>
</date>
, leg.
<collectorName id="3F6B53369D3E9279FD06FE3CCAAB29FE" box="[661,765,386,412]" pageId="34" pageNumber="35">M.Balke</collectorName>
// Sammlung NATURKUNDEMUSEUM ERFURT”;
<collectionCode id="F48FAE259D3E9279FF2EFE19C95429A3" box="[189,258,423,449]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="34" pageNumber="35">NME</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3E9278FF2EFE57C9FD2933" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
<paragraph id="922136E09D3E9279FF2EFE57C9182A61" blockId="34.[189,334,489,515]" box="[189,334,489,515]" pageId="34" pageNumber="35">
<heading id="C969818C9D3E9279FF2EFE57C9182A61" bold="true" box="[189,334,489,515]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="A0EAEAF29D3E9279FF2EFE57C9182A61" bold="true" box="[189,334,489,515]" pageId="34" pageNumber="35">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3E9279FF2EFDA4C9212A1F" blockId="34.[189,1399,537,637]" pageId="34" pageNumber="35">
MEASUREMENTS. Male
<typeStatus id="4D2588429D3E9279FE52FDA7CA7D2A51" box="[449,555,537,563]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
, total body length
<quantity id="55669B059D3E9279FC9AFDA7CB202A56" box="[777,886,537,564]" metricMagnitude="-3" metricUnit="m" metricValue="8.15" pageId="34" pageNumber="35" unit="mm" value="8.15">8.15 mm</quantity>
. Head
<quantity id="55669B059D3E9279FC5AFDA7CC652A56" box="[969,1075,537,564]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="34" pageNumber="35" unit="mm" value="1.75">1.75 mm</quantity>
long, across eyes 2.00 mm wide. Pronotum
<quantity id="55669B059D3E9279FEEDFD80C9B02A3B" box="[382,486,574,601]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="34" pageNumber="35" unit="mm" value="1.9">1.90 mm</quantity>
long, maximum width
<quantity id="55669B059D3E9279FD66FD80CB092A3B" box="[757,863,574,601]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="34" pageNumber="35" unit="mm" value="3.2">3.20 mm</quantity>
. Elytral length
<quantity id="55669B059D3E9279FB83FD80CC2D2A3B" box="[1040,1147,574,601]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="34" pageNumber="35" unit="mm" value="4.5">4.50 mm</quantity>
, maximum combined width
<quantity id="55669B059D3E9279FE94FDDDC9252A1F" box="[263,371,611,637]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="34" pageNumber="35" unit="mm" value="3.6">3.60 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D3E9279FF2EFD18CCC02F8B" blockId="34.[189,1399,678,2025]" pageId="34" pageNumber="35">
Dorsum and venter uniformly black, edges of labroclypeus, legs and antennae dark brown. Head transverse, glossy dorsally and ventrally, inconspicuously convex on frons. Male labroclypeus (
<figureCitation id="0AA52A659D3E9279FA98FD74CD382A87" box="[1291,1390,714,741]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="34" pageNumber="35">Fig. 108</figureCitation>
) broadly V-shaped emarginate on anterior margin, its lateral margins sinuous in both dorsal and lateral views. Anterolateral angles of labroclypeus almost right-angular, protruding anteriad. Canthus sinuous in dorsal view. Head punctures circular to irregularly ovoid, very deep and dense, variably large. Intervening spaces glossy, much smaller than punctures except on labroclypeus. At least labroclypeus and areas around either compound eye covered with microscopical velvety pubescence. Head setae minute, rise from anterior margin of each puncture, in most cases not surpassing length of corresponding punctures. Pronotum strongly transverse, glossy dorsally and laterally, sinuous on anterior margin, anterolateral angles and mesal part almost equally slightly protruding anteriad. Basal margin of pronotum slightly sinuous. Irregular shallow dorsal impressions present on pronotum: small circular one on either side around centre of lateral margin, longitudinal one delimiting lateral third of pronotal disc from central part, and single median longitudinal one (
<figureCitation id="0AA52A659D3E9279FC9AFBE0CB342C1A" box="[777,866,1118,1144]" captionStart="Figs 28–30" captionStartId="31.[189,241,1928,1954]" captionTargetBox="[347,1240,269,1901]" captionTargetId="figure-52@31.[345,1242,265,1911]" captionTargetPageId="31" captionText="Figs 28–30. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 28. M. boessnecki sp. nov., holotype, ♂ (SNSD). 29. M. brocki sp. nov., holotype, ♂ (BMNH). 30. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122136" httpUri="https://zenodo.org/record/4122136/files/figure.png" pageId="34" pageNumber="35">Fig. 30</figureCitation>
). Lateral margin of pronotum in dorsal view broadly rounded, stronger constricted postmedially towards base than anteriad, delicately crenulate all along (
<figureCitation id="0AA52A659D3E9279FE9CFB19C92E2CA3" box="[271,376,1191,1217]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="34" pageNumber="35">Fig. 208</figureCitation>
). Inconspicuous appressed scale-like seta present between every two crenulae. Lateral margin of pronotum sinuous in lateral view. Hypomeron slightly emarginate and long setose on anterior margin which is flange-like protruding. Antennal pocket deep. Median anterior process of prosternum extraordinarily long brushy setose, flange-like raised. Pronotal punctures irregularly ovoid, very deep, dense, mainly positioned in sinuous rows. Punctures generally smaller and more regularly circular in areas of dorsal impressions. Intervening spaces glossy, in part wrinkled, smaller than punctures. Lateral margins, antero- and posterolateral angles and dorsal impressions covered with delicate microscopical velvety pubescence. Pronotal setae similar to those on head, rarely surpassing midlength of corresponding punctures. Scutellar shield covered with microscopical velvety pubescence, narrowly rounded apically. Elytra cylindrical, subopaque, maximum width in median third. Large obtuse hump on each elytron near apex. Elytral punctures irregularly ovoid, very deep and dense, mainly positioned in sinuous rows in various directions, building conspicuous “dactyloscopic” pattern (
<figureCitation id="0AA52A659D3E9279FB9EF985CC2D2E37" box="[1037,1147,1595,1621]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="34" pageNumber="35">Fig. 297</figureCitation>
). Intervening spaces glossy, wrinkled, generally smaller than punctures. Elytral setae minute, rise from anterior margin of each puncture, not surpassing midlength of corresponding puncture. With few longer appressed scalelike clavate setae scattered over elytral disc; punctures bearing those longer setae partly or completely encircled with microscopical velvety pubescence (
<figureCitation id="0AA52A659D3E9279FC8AF973CBD22E8A" box="[793,900,1741,1768]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="34" pageNumber="35">Fig. 297</figureCitation>
). Male abdominal sternites medially with somewhat longer and curved scale-like setae. Male pygidium (
<figureCitation id="0AA52A659D3E9279FC36F94CCC5D2F6F" box="[933,1035,1778,1805]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="34" pageNumber="35">Fig. 463</figureCitation>
) with deep ovoid, rather small punctures. Intervening spaces glossy, generally larger than punctures, covered with microscopical velvety pubescence. Setae of pygidium scale-like, clavate, shorter on anterior, longer on median and posterior parts (surpassing length of corresponding punctures), becoming suberect and longer along distal margin. External margin of male protibia arched, with single acute distal tooth (Fig. 369). Male terminal protibial spur straight, short and pointed. Male lower meso- and metatibial terminal spurs large and strongly curved. Spiculum gastrale as in Fig. 540 Aedeagus as in
<figureCitation id="0AA52A659D3E9279FC7EF870CCC72F8B" box="[1005,1169,1998,2025]" captionStart="Figs 586–600" captionStartId="171.[189,241,1892,1918]" captionTargetBox="[400,1175,268,1865]" captionTargetId="figure-72@171.[391,1196,265,1868]" captionTargetPageId="171" captionText="Figs 586–600. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 586– 588. M. bintang sp. nov., holotype, ♂ (NME). 589–591. M. boessnecki sp. nov., holotype, ♂ (SNSD). 592–594. M. caperatus sp. nov., holotype, ♂ (NME). 595–597. M. ciliatus sp. nov., holotype, ♂ (IECA). 598–600. M. crypticus sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122234" httpUri="https://zenodo.org/record/4122234/files/figure.png" pageId="34" pageNumber="35">Figs 592–594</figureCitation>
.
</paragraph>
<paragraph id="922136E09D3F9278FF2EFEB6C9E62940" blockId="35.[189,432,264,290]" box="[189,432,264,290]" pageId="35" pageNumber="36">
<heading id="C969818C9D3F9278FF2EFEB6C9E62940" bold="true" box="[189,432,264,290]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="A0EAEAF29D3F9278FF2EFEB6C9E62940" bold="true" box="[189,432,264,290]" pageId="35" pageNumber="36">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3F9278FF2EFE89C9FD2933" blockId="35.[189,427,311,337]" box="[189,427,311,337]" pageId="35" pageNumber="36">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3F9278FF2EFEC4CB6329A0" box="[189,821,378,451]" pageId="35" pageNumber="36" type="biology_ecology">
<paragraph id="922136E09D3F9278FF2EFEC4C97629F6" blockId="35.[189,288,378,404]" box="[189,288,378,404]" pageId="35" pageNumber="36">
<heading id="C969818C9D3F9278FF2EFEC4C97629F6" bold="true" box="[189,288,378,404]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="A0EAEAF29D3F9278FF2EFEC4C97629F6" bold="true" box="[189,288,378,404]" pageId="35" pageNumber="36">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3F9278FF2EFE17CB6329A0" blockId="35.[189,821,424,451]" box="[189,821,424,451]" pageId="35" pageNumber="36">
Occurs in lowland rainforests at about
<quantity id="55669B059D3F9278FD17FE16CA9829A1" box="[644,718,424,451]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="35" pageNumber="36" unit="m" value="750.0">750 m</quantity>
altitude.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3F9278FF2EFE54CC582A51" box="[189,1038,490,564]" pageId="35" pageNumber="36" type="distribution">
<paragraph id="922136E09D3F9278FF2EFE54C9002A66" blockId="35.[189,342,490,516]" box="[189,342,490,516]" pageId="35" pageNumber="36">
<heading id="C969818C9D3F9278FF2EFE54C9002A66" bold="true" box="[189,342,490,516]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="A0EAEAF29D3F9278FF2EFE54C9002A66" bold="true" box="[189,342,490,516]" pageId="35" pageNumber="36">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3F9278FF2EFDA7CC582A51" blockId="35.[189,1038,537,564]" box="[189,1038,537,564]" pageId="35" pageNumber="36">
Hitherto only known from the Bird’s Neck isthmus area of New
<collectingCountry id="EA8976709D3F9278FC21FDA7CC5D2A51" box="[946,1035,537,563]" name="Guinea-Bissau" pageId="35" pageNumber="36">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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