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<document id="5A50C2C1EECC9FBCC72DDF0566027C38" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D39927DFDFBFABFCB962D8C" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius crypticus Telnov 2020, sp. nov." docType="treatment" docUuid="B63FE428-02B0-42E0-A347-510931841CAB" docUuidSource="ZooBank" docVersion="10" lastPageNumber="39" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallaces Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="38" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="46FF346B2C5A36D707BB2ABEE10CAC65">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallaces Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="FFCE6FA1FE0A922CC6145C746C1798FC">Telnov, Dmitry</mods:namePart>
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<mods:title id="769AF791B57E95B1B9FFB5D0555C07A6">European Journal of Taxonomy</mods:title>
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<mods:part id="32326E4847B1AFBFB6490590B44DC250">
<mods:date id="2D3F9A1330CE5569B34BD3AB4E87E009">2020</mods:date>
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<mods:number id="5B5EB7EDE3EEEB3CBC640F482F3CAF5E">2020-10-19</mods:number>
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<treatment id="1A3787F69D39927DFDFBFABFCB962D8C" ID-DOI="http://doi.org/10.5281/zenodo.4344349" ID-GBIF-Taxon="168648147" ID-Zenodo-Dep="4344349" LSID="urn:lsid:zoobank.org:act:B63FE428-02B0-42E0-A347-510931841CAB" httpUri="http://treatment.plazi.org/id/1A3787F69D39927DFDFBFABFCB962D8C" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<subSubSection id="DA84656B9D39927EFDFBFABFCB9A2D7E" box="[616,972,1281,1308]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="922136E09D39927EFDFBFABFCB9A2D7E" blockId="37.[372,1215,1281,1378]" box="[616,972,1281,1308]" pageId="37" pageNumber="38">
<heading id="C969818C9D39927EFDFBFABFCB9A2D7E" box="[616,972,1281,1308]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="38" reason="2">
<taxonomicName id="559E4D639D39927EFDFBFABFCB3C2D79" ID-CoL="7X3WK" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[616,874,1281,1308]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="crypticus" status="sp. nov.">
<emphasis id="A0EAEAF29D39927EFDFBFABFCB3C2D79" bold="true" box="[616,874,1281,1308]" italics="true" pageId="37" pageNumber="38">Maechidius crypticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D39927EFCE2FABCCB9A2D7E" box="[881,972,1282,1308]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D39927EFEE7FA9ACC3F2D00" pageId="37" pageNumber="38" type="description">
<paragraph id="922136E09D39927EFEE7FA9ACCE92D5D" blockId="37.[372,1215,1281,1378]" box="[372,1215,1316,1343]" pageId="37" pageNumber="38">
<uri id="E60F3AE29D39927EFEE7FA9ACCE92D5D" box="[372,1215,1316,1343]" pageId="37" pageNumber="38">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D39927EFD00FA9ACCE92D5D" box="[659,1215,1316,1343]" pageId="37" pageNumber="38">B63FE428-02B0-42E0-A347-510931841CAB</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D39927EFE59FAF9CC3F2D00" blockId="37.[372,1215,1281,1378]" box="[458,1129,1351,1378]" pageId="37" pageNumber="38">
<figureCitation id="0AA52A659D39927EFE59FAF9CA722D03" box="[458,548,1351,1377]" captionStart="Figs 3134" captionStartId="36.[189,241,1928,1954]" captionTargetBox="[337,1255,261,1900]" captionTargetId="figure-64@36.[325,1262,257,1921]" captionTargetPageId="36" captionText="Figs 3134. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 31. M. ciliatus sp. nov., holotype, ♂ (IECA). 32. M. crypticus sp. nov., paratype, ♂ (BMNH). 33. M. dani sp. nov., holotype, ♂ (IECA). 34. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122138" httpUri="https://zenodo.org/record/4122138/files/figure.png" pageId="37" pageNumber="38">Figs 32</figureCitation>
,
<figureCitation id="0AA52A659D39927EFDA3FAF9CAC12D03" box="[560,663,1351,1377]" captionStart="Figs 109120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="37" pageNumber="38">112113</figureCitation>
,
<figureCitation id="0AA52A659D39927EFD30FAF9CA842D03" box="[675,722,1351,1377]" captionStart="Figs 193207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193207. Papuan and Wallacean Maechidius Macleay, 1819, head (193196) and pronotum (197 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="37" pageNumber="38">207</figureCitation>
,
<figureCitation id="0AA52A659D39927EFD4DFAF9CB5B2D03" box="[734,781,1351,1377]" captionStart="Figs 286301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="37" pageNumber="38">299</figureCitation>
, 371, 465466, 541, 598600
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D39927EFF2EFA37CCBA2D97" pageId="37" pageNumber="38" type="etymology">
<paragraph id="922136E09D39927EFF2EFA37C9122DC1" blockId="37.[189,324,1417,1443]" box="[189,324,1417,1443]" pageId="37" pageNumber="38">
<heading id="C969818C9D39927EFF2EFA37C9122DC1" bold="true" box="[189,324,1417,1443]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="A0EAEAF29D39927EFF2EFA37C9122DC1" bold="true" box="[189,324,1417,1443]" pageId="37" pageNumber="38">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D39927EFF2EFA09CCBA2D97" blockId="37.[189,1398,1463,1525]" pageId="37" pageNumber="38">The name of this species is derived from the Greek “κρυπτός” (cryptic), referring to its strong external similarity to several Papuan congeners (see description for the list of these similar species).</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D39927EFF2EF9A5CC142F53" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="922136E09D39927EFF2EF9A5C93A2E57" blockId="37.[189,364,1563,1589]" box="[189,364,1563,1589]" pageId="37" pageNumber="38">
<heading id="C969818C9D39927EFF2EF9A5C93A2E57" bold="true" box="[189,364,1563,1589]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="A0EAEAF29D39927EFF2EF9A5C93A2E57" bold="true" box="[189,364,1563,1589]" pageId="37" pageNumber="38">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D39927EFF79F9F7C90A2EAF" ID-GBIF-Occurrence="2898878412" collectingDate="1965-01-04" collectingDateMax="1965-01-17" collectingDateMin="1965-01-04" collectionCode="NEW, BMNH" collectorName="M. E. Bacchus." country="Papua New Guinea" determinerName="G Frey." elevation="1219" location="Morobe Dist." municipality="Herzog Mts." pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="922136E09D39927EFF79F9F7C90A2E01" blockId="37.[189,1399,1609,1741]" box="[234,348,1609,1635]" pageId="37" pageNumber="38">
<emphasis id="A0EAEAF29D39927EFF79F9F7C90A2E01" bold="true" box="[234,348,1609,1635]" pageId="37" pageNumber="38">
<typeStatus id="4D2588429D39927EFF79F9F7C90A2E01" box="[234,348,1609,1635]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D39927EFF2EF9D2C90A2EAF" blockId="37.[189,1399,1609,1741]" pageId="37" pageNumber="38">
<collectingCountry id="EA8976709D39927EFF2EF9D2C98D2EE4" box="[189,475,1644,1671]" name="Papua New Guinea" pageId="37" pageNumber="38">PAPUA NEW GUINEA</collectingCountry>
<specimenCount id="8498FD699D39927EFE65F9D3CA582EE5" box="[502,526,1645,1671]" pageId="37" pageNumber="38" type="male"></specimenCount>
; “Stn. No. 137. [p] //
<collectionCode id="F48FAE259D39927EFC99F9D3CB1A2EE5" box="[778,844,1645,1671]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="37" pageNumber="38">NEW</collectionCode>
<collectingCountry id="EA8976709D39927EFCC7F9D2CB9F2EE4" box="[852,969,1644,1670]" name="Guinea-Bissau" pageId="37" pageNumber="38">GUINEA</collectingCountry>
:
<location id="9741603B9D39927EFC46F9D2CC202EE4" LSID="urn:lsid:plazi:treatment:1A3787F69D39927DFDFBFABFCB962D8C:9741603B9D39927EFC46F9D2CC202EE4" box="[981,1142,1644,1670]" country="Papua New Guinea" municipality="Herzog Mts." name="Morobe Dist." pageId="37" pageNumber="38">Morobe Dist.</location>
,
<collectingMunicipality id="7245AC9A9D39927EFB10F9D3CD4F2EE5" box="[1155,1305,1645,1671]" pageId="37" pageNumber="38">Herzog Mts.</collectingMunicipality>
, Vagau, C.
<quantity id="55669B059D39927EFF47F931C96E2ECB" box="[212,312,1679,1705]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="37" pageNumber="38" unit="ft" value="4000.0">
<elevation id="19B3D1D39D39927EFF47F931C96E2ECB" box="[212,312,1679,1705]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="37" pageNumber="38" unit="ft" value="4000.0">4,000 ft.</elevation>
</quantity>
<date id="E62010209D39927EFEAFF931C99E2EC8" box="[316,456,1679,1706]" pageId="37" pageNumber="38" value="1965-01-04" valueMax="1965-01-17" valueMin="1965-01-04">
<collectingDate id="F664E9C89D39927EFEAFF931C99E2EC8" box="[316,456,1679,1706]" pageId="37" pageNumber="38" value="1965-01-04" valueMax="1965-01-17" valueMin="1965-01-04">4-17.I.1965</collectingDate>
</date>
. [p]//
<collectorName id="3F6B53369D39927EFD99F92ECAE32ECB" box="[522,693,1679,1706]" pageId="37" pageNumber="38">M.E. Bacchus.</collectorName>
B.M. 1965-120 [p] //
<taxonomicName id="559E4D639D39927EFC3BF92ECC3A2ECB" authority="Hell." authorityName="Heller" authorityYear="1910" box="[936,1132,1679,1705]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pauxillus">
<emphasis id="A0EAEAF29D39927EFC3BF92ECC782ECB" box="[936,1070,1679,1705]" italics="true" pageId="37" pageNumber="38">P. pauxillus</emphasis>
Hell.
</taxonomicName>
[h] det.
<determinerName id="67FDDA249D39927EFB52F931CD4E2ECB" box="[1217,1304,1679,1705]" pageId="37" pageNumber="38">G Frey.</determinerName>
1967/68 [p]”;
<collectionCode id="F48FAE259D39927EFF6FF90DC90E2EAF" box="[252,344,1715,1741]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="37" pageNumber="38">BMNH</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="922136E09D39927EFF79F94DCA452F6C" blockId="37.[189,1090,1779,1842]" box="[234,531,1779,1806]" pageId="37" pageNumber="38">
<emphasis id="A0EAEAF29D39927EFF79F94DC9312F6F" bold="true" box="[234,359,1779,1805]" pageId="37" pageNumber="38">
<typeStatus id="4D2588429D39927EFF79F94DC9312F6F" box="[234,359,1779,1805]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="8498FD699D39927EFEEBF94DCA5D2F6C" box="[376,523,1779,1806]" pageId="37" pageNumber="38" type="generic">7 specimens</specimenCount>
)
</paragraph>
<paragraph id="922136E09D39927EFF2EF8A9CC142F53" blockId="37.[189,1090,1779,1842]" box="[189,1090,1815,1842]" pageId="37" pageNumber="38">
<materialsCitation id="22F63CBD9D39927EFF2EF8A9CC682F53" ID-GBIF-Occurrence="2898878573" box="[189,1086,1815,1842]" collectingDate="1965-01-04" collectingDateMax="1965-01-17" collectingDateMin="1965-01-04" collectionCode="BMNH" collectorName="M. E. Bacchus." country="Papua New Guinea" determinerName="G Frey." elevation="1219" location="PAPUA NEW GUINEA" municipality="Herzog Mts." pageId="37" pageNumber="38" specimenCount="7" specimenCount-female="5" specimenCount-male="2" typeStatus="holotype">
<collectingCountry id="EA8976709D39927EFF2EF8A9C98F2F53" box="[189,473,1815,1841]" name="Papua New Guinea" pageId="37" pageNumber="38">PAPUA NEW GUINEA</collectingCountry>
<specimenCount id="8498FD699D39927EFE61F8A9CA602F53" box="[498,566,1815,1841]" pageId="37" pageNumber="38" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="8498FD699D39927EFDD1F8A9CAD32F53" box="[578,645,1815,1842]" pageId="37" pageNumber="38" type="female">5 ♀♀</specimenCount>
; same labels as for holotype;
<collectionCode id="F48FAE259D39927EFC71F8A9CC682F53" box="[994,1086,1815,1841]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="37" pageNumber="38">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D39927DFF2EF8E6CA0A2D65" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="description">
<paragraph id="922136E09D39927EFF2EF8E6C9182F10" blockId="37.[189,334,1880,1906]" box="[189,334,1880,1906]" pageId="37" pageNumber="38">
<heading id="C969818C9D39927EFF2EF8E6C9182F10" bold="true" box="[189,334,1880,1906]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="A0EAEAF29D39927EFF2EF8E6C9182F10" bold="true" box="[189,334,1880,1906]" pageId="37" pageNumber="38">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D39927EFF2EF836CD202F8B" blockId="37.[189,1399,1927,2025]" pageId="37" pageNumber="38">
MEASUREMENTS. Male
<typeStatus id="4D2588429D39927EFE52F839CA7D2FC3" box="[449,555,1927,1953]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
, total body length
<quantity id="55669B059D39927EFC9AF839CB202FC0" box="[777,886,1927,1954]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="37" pageNumber="38" unit="mm" value="6.5">6.50 mm</quantity>
. Head
<quantity id="55669B059D39927EFC5AF839CC652FC0" box="[969,1075,1927,1954]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="37" pageNumber="38" unit="mm" value="1.2">1.20 mm</quantity>
long, across eyes
<quantity id="55669B059D39927EFA9EF839CD212FC0" box="[1293,1399,1927,1954]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="38" unit="mm" value="1.5">1.50 mm</quantity>
wide. Pronotum
<quantity id="55669B059D39927EFEEDF815C9B02FA7" box="[382,486,1963,1989]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="37" pageNumber="38" unit="mm" value="1.3">1.30 mm</quantity>
long, maximum width
<quantity id="55669B059D39927EFD66F815CB092FA7" box="[757,863,1963,1989]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="37" pageNumber="38" unit="mm" value="2.1">2.10 mm</quantity>
. Elytral length 4.00 mm, maximum combined width
<quantity id="55669B059D39927EFE94F870C9252F8B" box="[263,371,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="37" pageNumber="38" unit="mm" value="2.8">2.80 mm</quantity>
. Selected female
<typeStatus id="4D2588429D39927EFDD3F871CAFE2F8B" box="[576,680,1999,2025]" pageId="37" pageNumber="38" type="paratype">paratype</typeStatus>
, total body length
<quantity id="55669B059D39927EFC11F870CBB82F8B" box="[898,1006,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="7.85" pageId="37" pageNumber="38" unit="mm" value="7.85">7.85 mm</quantity>
. Head
<quantity id="55669B059D39927EFBACF870CCFE2F8B" box="[1087,1192,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="37" pageNumber="38" unit="mm" value="1.35">1.35 mm</quantity>
long, across eyes
</paragraph>
<paragraph id="922136E09D3A927DFF2EFEB7C9A4292A" blockId="38.[189,1398,265,328]" pageId="38" pageNumber="39">
<quantity id="55669B059D3A927DFF2EFEB7C9702941" box="[189,294,265,291]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="38" pageNumber="39" unit="mm" value="1.6">1.60 mm</quantity>
wide. Pronotum
<quantity id="55669B059D3A927DFE7CFEB7CA0E2941" box="[495,600,265,292]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="39" unit="mm" value="1.5">1.50 mm</quantity>
long, maximum width
<quantity id="55669B059D3A927DFCF9FEB7CB802941" box="[874,982,265,291]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="38" pageNumber="39" unit="mm" value="2.3">2.30 mm</quantity>
. Elytral length 5.00 mm, maximum combined width
<quantity id="55669B059D3A927DFE11FE90C9B8292A" box="[386,494,302,328]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="38" pageNumber="39" unit="mm" value="3.7">3.70 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D3A927DFF2EFECFCA402C0D" blockId="38.[189,1399,369,1135]" pageId="38" pageNumber="39">
With general features of
<taxonomicName id="559E4D639D3A927DFE63FECCCA3029E9" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[496,614,369,395]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="lapsus" status="sp. nov.">
<emphasis id="A0EAEAF29D3A927DFE63FECCCA3029E9" box="[496,614,369,395]" italics="true" pageId="38" pageNumber="39">M. lapsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D3A927DFDE2FECCCA8429EE" box="[625,722,370,396]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="559E4D639D3A927DFD71FECCCBF029E9" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[738,934,369,395]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="owenstanleyi" status="sp. nov.">
<emphasis id="A0EAEAF29D3A927DFD71FECCCBF029E9" box="[738,934,369,395]" italics="true" pageId="38" pageNumber="39">M. owenstanleyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D3A927DFC21FECCCC4729EE" box="[946,1041,370,396]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
(see descriptions below) and
<taxonomicName id="559E4D639D3A927DFF2EFE29C90E29D2" authorityName="Heller" authorityYear="1910" box="[189,344,406,432]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pauxillus">
<emphasis id="A0EAEAF29D3A927DFF2EFE29C90E29D2" box="[189,344,406,432]" italics="true" pageId="38" pageNumber="39">M. pauxillus</emphasis>
</taxonomicName>
comb. rest. Frons strongly humped in lateral view. Male and female labroclypeus broadly emarginate on anterior margin. Anterolateral angles of labroclypeus strongly protruding, acute (
<figureCitation id="0AA52A659D3A927DFF56FE5EC9392998" box="[197,367,480,506]" captionStart="Figs 109120" captionStartId="138.[189,241,1750,1776]" captionTargetBox="[365,1214,275,1722]" captionTargetId="figure-15@138.[365,1222,265,1730]" captionTargetPageId="138" captionText="Figs 109120. Papuan andWallacean MaechidiusMacleay, 1819,head, dorsal (109119) and frontal (120) views. 109. M. brocki sp. nov., holotype, ♂ (BMNH). 110. Same, paratype, ♀ (BMNH). 111. M. ciliatus sp. nov., holotype, ♂ (IECA). 112. M. crypticus sp. nov., paratype, ♂ (BMNH). 113. Same, paratype, ♀ (BMNH). 114. M. dani sp. nov., holotype, ♂ (IECA). 115. M. deltouri sp. nov., holotype, ♂ (BMNH). 116. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 117. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 118. M. esau Heller, 1914, lectotype, ♂ (SNSD). 119. Same, ♂ (lectotype of M. setosus Moser, 1920) (ZMHB). 120. M. gressitti Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122175" httpUri="https://zenodo.org/record/4122175/files/figure.png" pageId="38" pageNumber="39">Figs 112113</figureCitation>
). Lateral margins of labroclypeus in both sexes slightly sinuous in both dorsal and lateral views. Antennae 9-segmented, club 3-lamellate. Dorsal surface in part covered with microscopic velvety pubescence. Head and pronotum with moderately long, suberect elongate scale-like setae; each seta rises from anterior margin of deep irregularly oblong puncture. Lateral margin of pronotum obtuse angulate medially, deeply emarginate postmedially (
<figureCitation id="0AA52A659D3A927DFCAEFDCACBFD2AEC" box="[829,939,628,654]" captionStart="Figs 193207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193207. Papuan and Wallacean Maechidius Macleay, 1819, head (193196) and pronotum (197 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="38" pageNumber="39">Fig. 207</figureCitation>
). Scale-like setae longer and broader on sides and anterior part of pronotal disc than along median part, also with numerous much smaller setae not surpassing length of corresponding punctures. Hypomeron flange-like produced. Antennal pocket deep. With or without tracks of two longitudinal carinae on each elytron, carinae incomplete, with interruptions. Sutural carinae incomplete and interrupted. Elytral punctures irregular in shape, deep, somewhat irregularly clustered (
<figureCitation id="0AA52A659D3A927DFD14FC93CAB82B25" box="[647,750,813,839]" captionStart="Figs 286301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="38" pageNumber="39">Fig. 299</figureCitation>
) and generally smaller than those on pronotum. Setae between puncture clusters longer, scale-like, stronger erect, arranged in several irregular longitudinal rows. Setae in puncture clusters minute, not surpassing length of corresponding punctures. Male and female pygidium deeply oblongo-punctate, with more or less dense scale-like elongate setae which point obliquely to middle (in main part) (
<figureCitation id="0AA52A659D3A927DFDCAFC7ECB572BB9" box="[601,769,960,987]" captionStart="Figs 451468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="38" pageNumber="39">Figs 465466</figureCitation>
). Intervening spaces of pygidium in part covered with microscopical velvety pubescence. Protibia with inconspicuous longitudinal dorsal carina, in male with single inconspicuous distal tooth, in female with two acute teeth (Fig. 371). Metatibial terminal spurs in both sexes distinctly shorter than length of basal metatarsomere. Spiculum gastrale as in Fig. 541. Male aedeagus as in
<figureCitation id="0AA52A659D3A927DFEFDFBEACA442C0D" box="[366,530,1108,1135]" captionStart="Figs 586600" captionStartId="171.[189,241,1892,1918]" captionTargetBox="[400,1175,268,1865]" captionTargetId="figure-72@171.[391,1196,265,1868]" captionTargetPageId="171" captionText="Figs 586600. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 586 588. M. bintang sp. nov., holotype, ♂ (NME). 589591. M. boessnecki sp. nov., holotype, ♂ (SNSD). 592594. M. caperatus sp. nov., holotype, ♂ (NME). 595597. M. ciliatus sp. nov., holotype, ♂ (IECA). 598600. M. crypticus sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122234" httpUri="https://zenodo.org/record/4122234/files/figure.png" pageId="38" pageNumber="39">Figs 598600</figureCitation>
.
</paragraph>
<paragraph id="922136E09D3A927DFF2EFB29C9E62CD3" blockId="38.[189,432,1175,1201]" box="[189,432,1175,1201]" pageId="38" pageNumber="39">
<heading id="C969818C9D3A927DFF2EFB29C9E62CD3" bold="true" box="[189,432,1175,1201]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="A0EAEAF29D3A927DFF2EFB29C9E62CD3" bold="true" box="[189,432,1175,1201]" pageId="38" pageNumber="39">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3A927DFF2EFB76CA0A2D65" blockId="38.[189,1399,1224,1287]" pageId="38" pageNumber="39">Female is generally larger, with shorter lamellae of antennal club, straight and pointed protibial terminal spur and distally bidentate protibia.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3A927DFF2EFA8ECBC22D18" box="[189,916,1328,1403]" pageId="38" pageNumber="39" type="biology_ecology">
<paragraph id="922136E09D3A927DFF2EFA8EC9762D28" blockId="38.[189,288,1328,1354]" box="[189,288,1328,1354]" pageId="38" pageNumber="39">
<heading id="C969818C9D3A927DFF2EFA8EC9762D28" bold="true" box="[189,288,1328,1354]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="A0EAEAF29D3A927DFF2EFA8EC9762D28" bold="true" box="[189,288,1328,1354]" pageId="38" pageNumber="39">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3A927DFF2EFADECBC22D18" blockId="38.[189,916,1376,1403]" box="[189,916,1376,1403]" pageId="38" pageNumber="39">
Occurs in lower montane rainforests at about
<quantity id="55669B059D3A927DFD47FADECB7B2D19" box="[724,813,1376,1403]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="38" pageNumber="39" unit="m" value="1220.0">1220 m</quantity>
altitude.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D3A927DFF2EFA1DCB962D8C" box="[189,960,1443,1518]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="922136E09D3A927DFF2EFA1DC9002DDF" blockId="38.[189,342,1443,1469]" box="[189,342,1443,1469]" pageId="38" pageNumber="39">
<heading id="C969818C9D3A927DFF2EFA1DC9002DDF" bold="true" box="[189,342,1443,1469]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="A0EAEAF29D3A927DFF2EFA1DC9002DDF" bold="true" box="[189,342,1443,1469]" pageId="38" pageNumber="39">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D3A927DFF2EFA6ACB962D8C" blockId="38.[189,960,1492,1518]" box="[189,960,1492,1518]" pageId="38" pageNumber="39">
Hitherto known from the Herzog Mountains of East New
<collectingCountry id="EA8976709D3A927DFCF7FA6ACBEB2D8C" box="[868,957,1492,1518]" name="Guinea-Bissau" pageId="38" pageNumber="39">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>