199 lines
20 KiB
XML
199 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. & Tishechkin, Alexey K." docDate="2013" docId="043142ADDBA3A8CEB8F955C8F6F4840D" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus ibiscus Caterino & Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="156" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="153" updateTime="1668155382655" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Caterino, Michael S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>271</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>401</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152040976" LSID="urn:lsid:zoobank.org:act:BF51784E-E209-4038-B318-F88FEC9130B0" httpUri="http://treatment.plazi.org/id/043142ADDBA3A8CEB8F955C8F6F4840D" lastPageId="155" lastPageNumber="156" pageId="152" pageNumber="153">
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<subSubSection pageId="152" pageNumber="153" type="nomenclature">
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<paragraph pageId="152" pageNumber="153">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:BF51784E-E209-4038-B318-F88FEC9130B0" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus ibiscus" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="species" species="ibiscus">Operclipygus ibiscus</taxonomicName>
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<taxonomicNameLabel pageId="152" pageNumber="153">sp. n.</taxonomicNameLabel>
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Figs 40F, K41
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<normalizedToken originalValue="A–CMap">A-CMap</normalizedToken>
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15
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</paragraph>
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</subSubSection>
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<subSubSection pageId="152" pageNumber="153" type="type locality">
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<paragraph pageId="152" pageNumber="153">Type locality.</paragraph>
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<paragraph pageId="152" pageNumber="153">
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PANAMA:
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<normalizedToken originalValue="Colón">Colon</normalizedToken>
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: San Lorenzo Forest [
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="9.283334">9°17'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-79.96667">79°58'W</geoCoordinate>
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].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="152" pageNumber="153" type="type material">
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<paragraph pageId="152" pageNumber="153">Type material.</paragraph>
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<paragraph pageId="152" pageNumber="153">
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Holotype male: "PANAMA:
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<normalizedToken originalValue="Colón">Colon</normalizedToken>
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Pr., San Lorenzo Forest.
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="9.283334">9°17'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-79.96667">79°58'W</geoCoordinate>
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. Flight Intercept FIT-C3-12 12-13 May 2004 A. Tishechkin. IBISCA
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<normalizedToken originalValue="‘04”">'04"</normalizedToken>
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/ "LSAM 0111298" (FMNH). Paratypes: (103): same data as type, except as noted: 1: 23-24.ix.2003 (LSAM), 3: 27-28.ix.2003 (LSAM, FMNH), 2: 30-1.x.2003 (LSAM), 1: 1-3.x.2003 (LSAM), 4: 4-6.x.2003 (LSAM, MSCC, AKTC), 2: 8-9.x.2003 (LSAM, CHND), 1: 9-11.x.2003 (LSAM), 3: 13-14.x.2003 (LSAM, CMNC), 4: 14-17.x.2003 (LSAM), 3: 17-18.x.2003 (LSAM), 3: 17.x.2003, R. Didham, L. Fagan (LSAM), 1: 18-21.x.2003 (LSAM), 6: 19-21.x.2003 (LSAM), 4: 21-22.x.2003 (LSAM), 1: 24.x.2003, in rotting fruit of
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<taxonomicName genus="Bombacaceae" lsidName="Bombacaceae" pageId="152" pageNumber="153" rank="genus">Bombacaceae</taxonomicName>
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tree, L. Cizek (LSAM), 1: 25-26.x.2003 (LSAM), 1: 28-30.x.2003 (LSAM), 4: 29.x.2003, R. Didham, L. Fagan (GBFM, LSAM), 3: 11-12.v.2004 (LSAM), 2: 12-13.v.2004 (LSAM), 2: 14-15.v.2004 (LSAM), 6: 15-17.v.2004 (GBFM, LSAM), 1: 15-25.v.2004, R. Didham (LSAM), 6: 17-18.v.2004 (LSAM), 4: 18-19.v.2004 (LSAM), 3: 19-20.v.2004 (LSAM), 6: 20-21.v.2004 (LSAM), 9: 21-24.v.2004 (LSAM), 2: 24-25.v.2004 (LSAM), 1: 25.
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<normalizedToken originalValue="v–">v-</normalizedToken>
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5.vi.2004, R. Didham (LSAM), 8: 25-26.v.2004 (GBFM, LSAM), 1: 26-28.v.2004 (LSAM), 3: 26-29.v.2004 (LSAM), 1: 4-17.vi.2004, M. Rapp (LSAM).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="153" lastPageNumber="154" pageId="152" pageNumber="153" type="other material">
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<paragraph pageId="152" pageNumber="153">Other material.</paragraph>
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<paragraph lastPageId="153" lastPageNumber="154" pageId="152" pageNumber="153">
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PANAMA:
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<normalizedToken originalValue="Colón">Colon</normalizedToken>
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: 1: Parque. Nac. Soberania, Pipeline Rd. Km 6.1,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="9.116667">09°07'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-79.75">79°45'W</geoCoordinate>
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, 40m, 7-21.vi.1995, FIT, J. Ashe, R. Brooks (SEMC); Bocas del Toro: 1: Almirante, 23.iii.1959, decayed fibrous palm trunk, H.S. Dybas (FMNH); 1: Almirante, 1.iv.1959, forest litter, H.S. Dybas (FMNH);
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<normalizedToken originalValue="Chiriquí">Chiriqui</normalizedToken>
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: 2: 4km N Sta. Clara Hartmann's Finca, 1500m, 30.
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<normalizedToken originalValue="vi–">vi-</normalizedToken>
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13.vii.1982, FIT, B.D. Gill (BDGC);
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<normalizedToken originalValue="Coclé">Cocle</normalizedToken>
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: 3: 7.2km NE. El Cope,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="8.616667">08°37'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-80.583336">80°35'W</geoCoordinate>
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, 730m, 20.
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<normalizedToken originalValue="v–">v-</normalizedToken>
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7.vi.1995, FIT, J. Ashe, R. Brooks (SEMC);
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<normalizedToken originalValue="Panamá">Panama</normalizedToken>
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: 1: Barro Colorado Isl.,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="9.183333">09°11'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-79.85">79°51'W</geoCoordinate>
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, 8.vii.1994, FIT, D. Banks (SEMC), 1: 1.viii.1994, FIT, D. Banks (SEMC), 1: 24.viii.1994, FIT, D. Banks (SEMC), 1: 16.ii.1976,
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Pseudobombax" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudobombax" order="Malvales" pageId="152" pageNumber="153" phylum="Tracheophyta" rank="genus">Pseudobombax</taxonomicName>
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flower fall, A.F. Newton (FMNH), 1: 1-5.vii.2000, FIT, S. Chatzimanolis (SEMC),1: 22-25.vi.2000, FIT, S. Chatzimanolis (SEMC), 1:
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<normalizedToken originalValue="i–iii">i-iii</normalizedToken>
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.1944 (FMNH); 7: Chepo-Carti Rd., 400m, vi.1982, FIT, B.D. Gill (BDGC, AKTC), 1: vii.1982, FIT, B.D. Gill (CHSM); 8: Cerro Azul, ca., 2000ft, 21.ii.1976, flood debris, A.F. Newton (FMNH); 1: Nusagandi, Ina Igar trail, 18-21.v.1993, FIT/pitfall, primary forest, E.G. Riley (TAMU); 1: Old Plantation Rd. 6.9km S Gamboa,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="9.083333">09°05'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-79.666664">79°40'W</geoCoordinate>
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, 80m, 7-22.vi.1995, FIT, J. Ashe, R.
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<pageBreakToken pageId="153" pageNumber="154" start="start">Brooks</pageBreakToken>
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(SEMC). COSTA RICA: Heredia: 2: La Selva Biol. Stn.,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.433333">10°26'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-84.01667">84°01'W</geoCoordinate>
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, 21.vi.1998, FIT, C.E. Carlton & A.K. Tishechkin (LSAM), 3: 22.vi.1998 (LSAM), 3: 23.vi.1998 (LSAM), 2: 24.vi.1998 (LSAM), 2: 25.vi.1998 (LSAM), 5: 26.vi.1998 (LSAM), 5: 27.vi.1998 (LSAM), 1: 28.vi.1998 (LSAM), 4: 29.vi.1998 (LSAM), 1: 24.ii.1992, FIT, W. Bell (SEMC), 1: 16.vi.2012, OTS Beetle Course, DNA Extract MSC-2307 (SBMNH), 1: 19.iii.1992, FIT, W. Bell (SEMC), 1: 6.vii.2005, FIT, M. Ferro (AKTC), 1: 8.vii.2005, FIT, M. Ferro (AKTC); 1: 10kmW. Puerto Viejo, 170m, 2-5.iii.1991, FIT, H. & A. Howden (CHSM);
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<normalizedToken originalValue="Limón">Limon</normalizedToken>
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: 3: Area Cons. Tortuguero, Sector Cerro Cocori, Fca. de E. Rojas, 150m, ii.1993, E. Rojas (INBIO), 1: v.1991, E. Rojas (INBIO), 6: vi.1991, E. Rojas (INBIO), 1: ix.1991, E. Rojas (INBIO), 2: x.1991, E. Rojas (INBIO), 1: iii.1992, E. Rojas (INBIO), 6: iv.1992, E. Rojas (INBIO), 3: iv.1993, E. Rojas (INBIO), 2: ix.1993, E. Rojas (INBIO), 1: v.1993, E. Rojas (INBIO), 2: xii.1992, E. Rojas (INBIO), 3: 26.vii-2.viii.1992, E. Rojas (INBIO), 1: 9-30.xi.1992, E. Rojas (INBIO); 1: Area Cons. Tortuguero, Rio Sardinas, R.N.F.S. Barra del Colorado, 50m, v.1994, F.V. Araya (INBIO), 1: 10m, 1-14.ii.1994, F.V. Araya (INBIO); 1: Sardinas, Barra del Colorado, 15m, 28.
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<normalizedToken originalValue="i–">i-</normalizedToken>
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12.ii.1995, F. Araya, (INBIO), 2: 26.
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<normalizedToken originalValue="iv–">iv-</normalizedToken>
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3.v.1995, F. Araya, (INBIO), 2: 4-11.i.1995, F. Araya, (INBIO); 2: 3km N. del Puente Rio Suerte, Ruta Puerto Lindo, Cedrales de la Rita, 10m, viii.1996, FIT, E. Rojas (INBIO); 1: Amubri, 70m, 1-22.x.1994, G. Gallardo, (INBIO); Puntarenas: 1: Res. Biol. Carara, Est. Quebrada Bonita, 100m, ii.1995, R. Guzman, (INBIO), 1: 50m, iv.1994, R. Guzman, (INBIO); 1: Rancho Quemado, Peninsula de Osa, 200m, ix.1991, F. Quesada, (INBIO), 1: viii.1991, F. Quesada, (INBIO), 2: ix.1992, F. Quesada, (INBIO), 2: v.1992, F. Quesada y G. Varela, (INBIO), 2: vi.1992, F. Quesada, (INBIO), 1: vii.1992, M. Segura, (INBIO), 1: 12.v.1993, 24.v.1993, A. Gutierrez, (INBIO); 1: Est. Esquinas, Peninsula de Osa, x.1993, M. Segura, (INBIO); 1: Bosque Esquinas, Peninsula de Osa, 200m, vi.1994, M. Segura, (INBIO); 1: Reserva Forestal Golfo Dulce, Est. Agujas, Golfito, Sendero Zamia, 250-350m, 5-7.xi.1999, FIT, A. Azofeifa (INBIO); 1: Est. Biol. Las Cruces, Coto Brus,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="8.783334">8°47'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-82.95">82°57'W</geoCoordinate>
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, 1100m, 22-23.iii.2002, FIT, A.K. Tishechkin (LSAM), 1: 1100m, 2-8.iv.2002, FIT, A.K. Tishechkin (LSAM), 1: 1100m, 31.iii-1.iv.2002, FIT, A.K. Tishechkin (LSAM), 1: 1330m, 28-31.v.2004, FIT, J.S. Ashe, Z. Falin, I. Hinojosa (SEMC), 1: 1330m, 28-30.v.2004, FIT, J.S. Ashe, Z. Falin, I. Hinojosa (SEMC); 1: Est. Biol. Las Cruces, San Vito,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="8.766666">8°46'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-82.96667">82°58'W</geoCoordinate>
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, 4000ft, 19.iii.1973, Berlese, stream bed litter, Virgin forest, J. Wagner, J. Kethley (FMNH), 7: 17.viii-12.ix.1982, FIT, B.D. Gill (AKTC), 2: 1200m, 1-30.vii.1982, B.D. Gill (BDGC); 2: Est. Biol. Las Alturas, Coto Brus,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="8.95">8°57'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-82.86667">82°52'W</geoCoordinate>
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, 1550m, 30.iii-4.iv.2002, FIT, A.K. Tishechkin (LSAM), 1: 25-30.iii.2002, FIT, A. Cline & A. Tishechkin (LSAM); 1: Est. F.N. Aguas Buenas, Rincon, 7km W. Osa Penn., 50m, 21-25.vi.1997, FIT, S. & J. Peck (SEMC); 1: Estrella Valley, Pandora, 1-16.iii.1984, FIT, H. Howden & G. Manley (CHSM).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="155" lastPageNumber="156" pageId="153" pageNumber="154" type="description">
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<paragraph pageId="153" pageNumber="154">Diagnostic description.</paragraph>
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<paragraph lastPageId="155" lastPageNumber="156" pageId="153" pageNumber="154">
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This species is very similar to the preceding, differing in the following characters: length: 1.53-1.78 mm, width: 1.25-1.44 mm; frontal stria complete, rather evenly rounded from sides to front; lateral pronotal complete; pronotum finely punctate, lacking any larger lateral punctures; pronotum depressed, but
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<pageBreakToken pageId="154" pageNumber="155" start="start">lacking</pageBreakToken>
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distinct antescutellar impression; lateral metaventral stria extending obliquely laterad just behind postmesocoxal stria; 1st abdominal ventrite with the two lateral striae fine and very close at base, diverging posteriorly, outer stria frequently abbreviated or
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<pageBreakToken pageId="155" pageNumber="156" start="start">fragmented</pageBreakToken>
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; outer subhumeral stria complete, fine, occasionally fragmented basally, elytral striae 1-3 complete, 4th and 5th striae present in apical third, sutural stria present in apical two-thirds; elytral striae 3-5 weakly expanded at apices; marginal pygidial sulcus deep, well developed, somewhat crenulate; pygidium with fine, dense ground punctation interspersed with sparse, slightly coarser secondary punctures. Male genitalia similar in most respects to those of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fusistrius" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="fusistrius">Operclipygus fusistrius</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus diffluens" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="diffluens">Operclipygus diffluens</taxonomicName>
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(see Figs 40
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<normalizedToken originalValue="B–D">B-D</normalizedToken>
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), but T8 (Fig. 40F) with accessory sclerites present, submarginal desclerotization of T8 extending to near midpoint of segment, and tegmen (Fig. 40K) narrower and more elongate.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="155" pageNumber="156" type="remarks">
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<paragraph pageId="155" pageNumber="156">Remarks.</paragraph>
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<paragraph pageId="155" pageNumber="156">
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Among the species that exhibit a subapical desclerotization along the apical emargination of the male 8th tergite (Fig. 40F),
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus ibiscus" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="ibiscus">Operclipygus ibiscus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus novateutoniae" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="novateutoniae">Operclipygus novateutoniae</taxonomicName>
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are unusual in having a complete outer subhumeral stria. In addition, the absence of coarse punctures on the sides of the pronotum (Fig. 41B), weakly apically widened elytral striae (Fig. 41A), and very closely-set lateral striae of abdominal ventrite 1 will distinguish
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus ibiscus" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="ibiscus">Operclipygus ibiscus</taxonomicName>
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.
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus fusistrius" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="fusistrius">Operclipygus fusistrius</taxonomicName>
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is also very similar, and has a similar distribution, but has more distinctly expanded elytral striae and clearly separated lateral striae on ventrite 1. Because there is slight variation throughout the range of this species in propygidial punctation and degree of elytral stria widening, we restrict the type series to specimens from the type locality.
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</paragraph>
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<caption pageId="155" pageNumber="156">
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<paragraph pageId="155" pageNumber="156">
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Figure 41.
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hospes" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="hospes">Operclipygus hospes</taxonomicName>
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group. A Dorsal habitus of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus ibiscus" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="ibiscus">Operclipygus ibiscus</taxonomicName>
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B Pronotum of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus ibiscus" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="ibiscus">Operclipygus ibiscus</taxonomicName>
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C Pygidia of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus ibiscus" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="ibiscus">Operclipygus ibiscus</taxonomicName>
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D Dorsal habitus of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus novateutoniae" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="novateutoniae">Operclipygus novateutoniae</taxonomicName>
|
||
E Pygidia of
|
||
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus novateutoniae" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="novateutoniae">Operclipygus novateutoniae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="155" pageNumber="156" type="etymology">
|
||
<paragraph pageId="155" pageNumber="156">Etymology.</paragraph>
|
||
<paragraph pageId="155" pageNumber="156">
|
||
This
|
||
<normalizedToken originalValue="species’">species'</normalizedToken>
|
||
name recognizes the IBISCA (Investigating the Biodiversity of Soil and Canopy Arthropods) project, whose 2003-04 Panamanian survey yielded the type specimens of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |