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<document id="759F0DBC04BD282826357EAD0B8DBE43" ID-DOI="10.5852/ejt.2024.948.2637" ID-ISSN="2118-9773" ID-Zenodo-Dep="13511901" ID-ZooBank="9985FD07-5280-41D9-B982-B175085AE5F8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1724950631476" checkinUser="plazi" docAuthor="Gautam, R. K., Uniyal, V. P. &amp; Wood, Thomas J." docDate="2024" docId="03DE87B4FFA7FF900E936B51FC04F88F" docLanguage="en" docName="EJT.2024.948.1-59.pdf" docOrigin="European Journal of Taxonomy 948" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2637/12143" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Andrena (Melandrena) kedarnatha Wood &amp; Gautam 2024, sp. nov." docType="treatment" docUuid="19CF4125-2926-4B48-B6DA-327D82095F3C" docUuidSource="ZooBank" docVersion="2" lastPageNumber="14" masterDocId="FFE7FFCCFFADFF9D0F396A02FF8EFF80" masterDocTitle="A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand" masterLastPageNumber="59" masterPageNumber="1" pageNumber="11" updateTime="1724951248699" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="50E85B28579E8248D3C532F4BB6D4B33">A critical revision of the Andrena Fabricius, 1775 of India, with the description of two new species (Hymenoptera: Andrenidae) from Uttarakhand</mods:title>
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<mods:namePart id="18D4E92048049BC0F90F0C2033029195">Gautam, R. K.</mods:namePart>
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<mods:namePart id="ECBA7B49F3DF6E3AF584587F09839914">Uniyal, V. P.</mods:namePart>
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<mods:namePart id="FADEEA0DAE421D9F8AA6AE72B6BBEDC5">Wood, Thomas J.</mods:namePart>
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<taxonomicName id="4C774D21FFA7FF970E936B51FBA6FEED" authority="Wood &amp; Gautam" authorityName="Wood &amp; Gautam" authorityYear="2024" box="[426,1064,339,366]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kedarnatha" status="sp. nov." subGenus="Melandrena">
<emphasis id="B903EAB0FFA7FF970E936B51FCD7FEEE" bold="true" box="[426,857,339,366]" pageId="10" pageNumber="11">
<emphasis id="B903EAB0FFA7FF970E936B51FD9BFEEE" bold="true" box="[426,533,339,366]" italics="true" pageId="10" pageNumber="11">Andrena</emphasis>
(
<emphasis id="B903EAB0FFA7FF970D1F6B51FD32FEEE" bold="true" box="[550,700,339,366]" italics="true" pageId="10" pageNumber="11">Melandrena</emphasis>
) 
<emphasis id="B903EAB0FFA7FF970DF46B51FCD7FEEE" bold="true" box="[717,857,339,366]" italics="true" pageId="10" pageNumber="11">kedarnatha</emphasis>
</emphasis>
Wood &amp; Gautam
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<taxonomicNameLabel id="A23057CBFFA7FF970B096B56FB04FEEE" box="[1072,1162,340,366]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C36D6529FFA7FF970D906B98FC04FE34" box="[681,906,409,436]" pageId="10" pageNumber="11" type="description">
<paragraph id="8BC836A2FFA7FF970D906B98FC04FE34" blockId="10.[372,1215,339,436]" box="[681,906,409,436]" pageId="10" pageNumber="11">
<figureCitation id="134C2A27FFA7FF970D906B98FD7DFE34" box="[681,755,409,436]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[189,1398,1026,1924]" captionTargetId="figure-242@10.[189,1398,1026,1924]" captionTargetPageId="10" captionText="Fig. 5.Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Habitus, lateral view. B. Face, frontal view. C. Propodeum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13511961" httpUri="https://zenodo.org/record/13511961/files/figure.png" pageId="10" pageNumber="11">Figs 5</figureCitation>
, 
<figureCitation id="134C2A27FFA7FF970C396B98FC04FE34" box="[768,906,410,436]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="10" pageNumber="11">6A, C, E, G</figureCitation>
</paragraph>
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<subSubSection id="C36D6529FFA7FF960F846BD9FAA2FC1C" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
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<emphasis id="B903EAB0FFA7FF970F846BD9FEB9FE75" bold="true" box="[189,311,475,501]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</heading>
</paragraph>
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<taxonomicName id="4C774D21FFA7FF970F846808FD0DFDA4" authority="Wood &amp; Gautam" authorityName="Wood &amp; Gautam" authorityYear="2024" box="[189,643,522,548]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kedarnatha" status="sp. nov.">
<emphasis id="B903EAB0FFA7FF970F846808FE3FFDA4" box="[189,433,522,548]" italics="true" pageId="10" pageNumber="11">Andrena kedarnatha</emphasis>
Wood &amp; Gautam
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<taxonomicNameLabel id="A23057CBFFA7FF970DB26808FD69FDA4" box="[651,743,522,548]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
can be recognised as a 
<taxonomicName id="4C774D21FFA7FF970B3B6808FAABFDA4" authority="Perez, 1890" authorityName="Perez" authorityYear="1890" box="[1026,1317,522,548]" class="Insecta" genus="Melandrena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAB0FFA7FF970B3B6808FB1CFDA4" box="[1026,1170,522,548]" italics="true" pageId="10" pageNumber="11">Melandrena</emphasis>
Pérez, 1890
</taxonomicName>
due to its large body size and robust shape (
<quantity id="4C8F9B47FFA7FF970D49682FFD7AFDC7" box="[624,756,557,584]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="10" pageNumber="11" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15–16 mm</quantity>
), dark integument and predominantly dark pubescence with partially infuscate wings (
<figureCitation id="134C2A27FFA7FF970D0A6852FD1DFDEB" box="[563,659,592,619]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[189,1398,1026,1924]" captionTargetId="figure-242@10.[189,1398,1026,1924]" captionTargetPageId="10" captionText="Fig. 5.Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Habitus, lateral view. B. Face, frontal view. C. Propodeum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13511961" httpUri="https://zenodo.org/record/13511961/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
), long ocelloccipital distance (3 × diameter of lateral ocellus, 
<figureCitation id="134C2A27FFA7FF970F866871FEAFFD0D" box="[191,289,627,653]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="10" pageNumber="11">Fig. 6A</figureCitation>
), large propodeal triangle with roughly raised internal carinae (
<figureCitation id="134C2A27FFA7FF970B156871FB02FD0E" box="[1068,1164,627,654]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[189,1398,1026,1924]" captionTargetId="figure-242@10.[189,1398,1026,1924]" captionTargetPageId="10" captionText="Fig. 5.Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Habitus, lateral view. B. Face, frontal view. C. Propodeum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13511961" httpUri="https://zenodo.org/record/13511961/files/figure.png" pageId="10" pageNumber="11">Fig. 5C</figureCitation>
), and lack of other diagnostic characters. Due to the dark integument without metallic reflections, the almost entirely dark pubescence (without white bands or spots of pubescence), the broad facial foveae that occupy almost the entire distance between the compound eye and lateral ocellus (
<figureCitation id="134C2A27FFA7FF970CFE68DFFBABFD77" box="[967,1061,733,759]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="10" pageNumber="11">Fig. 6A</figureCitation>
), and the clearly and densely punctate terga (
<figureCitation id="134C2A27FFA7FF970E496902FE5CFC9B" box="[368,466,768,795]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[189,1398,1026,1924]" captionTargetId="figure-242@10.[189,1398,1026,1924]" captionTargetPageId="10" captionText="Fig. 5.Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Habitus, lateral view. B. Face, frontal view. C. Propodeum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13511961" httpUri="https://zenodo.org/record/13511961/files/figure.png" pageId="10" pageNumber="11">Figs 5D</figureCitation>
, 
<figureCitation id="134C2A27FFA7FF970EE46902FDBDFC9A" box="[477,563,768,794]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="10" pageNumber="11">6E, 6G</figureCitation>
), it can be placed close to 
<taxonomicName id="4C774D21FFA7FF970C5C6902FB52FC9A" authority="Morawitz, 1886" authorityName="Morawitz" authorityYear="1886" box="[869,1244,768,794]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cussariensis">
<emphasis id="B903EAB0FFA7FF970C5C6902FB99FC9A" box="[869,1047,768,794]" italics="true" pageId="10" pageNumber="11">A. cussariensis</emphasis>
Morawitz, 1886
</taxonomicName>
. This species was described from what is now Azerbaijan (
<typeStatus id="54CC8800FFA7FF970DE86921FC8BFCBD" box="[721,773,803,829]" pageId="10" pageNumber="11">type</typeStatus>
material illustrated by 
<bibRefCitation id="EFE64B53FFA7FF970B2F6921FAAEFCBD" author="Astafurova Y. A. &amp; Proshchalykin M. Y. &amp; Sidorov D. A. &amp; Osytshnjuk A. Z." box="[1046,1312,803,829]" pageId="10" pageNumber="11" pagination="1 - 78" refId="ref27013" refString="Astafurova Y. A., Proshchalykin M. Y., Sidorov D. A. &amp; Osytshnjuk A. Z. 2021. The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa 5037: 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">
Astafurova 
<emphasis id="B903EAB0FFA7FF970B986926FB52FCBD" box="[1185,1244,803,829]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2021
</bibRefCitation>
) and is found from Crimea and European 
<collectingCountry id="F3607632FFA7FF970D606944FD29FCE0" box="[601,679,838,864]" name="Russia" pageId="10" pageNumber="11">Russia</collectingCountry>
through 
<collectingCountry id="F3607632FFA7FF970C2A6944FCE6FCE0" box="[787,872,838,864]" name="Turkey" pageId="10" pageNumber="11">Turkey</collectingCountry>
, 
<collectingCountry id="F3607632FFA7FF970C4C6945FC29FCE1" box="[885,935,839,865]" name="Iran" pageId="10" pageNumber="11">Iran</collectingCountry>
, the Caucasus to Central Asia, Siberia, and 
<collectingCountry id="F3607632FFA7FF970FCD6968FEE8FC04" box="[244,358,874,900]" name="Mongolia" pageId="10" pageNumber="11">Mongolia</collectingCountry>
(
<bibRefCitation id="EFE64B53FFA7FF970E406968FD1AFC04" author="Osytshnjuk A. Z. &amp; Romasenko L. &amp; Banaszak J. &amp; Motyka E." box="[377,660,874,900]" pageId="10" pageNumber="11" refId="ref29506" refString="Osytshnjuk A. Z., Romasenko L., Banaszak J. &amp; Motyka E. 2008. Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, Poland" type="book" year="2008">
Osytshnjuk 
<emphasis id="B903EAB0FFA7FF970D326968FDC4FC03" box="[523,586,874,900]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2008
</bibRefCitation>
; 
<bibRefCitation id="EFE64B53FFA7FF970D9B6968FC37FC04" author="Astafurova Y. A. &amp; Proshchalykin M. Y. &amp; Sidorov D. A. &amp; Osytshnjuk A. Z." box="[674,953,874,900]" pageId="10" pageNumber="11" pagination="1 - 78" refId="ref27013" refString="Astafurova Y. A., Proshchalykin M. Y., Sidorov D. A. &amp; Osytshnjuk A. Z. 2021. The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa 5037: 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">
Astafurova 
<emphasis id="B903EAB0FFA7FF970C0B6968FCE1FC03" box="[818,879,874,900]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2021
</bibRefCitation>
; 
<bibRefCitation id="EFE64B53FFA7FF970CF06968FB74FC04" author="Wood T. J. &amp; Monfared A." box="[969,1274,874,900]" pageId="10" pageNumber="11" pagination="1 - 136" refId="ref30834" refString="Wood T. J. &amp; Monfared A. 2022. A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy 843: 1 - 136. https: // doi. org / 10.5852 / ejt. 2022.843.1947" type="journal article" year="2022">Wood &amp; Monfared 2022</bibRefCitation>
). 
<taxonomicName id="4C774D21FFA7FF970A2B6968FEC0FC27" authorityName="Morawitz" authorityYear="1886" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cussariensis">
<emphasis id="B903EAB0FFA7FF970A2B6968FEC0FC27" italics="true" pageId="10" pageNumber="11">Andrena cussariensis</emphasis>
</taxonomicName>
is currently not confidently recorded from 
<collectingCountry id="F3607632FFA7FF970C7F698FFC0CFC27" box="[838,898,909,935]" name="India" pageId="10" pageNumber="11">India</collectingCountry>
specifically or the Himalayan region more generally, and is not considered to be present (see Remarks).
</paragraph>
<caption id="DF08662AFFA7FF970F846DA8FB26F868" ID-DOI="http://doi.org/10.5281/zenodo.13511961" ID-Zenodo-Dep="13511961" httpUri="https://zenodo.org/record/13511961/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1962,1988]" targetBox="[189,1398,1026,1924]" targetPageId="10">
<paragraph id="8BC836A2FFA7FF970F846DA8FB26F868" blockId="10.[187,1398,1962,2024]" pageId="10" pageNumber="11">
<emphasis id="B903EAB0FFA7FF970F846DA8FE87F845" bold="true" box="[189,265,1962,1989]" pageId="10" pageNumber="11">Fig. 5.</emphasis>
<taxonomicName id="4C774D21FFA7FF970E356DA9FCE1F845" authority="Wood &amp; Gautam" authorityName="Wood &amp; Gautam" authorityYear="2024" box="[268,879,1963,1989]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kedarnatha" status="sp. nov." subGenus="Melandrena">
<emphasis id="B903EAB0FFA7FF970E356DA9FEFFF845" box="[268,369,1963,1989]" italics="true" pageId="10" pageNumber="11">Andrena</emphasis>
(
<emphasis id="B903EAB0FFA7FF970EBB6DA9FD9CF845" box="[386,530,1963,1989]" italics="true" pageId="10" pageNumber="11">Melandrena</emphasis>
) 
<emphasis id="B903EAB0FFA7FF970D196DA9FD29F845" box="[544,679,1963,1989]" italics="true" pageId="10" pageNumber="11">kedarnatha</emphasis>
Wood &amp; Gautam
</taxonomicName>
<taxonomicNameLabel id="A23057CBFFA7FF970C4A6DA9FC40F845" box="[883,974,1963,1989]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♀ (NHMUK). 
<emphasis id="B903EAB0FFA7FF970BCB6DA8FA89F844" bold="true" box="[1266,1287,1962,1988]" pageId="10" pageNumber="11">A</emphasis>
. Habitus, lateral view. 
<emphasis id="B903EAB0FFA7FF970E6A6DCCFEE9F868" bold="true" box="[339,359,1998,2024]" pageId="10" pageNumber="11">B</emphasis>
. Face, frontal view. 
<emphasis id="B903EAB0FFA7FF970D6E6DCCFDE2F868" bold="true" box="[599,620,1998,2024]" pageId="10" pageNumber="11">C</emphasis>
. Propodeum, dorsal view. 
<emphasis id="B903EAB0FFA7FF970C9F6DCCFC35F868" bold="true" box="[934,955,1998,2024]" pageId="10" pageNumber="11">D</emphasis>
. Terga, dorsal view.
</paragraph>
</caption>
<paragraph id="8BC836A2FFA6FF960F846B0BFAA2FC1C" blockId="11.[188,1401,265,925]" pageId="11" pageNumber="12">
Morphologically, 
<taxonomicName id="4C774D21FFA6FF960EB76B0BFC8CFEA3" authority="Wood &amp; Gautam" authorityName="Wood &amp; Gautam" authorityYear="2024" box="[398,770,265,291]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="kedarnatha" status="sp. nov.">
<emphasis id="B903EAB0FFA6FF960EB76B0BFDB8FEA3" box="[398,566,265,291]" italics="true" pageId="11" pageNumber="12">A. kedarnatha</emphasis>
Wood &amp; Gautam
</taxonomicName>
<taxonomicNameLabel id="A23057CBFFA6FF960C306B08FCEDFEA4" box="[777,867,266,292]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
can be separated due to its larger body length of 
<quantity id="4C8F9B47FFA6FF960FE56B2EFED4FEC7" box="[220,346,300,327]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="11" pageNumber="12" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15–16 mm</quantity>
(slightly but noticeably larger than the body length of 
<taxonomicName id="4C774D21FFA6FF960CEF6B2FFB09FEC6" authorityName="Morawitz" authorityYear="1886" box="[982,1159,301,327]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cussariensis">
<emphasis id="B903EAB0FFA6FF960CEF6B2FFB09FEC6" box="[982,1159,301,327]" italics="true" pageId="11" pageNumber="12">A. cussariensis</emphasis>
</taxonomicName>
which averages 
<quantity id="4C8F9B47FFA6FF960A736B2EFE81FEEA" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="11" pageNumber="12" unit="mm" value="13.5" valueMax="14.0" valueMin="13.0">13– 14 mm</quantity>
in length), its longer ocelloccipital distance of 3–3.5 × diameter of lateral ocellus with the vertex most strongly and coarsely punctate (
<figureCitation id="134C2A27FFA6FF960D4C6B70FD5AFE0C" box="[629,724,370,396]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="11" pageNumber="12">Fig. 6A</figureCitation>
; in 
<taxonomicName id="4C774D21FFA6FF960C386B71FC3DFE0C" authorityName="Morawitz" authorityYear="1886" box="[769,947,371,397]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cussariensis">
<emphasis id="B903EAB0FFA6FF960C386B71FC3DFE0C" box="[769,947,371,397]" italics="true" pageId="11" pageNumber="12">A. cussariensis</emphasis>
</taxonomicName>
with shorter ocelloccipital distance of 2–2.5 × diameter of lateral ocellus, with the vertex more weakly and finely punctate, 
<figureCitation id="134C2A27FFA6FF960BAC6B94FB60FE30" box="[1173,1262,406,432]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="11" pageNumber="12">Fig. 6B</figureCitation>
), by a more consistently dense punctation of the scutum with punctures separated by &lt;0.5–0.5 puncture diameters, not becoming slightly but noticeably sparser posteromedially (
<figureCitation id="134C2A27FFA6FF960CAA6BDEFC61FE76" box="[915,1007,476,502]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="11" pageNumber="12">Fig. 6C</figureCitation>
; in 
<taxonomicName id="4C774D21FFA6FF960B2C6BDFFB48FE76" authorityName="Morawitz" authorityYear="1886" box="[1045,1222,477,503]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cussariensis">
<emphasis id="B903EAB0FFA6FF960B2C6BDFFB48FE76" box="[1045,1222,477,503]" italics="true" pageId="11" pageNumber="12">A. cussariensis</emphasis>
</taxonomicName>
with the scutal punctures typically separated by 0.5 puncture diameters but in some specimens with punctures separated by 1–2 puncture diameters posteromedially, forming a small but distinct shining space, 
<figureCitation id="134C2A27FFA6FF960BFE6820FAABFDBC" box="[1223,1317,546,572]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="11" pageNumber="12">Fig. 6D</figureCitation>
), terga with slightly but comparatively larger and coarser punctures, most visible on the discs of T1–2 (
<figureCitation id="134C2A27FFA6FF960A216844FAFDFDE0" box="[1304,1395,582,608]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="11" pageNumber="12">Fig. 6E</figureCitation>
; in 
<taxonomicName id="4C774D21FFA6FF960FE4686BFE1FFD03" authorityName="Morawitz" authorityYear="1886" box="[221,401,617,643]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cussariensis">
<emphasis id="B903EAB0FFA6FF960FE4686BFE1FFD03" box="[221,401,617,643]" italics="true" pageId="11" pageNumber="12">A. cussariensis</emphasis>
</taxonomicName>
with comparatively smaller and finer punctures, 
<figureCitation id="134C2A27FFA6FF960CE7686BFBB6FD03" box="[990,1080,617,643]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="11" pageNumber="12">Fig. 6F</figureCitation>
), and T2 laterally with the shallow foveae broadened and filled with reddish-bristles, surface densely punctate with punctures separated by 0.5 puncture diameters (
<figureCitation id="134C2A27FFA6FF960DB968ADFD6EFD49" box="[640,736,687,713]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="11" pageNumber="12">Fig. 6G</figureCitation>
; in 
<taxonomicName id="4C774D21FFA6FF960C2B68B2FC48FD49" authorityName="Morawitz" authorityYear="1886" box="[786,966,688,714]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cussariensis">
<emphasis id="B903EAB0FFA6FF960C2B68B2FC48FD49" box="[786,966,688,714]" italics="true" pageId="11" pageNumber="12">A. cussariensis</emphasis>
</taxonomicName>
without a noticeable foveae present laterally on T2, surrounding surface comparatively sparsely and weakly punctate, 
<figureCitation id="134C2A27FFA6FF960B9E68D0FA86FD6C" box="[1191,1288,722,748]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="11" pageNumber="12">Fig. 6H</figureCitation>
). To best appreciate the final characters, comparative material is required. There is some variation in colouration between the older museum specimens which have the wing venation orange-brown, the tergal margins slightly lightened brownish, the tibial scopa with off-white to light brownish hairs whereas recently collected specimens are uniformly darker (dark venation, dark terga, and dark scopal hairs). This is considered to represent variation, as sculptural characters do not differ between these specimens.
</paragraph>
</subSubSection>
<subSubSection id="C36D6529FFA6FF960F8469C6FB96FBB0" pageId="11" pageNumber="12" type="etymology">
<paragraph id="8BC836A2FFA6FF960F8469C6FECBFC5E" blockId="11.[189,325,964,990]" box="[189,325,964,990]" pageId="11" pageNumber="12">
<heading id="D08081CEFFA6FF960F8469C6FECBFC5E" bold="true" box="[189,325,964,990]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B903EAB0FFA6FF960F8469C6FECBFC5E" bold="true" box="[189,325,964,990]" pageId="11" pageNumber="12">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A2FFA6FF960F8469F0FB96FBB0" blockId="11.[189,1399,1010,1072]" pageId="11" pageNumber="12">
Taken from the name of the Kedarnath temple (
<collectingRegion id="49B3F840FFA6FF960C2B69F0FC29FB8C" box="[786,935,1010,1036]" country="India" name="Uttarakhand" pageId="11" pageNumber="12">Uttarakhand</collectingRegion>
) which is found close to the modern sampling localities of this new species (
<figureCitation id="134C2A27FFA6FF960DA96E14FD59FBB0" box="[656,727,1046,1072]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[244,1364,494,1878]" captionTargetId="figure-44@15.[219,1369,459,1892]" captionTargetPageId="15" captionText="Fig. 7. Distribution map of northern Uttarakhand province in India showing the loci typici for Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov. and Andrena (Simandrena) tungnatha Wood &amp; Gautam sp. nov., as well as Kedarnath and Tungnath temples." figureDoi="http://doi.org/10.5281/zenodo.13511987" httpUri="https://zenodo.org/record/13511987/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
). It is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C36D6529FFA6FF960F846E55FA8FFA55" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BC836A2FFA6FF960F846E55FEE3FBF1" blockId="11.[189,365,1111,1137]" box="[189,365,1111,1137]" pageId="11" pageNumber="12">
<heading id="D08081CEFFA6FF960F846E55FEE3FBF1" bold="true" box="[189,365,1111,1137]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B903EAB0FFA6FF960F846E55FEE3FBF1" bold="true" box="[189,365,1111,1137]" pageId="11" pageNumber="12">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A2FFA6FF960FD36E84FED3FB20" blockId="11.[189,1398,1158,1255]" box="[234,349,1158,1184]" pageId="11" pageNumber="12">
<emphasis id="B903EAB0FFA6FF960FD36E84FED3FB20" bold="true" box="[234,349,1158,1184]" pageId="11" pageNumber="12">
<typeStatus id="54CC8800FFA6FF960FD36E84FED3FB20" box="[234,349,1158,1184]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC836A2FFA6FF960F846EABFE1CFB67" blockId="11.[189,1398,1158,1255]" pageId="11" pageNumber="12">
<materialsCitation id="3B1F3CFFFFA6FF960F846EABFE1CFB67" collectingDate="2022-05-08" collectionCode="WII" collectorName="R. K. Gautam" country="India" elevation="2250" latitude="30.6146" location="Chaumasi Village" longLatPrecision="6" longitude="79.0695" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1" stateProvince="Uttarakhand" typeStatus="holotype">
<collectingCountry id="F3607632FFA6FF960F846EABFE80FB43" box="[189,270,1193,1219]" name="India" pageId="11" pageNumber="12">INDIA</collectingCountry>
<emphasis id="B903EAB0FFA6FF960E2F6EABFEAEFB43" bold="true" box="[278,288,1193,1219]" pageId="11" pageNumber="12">•</emphasis>
<specimenCount id="9D71FD2BFFA6FF960E116EABFEB1FB43" box="[296,319,1193,1219]" count="1" pageId="11" pageNumber="12" type="female">♀</specimenCount>
; 
<collectingRegion id="49B3F840FFA6FF960E776EABFE6CFB43" box="[334,482,1193,1219]" country="India" name="Uttarakhand" pageId="11" pageNumber="12">Uttarakhand</collectingRegion>
, 
<location id="8EA86079FFA6FF960ED76EABFD4CFB43" LSID="urn:lsid:plazi:treatment:03DE87B4FFA7FF900E936B51FC04F88F:8EA86079FFA6FF960ED76EABFD4CFB43" box="[494,706,1193,1219]" country="India" latitude="30.6146" longLatPrecision="6" longitude="79.0695" name="Chaumasi Village" pageId="11" pageNumber="12" stateProvince="Uttarakhand">Chaumasi Village</location>
; 
<geoCoordinate id="EE435065FFA6FF960DF66EABFCD7FB43" box="[719,857,1193,1219]" degrees="30.6146" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="30.6146">30.6146° N</geoCoordinate>
, 
<geoCoordinate id="EE435065FFA6FF960C5C6EABFC63FB43" box="[869,1005,1193,1220]" degrees="79.0695" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="79.0695">79.0695° E</geoCoordinate>
; 
<elevation id="005AD191FFA6FF960CC36EABFB1DFB43" box="[1018,1171,1193,1220]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="11" pageNumber="12" unit="m" value="2250.0">
<quantity id="4C8F9B47FFA6FF960CC36EABFBDDFB43" box="[1018,1107,1193,1220]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="11" pageNumber="12" unit="m" value="2250.0">2250 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFC91062FFA6FF960B986EABFABDFB43" box="[1185,1331,1193,1219]" pageId="11" pageNumber="12" value="2022-05-08">
<collectingDate id="EF8DE98AFFA6FF960B986EABFABDFB43" box="[1185,1331,1193,1219]" pageId="11" pageNumber="12" value="2022-05-08">8 May 2022</collectingDate>
</date>
; 
<collectorName id="26825374FFA6FF960A066EABFE94FB66" pageId="11" pageNumber="12">R.K. Gautam</collectorName>
leg.; 
<collectionCode id="ED66AE67FFA6FF960E626ECFFE00FB67" box="[347,398,1229,1255]" pageId="11" pageNumber="12">WII</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC836A2FFA6FF960FD36F0CFEE7FAA8" blockId="11.[189,1398,1294,1426]" box="[234,361,1294,1320]" pageId="11" pageNumber="12">
<emphasis id="B903EAB0FFA6FF960FD36F0CFEE7FAA8" bold="true" box="[234,361,1294,1320]" pageId="11" pageNumber="12">
<typeStatus id="54CC8800FFA6FF960FD36F0CFEE7FAA8" box="[234,361,1294,1320]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC836A2FFA6FF960F846F33FEBAFA12" blockId="11.[189,1398,1294,1426]" pageId="11" pageNumber="12">
<materialsCitation id="3B1F3CFFFFA6FF960F846F33FE00FAEF" collectingDate="2022-05-08" collectionCode="WII" collectorName="R. K. Gautam" country="India" elevation="2250" latitude="30.6146" location="Chaumasi Village" longLatPrecision="6" longitude="79.0695" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1" stateProvince="Uttarakhand" typeStatus="paratype">
<collectingCountry id="F3607632FFA6FF960F846F33FE80FACB" box="[189,270,1329,1355]" name="India" pageId="11" pageNumber="12">INDIA</collectingCountry>
<emphasis id="B903EAB0FFA6FF960E2C6F33FE91FACB" bold="true" box="[277,287,1329,1355]" pageId="11" pageNumber="12">•</emphasis>
<specimenCount id="9D71FD2BFFA6FF960E1F6F33FEDCFACB" box="[294,338,1329,1355]" count="1" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
; 
<collectingRegion id="49B3F840FFA6FF960E666F33FE7DFACB" box="[351,499,1329,1355]" country="India" name="Uttarakhand" pageId="11" pageNumber="12">Uttarakhand</collectingRegion>
, 
<location id="8EA86079FFA6FF960EC76F33FD5FFACB" LSID="urn:lsid:plazi:treatment:03DE87B4FFA7FF900E936B51FC04F88F:8EA86079FFA6FF960EC76F33FD5FFACB" box="[510,721,1329,1355]" country="India" latitude="30.6146" longLatPrecision="6" longitude="79.0695" name="Chaumasi Village" pageId="11" pageNumber="12" stateProvince="Uttarakhand">Chaumasi Village</location>
; 
<geoCoordinate id="EE435065FFA6FF960DE56F33FCEBFACB" box="[732,869,1329,1355]" degrees="30.6146" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="30.6146">30.6146° N</geoCoordinate>
, 
<geoCoordinate id="EE435065FFA6FF960C496F33FC78FACB" box="[880,1014,1329,1356]" degrees="79.0695" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="79.0695">79.0695° E</geoCoordinate>
; 
<elevation id="005AD191FFA6FF960B3B6F33FB16FACB" box="[1026,1176,1329,1356]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="11" pageNumber="12" unit="m" value="2250.0">
<quantity id="4C8F9B47FFA6FF960B3B6F33FBD4FACB" box="[1026,1114,1329,1356]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="11" pageNumber="12" unit="m" value="2250.0">2250 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFC91062FFA6FF960B9C6F33FABAFACB" box="[1189,1332,1329,1355]" pageId="11" pageNumber="12" value="2022-05-08">
<collectingDate id="EF8DE98AFFA6FF960B9C6F33FABAFACB" box="[1189,1332,1329,1355]" pageId="11" pageNumber="12" value="2022-05-08">8 May 2022</collectingDate>
</date>
; 
<collectorName id="26825374FFA6FF960A066F33FE94FAEE" pageId="11" pageNumber="12">R.K. Gautam</collectorName>
leg.; 
<collectionCode id="ED66AE67FFA6FF960E666F57FE00FAEF" box="[351,398,1365,1391]" pageId="11" pageNumber="12">WII</collectionCode>
</materialsCitation>
<emphasis id="B903EAB0FFA6FF960EA36F56FE2AFAEE" bold="true" box="[410,420,1364,1390]" pageId="11" pageNumber="12">•</emphasis>
<materialsCitation id="3B1F3CFFFFA6FF960E956F56FEBEFA12" collectionCode="NHMUK" collectorName="Gulmarg &amp; Lt-Col. F. W. Thomson" country="India" location="Jammu and Kashmir" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1" stateProvince="Jammu and Kashmir" typeStatus="paratype">
<specimenCount id="9D71FD2BFFA6FF960E956F56FE55FAEF" box="[428,475,1364,1391]" count="1" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
; 
<collectorName id="26825374FFA6FF960ED56F56FDDAFAEE" box="[492,596,1364,1390]" pageId="11" pageNumber="12">Gulmarg</collectorName>
[
<collectingRegion id="49B3F840FFA6FF960D5D6F57FCEBFAEE" box="[612,869,1364,1391]" country="India" name="Jammu and Kashmir" pageId="11" pageNumber="12">Jammu and Kashmir</collectingRegion>
]; summer 1913; 
<collectorName id="26825374FFA6FF960B146F56FAB4FAEE" box="[1069,1338,1364,1391]" pageId="11" pageNumber="12">Lt-Col. F.W. Thomson</collectorName>
leg.; 
<collectionCode id="ED66AE67FFA6FF960F846F7AFEBEFA12" box="[189,304,1400,1426]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="11" pageNumber="12" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC836A2FFA6FF960F846FBBFA8FFA55" blockId="11.[189,1281,1464,1493]" box="[189,1281,1464,1493]" pageId="11" pageNumber="12">
<materialsCitation id="3B1F3CFFFFA6FF960F846FBBFB73FA55" box="[189,1277,1464,1493]" collectingDate="1993-10-04" collectionCode="OOLM" collectorName="E. Huttinger" country="Nepal" elevation="2850" location="Jumla" municipality="Jumla" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F3607632FFA6FF960F846FBBFE97FA53" box="[189,281,1465,1491]" name="Nepal" pageId="11" pageNumber="12">NEPAL</collectingCountry>
<emphasis id="B903EAB0FFA6FF960E196FBBFEA4FA53" bold="true" box="[288,298,1465,1491]" pageId="11" pageNumber="12">•</emphasis>
<specimenCount id="9D71FD2BFFA6FF960E0B6FBBFED1FA53" box="[306,351,1465,1491]" count="1" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
; W-Nepal, 
<location id="8EA86079FFA6FF960EDF6FBBFDA1FA53" LSID="urn:lsid:plazi:treatment:03DE87B4FFA7FF900E936B51FC04F88F:8EA86079FFA6FF960EDF6FBBFDA1FA53" box="[486,559,1465,1491]" country="Nepal" municipality="Jumla" name="Jumla" pageId="11" pageNumber="12">Jumla</location>
, 
<collectingMunicipality id="6BACACD8FFA6FF960D026FBBFD0BFA53" box="[571,645,1465,1491]" pageId="11" pageNumber="12">Jumla</collectingMunicipality>
; 
<elevation id="005AD191FFA6FF960DA96FBAFCA7FA53" box="[656,809,1464,1491]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="11" pageNumber="12" unit="m" value="2850.0">
<quantity id="4C8F9B47FFA6FF960DA96FBAFD67FA53" box="[656,745,1464,1491]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="11" pageNumber="12" unit="m" value="2850.0">2850 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFC91062FFA6FF960C0F6FBBFC4DFA53" box="[822,963,1465,1491]" pageId="11" pageNumber="12" value="1993-10-04">
<collectingDate id="EF8DE98AFFA6FF960C0F6FBBFC4DFA53" box="[822,963,1465,1491]" pageId="11" pageNumber="12" value="1993-10-04">4 Oct. 1993</collectingDate>
</date>
; 
<collectorName id="26825374FFA6FF960CE96FBBFBEEFA53" box="[976,1120,1465,1491]" pageId="11" pageNumber="12">E. Hüttinger</collectorName>
leg.; 
<collectionCode id="ED66AE67FFA6FF960B9A6FBBFB73FA55" box="[1187,1277,1465,1493]" pageId="11" pageNumber="12">OÖLM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D6529FFA6FF900F846FF8FEB7FB4F" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="description">
<paragraph id="8BC836A2FFA6FF960F846FF8FEC1F994" blockId="11.[189,335,1530,1556]" box="[189,335,1530,1556]" pageId="11" pageNumber="12">
<heading id="D08081CEFFA6FF960F846FF8FEC1F994" bold="true" box="[189,335,1530,1556]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B903EAB0FFA6FF960F846FF8FEC1F994" bold="true" box="[189,335,1530,1556]" pageId="11" pageNumber="12">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A2FFA6FF960FD36C2BFEC8F9C3" blockId="11.[189,609,1577,1639]" box="[234,326,1577,1603]" pageId="11" pageNumber="12">
<emphasis id="B903EAB0FFA6FF960FD36C2BFEC8F9C3" bold="true" box="[234,326,1577,1603]" pageId="11" pageNumber="12">Female</emphasis>
</paragraph>
<paragraph id="8BC836A2FFA6FF960F846C4EFDEFF9E7" blockId="11.[189,609,1577,1639]" box="[189,609,1612,1639]" pageId="11" pageNumber="12">
<smallCapsWord id="8D2EA07EFFA6FF960F846C4EFF70F9E4" baselines="1631,1631" box="[189,254,1612,1638]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="11" pageNumber="12">BODY</smallCapsWord>
. Length 
<quantity id="4C8F9B47FFA6FF960E5E6C4EFE68F9E6" box="[359,486,1612,1639]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="11" pageNumber="12" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15–16 mm</quantity>
(
<figureCitation id="134C2A27FFA6FF960ECC6C4EFDDAF9E7" box="[501,596,1612,1639]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[189,1398,1026,1924]" captionTargetId="figure-242@10.[189,1398,1026,1924]" captionTargetPageId="10" captionText="Fig. 5.Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Habitus, lateral view. B. Face, frontal view. C. Propodeum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13511961" httpUri="https://zenodo.org/record/13511961/files/figure.png" pageId="11" pageNumber="12">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8BC836A2FFA6FF960F846C8CFB1AF842" blockId="11.[189,1400,1677,1986]" pageId="11" pageNumber="12">
<smallCapsWord id="8D2EA07EFFA6FF960F846C8CFF72F926" baselines="1697,1697" box="[189,252,1678,1704]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="11" pageNumber="12">HEAD</smallCapsWord>
. Dark, 1.2 × as wide as long (
<figureCitation id="134C2A27FFA6FF960D676C8FFD35F928" box="[606,699,1677,1704]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[189,1398,1026,1924]" captionTargetId="figure-242@10.[189,1398,1026,1924]" captionTargetPageId="10" captionText="Fig. 5.Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Habitus, lateral view. B. Face, frontal view. C. Propodeum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13511961" httpUri="https://zenodo.org/record/13511961/files/figure.png" pageId="11" pageNumber="12">Fig. 5B</figureCitation>
). Clypeus domed, densely punctate, punctures separated by 0.5 puncture diameters, underlying surface dull. Process of labrum broadly trapezoidal, 2.5 × as wide as long, lateral margins weakly emarginate, apical margin truncate. Gena broad, 1.2–1.5 × diameter of compound eye; ocelloccipital distance 3–3.5 × diameter of lateral ocellus. Vertex strongly and coarsely punctate, punctures behind ocellar triangle separated by 0.5 puncture diameters (
<figureCitation id="134C2A27FFA6FF960B9D6D18FA87F8B4" box="[1188,1289,1818,1844]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="11" pageNumber="12">Fig. 6A</figureCitation>
). Foveae dorsally broad, occupying almost entire space between compound eye and lateral ocellus, separated from lateral ocellus by distance subequal to its diameter; foveae ventrally strongly passing level of antennal insertions ventrally, filled with dark brown hairs. Head with black to dark brown hairs, none equalling length of scape. Antennae dark, A3 equalling A4+5, shorter than A4+5+6.
</paragraph>
<caption id="DF08662AFFA1FF910F846D66FB80F868" ID-DOI="http://doi.org/10.5281/zenodo.13511969" ID-Zenodo-Dep="13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1892,1918]" targetBox="[263,1324,265,1855]" targetPageId="12">
<paragraph id="8BC836A2FFA1FF910F846D66FB80F868" blockId="12.[189,1399,1892,2025]" pageId="12" pageNumber="13">
<emphasis id="B903EAB0FFA1FF910F846D66FE04F8FE" bold="true" box="[189,394,1892,1919]" pageId="12" pageNumber="13">Fig. 6. A, C, E, G</emphasis>
. 
<taxonomicName id="4C774D21FFA1FF910EAF6D67FC79F8FF" authority="Wood &amp; Gautam" authorityName="Wood &amp; Gautam" authorityYear="2024" box="[406,1015,1893,1919]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kedarnatha" status="sp. nov." subGenus="Melandrena">
<emphasis id="B903EAB0FFA1FF910EAF6D67FE75F8FF" box="[406,507,1893,1919]" italics="true" pageId="12" pageNumber="13">Andrena</emphasis>
(
<emphasis id="B903EAB0FFA1FF910D336D67FD14F8FF" box="[522,666,1893,1919]" italics="true" pageId="12" pageNumber="13">Melandrena</emphasis>
) 
<emphasis id="B903EAB0FFA1FF910D916D67FCA1F8FF" box="[680,815,1893,1919]" italics="true" pageId="12" pageNumber="13">kedarnatha</emphasis>
Wood &amp; Gautam
</taxonomicName>
<taxonomicNameLabel id="A23057CBFFA1FF910CC56D67FBD9F8FF" box="[1020,1111,1893,1919]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♀ (NHMUK). 
<emphasis id="B903EAB0FFA1FF910F846D8AFF5CF822" bold="true" box="[189,210,1928,1954]" pageId="12" pageNumber="13">A</emphasis>
. Vertex, dorsal view. 
<emphasis id="B903EAB0FFA1FF910EEF6D8AFE64F822" bold="true" box="[470,490,1928,1954]" pageId="12" pageNumber="13">C</emphasis>
. Scutum, dorsal view. 
<emphasis id="B903EAB0FFA1FF910DC26D8AFC80F822" bold="true" box="[763,782,1928,1954]" pageId="12" pageNumber="13">E</emphasis>
. T1–2, dorsal view. 
<emphasis id="B903EAB0FFA1FF910B3A6D8AFB97F822" bold="true" box="[1027,1049,1928,1954]" pageId="12" pageNumber="13">G</emphasis>
. T2, lateral view detail. – 
<emphasis id="B903EAB0FFA1FF910A656D8AFEAFF845" bold="true" pageId="12" pageNumber="13">B, D, F, H</emphasis>
. 
<taxonomicName id="4C774D21FFA1FF910E0D6DA9FC3BF845" authority="Morawitz, 1886" authorityName="Morawitz" authorityYear="1886" box="[308,949,1963,1989]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cussariensis" subGenus="Melandrena">
<emphasis id="B903EAB0FFA1FF910E0D6DA9FE17F845" box="[308,409,1963,1989]" italics="true" pageId="12" pageNumber="13">Andrena</emphasis>
(
<emphasis id="B903EAB0FFA1FF910E966DA9FDB1F845" box="[431,575,1963,1989]" italics="true" pageId="12" pageNumber="13">Melandrena</emphasis>
) 
<emphasis id="B903EAB0FFA1FF910D6C6DAEFD68F845" box="[597,742,1964,1989]" italics="true" pageId="12" pageNumber="13">cussariensis</emphasis>
Morawitz, 1886
</taxonomicName>
, ♀ (TJWC). 
<emphasis id="B903EAB0FFA1FF910B646DA9FBFFF845" bold="true" box="[1117,1137,1963,1989]" pageId="12" pageNumber="13">B</emphasis>
. Vertex, dorsal view. 
<emphasis id="B903EAB0FFA1FF910F846DCCFF5CF868" bold="true" box="[189,210,1998,2024]" pageId="12" pageNumber="13">D</emphasis>
. Scutum, dorsal view. 
<emphasis id="B903EAB0FFA1FF910EE76DCCFE7EF868" bold="true" box="[478,496,1998,2024]" pageId="12" pageNumber="13">F</emphasis>
. T1–2, dorsal view. 
<emphasis id="B903EAB0FFA1FF910DD96DCCFD79F868" bold="true" box="[736,759,1998,2024]" pageId="12" pageNumber="13">H</emphasis>
. T2, lateral view detail.
</paragraph>
</caption>
<paragraph id="8BC836A2FFA0FF900F846B08FC7EFC90" blockId="13.[188,1400,265,784]" pageId="13" pageNumber="14">
<smallCapsWord id="8D2EA07EFFA0FF900F846B08FEB4FEA2" baselines="285,285" box="[189,314,266,292]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Mesosoma" pageId="13" pageNumber="14">MESOSOMA</smallCapsWord>
. Scutum and scutellum densely and regularly punctate, punctures separated by &lt;0.5 puncture diameters, almost confluent, interspaces weakly shining medially, dull anteriorly and laterally (
<figureCitation id="134C2A27FFA0FF900A356B2EFAE7FEC6" box="[1292,1385,300,326]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="13" pageNumber="14">Fig. 6C</figureCitation>
). Pronotum rounded. Mesepisternum densely but shallowly punctate, punctures separated by 0.5 puncture diameters, interspaces microreticulate, dull. Dorsolateral parts of propodeum densely and shallowly punctate, punctures separated by 0.5 puncture diameters, interspaces raised to form network of finely raised rugae; propodeal triangle large, laterally delineated by finely raised carinae, internal surface with dense network of raised carinae (
<figureCitation id="134C2A27FFA0FF900DA96BDEFD7FFE77" box="[656,753,476,503]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[189,1398,1026,1924]" captionTargetId="figure-242@10.[189,1398,1026,1924]" captionTargetPageId="10" captionText="Fig. 5.Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Habitus, lateral view. B. Face, frontal view. C. Propodeum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13511961" httpUri="https://zenodo.org/record/13511961/files/figure.png" pageId="13" pageNumber="14">Fig. 5C</figureCitation>
). Mesosoma with abundant but relatively short black pubescence, longest on mesepisternum, becoming shorter but not squamous on scutum and scutellum. Propodeal corbicula incomplete, dorsal fringe composed of abundant and strongly plumose black hairs, internal surface covered with latitudinal weakly raised rugae, with abundant simple black hairs. Legs dark, apical tarsal segments lightened reddish brown; pubescence black to dark brown. Flocculus incomplete, weakly produced, composed of plumose black hairs; femoral and tibial scopae composed of off-white to light brownish or predominantly black simple hairs. Hind tarsal claws with strong inner tooth. Wings infuscate over majority of surface, becoming marginally weaker away from venation; stigma and venation dark orange-brown to black, nervulus interstitial.
</paragraph>
<paragraph id="8BC836A2FFA0FF900F84693AFEF9FBEB" blockId="13.[189,1400,823,1131]" pageId="13" pageNumber="14">
<smallCapsWord id="8D2EA07EFFA0FF900F84693AFEB7FCD0" baselines="843,843" box="[189,313,824,850]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Metasoma" pageId="13" pageNumber="14">METASOMA</smallCapsWord>
. Tergal discs dark, apical margins narrowly lightened dark brown or almost entirely dark (
<figureCitation id="134C2A27FFA0FF900FFF6958FEABFCF5" box="[198,293,858,885]" captionStart="Fig" captionStartId="10.[189,232,1962,1988]" captionTargetBox="[189,1398,1026,1924]" captionTargetId="figure-242@10.[189,1398,1026,1924]" captionTargetPageId="10" captionText="Fig. 5.Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Habitus, lateral view. B. Face, frontal view. C. Propodeum, dorsal view. D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13511961" httpUri="https://zenodo.org/record/13511961/files/figure.png" pageId="13" pageNumber="14">Fig. 5D</figureCitation>
). Tergal discs densely but slightly irregularly punctate, punctures typically separated by 0.5–1 puncture diameters (
<figureCitation id="134C2A27FFA0FF900E89697CFD83FC17" box="[432,525,893,920]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="13" pageNumber="14">Fig. 6E</figureCitation>
), up to 2 puncture diameters medially; punctures becoming progressively finer and weaker on marginal areas from T1–4, on T4 almost disappearing into underlying sculpture. Tergal discs obscurely shagreened, shining to weakly shining. T2 laterally with shallow but distinct broad foveae, foveae subtly impressed, covered with reddish-brown to black bristles (
<figureCitation id="134C2A27FFA0FF900B8E69E5FA99FB81" box="[1207,1303,999,1025]" captionStart="Fig" captionStartId="12.[189,232,1892,1918]" captionTargetBox="[263,1324,265,1855]" captionTargetId="figure-20@12.[263,1324,265,1856]" captionTargetPageId="12" captionText="Fig. 6. A, C, E, G. Andrena (Melandrena) kedarnatha Wood &amp; Gautam sp. nov., paratype, ♀ (NHMUK). A. Vertex, dorsal view. C. Scutum, dorsal view. E. T1–2, dorsal view. G. T2, lateral view detail. – B, D, F, H. Andrena (Melandrena) cussariensis Morawitz, 1886, ♀ (TJWC). B. Vertex, dorsal view. D. Scutum, dorsal view. F. T1–2, dorsal view. H. T2, lateral view detail." figureDoi="http://doi.org/10.5281/zenodo.13511969" httpUri="https://zenodo.org/record/13511969/files/figure.png" pageId="13" pageNumber="14">Fig. 6G</figureCitation>
). Tergal discs with scattered short dark hairs, not forming apical hairbands. Apical fringe of T5 and hairs flanking pygidial plate dark brown. Pygidial plate large, rounded triangular, surface finely and evenly shagreened, weakly shining.
</paragraph>
<paragraph id="8BC836A2FFA0FF900FD36E90FEA4FB2C" blockId="13.[189,313,1170,1231]" box="[234,298,1170,1196]" pageId="13" pageNumber="14">
<emphasis id="B903EAB0FFA0FF900FD36E90FEA4FB2C" bold="true" box="[234,298,1170,1196]" pageId="13" pageNumber="14">Male</emphasis>
</paragraph>
<paragraph id="8BC836A2FFA0FF900F846EB7FEB7FB4F" blockId="13.[189,313,1170,1231]" box="[189,313,1205,1231]" pageId="13" pageNumber="14">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C36D6529FFA0FF900F846EF5FAA2F920" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BC836A2FFA0FF900F846EF5FEBEFA91" blockId="13.[189,304,1271,1297]" box="[189,304,1271,1297]" pageId="13" pageNumber="14">
<heading id="D08081CEFFA0FF900F846EF5FEBEFA91" bold="true" box="[189,304,1271,1297]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B903EAB0FFA0FF900F846EF5FEBEFA91" bold="true" box="[189,304,1271,1297]" pageId="13" pageNumber="14">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A2FFA0FF900F846F27FAA2F920" blockId="13.[186,1400,1317,1696]" pageId="13" pageNumber="14">
<taxonomicName id="4C774D21FFA0FF900F846F27FE32FABF" authorityName="Morawitz" authorityYear="1886" box="[189,444,1317,1343]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cussariensis">
<emphasis id="B903EAB0FFA0FF900F846F27FE32FABF" box="[189,444,1317,1343]" italics="true" pageId="13" pageNumber="14">Andrena cussariensis</emphasis>
</taxonomicName>
was listed for the Indian fauna by 
<bibRefCitation id="EFE64B53FFA0FF900C646F24FBD2FAC0" author="Meena L. K. &amp; Dey D." box="[861,1116,1318,1344]" pageId="13" pageNumber="14" pagination="443 - 452" refId="ref29114" refString="Meena L. K. &amp; Dey D. 2019. Annotated checklist of non- Apis bees of the genus Andrena (Hymenoptera: Andrenidae) from India. Indian Journal of Entomology 81: 443 - 452 https: // doi. org / 10.5958 / 0974 - 8172.2019.00095.6" type="journal article" year="2019">Meena &amp; Dey (2019)</bibRefCitation>
who mentioned it from Kohtak [= Kohat, 
<collectingCountry id="F3607632FFA0FF900EA06F4BFD8DFAE3" box="[409,515,1353,1379]" name="Pakistan" pageId="13" pageNumber="14">Pakistan</collectingCountry>
], Musooree [= Mussoorie], Shimla, and 
<collectingRegion id="49B3F840FFA0FF900CDD6F4BFBB7FAE3" box="[996,1081,1353,1379]" country="Pakistan" name="Punjab" pageId="13" pageNumber="14">Punjab</collectingRegion>
. The listing of this species from 
<collectingCountry id="F3607632FFA0FF900FC56F6EFEB6FA06" box="[252,312,1388,1414]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
is complicated, and requires dissection. The listing from Kohat refers to the record of 
<bibRefCitation id="EFE64B53FFA0FF900A3F6F6EFE87FA29" author="Cockerell T. D. A." pageId="13" pageNumber="14" pagination="282 - 290" refId="ref27877" refString="Cockerell T. D. A. 1917. XXV. - Descriptions and records of bees. - LXXIV. Annals and Magazine of Natural History 19: 282 - 290. https: // doi. org / 10.1080 / 00222931709486938" type="journal article" year="1917">Cockerell (1917)</bibRefCitation>
who reported 
<taxonomicName id="4C774D21FFA0FF900E8B6F8DFDEAFA29" authorityName="Morawitz" authorityYear="1886" box="[434,612,1423,1449]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cussariensis">
<emphasis id="B903EAB0FFA0FF900E8B6F8DFDEAFA29" box="[434,612,1423,1449]" italics="true" pageId="13" pageNumber="14">A. cussariensis</emphasis>
</taxonomicName>
from “Kohat, N.W. Provinces, 
<collectingCountry id="F3607632FFA0FF900CE96F8DFB9FFA29" box="[976,1041,1423,1449]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
”, and described the subspecies 
<taxonomicName id="4C774D21FFA0FF900F856FB1FD39FA4C" authority="Cockerell, 1917" authorityName="Cockerell" authorityYear="1917" box="[188,695,1458,1485]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="cussariensis" subSpecies="kohatensis">
<emphasis id="B903EAB0FFA0FF900F856FB1FE7FFA4C" box="[188,497,1458,1485]" italics="true" pageId="13" pageNumber="14">A. cussariensis kohatensis</emphasis>
<bibRefCitation id="EFE64B53FFA0FF900EC06FB0FD39FA4C" author="Cockerell T. D. A." box="[505,695,1458,1484]" pageId="13" pageNumber="14" pagination="282 - 290" refId="ref27877" refString="Cockerell T. D. A. 1917. XXV. - Descriptions and records of bees. - LXXIV. Annals and Magazine of Natural History 19: 282 - 290. https: // doi. org / 10.1080 / 00222931709486938" type="journal article" year="1917">Cockerell, 1917</bibRefCitation>
</taxonomicName>
. Kohat is today found in the state of 
<collectingCountry id="F3607632FFA0FF900B586FB0FB46FA4C" box="[1121,1224,1458,1484]" name="Pakistan" pageId="13" pageNumber="14">Pakistan</collectingCountry>
. Moreover, the material identified by Cockerell actually belongs to 
<taxonomicName id="4C774D21FFA0FF900C246FD4FC19FA70" baseAuthorityName="Wood" baseAuthorityYear="2024" box="[797,919,1494,1520]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fuscosa">
<emphasis id="B903EAB0FFA0FF900C246FD4FC19FA70" box="[797,919,1494,1520]" italics="true" pageId="13" pageNumber="14">A. fuscosa</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C774D21FFA0FF900CEE6FD4FA80FA6F" authorityName="Cockerell" authorityYear="1917" box="[983,1294,1493,1520]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="cussariensis" subSpecies="kohatensis">
<emphasis id="B903EAB0FFA0FF900CEE6FD4FA80FA6F" box="[983,1294,1493,1520]" italics="true" pageId="13" pageNumber="14">A. cussariensis kohatensis</emphasis>
</taxonomicName>
has now been synonymised with 
<taxonomicName id="4C774D21FFA0FF900EE26FFBFC86F993" authority="(Wood 2024 a)" baseAuthorityName="Wood" baseAuthorityYear="2024" box="[475,776,1529,1555]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fuscosa">
<emphasis id="B903EAB0FFA0FF900EE26FFBFDDBF993" box="[475,597,1529,1555]" italics="true" pageId="13" pageNumber="14">A. fuscosa</emphasis>
(
<bibRefCitation id="EFE64B53FFA0FF900D5D6FFBFD71F993" author="Wood T. J." box="[612,767,1529,1555]" pageId="13" pageNumber="14" pagination="1 - 150" refId="ref30750" refString="Wood T. J. 2024 a. Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae). Zootaxa 5483: 1 - 150. https: // doi. org / 10.11646 / zootaxa. 5483.1.1" type="journal article" year="2024">Wood 2024a</bibRefCitation>
)
</taxonomicName>
. Records from low-elevation areas of 
<collectingCountry id="F3607632FFA0FF900BF06FFBFA8BF993" box="[1225,1285,1529,1555]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
therefore would therefore seem most likely to be 
<taxonomicName id="4C774D21FFA0FF900D9B6C1EFC91F9B6" baseAuthorityName="Wood" baseAuthorityYear="2024" box="[674,799,1564,1590]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fuscosa">
<emphasis id="B903EAB0FFA0FF900D9B6C1EFC91F9B6" box="[674,799,1564,1590]" italics="true" pageId="13" pageNumber="14">A. fuscosa</emphasis>
</taxonomicName>
. It is impossible to say whether the records from Mussoorie refer to 
<taxonomicName id="4C774D21FFA0FF900E9A6C3DFCAEF9D9" authority="Wood &amp; Gautam" authorityName="Wood &amp; Gautam" authorityYear="2024" box="[419,800,1599,1625]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kedarnatha" status="sp. nov.">
<emphasis id="B903EAB0FFA0FF900E9A6C3DFDC5F9D9" box="[419,587,1599,1625]" italics="true" pageId="13" pageNumber="14">A. kedarnatha</emphasis>
Wood &amp; Gautam
</taxonomicName>
<taxonomicNameLabel id="A23057CBFFA0FF900C106C42FC08F9DA" box="[809,902,1600,1626]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
or not without examination of specimens (which are not precisely cited or referenced), and so the presence of 
<taxonomicName id="4C774D21FFA0FF900CFE6C61FBF9F9FC" authorityName="Morawitz" authorityYear="1886" box="[967,1143,1635,1661]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cussariensis">
<emphasis id="B903EAB0FFA0FF900CFE6C61FBF9F9FC" box="[967,1143,1635,1661]" italics="true" pageId="13" pageNumber="14">A. cussariensis</emphasis>
</taxonomicName>
in 
<collectingCountry id="F3607632FFA0FF900BA56C60FB57F9FC" box="[1180,1241,1634,1660]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
is considered to be unproven given its known global distribution and the risk of confusion with 
<taxonomicName id="4C774D21FFA0FF900B446C84FAABF91F" authorityName="Wood &amp; Gautam" authorityYear="2024" box="[1149,1317,1669,1696]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kedarnatha">
<emphasis id="B903EAB0FFA0FF900B446C84FAABF91F" box="[1149,1317,1669,1696]" italics="true" pageId="13" pageNumber="14">A. kedarnatha</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D6529FFA0FF900F846CC5FC04F88F" box="[189,906,1735,1808]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BC836A2FFA0FF900F846CC5FED6F961" blockId="13.[189,344,1735,1761]" box="[189,344,1735,1761]" pageId="13" pageNumber="14">
<heading id="D08081CEFFA0FF900F846CC5FED6F961" bold="true" box="[189,344,1735,1761]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B903EAB0FFA0FF900F846CC5FED6F961" bold="true" box="[189,344,1735,1761]" pageId="13" pageNumber="14">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A2FFA0FF900F846CF7FC04F88F" blockId="13.[189,906,1781,1808]" box="[189,906,1781,1808]" pageId="13" pageNumber="14">
Northern 
<collectingCountry id="F3607632FFA0FF900E176CF7FEE4F88F" box="[302,362,1781,1807]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
(Jammu &amp; Kashmir, 
<collectingRegion id="49B3F840FFA0FF900D516CF7FD75F88F" box="[616,763,1781,1807]" country="India" name="Uttarakhand" pageId="13" pageNumber="14">Uttarakhand</collectingRegion>
) and 
<collectingCountry id="F3607632FFA0FF900C056CF7FC08F88F" box="[828,902,1781,1807]" name="Nepal" pageId="13" pageNumber="14">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>