treatments-xml/data/03/DE/87/03DE87B0FFECFF90FE1BFD31A64EFD0C.xml

<document id="6A3A339DE9C06C67F3A7E1CCBA6DC94A" ID-DOI="10.5852/ejt.2024.919.2407" ID-ISSN="2118-9773" ID-Zenodo-Dep="10559520" ID-ZooBank="72B7907F-F901-4DDD-B2DD-63D03253837E" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1706085863884" checkinUser="plazi" docAuthor="Constant, Jérôme &amp; Pham, Hong Thai" docDate="2024" docId="03DE87B0FFECFF90FE1BFD31A64EFD0C" docLanguage="en" docName="EJT.2024.919.1-87.pdf" docOrigin="European Journal of Taxonomy 919 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2407/10617" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Melichergithus gravidus Constant &amp; Pham 2024, gen. et comb. nov." docType="treatment" docVersion="5" lastPageNumber="37" masterDocId="FFE7FFC8FFF7FFB4FFABFFD4A706FFDA" masterDocTitle="Sixteen issid planthopper species in one day in Dong Son-Ky Thuong Nature Reserve in North Vietnam: Eight new species, one new genus and additional new records (Hemiptera: Fulgoromorpha: Issidae)" masterLastPageNumber="87" masterPageNumber="1" pageNumber="28" updateTime="1706193695395" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E7E7AA930876BAD293AE62FE862CEC52">Sixteen issid planthopper species in one day in Dong Son-Ky Thuong Nature Reserve in North Vietnam: Eight new species, one new genus and additional new records (Hemiptera: Fulgoromorpha: Issidae)</mods:title>
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<mods:namePart id="B18F0360F26184E29A651D3263E487BF">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="56E54C8F100F3F74E2487EBFDDDEE91E">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier street 29, B- 1000 Brussels, Belgium.</mods:affiliation>
<mods:nameIdentifier id="157DD66E525939CF93214049AD638F8F" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:namePart id="D8D7A406AA7C8A248000DF470D35A5FB">Pham, Hong Thai</mods:namePart>
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<mods:affiliation id="46E79BEE5B5862FB985FB64FD1FD3577">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Street, Hanoi, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, Hanoi, Vietnam.</mods:affiliation>
<mods:nameIdentifier id="A07AABF52E33A856DDC8BC7F746392FB" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:date id="F246EC3B520D29BA19DC15322027DF84">2024</mods:date>
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<taxonomicName id="4C774D25FFECFFAFFE1BFD31A4A3FD25" authority="(Melichar, 1906)" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[432,933,741,768]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gravidus" status="gen. et comb. nov.">
<emphasis id="B903EAB4FFECFFAFFE1BFD31A5DEFCDA" bold="true" box="[432,728,741,768]" italics="true" pageId="27" pageNumber="28">Melichergithus gravidus</emphasis>
(
<bibRefCitation id="EFE64B57FFECFFAFFD4CFD31A49BFD25" author="Melichar L." box="[743,925,741,767]" pageId="27" pageNumber="28" pagination="1 - 327" refId="ref35004" refString="Melichar L. 1906. Monographie der Issiden (Homoptera). Abhandlungen der K. K. Zoologischbotanischen Gesellschaft in Wien 3: 1 - 327." type="journal article" year="1906">Melichar, 1906</bibRefCitation>
)
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<taxonomicNameLabel id="A23057CFFFECFFAFFC07FD32A382FCDA" box="[940,1156,741,768]" pageId="27" pageNumber="28" rank="species">gen. et comb. nov.</taxonomicNameLabel>
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<subSubSection id="C36D652DFFECFFAFFD62FCDCA46CFCF9" box="[713,874,776,803]" pageId="27" pageNumber="28" type="description">
<paragraph id="8BC836A6FFECFFAFFD62FCDCA46CFCF9" blockId="27.[432,1156,741,803]" box="[713,874,776,803]" pageId="27" pageNumber="28">
<figureCitation id="134C2A23FFECFFAFFD62FCDCA414FCF9" box="[713,786,776,803]" captionStart="Fig" captionStartId="2.[189,232,1998,2024]" captionTargetBox="[218,1369,265,1958]" captionTargetId="figure-18@2.[218,1369,265,1958]" captionTargetPageId="2" captionText="Fig. 1. Dong Son-Ky Thuong Nature Reserve. A. Topographic map. B. Sampled habitat." figureDoi="http://doi.org/10.5281/zenodo.10559522" httpUri="https://zenodo.org/record/10559522/files/figure.png" pageId="27" pageNumber="28">Figs 1</figureCitation>
, 
<figureCitation id="134C2A23FFECFFAFFC8AFCDDA46CFCF9" box="[801,874,777,803]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="29.[189,232,1928,1954]" captionStartId-1="30.[189,232,1928,1954]" captionStartId-2="31.[189,232,1892,1918]" captionStartId-3="32.[189,232,1928,1954]" captionStartId-4="33.[189,232,1857,1883]" captionStartId-5="34.[189,232,1857,1883]" captionStartId-6="35.[189,232,1928,1954]" captionTargetBox-0="[189,1398,360,1788]" captionTargetBox-1="[250,1340,271,1881]" captionTargetBox-2="[189,1393,487,1831]" captionTargetBox-3="[193,1386,305,1841]" captionTargetBox-4="[270,1316,275,1809]" captionTargetBox-5="[232,1356,277,1790]" captionTargetBox-6="[242,1346,265,1888]" captionTargetId-0="figure-11@29.[189,1398,360,1788]" captionTargetId-1="figure-18@30.[243,1344,265,1885]" captionTargetId-2="figure-63@31.[189,1398,481,1843]" captionTargetId-3="figure-18@32.[189,1398,295,1846]" captionTargetId-4="figure-12@33.[267,1321,265,1815]" captionTargetId-5="figure-18@34.[224,1362,265,1805]" captionTargetId-6="figure-12@35.[242,1346,265,1888]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionTargetPageId-2="31" captionTargetPageId-3="32" captionTargetPageId-4="33" captionTargetPageId-5="34" captionTargetPageId-6="35" captionText-0="Fig. 13. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., syntypes, ♀♀ (NHMW – photographs by H. Brückner). A–D. Syntype 1. A. Habitus, dorsal view. B. Habitus, frontal view. C. Habitus, lateral view. D. Labels. E–F. Syntype 2. E. Habitus, dorsal view. F. Labels." captionText-1="Fig. 14. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., dissected ♂, black morph from Dong Son-Ky Thuong Nature Reserve (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Right hind wing." captionText-2="Fig. 15. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., dissected ♂, typical morph from Ngoc Son-Ngo Luong Nature Reserve (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Anterolateral view. F. Right hind wing." captionText-3="Fig. 16. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., dissected ♂, typical morph from Mai Chau (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Anterolateral view. F. Right hind wing." captionText-4="Fig. 17. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, black morph from Dong Son- Ky Thuong Nature Reserve (RBINS), terminalia. A–C. Pygofer, anal tube and gonostyli. A. Lateral view. B. Caudal view. C. Dorsal view. D–I. Aedeagus and connective. D. Left lateral view. E. Right lateral view. F. Caudal view. G. Ventral view. H. Dorsal view. I. Anterodorsal view. Abbreviations: see Material and methods." captionText-5="Fig. 18. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, typical morph from Ngoc Son- Ngo Luong Nature Reserve (RBINS), terminalia.A–D. Pygofer, anal tube and gonostyli.A. Lateral view. B. Laterocaudal view. C. Dorsocaudal view. D. Dorsal view. E–J. Aedeagus and connective. E. Dorsal view. F. Left lateral view. G. Right lateral view. H. Left lateroventral view. I. Right laterocaudal view. J. Left laterocaudal view. Abbreviations: see Material and methods." captionText-6="Fig. 19. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov. A. Distribution map. B. Live specimen on fern, Xuan Lien Nature Reserve, 11 Jul. 2023. C. Live specimen on fern, Cuc Phuong National Park, 20 Jul. 2011. D. Habitat in Cuc Phuong National Park, 20 Jul. 2011." figureDoi-0="http://doi.org/10.5281/zenodo.10559560" figureDoi-1="http://doi.org/10.5281/zenodo.10559562" figureDoi-2="http://doi.org/10.5281/zenodo.10559566" figureDoi-3="http://doi.org/10.5281/zenodo.10559568" figureDoi-4="http://doi.org/10.5281/zenodo.10559570" figureDoi-5="http://doi.org/10.5281/zenodo.10559575" figureDoi-6="http://doi.org/10.5281/zenodo.10559577" httpUri-0="https://zenodo.org/record/10559560/files/figure.png" httpUri-1="https://zenodo.org/record/10559562/files/figure.png" httpUri-2="https://zenodo.org/record/10559566/files/figure.png" httpUri-3="https://zenodo.org/record/10559568/files/figure.png" httpUri-4="https://zenodo.org/record/10559570/files/figure.png" httpUri-5="https://zenodo.org/record/10559575/files/figure.png" httpUri-6="https://zenodo.org/record/10559577/files/figure.png" pageId="27" pageNumber="28">13–19</figureCitation>
</paragraph>
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<paragraph id="8BC836A6FFECFFAFFF16FC9EA37AFCBE" blockId="27.[189,1148,842,869]" box="[189,1148,842,869]" pageId="27" pageNumber="28">
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<treatmentCitation id="0AD610B7FFECFFAFFF16FC9EA583FCBF" author="Melichar L." box="[189,645,842,869]" page="59" pageId="27" pageNumber="28" year="1906">
<taxonomicName id="4C774D25FFECFFAFFF16FC9EA583FCBF" ID-CoL="3FWFR" authority="Melichar, 1906: 59" authorityName="Melichar" authorityPageNumber="59" authorityYear="1906" box="[189,645,842,869]" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gravidus">
<emphasis id="B903EAB4FFECFFAFFF16FC9EA6A6FCBE" box="[189,416,842,868]" italics="true" pageId="27" pageNumber="28">Gergithus gravidus</emphasis>
<bibRefCitation id="EFE64B57FFECFFAFFE0CFC9EA583FCBF" author="Melichar L." box="[423,645,842,869]" pageId="27" pageNumber="28" pagination="1 - 327" refId="ref35004" refString="Melichar L. 1906. Monographie der Issiden (Homoptera). Abhandlungen der K. K. Zoologischbotanischen Gesellschaft in Wien 3: 1 - 327." type="journal article" year="1906">Melichar, 1906: 59</bibRefCitation>
</taxonomicName>
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(keyed), 61 (described). 
<typeStatus id="54CC8804FFECFFAFFC07FC9EA4F4FCBE" box="[940,1010,842,868]" pageId="27" pageNumber="28">Types</typeStatus>
in NHMW.
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</paragraph>
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<treatmentCitationGroup id="AB671188FFECFFAFFF16FC5FA3CBFBE8" pageId="27" pageNumber="28">
<taxonomicName id="4C774D25FFECFFAFFF16FC5FA6A4FC7F" ID-CoL="3FWFR" authorityName="Melichar" authorityYear="1906" box="[189,418,907,933]" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gravidus">
<emphasis id="B903EAB4FFECFFAFFF16FC5FA6A4FC7F" box="[189,418,907,933]" italics="true" pageId="27" pageNumber="28">Gergithus gravidus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AD610B7FFECFFAFFE6FFC5FA5B3FC7F" author="Lallemand V." box="[452,693,907,933]" page="70" pageId="27" pageNumber="28" year="1942">
<bibRefCitation id="EFE64B57FFECFFAFFE6FFC5FA5B3FC7F" author="Lallemand V." box="[452,693,907,933]" pageId="27" pageNumber="28" pagination="69 - 77" refId="ref34845" refString="Lallemand V. 1942. Notes sur quelques especes recueillies par le R. Piel (Musee Heude, Shanghai) et le R. P. de Cooman (Hoa Binh, Tonkin). Notes d'Entomologie chinoise. Musee Heude. Chang-Hai 9 (4): 69 - 77." type="journal article" year="1942">Lallemand 1942: 70</bibRefCitation>
</treatmentCitation>
(listed). — 
<treatmentCitation id="0AD610B7FFECFFAFFCE0FC5FA322FC7F" author="Fennah R. G." box="[843,1060,907,933]" page="267" pageId="27" pageNumber="28" year="1978">
<bibRefCitation id="EFE64B57FFECFFAFFCE0FC5FA322FC7F" author="Fennah R. G." box="[843,1060,907,933]" pageId="27" pageNumber="28" pagination="207 - 279" refId="ref33692" refString="Fennah R. G. 1978. Fulgoroidea (Homoptera) from Vietnam. Annales Zoologici 34 (9): 207 - 279." type="journal article" year="1978">Fennah 1978: 267</bibRefCitation>
</treatmentCitation>
(recorded from Cuc Phong, Ninh Binh Province). — 
<treatmentCitation id="0AD610B7FFECFFAFFDBFFC7AA403FC12" author="Che Y. &amp; Zhang Y. - L. &amp; Wang Y. - L." box="[532,773,942,969]" page="612" pageId="27" pageNumber="28" year="2007">
<bibRefCitation id="EFE64B57FFECFFAFFDBFFC7AA403FC12" author="Che Y. &amp; Zhang Y. - L. &amp; Wang Y. - L." box="[532,773,942,969]" pageId="27" pageNumber="28" pagination="611 - 627" refId="ref32906" refString="Che Y. - L, Zhang Y. - L. &amp; Wang Y. - L. 2007. Seven new species and one new record of Gergithus Stal (Hemiptera: Fulgoroidea: Issidae) from China. Proceedings of the Entomological Society of Washington 109 (3): 611 - 627." type="journal article" year="2007">
Che 
<emphasis id="B903EAB4FFECFFAFFDE7FC7BA579FC12" box="[588,639,942,968]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. 2007: 612
</bibRefCitation>
</treatmentCitation>
(recorded from 
<collectingCountry id="F3607636FFECFFAFFC6EFC7AA308FC12" box="[965,1038,942,968]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
, 
<collectingRegion id="49B3F844FFECFFAFFBB0FC7AA385FC12" box="[1051,1155,942,968]" country="China" name="Guangxi" pageId="27" pageNumber="28">Guangxi</collectingRegion>
, described), figs 1–4 (head, thorax, wings). — 
<treatmentCitation id="0AD610B7FFECFFAFFD8DFC06A445FC36" author="Meng R. &amp; Wang Y. - L." box="[550,835,978,1004]" page="3" pageId="27" pageNumber="28" year="2012">
<bibRefCitation id="EFE64B57FFECFFAFFD8DFC06A445FC36" author="Meng R. &amp; Wang Y. - L." box="[550,835,978,1004]" pageId="27" pageNumber="28" pagination="1 - 18" refId="ref35032" refString="Meng R. &amp; Wang Y. - L. 2012. Two new species of the genus Gergithus Stal (Hemiptera: Fulgoromorpha: Issidae) from China, with a redescription of G. bimaculatus Zhang and Che, and G. tessellatus Matsumura. Zootaxa 3247: 1 - 18. https: // doi. org / 10.11646 / zootaxa. 3247.1.1" type="journal article" year="2012">Meng &amp; Wang 2012: 3</bibRefCitation>
</treatmentCitation>
(keyed). — 
<treatmentCitation id="0AD610B7FFECFFAFFC49FC06A270FC36" author="Gnezdilov V. M. &amp; Constant J." box="[994,1398,977,1004]" page="573" pageId="27" pageNumber="28" year="2012">
<bibRefCitation id="EFE64B57FFECFFAFFC49FC06A270FC36" author="Gnezdilov V. M. &amp; Constant J." box="[994,1398,977,1004]" pageId="27" pageNumber="28" pagination="571 - 576" refId="ref34367" refString="Gnezdilov V. M. &amp; Constant J. 2012. Review of the family Issidae (Hemiptera: Fulgoromorpha) in Vietnam with description of a new species. Annales Zoologici 62 (4): 571 - 576. https: // doi. org / 10.3161 / 000345412 X 659632" type="journal article" year="2012">Gnezdilov &amp; Constant 2012: 573</bibRefCitation>
</treatmentCitation>
(listed from 
<collectingCountry id="F3607636FFECFFAFFED1FC21A6E5FBD5" box="[378,483,1013,1039]" name="Vietnam" pageId="27" pageNumber="28">Vietnam</collectingCountry>
). — 
<treatmentCitation id="0AD610B7FFECFFAFFDB2FC21A400FBCA" author="Chen X. - S. &amp; Zhang Z. - G. &amp; Chang Z. - M." box="[537,774,1013,1040]" page="52" pageId="27" pageNumber="28" year="2014">
<bibRefCitation id="EFE64B57FFECFFAFFDB2FC21A400FBCA" author="Chen X. - S. &amp; Zhang Z. - G. &amp; Chang Z. - M." box="[537,774,1013,1040]" pageId="27" pageNumber="28" refId="ref32963" refString="Chen X. - S., Zhang Z. - G. &amp; Chang Z. - M. 2014. Issidae and Caliscelidae from China. Guizhou Science and Technology Publishing House, Guiyang." type="book" year="2014">
Chen 
<emphasis id="B903EAB4FFECFFAFFDF6FC22A596FBD5" box="[605,656,1013,1039]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. 2014: 52
</bibRefCitation>
</treatmentCitation>
(described, recorded from 
<collectingCountry id="F3607636FFECFFAFFBE9FC21A38DFBD5" box="[1090,1163,1013,1039]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
, 
<collectingRegion id="49B3F844FFECFFAFFB3CFC21A207FBD5" box="[1175,1281,1013,1039]" country="China" name="Guangxi" pageId="27" pageNumber="28">Guangxi</collectingRegion>
), fig. 2-15 (specimen, details, terminalia). — 
<treatmentCitation id="0AD610B7FFECFFAFFD23FBCCA4BCFBE8" author="Gnezdilov V. M. &amp; Bourgoin T. &amp; Soulier-Perkins A." box="[648,954,1048,1075]" page="82" pageId="27" pageNumber="28" year="2014">
<bibRefCitation id="EFE64B57FFECFFAFFD23FBCCA4BCFBE8" author="Gnezdilov V. M. &amp; Bourgoin T. &amp; Soulier-Perkins A." box="[648,954,1048,1075]" pageId="27" pageNumber="28" pagination="80 - 96" refId="ref34543" refString="Gnezdilov V. M., Bourgoin T. &amp; Soulier-Perkins A. 2014. Vietnamese Issidae (Hemiptera, Fulgoroidea): new taxa, new records and new distribution data. Zootaxa 3847 (1): 80 - 96. https: // doi. org / 10.11646 / zootaxa. 3847.1.4" type="journal article" year="2014">
Gnezdilov 
<emphasis id="B903EAB4FFECFFAFFCA5FBCDA447FBE8" box="[782,833,1048,1074]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. 2014: 82
</bibRefCitation>
</treatmentCitation>
(listed from 
<collectingCountry id="F3607636FFECFFAFFBF3FBCCA3C7FBE8" box="[1112,1217,1048,1074]" name="Vietnam" pageId="27" pageNumber="28">Vietnam</collectingCountry>
).
</treatmentCitationGroup>
<treatmentCitationGroup id="AB671188FFECFFAFFB7DFBCDA32DFB8F" pageId="27" pageNumber="28">
— 
<bibRefCitation id="EFE64B57FFECFFAFFB57FBCCA633FB8F" author="Gnezdilov V. M." pageId="27" pageNumber="28" pagination="1338 - 1352" refId="ref34143" refString="Gnezdilov V. M. 2017 b. Addenda to the revisions of the genera Gergithus Stal and Hemisphaerius Schaum (Hemiptera, Auchenorrhyncha, Fulgoroidea: Issidae). Entomological Review 97 (9): 1338 - 1352. https: // doi. org / 10.1134 / S 0013873817090123" type="journal article" year="2017">Gnezdilov 2017b</bibRefCitation>
(transferred back from 
<taxonomicName id="4C774D25FFECFFAFFDE9FBEFA321FB8F" ID-CoL="4PML" authority="Meng, Webb &amp; Wang, 2017" authorityName="Meng, Webb &amp; Wang" authorityYear="2017" box="[578,1063,1083,1110]" class="Insecta" family="Issidae" genus="Gnezdilovius" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAB4FFECFFAFFDE9FBEFA5DDFB8F" box="[578,731,1083,1109]" italics="true" pageId="27" pageNumber="28">Gnezdilovius</emphasis>
<bibRefCitation id="EFE64B57FFECFFAFFD49FBE8A321FB8F" author="Meng R. &amp; Webb M. D. &amp; Wang Y. - L." box="[738,1063,1083,1110]" pageId="27" pageNumber="28" pagination="1 - 25" refId="ref35158" refString="Meng R., Webb M. D. &amp; Wang Y. - L. 2017. Nomenclatural changes in the planthopper tribe Hemisphaeriini (Hemiptera: Fulgoromorpha: Issidae), with the description of a new genus and a new species. European Journal of Taxonomy 298: 1 - 25. https: // doi. org / 10.5852 / ejt. 2017.298" type="journal article" year="2017">Meng, Webb &amp; Wang, 2017</bibRefCitation>
</taxonomicName>
,
</treatmentCitationGroup>
compared to representatives of 
<taxonomicName id="4C774D25FFECFFAFFEA1FB8AA5DDFBA2" ID-CoL="5JTW" authority="Constant &amp; Pham, 2016" authorityName="Constant &amp; Pham" authorityYear="2016" box="[266,731,1118,1145]" class="Insecta" family="Issidae" genus="Maculergithus" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAB4FFECFFAFFEA1FB8AA6B2FBA2" box="[266,436,1118,1144]" italics="true" pageId="27" pageNumber="28">Maculergithus</emphasis>
<bibRefCitation id="EFE64B57FFECFFAFFE10FB8BA5DDFBA2" author="Constant J. &amp; Pham H. T." box="[443,731,1118,1145]" pageId="27" pageNumber="28" pagination="1 - 16" refId="ref33300" refString="Constant J. &amp; Pham H. T. 2016. Maculergithus, a new subgenus in Gergithus Schumacher, 1915 with two new species from northern Vietnam (Hemiptera: Fulgoromorpha: Issidae). European Journal of Taxonomy 198: 1 - 16. https: // doi. org / 10.5852 / ejt. 2016.198" type="journal article" year="2016">Constant &amp; Pham, 2016</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC836A6FFECFFAFFF16FB56A32DFB38" blockId="27.[189,1399,907,1251]" pageId="27" pageNumber="28">
<taxonomicName id="4C774D25FFECFFAFFF16FB56A6CFFB46" ID-CoL="3GMDL" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[189,457,1154,1180]" class="Insecta" family="Issidae" genus="Gnezdilovius" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gravidus">
<emphasis id="B903EAB4FFECFFAFFF16FB56A6CFFB46" box="[189,457,1154,1180]" italics="true" pageId="27" pageNumber="28">Gnezdilovius gravidus</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EFE64B57FFECFFAFFE45FB56A5F5FB46" author="Meng R. &amp; Webb M. D. &amp; Wang Y. - L." box="[494,755,1154,1180]" pageId="27" pageNumber="28" pagination="1 - 25" refId="ref35158" refString="Meng R., Webb M. D. &amp; Wang Y. - L. 2017. Nomenclatural changes in the planthopper tribe Hemisphaeriini (Hemiptera: Fulgoromorpha: Issidae), with the description of a new genus and a new species. European Journal of Taxonomy 298: 1 - 25. https: // doi. org / 10.5852 / ejt. 2017.298" type="journal article" year="2017">
Meng 
<emphasis id="B903EAB4FFECFFAFFD96FB56A573FB46" box="[573,629,1154,1180]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. 2017: 18
</bibRefCitation>
(transferred to 
<taxonomicName id="4C774D25FFECFFAFFC1EFB56A348FB46" ID-CoL="4PML" authorityName="Meng, Webb &amp; Wang" authorityYear="2017" box="[949,1102,1154,1180]" class="Insecta" family="Issidae" genus="Gnezdilovius" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAB4FFECFFAFFC1EFB56A348FB46" box="[949,1102,1154,1180]" italics="true" pageId="27" pageNumber="28">Gnezdilovius</emphasis>
</taxonomicName>
). — 
<bibRefCitation id="EFE64B57FFECFFAFFB38FB56A610FB65" author="Zhang Y. - L. &amp; Che Y. - L. &amp; Meng R. &amp; Wang Y. - L." pageId="27" pageNumber="28" refId="ref36146" refString="Zhang Y. - L., Che Y. - L., Meng R. &amp; Wang Y. - L. 2020. Hemiptera. Caliscelidae. Issidae. Insecta. Fauna Sinica. Vol. 70. Science Press, Beijing." type="book" year="2020">
Zhang 
<emphasis id="B903EAB4FFECFFAFFB41FB56A227FB46" box="[1258,1313,1154,1180]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. 2020: 204
</bibRefCitation>
(keyed in Chinese, transferred back from 
<taxonomicName id="4C774D25FFECFFAFFCA7FB71A479FB65" ID-CoL="4N6H" authorityName="Stål" authorityYear="1870" box="[780,895,1189,1215]" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAB4FFECFFAFFCA7FB71A479FB65" box="[780,895,1189,1215]" italics="true" pageId="27" pageNumber="28">Gergithus</emphasis>
</taxonomicName>
), 219 (described), 547 (keyed in English), fig. 72 (head, thorax, wings, female terminalia), pl. XVI, a–c (habitus).
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFECFFAFFF16FADDA4E0F9FA" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8BC836A6FFECFFAFFF16FADDA630FAF9" blockId="27.[189,310,1289,1315]" box="[189,310,1289,1315]" pageId="27" pageNumber="28">
<heading id="D08081CAFFECFFAFFF16FADDA630FAF9" bold="true" box="[189,310,1289,1315]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B903EAB4FFECFFAFFF16FADDA630FAF9" bold="true" box="[189,310,1289,1315]" pageId="27" pageNumber="28">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFECFFAFFF16FAECA6E6FAAC" blockId="27.[189,1398,1336,1398]" pageId="27" pageNumber="28">
<taxonomicName id="4C774D25FFECFFAFFF16FAECA5C5FA88" authority="(Melichar, 1906)" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[189,707,1336,1362]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gravidus" status="gen. et comb. nov.">
<emphasis id="B903EAB4FFECFFAFFF16FAECA6E3FA88" box="[189,485,1336,1362]" italics="true" pageId="27" pageNumber="28">Melichergithus gravidus</emphasis>
(
<bibRefCitation id="EFE64B57FFECFFAFFE56FAECA5BDFA88" author="Melichar L." box="[509,699,1336,1362]" pageId="27" pageNumber="28" pagination="1 - 327" refId="ref35004" refString="Melichar L. 1906. Monographie der Issiden (Homoptera). Abhandlungen der K. K. Zoologischbotanischen Gesellschaft in Wien 3: 1 - 327." type="journal article" year="1906">Melichar, 1906</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFECFFAFFD78FAEDA4C0FA89" box="[723,966,1336,1363]" pageId="27" pageNumber="28" rank="species">gen. et comb. nov.</taxonomicNameLabel>
is the only species in the genus 
<taxonomicName id="4C774D25FFECFFAFFF16FA8FA668FAAF" authority="Constant &amp; Pham, 2024" authorityName="Constant &amp; Pham" authorityYear="2024" box="[189,366,1371,1397]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B903EAB4FFECFFAFFF16FA8FA668FAAF" box="[189,366,1371,1397]" italics="true" pageId="27" pageNumber="28">Melichergithus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFECFFAFFEDEFA88A6E6FAAC" box="[373,480,1372,1398]" pageId="27" pageNumber="28" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC836A6FFECFFAFFF16FA49A4E0F9FA" blockId="27.[189,1400,1437,1569]" pageId="27" pageNumber="28">
The characters of the male terminalia (
<figureCitation id="134C2A23FFECFFAFFD20FA49A415FA6D" box="[651,787,1437,1463]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="33.[189,232,1857,1883]" captionStartId-1="34.[189,232,1857,1883]" captionTargetBox-0="[270,1316,275,1809]" captionTargetBox-1="[232,1356,277,1790]" captionTargetId-0="figure-12@33.[267,1321,265,1815]" captionTargetId-1="figure-18@34.[224,1362,265,1805]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionText-0="Fig. 17. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, black morph from Dong Son- Ky Thuong Nature Reserve (RBINS), terminalia. A–C. Pygofer, anal tube and gonostyli. A. Lateral view. B. Caudal view. C. Dorsal view. D–I. Aedeagus and connective. D. Left lateral view. E. Right lateral view. F. Caudal view. G. Ventral view. H. Dorsal view. I. Anterodorsal view. Abbreviations: see Material and methods." captionText-1="Fig. 18. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, typical morph from Ngoc Son- Ngo Luong Nature Reserve (RBINS), terminalia.A–D. Pygofer, anal tube and gonostyli.A. Lateral view. B. Laterocaudal view. C. Dorsocaudal view. D. Dorsal view. E–J. Aedeagus and connective. E. Dorsal view. F. Left lateral view. G. Right lateral view. H. Left lateroventral view. I. Right laterocaudal view. J. Left laterocaudal view. Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.10559570" figureDoi-1="http://doi.org/10.5281/zenodo.10559575" httpUri-0="https://zenodo.org/record/10559570/files/figure.png" httpUri-1="https://zenodo.org/record/10559575/files/figure.png" pageId="27" pageNumber="28">Figs 17–18</figureCitation>
) must be used for species recognition, for example the elongate, laminate lateroventral processes of the periandrium, the long sinuate shaft-shaped lateral processes of the periandrium, the long lateral spinose processes of the ventral lobe of the periandrium curved cephalad or the more or less round anal tube (in dorsal view).
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFECFFA8FF16F993A4B3F80D" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BC836A6FFECFFAFFF16F993A6ACF9BB" blockId="27.[189,426,1607,1633]" box="[189,426,1607,1633]" pageId="27" pageNumber="28">
<heading id="D08081CAFFECFFAFFF16F993A6ACF9BB" bold="true" box="[189,426,1607,1633]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B903EAB4FFECFFAFFF16F993A6ACF9BB" bold="true" box="[189,426,1607,1633]" pageId="27" pageNumber="28">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B1F3CFBFFECFFAFFF41F9A2A4F8F920" collectionCode="NHMW" collectorName="H. Fruhstorfer" country="Vietnam" determinerName="Melichar" latitude="21.838057" location="Mount Mau Son" longLatPrecision="21" longitude="106.91361" pageId="27" pageNumber="28" specimenCount="2" specimenCount-female="2" stateProvince="Lang Son Province" typeStatus="syntype">
<paragraph id="8BC836A6FFECFFAFFF41F9A2A65CF94A" blockId="27.[189,1399,1654,1786]" box="[234,346,1654,1680]" pageId="27" pageNumber="28">
<emphasis id="B903EAB4FFECFFAFFF41F9A2A65CF94A" bold="true" box="[234,346,1654,1680]" pageId="27" pageNumber="28">
<typeStatus id="54CC8804FFECFFAFFF41F9A2A65CF94A" box="[234,346,1654,1680]" pageId="27" pageNumber="28" type="syntype">Syntypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC836A6FFECFFAFFF16F94EA304F920" blockId="27.[189,1399,1654,1786]" pageId="27" pageNumber="28">
<collectingCountry id="F3607636FFECFFAFFF16F94EA642F96E" box="[189,324,1690,1716]" name="Vietnam" pageId="27" pageNumber="28">VIETNAM</collectingCountry>
• 
<specimenCount id="9D71FD2FFFECFFAFFEC9F94EA6A1F96E" box="[354,423,1690,1716]" count="2" pageId="27" pageNumber="28" type="female">2 ♀♀</specimenCount>
(examined from photographs – 
<figureCitation id="134C2A23FFECFFAFFC86F94DA483F96E" box="[813,901,1689,1716]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,360,1788]" captionTargetId="figure-11@29.[189,1398,360,1788]" captionTargetPageId="29" captionText="Fig. 13. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., syntypes, ♀♀ (NHMW – photographs by H. Brückner). A–D. Syntype 1. A. Habitus, dorsal view. B. Habitus, frontal view. C. Habitus, lateral view. D. Labels. E–F. Syntype 2. E. Habitus, dorsal view. F. Labels." figureDoi="http://doi.org/10.5281/zenodo.10559560" httpUri="https://zenodo.org/record/10559560/files/figure.png" pageId="27" pageNumber="28">Fig. 13</figureCitation>
); [
<collectingRegion id="49B3F844FFECFFAFFC0EF94EA392F969" box="[933,1172,1689,1716]" country="Vietnam" name="Lang Son" pageId="27" pageNumber="28">Lang Son Province</collectingRegion>
], 
<location id="8EA8607DFFECFFAFFB02F94EA274F96E" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFECFFAFFB02F94EA274F96E" box="[1193,1394,1690,1716]" country="Vietnam" latitude="21.838057" longLatPrecision="21" longitude="106.91361" name="Mount Mau Son" pageId="27" pageNumber="28" stateProvince="Lang Son Province">Mount Mau Son</location>
; [
<geoCoordinate id="EE435061FFECFFAFFF6FF96FA658F90D" box="[196,350,1723,1751]" degrees="21" direction="north" minutes="50" orientation="latitude" pageId="27" pageNumber="28" precision="15" seconds="17" value="21.838057">21°50′17″ N</geoCoordinate>
, 
<geoCoordinate id="EE435061FFECFFAFFEC0F96FA509F90D" box="[363,527,1723,1751]" degrees="106" direction="east" minutes="54" orientation="longitude" pageId="27" pageNumber="28" precision="15" seconds="49" value="106.91361">106°54′49″ E</geoCoordinate>
]; Apr.–May [1900]; 
<collectorName id="26825370FFECFFAFFCAFF969A4ABF90D" box="[772,941,1725,1751]" pageId="27" pageNumber="28">H. Fruhstorfer</collectorName>
leg.; “Tonkin, Montes Mauson; April, 
<date id="FFC91066FFECFFAFFF16F934A605F920" box="[189,259,1760,1786]" pageId="27" pageNumber="28">Mai 2</date>
-3000’, 
<collectorName id="26825370FFECFFAFFEF4F934A50BF920" box="[351,525,1760,1786]" pageId="27" pageNumber="28">H. Fruhstorfer</collectorName>
”, 
<taxonomicName id="4C774D25FFECFFAFFD88F934A591F920" authority="Constant &amp; Pham, 2024" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[547,663,1760,1786]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gravidus" status="sp. nov.">
“ 
<emphasis id="B903EAB4FFECFFAFFD9BF934A591F920" box="[560,663,1760,1786]" italics="true" pageId="27" pageNumber="28">gravidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFECFFAFFD35F934A5DAF920" box="[670,732,1760,1786]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
det. 
<determinerName id="7E14DA62FFECFFAFFCBDF934A480F920" box="[790,902,1760,1786]" pageId="27" pageNumber="28">Melichar</determinerName>
”; 
<collectionCode id="ED66AE63FFECFFAFFC36F934A4F8F920" box="[925,1022,1760,1786]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="27" pageNumber="28" type="Museum">NHMW</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8BC836A6FFECFFAFFF41F8F5A6E7F8E1" blockId="27.[189,1399,1825,1993]" box="[234,481,1825,1851]" pageId="27" pageNumber="28">
<emphasis id="B903EAB4FFECFFAFFF41F8F5A6E7F8E1" bold="true" box="[234,481,1825,1851]" pageId="27" pageNumber="28">Additional material</emphasis>
</paragraph>
<paragraph id="8BC836A6FFECFFA8FF16F891A242FCA0" blockId="27.[189,1399,1825,1993]" lastBlockId="28.[188,1399,265,890]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<materialsCitation id="3B1F3CFBFFECFFAFFF16F891A4C3F87F" collectingDate="2022-08-27" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel &amp; L. Semeraro" country="Vietnam" elevation="550" latitude="21.141388" location="Dong Son-Ky Thuong Nature Reserve" longLatPrecision="21" longitude="107.08139" municipality="Dong" pageId="27" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Quang Ninh Province">
<collectingCountry id="F3607636FFECFFAFFF16F891A642F885" box="[189,324,1861,1887]" name="Vietnam" pageId="27" pageNumber="28">VIETNAM</collectingCountry>
• 
<specimenCount id="9D71FD2FFFECFFAFFECAF891A689F885" box="[353,399,1861,1887]" count="1" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
(dissected, 
<figureCitation id="134C2A23FFECFFAFFDB5F890A57CF885" box="[542,634,1860,1887]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[250,1340,271,1881]" captionTargetId="figure-18@30.[243,1344,265,1885]" captionTargetPageId="30" captionText="Fig. 14. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., dissected ♂, black morph from Dong Son-Ky Thuong Nature Reserve (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559562" httpUri="https://zenodo.org/record/10559562/files/figure.png" pageId="27" pageNumber="28">Figs 14</figureCitation>
, 
<figureCitation id="134C2A23FFECFFAFFD23F891A5AEF885" box="[648,680,1861,1887]" captionStart="Fig" captionStartId="33.[189,232,1857,1883]" captionTargetBox="[270,1316,275,1809]" captionTargetId="figure-12@33.[267,1321,265,1815]" captionTargetPageId="33" captionText="Fig. 17. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, black morph from Dong Son- Ky Thuong Nature Reserve (RBINS), terminalia. A–C. Pygofer, anal tube and gonostyli. A. Lateral view. B. Caudal view. C. Dorsal view. D–I. Aedeagus and connective. D. Left lateral view. E. Right lateral view. F. Caudal view. G. Ventral view. H. Dorsal view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559570" httpUri="https://zenodo.org/record/10559570/files/figure.png" pageId="27" pageNumber="28">17</figureCitation>
), 
<specimenCount id="9D71FD2FFFECFFAFFD6BF891A5E9F885" box="[704,751,1861,1887]" count="1" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
; 
<collectingRegion id="49B3F844FFECFFAFFD54F891A300F884" box="[767,1030,1860,1887]" country="Vietnam" name="Quang Ninh" pageId="27" pageNumber="28">Quang Ninh Province</collectingRegion>
, 
<location id="8EA8607DFFECFFAFFBBFF891A619F858" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFECFFAFFBBFF891A619F858" country="Vietnam" latitude="21.141388" longLatPrecision="21" longitude="107.08139" municipality="Dong" name="Dong Son-Ky Thuong Nature Reserve" pageId="27" pageNumber="28" stateProvince="Quang Ninh Province">Dong Son-Ky Thuong Nature Reserve</location>
; 
<geoCoordinate id="EE435061FFECFFAFFE9AF8B2A6CAF858" box="[305,460,1894,1922]" degrees="21" direction="north" minutes="08" orientation="latitude" pageId="27" pageNumber="28" precision="15" seconds="29" value="21.141388">21°08′29″ N</geoCoordinate>
, 
<geoCoordinate id="EE435061FFECFFAFFE75F8B2A583F858" box="[478,645,1894,1922]" degrees="107" direction="east" minutes="04" orientation="longitude" pageId="27" pageNumber="28" precision="15" seconds="53" value="107.08139">107°04′53″ E</geoCoordinate>
; 
<date id="FFC91066FFECFFAFFD33F8BCA440F858" box="[664,838,1896,1922]" pageId="27" pageNumber="28" value="2022-08-27">
<collectingDate id="EF8DE98EFFECFFAFFD33F8BCA440F858" box="[664,838,1896,1922]" pageId="27" pageNumber="28" value="2022-08-27">27 Aug. 2022</collectingDate>
</date>
; 
<elevation id="005AD195FFECFFAFFCF3F8B3A4EAF858" box="[856,1004,1895,1922]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="27" pageNumber="28" unit="m" value="550.0">
<quantity id="4C8F9B43FFECFFAFFCF3F8B3A4AEF858" box="[856,936,1895,1922]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="27" pageNumber="28" unit="m" value="550.0">550 m</quantity>
a.s.l.
</elevation>
; secondary forest; GTI Project; 
<collectorName id="26825370FFECFFAFFF16F85FA647F87F" box="[189,321,1931,1957]" pageId="27" pageNumber="28">J. Constant</collectorName>
, 
<collectorName id="26825370FFECFFAFFEE6F85FA6CFF87F" box="[333,457,1931,1957]" pageId="27" pageNumber="28">J. Bresseel</collectorName>
and 
<collectorName id="26825370FFECFFAFFDA9F85FA594F87F" box="[514,658,1931,1957]" pageId="27" pageNumber="28">L. Semeraro</collectorName>
leg.; I.G.: 34.518; 
<collectionCode id="ED66AE63FFECFFAFFCC4F85FA4C3F87F" box="[879,965,1931,1957]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="27" pageNumber="28" type="Museum">RBINS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFECFFA8FC76F85FA582FEF9" collectingDate="1939-07" collectionCode="RBINS" collectorName="A. de Cooman" country="Vietnam" determinerName="V. Lallemand" lastPageId="28" lastPageNumber="29" latitude="20.813334" location="Tonkin" longLatPrecision="21" longitude="105.33833" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" stateProvince="Hoa Binh">
<specimenCount id="9D71FD2FFFECFFAFFC76F85FA30FF87F" box="[989,1033,1931,1957]" count="1" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
(dissected); 
<location id="8EA8607DFFECFFAFFB36F85FA3F5F87F" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFECFFAFFB36F85FA3F5F87F" box="[1181,1267,1931,1957]" country="Vietnam" latitude="20.813334" longLatPrecision="21" longitude="105.33833" name="Tonkin" pageId="27" pageNumber="28" stateProvince="Hoa Binh">Tonkin</location>
, 
<collectingRegion id="49B3F844FFECFFAFFB55F85FA274F87F" box="[1278,1394,1931,1957]" country="Vietnam" name="Hoa Binh" pageId="27" pageNumber="28">Hoa Binh</collectingRegion>
; [
<geoCoordinate id="EE435061FFECFFAFFF6FF879A65DF813" box="[196,347,1965,1993]" degrees="20" direction="north" minutes="48" orientation="latitude" pageId="27" pageNumber="28" precision="15" seconds="48" value="20.813334">20°48′48″ N</geoCoordinate>
, 
<geoCoordinate id="EE435061FFECFFAFFECDF879A501F813" box="[358,519,1965,1993]" degrees="105" direction="east" minutes="20" orientation="longitude" pageId="27" pageNumber="28" precision="15" seconds="18" value="105.33833">105°20′18″ E</geoCoordinate>
]; 
<date id="FFC91066FFECFFAFFDB0F87AA58FF812" box="[539,649,1966,1992]" pageId="27" pageNumber="28" value="1939-07">
<collectingDate id="EF8DE98EFFECFFAFFDB0F87AA58FF812" box="[539,649,1966,1992]" pageId="27" pageNumber="28" value="1939-07">Jul. 1939</collectingDate>
</date>
; 
<collectorName id="26825370FFECFFAFFD38F87AA43CF812" box="[659,826,1966,1992]" pageId="27" pageNumber="28">A. de Cooman</collectorName>
leg.; R.I.Sc.N.B. I.G.18.385; 
<taxonomicName id="4C774D25FFECFFA8FB3EF87AA7F2FEF9" authority="Mel." authorityName="Melichar" authorityYear="1906" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" lastPageId="28" lastPageNumber="29" order="Hemiptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gravidus">
<emphasis id="B903EAB4FFECFFAFFB3EF87AA271F812" box="[1173,1399,1966,1992]" italics="true" pageId="27" pageNumber="28">Gergithus gravidus</emphasis>
Mel.
</taxonomicName>
det. 
<determinerName id="7E14DA62FFEBFFA8FE9FFEDEA6D3FEF9" box="[308,469,265,292]" pageId="28" pageNumber="29">V. Lallemand</determinerName>
1940; 
<collectionCode id="ED66AE63FFEBFFA8FD85FEDDA582FEF9" box="[558,644,265,291]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="28" pageNumber="29" type="Museum">RBINS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FD0EFEDDA5CEFE9C" collectionCode="RBINS" collectorName="R. I. Sc." country="Vietnam" determinerName="H. Synave" location="Tonkin" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Hoa Binh">
<specimenCount id="9D71FD2FFFEBFFA8FD0EFEDDA5D3FEFE" box="[677,725,265,292]" count="1" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; 
<location id="8EA8607DFFEBFFA8FD4CFEDDA43BFEF9" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFEBFFA8FD4CFEDDA43BFEF9" box="[743,829,265,291]" country="Vietnam" name="Tonkin" pageId="28" pageNumber="29" stateProvince="Hoa Binh">Tonkin</location>
, 
<collectingRegion id="49B3F844FFEBFFA8FCE7FEDEA4C3FEF9" box="[844,965,265,292]" country="Vietnam" name="Hoa Binh" pageId="28" pageNumber="29">Hoa Binh</collectingRegion>
; R.I.Sc.N.B. I.G.23.285; 
<determinerName id="7E14DA62FFEBFFA8FB52FEDEA270FEF9" box="[1273,1398,265,292]" pageId="28" pageNumber="29">H. Synave</determinerName>
det. 195. 
<taxonomicName id="4C774D25FFEBFFA8FE9FFEF8A567FE9C" authority="Mel." authorityName="Melichar" authorityYear="1906" box="[308,609,300,326]" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="gravidus">
<emphasis id="B903EAB4FFEBFFA8FE9FFEF8A51DFE9C" box="[308,539,300,326]" italics="true" pageId="28" pageNumber="29">Gergithus gravidus</emphasis>
Mel.
</taxonomicName>
; 
<collectionCode id="ED66AE63FFEBFFA8FDD9FEF8A5CEFE9C" box="[626,712,300,326]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="28" pageNumber="29" type="Museum">RBINS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FD40FEF8A4F2FEB3" collectionCode="MNHN" collectorName="A. de Cooman" country="Vietnam" location="Tonkin" pageId="28" pageNumber="29" specimenCode="EH24639, 24640, 24643, 24644" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Hoa Binh">
<specimenCount id="9D71FD2FFFEBFFA8FD40FEF8A41AFE9D" box="[747,796,300,327]" count="1" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
(dissected), 
<specimenCount id="9D71FD2FFFEBFFA8FC10FEF8A305FE9D" box="[955,1027,300,327]" count="3" pageId="28" pageNumber="29" type="female">3 ♀♀</specimenCount>
; 
<location id="8EA8607DFFEBFFA8FBBDFEF8A36AFE9C" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFEBFFA8FBBDFEF8A36AFE9C" box="[1046,1132,300,326]" country="Vietnam" name="Tonkin" pageId="28" pageNumber="29" stateProvince="Hoa Binh">Tonkin</location>
, region of 
<collectingRegion id="49B3F844FFEBFFA8FB53FEF9A274FE9C" box="[1272,1394,300,327]" country="Vietnam" name="Hoa Binh" pageId="28" pageNumber="29">Hoa Binh</collectingRegion>
; 1928; 
<collectorName id="26825370FFEBFFA8FEA3FE84A6B0FEB0" box="[264,438,335,362]" pageId="28" pageNumber="29">A. de Cooman</collectorName>
leg.; 
<collectionCode id="ED66AE63FFEBFFA8FE56FE84A551FEB0" box="[509,599,336,362]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">MNHN</collectionCode>
(
<specimenCode id="DBD19EDDFFEBFFA8FDC2FE9BA5DBFEB3" box="[617,733,335,361]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">EH24639</specimenCode>
, 
<specimenCode id="DBD19EDDFFEBFFA8FD47FE9BA43EFEB3" box="[748,824,335,361]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">24640</specimenCode>
, 
<specimenCode id="DBD19EDDFFEBFFA8FCEDFE9BA494FEB3" box="[838,914,335,361]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">24643</specimenCode>
, 
<specimenCode id="DBD19EDDFFEBFFA8FC0AFE9BA4EAFEB3" box="[929,1004,335,361]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="29">24644</specimenCode>
)
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FBBAFE84A4D1FE6A" collectingDate="2016-07-25" collectingDateMax="2016-07-30" collectingDateMin="2016-07-25" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel" country="Vietnam" latitude="20.437777" location="Ngoc Son-Ngo Luong Nature Reserve" longLatPrecision="21" longitude="105.3375" pageId="28" pageNumber="29" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Hoa Binh Province">
<specimenCount id="9D71FD2FFFEBFFA8FBBAFE84A350FEB0" box="[1041,1110,336,362]" count="4" pageId="28" pageNumber="29" type="male">4 ♂♂</specimenCount>
(dissected), 
<specimenCount id="9D71FD2FFFEBFFA8FB44FE9BA233FEB0" box="[1263,1333,335,362]" count="6" pageId="28" pageNumber="29" type="female">6 ♀♀</specimenCount>
; 
<collectingRegion id="49B3F844FFEBFFA8FAEDFE84A66DFE56" country="Vietnam" name="Hoa Binh" pageId="28" pageNumber="29">Hoa Binh Province</collectingRegion>
, 
<location id="8EA8607DFFEBFFA8FED2FEA7A441FE57" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFEBFFA8FED2FEA7A441FE57" box="[377,839,371,397]" country="Vietnam" latitude="20.437777" longLatPrecision="21" longitude="105.3375" name="Ngoc Son-Ngo Luong Nature Reserve" pageId="28" pageNumber="29" stateProvince="Hoa Binh Province">Ngoc Son-Ngo Luong Nature Reserve</location>
; 
<geoCoordinate id="EE435061FFEBFFA8FCFEFEA5A4EBFE57" box="[853,1005,369,397]" degrees="20" direction="north" minutes="26" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="16" value="20.437777">20°26′16″ N</geoCoordinate>
, 
<geoCoordinate id="EE435061FFEBFFA8FC50FEA5A399FE57" box="[1019,1183,369,397]" degrees="105" direction="east" minutes="20" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="15" value="105.3375">105°20′15″ E</geoCoordinate>
; 
<date id="FFC91066FFEBFFA8FB05FEA6A274FE56" box="[1198,1394,370,397]" pageId="28" pageNumber="29" value="2016-07-25" valueMax="2016-07-30" valueMin="2016-07-25">
<collectingDate id="EF8DE98EFFEBFFA8FB05FEA6A274FE56" box="[1198,1394,370,397]" pageId="28" pageNumber="29" value="2016-07-25" valueMax="2016-07-30" valueMin="2016-07-25">25–30 Jul. 2016</collectingDate>
</date>
; GTI Project; 
<collectorName id="26825370FFEBFFA8FEF0FE42A6D9FE6A" box="[347,479,406,432]" pageId="28" pageNumber="29">J. Constant</collectorName>
and 
<collectorName id="26825370FFEBFFA8FDB7FE42A59DFE6A" box="[540,667,406,432]" pageId="28" pageNumber="29">J. Bresseel</collectorName>
leg.; I.G.: 33.282; 
<collectionCode id="ED66AE63FFEBFFA8FC2AFE42A4D1FE6A" box="[897,983,406,432]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="28" pageNumber="29" type="Museum">RBINS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FC5FFE42A6B2FE2C" collectingDate="2011-07-19" collectingDateMax="2011-07-23" collectingDateMin="2011-07-19" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel" country="Vietnam" latitude="20.316668" location="Cuc Phuong National Park" longLatPrecision="21" longitude="105.60833" pageId="28" pageNumber="29" specimenCount="2" specimenCount-female="2" stateProvince="Ninh Binh Province">
<specimenCount id="9D71FD2FFFEBFFA8FC5FFE42A33FFE6A" box="[1012,1081,406,432]" count="2" pageId="28" pageNumber="29" type="female">2 ♀♀</specimenCount>
; 
<collectingRegion id="49B3F844FFEBFFA8FBE2FE42A23CFE6A" box="[1097,1338,406,432]" country="Vietnam" name="Ninh Binh" pageId="28" pageNumber="29">Ninh Binh Province</collectingRegion>
, 
<location id="8EA8607DFFEBFFA8FAECFE42A6CEFE09" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFEBFFA8FAECFE42A6CEFE09" country="Vietnam" latitude="20.316668" longLatPrecision="21" longitude="105.60833" name="Cuc Phuong National Park" pageId="28" pageNumber="29" stateProvince="Ninh Binh Province">Cuc Phuong National Park</location>
; 
<geoCoordinate id="EE435061FFEBFFA8FE7EFE63A56AFE0E" box="[469,620,439,468]" degrees="20" direction="north" minutes="19" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="00" value="20.316668">20°19′00″ N</geoCoordinate>
, 
<geoCoordinate id="EE435061FFEBFFA8FDD1FE63A41BFE0E" box="[634,797,439,468]" degrees="105" direction="east" minutes="36" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="30" value="105.60833">105°36′30″ E</geoCoordinate>
; 
<date id="FFC91066FFEBFFA8FC80FE6DA4EBFE09" box="[811,1005,441,467]" pageId="28" pageNumber="29" value="2011-07-19" valueMax="2011-07-23" valueMin="2011-07-19">
<collectingDate id="EF8DE98EFFEBFFA8FC80FE6DA4EBFE09" box="[811,1005,441,467]" pageId="28" pageNumber="29" value="2011-07-19" valueMax="2011-07-23" valueMin="2011-07-19">19–23 Jul. 2011</collectingDate>
</date>
; 
<collectorName id="26825370FFEBFFA8FC51FE6EA378FE09" box="[1018,1150,441,468]" pageId="28" pageNumber="29">J. Constant</collectorName>
and 
<collectorName id="26825370FFEBFFA8FB10FE6EA23CFE09" box="[1211,1338,441,468]" pageId="28" pageNumber="29">J. Bresseel</collectorName>
leg.; I.G.: 31.933; 
<collectionCode id="ED66AE63FFEBFFA8FEF5FE08A6B2FE2C" box="[350,436,476,502]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="28" pageNumber="29" type="Museum">RBINS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FE7AFE08A5EFFDC0" collectingDate="2001-06-30" collectionCode="VNMN" collectorName="H. T. Ta" country="Vietnam" elevation="900" latitude="20.679722" location="Mai Chau" longLatPrecision="21" longitude="105.01" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Hoa Binh Province">
<specimenCount id="9D71FD2FFFEBFFA8FE7AFE08A6F9FE2D" box="[465,511,476,503]" count="1" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
(dissected); 
<collectingRegion id="49B3F844FFEBFFA8FD32FE09A486FE2C" box="[665,896,476,503]" country="Vietnam" name="Hoa Binh" pageId="28" pageNumber="29">Hoa Binh Province</collectingRegion>
, 
<location id="8EA8607DFFEBFFA8FC25FE08A30EFE2C" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFEBFFA8FC25FE08A30EFE2C" box="[910,1032,476,502]" country="Vietnam" latitude="20.679722" longLatPrecision="21" longitude="105.01" name="Mai Chau" pageId="28" pageNumber="29" stateProvince="Hoa Binh Province">Mai Chau</location>
; [
<geoCoordinate id="EE435061FFEBFFA8FBB7FE0FA3B1FE2D" box="[1052,1207,475,503]" degrees="20" direction="north" minutes="40" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="47" value="20.679722">20°40′47″ N</geoCoordinate>
, 
<geoCoordinate id="EE435061FFEBFFA8FB6FFE0FA26FFE2D" box="[1220,1385,475,503]" degrees="105" direction="east" minutes="00" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="36" value="105.01">105°00′36″ E</geoCoordinate>
]; 
<date id="FFC91066FFEBFFA8FF16FDD4A650FDC0" box="[189,342,512,538]" pageId="28" pageNumber="29" value="2001-06-30">
<collectingDate id="EF8DE98EFFEBFFA8FF16FDD4A650FDC0" box="[189,342,512,538]" pageId="28" pageNumber="29" value="2001-06-30">30 Jun. 2001</collectingDate>
</date>
; 
<elevation id="005AD195FFEBFFA8FEC9FDD4A6EFFDC3" box="[354,489,511,538]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="28" pageNumber="29" unit="m" value="900.0">
<quantity id="4C8F9B43FFEBFFA8FEC9FDD4A6AAFDC0" box="[354,428,512,538]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="28" pageNumber="29" unit="m" value="900.0">900 m</quantity>
a.s.l.
</elevation>
; 
<collectorName id="26825370FFEBFFA8FE53FDD4A549FDC0" box="[504,591,512,538]" pageId="28" pageNumber="29">H.T. Ta</collectorName>
leg.; 
<collectionCode id="ED66AE63FFEBFFA8FD24FDD4A5EFFDC0" box="[655,745,512,538]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="28" pageNumber="29">VNMN</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FCAAFDD4A270FDE7" collectingDate="2015-07-25" collectingDateMax="2015-07-29" collectingDateMin="2015-07-25" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel" country="Vietnam" latitude="21.06778" location="Ba Vi National Park" longLatPrecision="21" longitude="105.35833" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Hanoi Province">
<specimenCount id="9D71FD2FFFEBFFA8FCAAFDD4A42AFDC0" box="[769,812,512,538]" count="1" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; 
<collectingRegion id="49B3F844FFEBFFA8FC90FE2BA4F3FDC3" box="[827,1013,511,537]" country="Vietnam" name="Ha Noi" pageId="28" pageNumber="29">Hanoi Province</collectingRegion>
, 
<location id="8EA8607DFFEBFFA8FBABFDD4A3E9FDC3" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFEBFFA8FBABFDD4A3E9FDC3" box="[1024,1263,511,538]" country="Vietnam" latitude="21.06778" longLatPrecision="21" longitude="105.35833" name="Ba Vi National Park" pageId="28" pageNumber="29" stateProvince="Hanoi Province">Ba Vi National Park</location>
; 
<geoCoordinate id="EE435061FFEBFFA8FB51FE2AA274FDC0" box="[1274,1394,510,538]" degrees="21" direction="north" minutes="4" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="4" value="21.06778">21°4′4″ N</geoCoordinate>
, 
<geoCoordinate id="EE435061FFEBFFA8FF16FDF5A666FDE7" box="[189,352,545,573]" degrees="105" direction="east" minutes="21" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="30" value="105.35833">105°21′30″ E</geoCoordinate>
; 
<date id="FFC91066FFEBFFA8FEC4FDF6A534FDE7" box="[367,562,546,573]" pageId="28" pageNumber="29" value="2015-07-25" valueMax="2015-07-29" valueMin="2015-07-25">
<collectingDate id="EF8DE98EFFEBFFA8FEC4FDF6A534FDE7" box="[367,562,546,573]" pageId="28" pageNumber="29" value="2015-07-25" valueMax="2015-07-29" valueMin="2015-07-25">25–29 Jul. 2015</collectingDate>
</date>
; day collecting; 
<collectorName id="26825370FFEBFFA8FD52FDF7A47BFDE7" box="[761,893,547,573]" pageId="28" pageNumber="29">J. Constant</collectorName>
and 
<collectorName id="26825370FFEBFFA8FC11FDF7A33FFDE6" box="[954,1081,546,573]" pageId="28" pageNumber="29">J. Bresseel</collectorName>
leg.; I.G.: 33.092; 
<collectionCode id="ED66AE63FFEBFFA8FA8BFDF7A270FDE7" box="[1312,1398,547,573]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="28" pageNumber="29" type="Museum">RBINS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FF78FD92A4C7FD59" collectingDate="2022-08-12" collectingDateMax="2022-08-18" collectingDateMin="2022-08-12" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel &amp; L. Semeraro" country="Vietnam" latitude="21.139723" location="Xuan Son National Park" longLatPrecision="21" longitude="104.94139" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Phu Tho Province">
<specimenCount id="9D71FD2FFFEBFFA8FF78FD92A602FDBA" box="[211,260,582,608]" count="1" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; 
<collectingRegion id="49B3F844FFEBFFA8FEB3FD92A6F1FDBA" box="[280,503,582,608]" country="Vietnam" name="Phu Tho" pageId="28" pageNumber="29">Phu Tho Province</collectingRegion>
, 
<location id="8EA8607DFFEBFFA8FDA3FD92A431FDBA" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFEBFFA8FDA3FD92A431FDBA" box="[520,823,582,608]" country="Vietnam" latitude="21.139723" longLatPrecision="21" longitude="104.94139" name="Xuan Son National Park" pageId="28" pageNumber="29" stateProvince="Phu Tho Province">Xuan Son National Park</location>
; 
<date id="FFC91066FFEBFFA8FCECFD92A319FDBA" box="[839,1055,582,608]" pageId="28" pageNumber="29" value="2022-08-12" valueMax="2022-08-18" valueMin="2022-08-12">
<collectingDate id="EF8DE98EFFEBFFA8FCECFD92A319FDBA" box="[839,1055,582,608]" pageId="28" pageNumber="29" value="2022-08-12" valueMax="2022-08-18" valueMin="2022-08-12">12–18 Aug. 2022</collectingDate>
</date>
; 
<geoCoordinate id="EE435061FFEBFFA8FB84FD90A3BDFDBA" box="[1071,1211,580,608]" degrees="21" direction="north" minutes="8" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="23" value="21.139723">21°8′23″ N</geoCoordinate>
, 
<geoCoordinate id="EE435061FFEBFFA8FB60FD90A277FDBA" box="[1227,1393,580,608]" degrees="104" direction="east" minutes="56" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="29" value="104.94139">104°56′29″ E</geoCoordinate>
; 
<collectorName id="26825370FFEBFFA8FF16FDBEA647FD59" box="[189,321,617,644]" pageId="28" pageNumber="29">J. Constant</collectorName>
, 
<collectorName id="26825370FFEBFFA8FEE7FDBEA6CEFD59" box="[332,456,617,644]" pageId="28" pageNumber="29">J. Bresseel</collectorName>
and 
<collectorName id="26825370FFEBFFA8FDABFDBEA589FD59" box="[512,655,617,644]" pageId="28" pageNumber="29">L. Semeraro</collectorName>
leg.; I.G.: 34.518; 
<collectionCode id="ED66AE63FFEBFFA8FCC0FDBDA4C7FD59" box="[875,961,617,643]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="28" pageNumber="29" type="Museum">RBINS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FC72FDBDA579FD7D" collectingDate="2004-06-12" collectingDateMax="2022-08-18" collectingDateMin="2022-08-12" collectionCode="VNMN" collectorName="H. T. Pham" country="Vietnam" latitude="21.139723" location="Xuan Son National Park" longLatPrecision="21" longitude="104.94139" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Phu Tho">
<specimenCount id="9D71FD2FFFEBFFA8FC72FDBDA302FD5E" box="[985,1028,617,644]" count="1" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; same locality as for preceding; 
<date id="FFC91066FFEBFFA8FF17FD58A65EFD7C" box="[188,344,652,679]" pageId="28" pageNumber="29" value="2004-06-12">
<collectingDate id="EF8DE98EFFEBFFA8FF17FD58A65EFD7C" box="[188,344,652,679]" pageId="28" pageNumber="29" value="2004-06-12">12 Jun. 2004</collectingDate>
</date>
; 
<collectorName id="26825370FFEBFFA8FECFFD59A6E5FD7C" box="[356,483,652,679]" pageId="28" pageNumber="29">H.T. Pham</collectorName>
leg.; 
<collectionCode id="ED66AE63FFEBFFA8FD8EFD59A579FD7D" box="[549,639,653,679]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="28" pageNumber="29">VNMN</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FD33FD58A43FFD10" collectingDate="2004-10-16" collectionCode="VNMN" collectorName="H. T. Pham" country="Vietnam" latitude="21.839724" location="Than Xa" longLatPrecision="21" longitude="105.90389" municipality="Vo Nhai" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Thai Nguyen Province">
<specimenCount id="9D71FD2FFFEBFFA8FD33FD58A5C2FD7D" box="[664,708,652,679]" count="1" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; 
<collectingRegion id="49B3F844FFEBFFA8FD7FFD58A4E7FD7C" box="[724,993,652,679]" country="Vietnam" name="Thai Nguyen" pageId="28" pageNumber="29">Thai Nguyen Province</collectingRegion>
, 
<collectingMunicipality id="6BACACDCFFEBFFA8FC46FD59A356FD7C" box="[1005,1104,652,679]" pageId="28" pageNumber="29">Vo Nhai</collectingMunicipality>
, 
<location id="8EA8607DFFEBFFA8FBF7FD58A3C0FD7D" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFEBFFA8FBF7FD58A3C0FD7D" box="[1116,1222,652,679]" country="Vietnam" latitude="21.839724" longLatPrecision="21" longitude="105.90389" municipality="Vo Nhai" name="Than Xa" pageId="28" pageNumber="29" stateProvince="Thai Nguyen Province">Than Xa</location>
; [
<geoCoordinate id="EE435061FFEBFFA8FB77FD5FA274FD7D" box="[1244,1394,651,679]" degrees="21" direction="north" minutes="50" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="23" value="21.839724">21°50′23″ N</geoCoordinate>
, 
<geoCoordinate id="EE435061FFEBFFA8FF16FD7AA666FD10" box="[189,352,686,714]" degrees="105" direction="east" minutes="54" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="14" value="105.90389">105°54′14″ E</geoCoordinate>
]; 
<date id="FFC91066FFEBFFA8FEDEFD7BA514FD10" box="[373,530,687,714]" pageId="28" pageNumber="29" value="2004-10-16">
<collectingDate id="EF8DE98EFFEBFFA8FEDEFD7BA514FD10" box="[373,530,687,714]" pageId="28" pageNumber="29" value="2004-10-16">16 Oct. 2004</collectingDate>
</date>
; 
<collectorName id="26825370FFEBFFA8FDB5FD64A59BFD13" box="[542,669,687,714]" pageId="28" pageNumber="29">H.T. Pham</collectorName>
leg.; 
<collectionCode id="ED66AE63FFEBFFA8FD74FD64A43FFD10" box="[735,825,688,714]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="28" pageNumber="29">VNMN</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FCF9FD7BA613FCCA" collectingDate="2011-07-25" collectingDateMax="2011-07-30" collectingDateMin="2011-07-25" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel" country="Vietnam" latitude="21.516666" location="Tam Dao National Park" pageId="28" pageNumber="29" specimenCount="5" specimenCount-female="5" stateProvince="Vinh Phuc Province">
<specimenCount id="9D71FD2FFFEBFFA8FCF9FD7BA493FD10" box="[850,917,687,714]" count="5" pageId="28" pageNumber="29" type="female">5 ♀♀</specimenCount>
; 
<collectingRegion id="49B3F844FFEBFFA8FC0FFD7BA394FD13" box="[932,1170,687,713]" country="Vietnam" name="Vinh Phuc" pageId="28" pageNumber="29">Vinh Phuc Province</collectingRegion>
, 
<location id="8EA8607DFFEBFFA8FB36FD64A7F1FD36" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFEBFFA8FB36FD64A7F1FD36" country="Vietnam" latitude="21.516666" name="Tam Dao National Park" pageId="28" pageNumber="29" stateProvince="Vinh Phuc Province">Tam Dao National Park</location>
; 21°31′ N, 105°33″ E; 
<date id="FFC91066FFEBFFA8FDA5FD06A5D7FD37" box="[526,721,722,749]" pageId="28" pageNumber="29" value="2011-07-25" valueMax="2011-07-30" valueMin="2011-07-25">
<collectingDate id="EF8DE98EFFEBFFA8FDA5FD06A5D7FD37" box="[526,721,722,749]" pageId="28" pageNumber="29" value="2011-07-25" valueMax="2011-07-30" valueMin="2011-07-25">25–30 Jul. 2011</collectingDate>
</date>
; day time; 
<collectorName id="26825370FFEBFFA8FCF3FD07A4DAFD37" box="[856,988,723,749]" pageId="28" pageNumber="29">J. Constant</collectorName>
and 
<collectorName id="26825370FFEBFFA8FBB2FD07A39EFD36" box="[1049,1176,722,749]" pageId="28" pageNumber="29">J. Bresseel</collectorName>
leg.; I.G.: 31.933; 
<collectionCode id="ED66AE63FFEBFFA8FF14FD22A613FCCA" box="[191,277,758,784]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="28" pageNumber="29" type="Museum">RBINS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FE81FD22A614FC8C" collectingDate="2023-07-10" collectingDateMax="2023-07-11" collectingDateMin="2023-07-10" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel &amp; L. Semeraro" country="Vietnam" latitude="19.861668" location="Hon Can station" longLatPrecision="21" longitude="105.21388" municipality="Xuan Lien Nature Reserve" pageId="28" pageNumber="29" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Thanh Hoa Province">
<specimenCount id="9D71FD2FFFEBFFA8FE81FD22A653FCCA" box="[298,341,758,784]" count="1" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
(dissected), 
<specimenCount id="9D71FD2FFFEBFFA8FE43FD22A52CFCCA" box="[488,554,758,784]" count="2" pageId="28" pageNumber="29" type="female">2 ♀♀</specimenCount>
; 
<collectingRegion id="49B3F844FFEBFFA8FD9CFD22A42CFCCA" box="[567,810,758,784]" country="Vietnam" name="Thanh Hoa" pageId="28" pageNumber="29">Thanh Hoa Province</collectingRegion>
, 
<collectingMunicipality id="6BACACDCFFEBFFA8FC9EFD22A368FCCA" box="[821,1134,758,784]" pageId="28" pageNumber="29">Xuan Lien Nature Reserve</collectingMunicipality>
, near 
<location id="8EA8607DFFEBFFA8FB19FD22A274FCCA" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFEBFFA8FB19FD22A274FCCA" box="[1202,1394,758,784]" country="Vietnam" latitude="19.861668" longLatPrecision="21" longitude="105.21388" municipality="Xuan Lien Nature Reserve" name="Hon Can station" pageId="28" pageNumber="29" stateProvince="Thanh Hoa Province">Hón Can station</location>
; 
<geoCoordinate id="EE435061FFEBFFA8FF16FCC3A654FCEE" box="[189,338,791,820]" degrees="19" direction="north" minutes="51" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="42" value="19.861668">19°51′42″ N</geoCoordinate>
, 
<geoCoordinate id="EE435061FFEBFFA8FEF7FCC3A6FBFCEE" box="[348,509,791,820]" degrees="105" direction="east" minutes="12" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="50" value="105.21388">105°12′50″ E</geoCoordinate>
; 
<date id="FFC91066FFEBFFA8FDA3FCCDA5C2FCE9" box="[520,708,793,819]" pageId="28" pageNumber="29" value="2023-07-10" valueMax="2023-07-11" valueMin="2023-07-10">
<collectingDate id="EF8DE98EFFEBFFA8FDA3FCCDA5C2FCE9" box="[520,708,793,819]" pageId="28" pageNumber="29" value="2023-07-10" valueMax="2023-07-11" valueMin="2023-07-10">10–11 Jul. 2023</collectingDate>
</date>
; 
<collectorName id="26825370FFEBFFA8FD7BFCCEA452FCE9" box="[720,852,793,820]" pageId="28" pageNumber="29">J. Constant</collectorName>
, 
<collectorName id="26825370FFEBFFA8FCF5FCCEA4DCFCE9" box="[862,986,793,820]" pageId="28" pageNumber="29">J. Bresseel</collectorName>
and 
<collectorName id="26825370FFEBFFA8FBB9FCCEA3A7FCE9" box="[1042,1185,793,820]" pageId="28" pageNumber="29">L. Semeraro</collectorName>
leg.; I.G.: 34.661; 
<collectionCode id="ED66AE63FFEBFFA8FF17FCE8A614FC8C" box="[188,274,828,854]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="28" pageNumber="29" type="Museum">RBINS</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B1F3CFBFFEBFFA8FE9BFCE8A246FCA0" collectingDate="2023-07-16" collectingDateMax="2023-07-18" collectingDateMin="2023-07-16" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel &amp; L. Semeraro" country="Vietnam" latitude="18.289165" location="Co station" longLatPrecision="21" longitude="105.375" municipality="Vu Quang National Park" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Ha Tinh Province">
<specimenCount id="9D71FD2FFFEBFFA8FE9BFCE8A658FC8D" box="[304,350,828,855]" count="1" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
(dissected); 
<collectingRegion id="49B3F844FFEBFFA8FE53FCE9A5C8FC8C" box="[504,718,828,855]" country="Vietnam" name="Ha Tinh" pageId="28" pageNumber="29">Ha Tinh Province</collectingRegion>
, 
<collectingMunicipality id="6BACACDCFFEBFFA8FD70FCE9A300FC8C" box="[731,1030,828,855]" pageId="28" pageNumber="29">Vu Quang National Park</collectingMunicipality>
, near 
<location id="8EA8607DFFEBFFA8FBE4FCE8A3CAFC8C" LSID="urn:lsid:plazi:treatment:03DE87B0FFECFF90FE1BFD31A64EFD0C:8EA8607DFFEBFFA8FBE4FCE8A3CAFC8C" box="[1103,1228,828,854]" country="Vietnam" latitude="18.289165" longLatPrecision="21" longitude="105.375" municipality="Vu Quang National Park" name="Co station" pageId="28" pageNumber="29" stateProvince="Ha Tinh Province">Co station</location>
; 
<geoCoordinate id="EE435061FFEBFFA8FB71FCEFA274FC8D" box="[1242,1394,827,855]" degrees="18" direction="north" minutes="17" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="21" value="18.289165">18°17′21″ N</geoCoordinate>
, 
<geoCoordinate id="EE435061FFEBFFA8FF16FC8AA658FCA0" box="[189,350,862,890]" degrees="105" direction="east" minutes="22" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="30" value="105.375">105°22′30″ E</geoCoordinate>
; 
<date id="FFC91066FFEBFFA8FEC0FC8BA52CFCA0" box="[363,554,863,890]" pageId="28" pageNumber="29" value="2023-07-16" valueMax="2023-07-18" valueMin="2023-07-16">
<collectingDate id="EF8DE98EFFEBFFA8FEC0FC8BA52CFCA0" box="[363,554,863,890]" pageId="28" pageNumber="29" value="2023-07-16" valueMax="2023-07-18" valueMin="2023-07-16">16–18 Jul. 2023</collectingDate>
</date>
; 
<collectorName id="26825370FFEBFFA8FD9DFCB4A5BAFCA0" box="[566,700,864,890]" pageId="28" pageNumber="29">J. Constant</collectorName>
, 
<collectorName id="26825370FFEBFFA8FD63FCB4A443FCA3" box="[712,837,863,890]" pageId="28" pageNumber="29">J. Bresseel</collectorName>
and 
<collectorName id="26825370FFEBFFA8FCD5FCB4A309FCA0" box="[894,1039,864,890]" pageId="28" pageNumber="29">L. Semeraro</collectorName>
leg. I.G.: 34.661; 
<collectionCode id="ED66AE63FFEBFFA8FB4DFCB4A246FCA0" box="[1254,1344,864,890]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="28" pageNumber="29" type="Museum">RBINS</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC836A6FFEBFFA8FF16FC74A517FC61" blockId="28.[189,529,928,955]" box="[189,529,928,955]" pageId="28" pageNumber="29">
<heading id="D08081CAFFEBFFA8FF16FC74A517FC61" bold="true" box="[189,529,928,955]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B903EAB4FFEBFFA8FF16FC74A517FC61" bold="true" box="[189,529,928,955]" pageId="28" pageNumber="29">Supplementary description</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFEBFFA8FF16FC04A5C5FBD7" blockId="28.[189,1398,975,1037]" pageId="28" pageNumber="29">
<smallCapsWord id="8D2EA07AFFEBFFA8FF16FC04A668FC32" baselines="995,995" box="[189,366,976,1002]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="28" pageNumber="29">MEASUREMENTS</smallCapsWord>
<smallCapsWord id="8D2EA07AFFEBFFA8FEDDFC01A6A5FC32" baselines="995" box="[374,419,981,1000]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="28" pageNumber="29">AND</smallCapsWord>
<smallCapsWord id="8D2EA07AFFEBFFA8FE07FC01A6F3FC32" baselines="995" box="[428,501,981,1000]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="ratios" pageId="28" pageNumber="29">RATIOS</smallCapsWord>
. LT: 
<specimenCount id="9D71FD2FFFEBFFA8FD9FFC04A54CFC30" box="[564,586,976,1002]" count="1" pageId="28" pageNumber="29" type="male">♂</specimenCount>
(n = 1): 
<quantity id="4C8F9B43FFEBFFA8FD1FFC1BA414FC30" box="[692,786,975,1002]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="28" pageNumber="29" unit="mm" value="5.4">5.4 mm</quantity>
, 
<specimenCount id="9D71FD2FFFEBFFA8FCB5FC04A432FC30" box="[798,820,976,1002]" count="1" pageId="28" pageNumber="29" type="female">♀</specimenCount>
(n = 1): 
<quantity id="4C8F9B43FFEBFFA8FC35FC1BA4FDFC30" box="[926,1019,975,1002]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="28" pageNumber="29" unit="mm" value="5.9">5.9 mm</quantity>
; LT/BB = 1.23; LTg/BTg = 1.5; LW/BW = 2.0; BV/LV = 2.3; LF/BF = 0.85.
</paragraph>
<paragraph id="8BC836A6FFEBFFA8FF16FBE0A518FB62" blockId="28.[189,1398,1076,1208]" pageId="28" pageNumber="29">
<smallCapsWord id="8D2EA07AFFEBFFA8FF16FBE0A61EFB97" baselines="1095,1096" box="[189,280,1076,1102]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="28" pageNumber="29">COLOUR</smallCapsWord>
. Variable with 2 main morphs: (1) black and bright yellow, with yellow transverse bands on frons, yellow marking on disc of tegmen and several, alternate black and yellow concentric lines along costal margin of tegmina (
<figureCitation id="134C2A23FFEBFFA8FDE0FBAEA5ACFB4F" box="[587,682,1146,1173]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[189,1398,360,1788]" captionTargetId="figure-11@29.[189,1398,360,1788]" captionTargetPageId="29" captionText="Fig. 13. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., syntypes, ♀♀ (NHMW – photographs by H. Brückner). A–D. Syntype 1. A. Habitus, dorsal view. B. Habitus, frontal view. C. Habitus, lateral view. D. Labels. E–F. Syntype 2. E. Habitus, dorsal view. F. Labels." figureDoi="http://doi.org/10.5281/zenodo.10559560" httpUri="https://zenodo.org/record/10559560/files/figure.png" pageId="28" pageNumber="29">Figs 13</figureCitation>
, 
<figureCitation id="134C2A23FFEBFFA8FD11FBAEA401FB4F" box="[698,775,1146,1173]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="31.[189,232,1892,1918]" captionStartId-1="32.[189,232,1928,1954]" captionTargetBox-0="[189,1393,487,1831]" captionTargetBox-1="[193,1386,305,1841]" captionTargetId-0="figure-63@31.[189,1398,481,1843]" captionTargetId-1="figure-18@32.[189,1398,295,1846]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="Fig. 15. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., dissected ♂, typical morph from Ngoc Son-Ngo Luong Nature Reserve (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Anterolateral view. F. Right hind wing." captionText-1="Fig. 16. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., dissected ♂, typical morph from Mai Chau (VNMN). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Anterolateral view. F. Right hind wing." figureDoi-0="http://doi.org/10.5281/zenodo.10559566" figureDoi-1="http://doi.org/10.5281/zenodo.10559568" httpUri-0="https://zenodo.org/record/10559566/files/figure.png" httpUri-1="https://zenodo.org/record/10559568/files/figure.png" pageId="28" pageNumber="29">15–16</figureCitation>
); (2) black, with one yellow transverse band along frontoclypeal suture (
<figureCitation id="134C2A23FFEBFFA8FE13FB4AA516FB62" box="[440,528,1182,1208]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[250,1340,271,1881]" captionTargetId="figure-18@30.[243,1344,265,1885]" captionTargetPageId="30" captionText="Fig. 14. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., dissected ♂, black morph from Dong Son-Ky Thuong Nature Reserve (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559562" httpUri="https://zenodo.org/record/10559562/files/figure.png" pageId="28" pageNumber="29">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8BC836A6FFEBFFA8FF16FB34A501FAB9" blockId="28.[189,1399,1247,1379]" pageId="28" pageNumber="29">
<smallCapsWord id="8D2EA07AFFEBFFA8FF16FB34A7F0FB22" baselines="1267,1267" box="[189,246,1248,1274]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="28" pageNumber="29">LEGS</smallCapsWord>
. With pro- and mesofemora ventrally with 3 contrasting black transverse markings and black line along anterior margin; pro- and mesotibiae with black line along outer margin; metafemora black with yellow markings; metatibiae black with more or less developed yellowish white portion in distal ⅓ sometimes extending basad.
</paragraph>
<paragraph id="8BC836A6FFEBFFA8FF16FA5FA4B3F80D" blockId="28.[188,1400,1418,2008]" pageId="28" pageNumber="29">
<smallCapsWord id="8D2EA07AFFEBFFA8FF16FA5FA7F9FA79" baselines="1438,1438" box="[189,255,1419,1445]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="28" pageNumber="29">MALE</smallCapsWord>
<smallCapsWord id="8D2EA07AFFEBFFA8FEA1FA44A68DFA79" baselines="1438" box="[266,395,1424,1443]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="terminalia" pageId="28" pageNumber="29">TERMINALIA</smallCapsWord>
. Pygofer (
<emphasis id="B903EAB4FFEBFFA8FDA4FA5FA528FA7E" box="[527,558,1419,1444]" italics="true" pageId="28" pageNumber="29">Py</emphasis>
– 
<figureCitation id="134C2A23FFEBFFA8FDF9FA5EA5E0FA7E" box="[594,742,1418,1444]" captionStart="Fig" captionStartId="33.[189,232,1857,1883]" captionTargetBox="[270,1316,275,1809]" captionTargetId="figure-12@33.[267,1321,265,1815]" captionTargetPageId="33" captionText="Fig. 17. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, black morph from Dong Son- Ky Thuong Nature Reserve (RBINS), terminalia. A–C. Pygofer, anal tube and gonostyli. A. Lateral view. B. Caudal view. C. Dorsal view. D–I. Aedeagus and connective. D. Left lateral view. E. Right lateral view. F. Caudal view. G. Ventral view. H. Dorsal view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559570" httpUri="https://zenodo.org/record/10559570/files/figure.png" pageId="28" pageNumber="29">Figs 17A–C</figureCitation>
, 
<figureCitation id="134C2A23FFEBFFA8FD5EFA5EA44BFA7E" box="[757,845,1418,1444]" captionStart="Fig" captionStartId="34.[189,232,1857,1883]" captionTargetBox="[232,1356,277,1790]" captionTargetId="figure-18@34.[224,1362,265,1805]" captionTargetPageId="34" captionText="Fig. 18. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, typical morph from Ngoc Son- Ngo Luong Nature Reserve (RBINS), terminalia.A–D. Pygofer, anal tube and gonostyli.A. Lateral view. B. Laterocaudal view. C. Dorsocaudal view. D. Dorsal view. E–J. Aedeagus and connective. E. Dorsal view. F. Left lateral view. G. Right lateral view. H. Left lateroventral view. I. Right laterocaudal view. J. Left laterocaudal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559575" httpUri="https://zenodo.org/record/10559575/files/figure.png" pageId="28" pageNumber="29">18A–D</figureCitation>
) short, curved, about 3 times as high as long at mid-height in lateral view, with posterior margin somewhat angularly rounded in middle in lateral view; suboval in caudal view. Gonostyli (
<emphasis id="B903EAB4FFEBFFA8FD0EFA05A5BCFA31" box="[677,698,1489,1515]" italics="true" pageId="28" pageNumber="29">G</emphasis>
– 
<figureCitation id="134C2A23FFEBFFA8FD73FA04A46FFA31" box="[728,873,1488,1515]" captionStart="Fig" captionStartId="33.[189,232,1857,1883]" captionTargetBox="[270,1316,275,1809]" captionTargetId="figure-12@33.[267,1321,265,1815]" captionTargetPageId="33" captionText="Fig. 17. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, black morph from Dong Son- Ky Thuong Nature Reserve (RBINS), terminalia. A–C. Pygofer, anal tube and gonostyli. A. Lateral view. B. Caudal view. C. Dorsal view. D–I. Aedeagus and connective. D. Left lateral view. E. Right lateral view. F. Caudal view. G. Ventral view. H. Dorsal view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559570" httpUri="https://zenodo.org/record/10559570/files/figure.png" pageId="28" pageNumber="29">Figs 17A–B</figureCitation>
, 
<figureCitation id="134C2A23FFEBFFA8FCDEFA05A4CAFA31" box="[885,972,1489,1515]" captionStart="Fig" captionStartId="34.[189,232,1857,1883]" captionTargetBox="[232,1356,277,1790]" captionTargetId="figure-18@34.[224,1362,265,1805]" captionTargetPageId="34" captionText="Fig. 18. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, typical morph from Ngoc Son- Ngo Luong Nature Reserve (RBINS), terminalia.A–D. Pygofer, anal tube and gonostyli.A. Lateral view. B. Laterocaudal view. C. Dorsocaudal view. D. Dorsal view. E–J. Aedeagus and connective. E. Dorsal view. F. Left lateral view. G. Right lateral view. H. Left lateroventral view. I. Right laterocaudal view. J. Left laterocaudal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559575" httpUri="https://zenodo.org/record/10559575/files/figure.png" pageId="28" pageNumber="29">18A–C</figureCitation>
) convex, with ventral margin nearly straight and posterior margin rounded in lateral view; elongate in ventral view; capitulum (
<emphasis id="B903EAB4FFEBFFA8FAA8FA20A219F9D7" box="[1283,1311,1524,1549]" italics="true" pageId="28" pageNumber="29">ca</emphasis>
) rather short, with a wide neck with basal hump on posterior margin, with apical tooth directed mesad and lateral laminate process at mid-height. Anal tube (
<emphasis id="B903EAB4FFEBFFA8FCBBF9EEA437F98E" box="[784,817,1594,1620]" italics="true" pageId="28" pageNumber="29">An</emphasis>
– 
<figureCitation id="134C2A23FFEBFFA8FCFBF9EEA4E4F98E" box="[848,994,1594,1620]" captionStart="Fig" captionStartId="33.[189,232,1857,1883]" captionTargetBox="[270,1316,275,1809]" captionTargetId="figure-12@33.[267,1321,265,1815]" captionTargetPageId="33" captionText="Fig. 17. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, black morph from Dong Son- Ky Thuong Nature Reserve (RBINS), terminalia. A–C. Pygofer, anal tube and gonostyli. A. Lateral view. B. Caudal view. C. Dorsal view. D–I. Aedeagus and connective. D. Left lateral view. E. Right lateral view. F. Caudal view. G. Ventral view. H. Dorsal view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559570" httpUri="https://zenodo.org/record/10559570/files/figure.png" pageId="28" pageNumber="29">Figs 17A–C</figureCitation>
, 
<figureCitation id="134C2A23FFEBFFA8FC5BF9EEA34EF98E" box="[1008,1096,1594,1620]" captionStart="Fig" captionStartId="34.[189,232,1857,1883]" captionTargetBox="[232,1356,277,1790]" captionTargetId="figure-18@34.[224,1362,265,1805]" captionTargetPageId="34" captionText="Fig. 18. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, typical morph from Ngoc Son- Ngo Luong Nature Reserve (RBINS), terminalia.A–D. Pygofer, anal tube and gonostyli.A. Lateral view. B. Laterocaudal view. C. Dorsocaudal view. D. Dorsal view. E–J. Aedeagus and connective. E. Dorsal view. F. Left lateral view. G. Right lateral view. H. Left lateroventral view. I. Right laterocaudal view. J. Left laterocaudal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559575" httpUri="https://zenodo.org/record/10559575/files/figure.png" pageId="28" pageNumber="29">18A–D</figureCitation>
) dorsoventrally flattened, ventrally slightly depressed medially, 1.2 times as wide as long in midline in dorsal view and with anal opening in basal half. Aedeagus (
<figureCitation id="134C2A23FFEBFFA8FDE3F954A5D4F941" box="[584,722,1664,1691]" captionStart="Fig" captionStartId="33.[189,232,1857,1883]" captionTargetBox="[270,1316,275,1809]" captionTargetId="figure-12@33.[267,1321,265,1815]" captionTargetPageId="33" captionText="Fig. 17. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, black morph from Dong Son- Ky Thuong Nature Reserve (RBINS), terminalia. A–C. Pygofer, anal tube and gonostyli. A. Lateral view. B. Caudal view. C. Dorsal view. D–I. Aedeagus and connective. D. Left lateral view. E. Right lateral view. F. Caudal view. G. Ventral view. H. Dorsal view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559570" httpUri="https://zenodo.org/record/10559570/files/figure.png" pageId="28" pageNumber="29">Figs 17D–I</figureCitation>
, 
<figureCitation id="134C2A23FFEBFFA8FD4BF955A42CF941" box="[736,810,1665,1691]" captionStart="Fig" captionStartId="34.[189,232,1857,1883]" captionTargetBox="[232,1356,277,1790]" captionTargetId="figure-18@34.[224,1362,265,1805]" captionTargetPageId="34" captionText="Fig. 18. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov., ♂, typical morph from Ngoc Son- Ngo Luong Nature Reserve (RBINS), terminalia.A–D. Pygofer, anal tube and gonostyli.A. Lateral view. B. Laterocaudal view. C. Dorsocaudal view. D. Dorsal view. E–J. Aedeagus and connective. E. Dorsal view. F. Left lateral view. G. Right lateral view. H. Left lateroventral view. I. Right laterocaudal view. J. Left laterocaudal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559575" httpUri="https://zenodo.org/record/10559575/files/figure.png" pageId="28" pageNumber="29">18E–J</figureCitation>
) asymmetrical, strongly upcurved in lateral view. Ventral lobe of periandrium (
<emphasis id="B903EAB4FFEBFFA8FDB6F970A534F964" box="[541,562,1700,1726]" italics="true" pageId="28" pageNumber="29">vl</emphasis>
) laminate with 2 longitudinal rows of minute teeth hooked cephalad, and with elongate, thin, lateroventral processes (
<emphasis id="B903EAB4FFEBFFA8FCADF913A42CF93B" box="[774,810,1735,1761]" italics="true" pageId="28" pageNumber="29">lvp</emphasis>
) sinuately curved dorsocephalad, left one before midlength of aedeagus, right one after midlength; lateral margins minutely denticulate; sides roundly projecting dorsad; median portion developed into a lobe obliquely weakly sinuate apically. Lateral lobes of periandrium (
<emphasis id="B903EAB4FFEBFFA8FED6F8E4A68BF890" box="[381,397,1840,1866]" italics="true" pageId="28" pageNumber="29">ll</emphasis>
) laminate, moderately curved posteroventrad, reaching about half length of aedeagus and covering base of lateral elongate processes of periandrium; latter shaft-like, directed caudad in basal half then strongly curved dorsocephalad. Connective (
<emphasis id="B903EAB4FFEBFFA8FCDDF8ACA494F84B" box="[886,914,1912,1937]" italics="true" pageId="28" pageNumber="29">co</emphasis>
) well developed, corpus connective long, regularly and rather weakly curved in lateral view, tectiductus (
<emphasis id="B903EAB4FFEBFFA8FC02F84FA4B8F86E" box="[937,958,1947,1972]" italics="true" pageId="28" pageNumber="29">te</emphasis>
) well developed, reaching about half length of corpus connective, conical with wide anterior foramen.
</paragraph>
</subSubSection>
<caption id="DF08662EFFEAFFA9FF16F85CA3C7F832" ID-DOI="http://doi.org/10.5281/zenodo.10559560" ID-Zenodo-Dep="10559560" httpUri="https://zenodo.org/record/10559560/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1928,1954]" targetBox="[189,1398,360,1788]" targetPageId="29">
<paragraph id="8BC836A6FFEAFFA9FF16F85CA3C7F832" blockId="29.[189,1399,1928,2025]" pageId="29" pageNumber="30">
<emphasis id="B903EAB4FFEAFFA9FF16F85CA626F878" bold="true" box="[189,288,1928,1954]" pageId="29" pageNumber="30">Fig. 13.</emphasis>
<taxonomicName id="4C774D25FFEAFFA9FE85F85CA42AF878" authority="(Melichar, 1906)" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[302,812,1928,1954]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="gravidus" status="gen. et comb. nov.">
<emphasis id="B903EAB4FFEAFFA9FE85F85CA555F878" box="[302,595,1928,1954]" italics="true" pageId="29" pageNumber="30">Melichergithus gravidus</emphasis>
(
<bibRefCitation id="EFE64B57FFEAFFA9FDC3F85CA422F878" author="Melichar L." box="[616,804,1928,1954]" pageId="29" pageNumber="30" pagination="1 - 327" refId="ref35004" refString="Melichar L. 1906. Monographie der Issiden (Homoptera). Abhandlungen der K. K. Zoologischbotanischen Gesellschaft in Wien 3: 1 - 327." type="journal article" year="1906">Melichar, 1906</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFEAFFA9FC90F85CA32EF878" box="[827,1064,1928,1954]" pageId="29" pageNumber="30" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, syntypes, ♀♀ (NHMW – photographs by H. Brückner). 
<emphasis id="B903EAB4FFEAFFA9FD9CF87FA576F81F" bold="true" box="[567,624,1963,1989]" pageId="29" pageNumber="30">A–D</emphasis>
. Syntype 1. 
<emphasis id="B903EAB4FFEAFFA9FCB9F87FA421F81F" bold="true" box="[786,807,1963,1989]" pageId="29" pageNumber="30">A</emphasis>
. Habitus, dorsal view. 
<emphasis id="B903EAB4FFEAFFA9FBE3F87FA35AF81F" bold="true" box="[1096,1116,1963,1989]" pageId="29" pageNumber="30">B</emphasis>
. Habitus, frontal view. 
<emphasis id="B903EAB4FFEAFFA9FF16F81AA7D4F832" bold="true" box="[189,210,1998,2024]" pageId="29" pageNumber="30">C</emphasis>
. Habitus, lateral view. 
<emphasis id="B903EAB4FFEAFFA9FE4AF81AA6F0F832" bold="true" box="[481,502,1998,2024]" pageId="29" pageNumber="30">D</emphasis>
. Labels. 
<emphasis id="B903EAB4FFEAFFA9FDC9F81AA590F832" bold="true" box="[610,662,1998,2024]" pageId="29" pageNumber="30">E–F</emphasis>
. Syntype 2. 
<emphasis id="B903EAB4FFEAFFA9FC82F81AA43BF832" bold="true" box="[809,829,1998,2024]" pageId="29" pageNumber="30">E</emphasis>
. Habitus, dorsal view. 
<emphasis id="B903EAB4FFEAFFA9FBE1F81AA35AF832" bold="true" box="[1098,1116,1998,2024]" pageId="29" pageNumber="30">F</emphasis>
. Labels.
</paragraph>
</caption>
<caption id="DF08662EFFE9FFAAFF16F85CA3ADF832" ID-DOI="http://doi.org/10.5281/zenodo.10559562" ID-Zenodo-Dep="10559562" httpUri="https://zenodo.org/record/10559562/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1928,1954]" targetBox="[250,1340,271,1881]" targetPageId="30">
<paragraph id="8BC836A6FFE9FFAAFF16F85CA3ADF832" blockId="30.[189,1399,1928,2025]" pageId="30" pageNumber="31">
<emphasis id="B903EAB4FFE9FFAAFF16F85CA61DF878" bold="true" box="[189,283,1928,1954]" pageId="30" pageNumber="31">Fig. 14.</emphasis>
<taxonomicName id="4C774D25FFE9FFAAFE8FF85CA414F878" authority="(Melichar, 1906)" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[292,786,1928,1954]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="gravidus" status="gen. et comb. nov.">
<emphasis id="B903EAB4FFE9FFAAFE8FF85CA543F878" box="[292,581,1928,1954]" italics="true" pageId="30" pageNumber="31">Melichergithus gravidus</emphasis>
(
<bibRefCitation id="EFE64B57FFE9FFAAFDFEF85CA40CF878" author="Melichar L." box="[597,778,1928,1954]" pageId="30" pageNumber="31" pagination="1 - 327" refId="ref35004" refString="Melichar L. 1906. Monographie der Issiden (Homoptera). Abhandlungen der K. K. Zoologischbotanischen Gesellschaft in Wien 3: 1 - 327." type="journal article" year="1906">Melichar, 1906</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFE9FFAAFCB0F85CA4FFF878" box="[795,1017,1928,1954]" pageId="30" pageNumber="31" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, dissected ♂, black morph from Dong Son-Ky Thuong Nature Reserve (RBINS). 
<emphasis id="B903EAB4FFE9FFAAFCBDF87FA42DF81F" bold="true" box="[790,811,1963,1989]" pageId="30" pageNumber="31">A</emphasis>
. Habitus, dorsal view. 
<emphasis id="B903EAB4FFE9FFAAFBE3F87FA35AF81F" bold="true" box="[1096,1116,1963,1989]" pageId="30" pageNumber="31">B</emphasis>
. Habitus, ventral view. 
<emphasis id="B903EAB4FFE9FFAAFF16F81AA7D4F832" bold="true" box="[189,210,1998,2024]" pageId="30" pageNumber="31">C</emphasis>
. Habitus, lateral view. 
<emphasis id="B903EAB4FFE9FFAAFE4AF81AA6F0F832" bold="true" box="[481,502,1998,2024]" pageId="30" pageNumber="31">D</emphasis>
. Habitus, perpendicular view of frons. 
<emphasis id="B903EAB4FFE9FFAAFC68F81AA4D1F832" bold="true" box="[963,983,1998,2024]" pageId="30" pageNumber="31">E</emphasis>
. Right hind wing.
</paragraph>
</caption>
<subSubSection id="C36D652DFFE8FF90FF16FEDCA4CBFEB3" lastPageId="36" lastPageNumber="37" pageId="31" pageNumber="32" type="biology_ecology">
<paragraph id="8BC836A6FFE8FFABFF16FEDCA61AFEF8" blockId="31.[189,284,264,290]" box="[189,284,264,290]" pageId="31" pageNumber="32">
<heading id="D08081CAFFE8FFABFF16FEDCA61AFEF8" bold="true" box="[189,284,264,290]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B903EAB4FFE8FFABFF16FEDCA61AFEF8" bold="true" box="[189,284,264,290]" pageId="31" pageNumber="32">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFE8FF90FF16FEE3A4CBFEB3" blockId="31.[189,1400,311,408]" lastBlockId="36.[188,1399,265,361]" lastPageId="36" lastPageNumber="37" pageId="31" pageNumber="32">
<taxonomicName id="4C774D25FFE8FFABFF16FEE3A6D8FE8B" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[189,478,311,337]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="gravidus" status="gen. et comb. nov.">
<emphasis id="B903EAB4FFE8FFABFF16FEE3A6D8FE8B" box="[189,478,311,337]" italics="true" pageId="31" pageNumber="32">Melichergithus gravidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFE8FFABFE4CFEECA5C3FE88" box="[487,709,311,338]" pageId="31" pageNumber="32" rank="species">gen. et comb. nov.</taxonomicNameLabel>
was collected in August on lower vegetation and bushes, in moist evergreen tropical forest at about 
<quantity id="4C8F9B43FFE8FFABFD10FE8EA401FEAF" box="[699,775,346,373]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="31" pageNumber="32" unit="m" value="550.0">550 m</quantity>
in altitude in Dong Son-Ky Thuong Nature Reserve (
<figureCitation id="134C2A23FFE8FFABFF6CFEA9A624FE42" box="[199,290,381,408]" captionStart="Fig" captionStartId="2.[189,232,1998,2024]" captionTargetBox="[218,1369,265,1958]" captionTargetId="figure-18@2.[218,1369,265,1958]" captionTargetPageId="2" captionText="Fig. 1. Dong Son-Ky Thuong Nature Reserve. A. Topographic map. B. Sampled habitat." figureDoi="http://doi.org/10.5281/zenodo.10559522" httpUri="https://zenodo.org/record/10559522/files/figure.png" pageId="31" pageNumber="32">Fig. 1B</figureCitation>
). In other places it was also collected in moist evergreen tropical forest, at altitudes from about 
<quantity id="4C8F9B43FFD3FF90FF16FEDDA601FEF9" box="[189,263,265,292]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="36" pageNumber="37" unit="m" value="150.0">150 m</quantity>
in Cuc Phuong National Park to over 
<quantity id="4C8F9B43FFD3FF90FD6AFEDDA41FFEF9" box="[705,793,265,291]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="1000.0">1000 m</quantity>
in Tam Dao National Park in the months from April to August, and in October. The specimens were often observed waving their forelegs (
<figureCitation id="134C2A23FFD3FF90FB66FEF8A23CFE9C" box="[1229,1338,300,326]" captionStart="Fig" captionStartId="35.[189,232,1928,1954]" captionTargetBox="[242,1346,265,1888]" captionTargetId="figure-12@35.[242,1346,265,1888]" captionTargetPageId="35" captionText="Fig. 19. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov. A. Distribution map. B. Live specimen on fern, Xuan Lien Nature Reserve, 11 Jul. 2023. C. Live specimen on fern, Cuc Phuong National Park, 20 Jul. 2011. D. Habitat in Cuc Phuong National Park, 20 Jul. 2011." figureDoi="http://doi.org/10.5281/zenodo.10559577" httpUri="https://zenodo.org/record/10559577/files/figure.png" pageId="36" pageNumber="37">Fig. 19B</figureCitation>
), the conspicuous markings of which might be used for communication.
</paragraph>
</subSubSection>
<caption id="DF08662EFFE8FFABFF16F8B0A606F832" ID-DOI="http://doi.org/10.5281/zenodo.10559566" ID-Zenodo-Dep="10559566" httpUri="https://zenodo.org/record/10559566/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,232,1892,1918]" targetBox="[189,1393,487,1831]" targetPageId="31">
<paragraph id="8BC836A6FFE8FFABFF16F8B0A606F832" blockId="31.[189,1399,1892,2024]" pageId="31" pageNumber="32">
<emphasis id="B903EAB4FFE8FFABFF16F8B0A61FF8A5" bold="true" box="[189,281,1892,1919]" pageId="31" pageNumber="32">Fig. 15.</emphasis>
<taxonomicName id="4C774D25FFE8FFABFE8AF8B1A40AF8A5" authority="(Melichar, 1906)" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[289,780,1893,1919]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="gravidus" status="gen. et comb. nov.">
<emphasis id="B903EAB4FFE8FFABFE8AF8B1A546F8A5" box="[289,576,1893,1919]" italics="true" pageId="31" pageNumber="32">Melichergithus gravidus</emphasis>
(
<bibRefCitation id="EFE64B57FFE8FFABFDE4F8B1A402F8A5" author="Melichar L." box="[591,772,1893,1919]" pageId="31" pageNumber="32" pagination="1 - 327" refId="ref35004" refString="Melichar L. 1906. Monographie der Issiden (Homoptera). Abhandlungen der K. K. Zoologischbotanischen Gesellschaft in Wien 3: 1 - 327." type="journal article" year="1906">Melichar, 1906</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFE8FFABFCBFF8B1A4E8F8A5" box="[788,1006,1893,1919]" pageId="31" pageNumber="32" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, dissected ♂, typical morph from Ngoc Son-Ngo Luong Nature Reserve (RBINS). 
<emphasis id="B903EAB4FFE8FFABFCBDF85CA42DF878" bold="true" box="[790,811,1928,1954]" pageId="31" pageNumber="32">A</emphasis>
. Habitus, dorsal view. 
<emphasis id="B903EAB4FFE8FFABFBE3F85CA35AF878" bold="true" box="[1096,1116,1928,1954]" pageId="31" pageNumber="32">B</emphasis>
. Habitus, ventral view. 
<emphasis id="B903EAB4FFE8FFABFF16F87FA7D4F81F" bold="true" box="[189,210,1963,1989]" pageId="31" pageNumber="32">C</emphasis>
. Habitus, lateral view. 
<emphasis id="B903EAB4FFE8FFABFE4DF87FA6FCF81F" bold="true" box="[486,506,1963,1989]" pageId="31" pageNumber="32">D</emphasis>
. Habitus, perpendicular view of frons. 
<emphasis id="B903EAB4FFE8FFABFC66F87FA4E6F81F" bold="true" box="[973,992,1963,1989]" pageId="31" pageNumber="32">E</emphasis>
. Anterolateral view. 
<emphasis id="B903EAB4FFE8FFABFB7CF87FA3EFF81F" bold="true" box="[1239,1257,1963,1989]" pageId="31" pageNumber="32">F</emphasis>
. Right hind wing.
</paragraph>
</caption>
<caption id="DF08662EFFD7FF94FF16F85CA388F832" ID-DOI="http://doi.org/10.5281/zenodo.10559568" ID-Zenodo-Dep="10559568" httpUri="https://zenodo.org/record/10559568/files/figure.png" pageId="32" pageNumber="33" startId="32.[189,232,1928,1954]" targetBox="[193,1386,305,1841]" targetPageId="32">
<paragraph id="8BC836A6FFD7FF94FF16F85CA388F832" blockId="32.[189,1399,1928,2025]" pageId="32" pageNumber="33">
<emphasis id="B903EAB4FFD7FF94FF16F85CA618F878" bold="true" box="[189,286,1928,1954]" pageId="32" pageNumber="33">Fig. 16.</emphasis>
<taxonomicName id="4C774D25FFD7FF94FE80F85CA423F878" authority="(Melichar, 1906)" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[299,805,1928,1954]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="gravidus" status="gen. et comb. nov.">
<emphasis id="B903EAB4FFD7FF94FE80F85CA549F878" box="[299,591,1928,1954]" italics="true" pageId="32" pageNumber="33">Melichergithus gravidus</emphasis>
(
<bibRefCitation id="EFE64B57FFD7FF94FDC8F85CA41BF878" author="Melichar L." box="[611,797,1928,1954]" pageId="32" pageNumber="33" pagination="1 - 327" refId="ref35004" refString="Melichar L. 1906. Monographie der Issiden (Homoptera). Abhandlungen der K. K. Zoologischbotanischen Gesellschaft in Wien 3: 1 - 327." type="journal article" year="1906">Melichar, 1906</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFD7FF94FC99F85CA31DF878" box="[818,1051,1928,1954]" pageId="32" pageNumber="33" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, dissected ♂, typical morph from Mai Chau (VNMN). 
<emphasis id="B903EAB4FFD7FF94FE56F87FA517F81F" bold="true" box="[509,529,1963,1989]" pageId="32" pageNumber="33">A</emphasis>
. Habitus, dorsal view. 
<emphasis id="B903EAB4FFD7FF94FC8DF87FA43EF81F" bold="true" box="[806,824,1963,1989]" pageId="32" pageNumber="33">B</emphasis>
. Habitus, ventral view. 
<emphasis id="B903EAB4FFD7FF94FBFDF87FA36CF81F" bold="true" box="[1110,1130,1963,1989]" pageId="32" pageNumber="33">C</emphasis>
. Habitus, lateral view. 
<emphasis id="B903EAB4FFD7FF94FF14F81AA7D5F832" bold="true" box="[191,211,1998,2024]" pageId="32" pageNumber="33">D</emphasis>
. Habitus, perpendicular view of frons. 
<emphasis id="B903EAB4FFD7FF94FD35F81AA5B4F832" bold="true" box="[670,690,1998,2024]" pageId="32" pageNumber="33">E</emphasis>
. Anterolateral view. 
<emphasis id="B903EAB4FFD7FF94FC0EF81AA4B1F832" bold="true" box="[933,951,1998,2024]" pageId="32" pageNumber="33">F</emphasis>
. Right hind wing.
</paragraph>
</caption>
<caption id="DF08662EFFD6FF95FF16F895A6C5F832" ID-DOI="http://doi.org/10.5281/zenodo.10559570" ID-Zenodo-Dep="10559570" httpUri="https://zenodo.org/record/10559570/files/figure.png" pageId="33" pageNumber="34" startId="33.[189,232,1857,1883]" targetBox="[270,1316,275,1809]" targetPageId="33">
<paragraph id="8BC836A6FFD6FF95FF16F895A6C5F832" blockId="33.[188,1398,1857,2024]" pageId="33" pageNumber="34">
<emphasis id="B903EAB4FFD6FF95FF16F895A61FF881" bold="true" box="[189,281,1857,1883]" pageId="33" pageNumber="34">Fig. 17.</emphasis>
<taxonomicName id="4C774D25FFD6FF95FE8BF895A40DF881" authority="(Melichar, 1906)" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[288,779,1857,1883]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="gravidus" status="gen. et comb. nov.">
<emphasis id="B903EAB4FFD6FF95FE8BF895A539F881" box="[288,575,1857,1883]" italics="true" pageId="33" pageNumber="34">Melichergithus gravidus</emphasis>
(
<bibRefCitation id="EFE64B57FFD6FF95FDE6F895A405F881" author="Melichar L." box="[589,771,1857,1883]" pageId="33" pageNumber="34" pagination="1 - 327" refId="ref35004" refString="Melichar L. 1906. Monographie der Issiden (Homoptera). Abhandlungen der K. K. Zoologischbotanischen Gesellschaft in Wien 3: 1 - 327." type="journal article" year="1906">Melichar, 1906</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFD6FF95FCB9F896A4EAF886" box="[786,1004,1857,1884]" pageId="33" pageNumber="34" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, ♂, black morph from Dong SonKy Thuong Nature Reserve (RBINS), terminalia. 
<emphasis id="B903EAB4FFD6FF95FCB3F8B0A457F8A4" bold="true" box="[792,849,1892,1918]" pageId="33" pageNumber="34">A–C</emphasis>
. Pygofer, anal tube and gonostyli. 
<emphasis id="B903EAB4FFD6FF95FB57F8B0A217F8A4" bold="true" box="[1276,1297,1892,1918]" pageId="33" pageNumber="34">A</emphasis>
. Lateral view. 
<emphasis id="B903EAB4FFD6FF95FEAEF85CA61FF878" bold="true" box="[261,281,1928,1954]" pageId="33" pageNumber="34">B</emphasis>
. Caudal view. 
<emphasis id="B903EAB4FFD6FF95FE7AF85CA6E2F878" bold="true" box="[465,484,1928,1954]" pageId="33" pageNumber="34">C</emphasis>
. Dorsal view. 
<emphasis id="B903EAB4FFD6FF95FD3CF85CA5C2F878" bold="true" box="[663,708,1928,1954]" pageId="33" pageNumber="34">D–I</emphasis>
. Aedeagus and connective. 
<emphasis id="B903EAB4FFD6FF95FBBCF85CA32CF878" bold="true" box="[1047,1066,1928,1954]" pageId="33" pageNumber="34">D</emphasis>
. Left lateral view. 
<emphasis id="B903EAB4FFD6FF95FAB8F85CA222F878" bold="true" box="[1299,1316,1928,1954]" pageId="33" pageNumber="34">E</emphasis>
. Right lateral view. 
<emphasis id="B903EAB4FFD6FF95FEFFF87FA660F81F" bold="true" box="[340,358,1963,1989]" pageId="33" pageNumber="34">F</emphasis>
. Caudal view. 
<emphasis id="B903EAB4FFD6FF95FDBEF87FA52AF81F" bold="true" box="[533,556,1963,1989]" pageId="33" pageNumber="34">G</emphasis>
. Ventral view. 
<emphasis id="B903EAB4FFD6FF95FD77F87FA5F5F81F" bold="true" box="[732,755,1963,1989]" pageId="33" pageNumber="34">H</emphasis>
. Dorsal view. 
<emphasis id="B903EAB4FFD6FF95FC35F87FA4AFF81F" bold="true" box="[926,937,1963,1989]" pageId="33" pageNumber="34">I</emphasis>
. Anterodorsal view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF08662EFFD5FF96FF16F895A4D1F832" ID-DOI="http://doi.org/10.5281/zenodo.10559575" ID-Zenodo-Dep="10559575" httpUri="https://zenodo.org/record/10559575/files/figure.png" pageId="34" pageNumber="35" startId="34.[189,232,1857,1883]" targetBox="[232,1356,277,1790]" targetPageId="34">
<paragraph id="8BC836A6FFD5FF96FF16F895A4D1F832" blockId="34.[187,1399,1857,2025]" pageId="34" pageNumber="35">
<emphasis id="B903EAB4FFD5FF96FF16F895A61EF881" bold="true" box="[189,280,1857,1883]" pageId="34" pageNumber="35">Fig. 18.</emphasis>
<taxonomicName id="4C774D25FFD5FF96FEB6F895A400F881" authority="(Melichar, 1906)" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[285,774,1857,1883]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="gravidus" status="gen. et comb. nov.">
<emphasis id="B903EAB4FFD5FF96FEB6F895A53DF881" box="[285,571,1857,1883]" italics="true" pageId="34" pageNumber="35">Melichergithus gravidus</emphasis>
(
<bibRefCitation id="EFE64B57FFD5FF96FDE2F895A5F8F881" author="Melichar L." box="[585,766,1857,1883]" pageId="34" pageNumber="35" pagination="1 - 327" refId="ref35004" refString="Melichar L. 1906. Monographie der Issiden (Homoptera). Abhandlungen der K. K. Zoologischbotanischen Gesellschaft in Wien 3: 1 - 327." type="journal article" year="1906">Melichar, 1906</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFD5FF96FCA7F896A4E5F886" box="[780,995,1857,1884]" pageId="34" pageNumber="35" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, ♂, typical morph from Ngoc SonNgo Luong Nature Reserve (RBINS), terminalia. 
<emphasis id="B903EAB4FFD5FF96FD5CF8B0A436F8A4" bold="true" box="[759,816,1892,1918]" pageId="34" pageNumber="35">A–D</emphasis>
. Pygofer, anal tube and gonostyli. 
<emphasis id="B903EAB4FFD5FF96FB17F8B0A3D7F8A4" bold="true" box="[1212,1233,1892,1918]" pageId="34" pageNumber="35">A</emphasis>
. Lateral view. 
<emphasis id="B903EAB4FFD5FF96FF10F85CA7C9F878" bold="true" box="[187,207,1928,1954]" pageId="34" pageNumber="35">B</emphasis>
. Laterocaudal view. 
<emphasis id="B903EAB4FFD5FF96FE6CF85CA6DAF878" bold="true" box="[455,476,1928,1954]" pageId="34" pageNumber="35">C</emphasis>
. Dorsocaudal view. 
<emphasis id="B903EAB4FFD5FF96FD66F85CA5E4F878" bold="true" box="[717,738,1928,1954]" pageId="34" pageNumber="35">D</emphasis>
. Dorsal view. 
<emphasis id="B903EAB4FFD5FF96FC26F85CA4BBF878" bold="true" box="[909,957,1928,1954]" pageId="34" pageNumber="35">E–J</emphasis>
. Aedeagus and connective. 
<emphasis id="B903EAB4FFD5FF96FAADF85CA21FF878" bold="true" box="[1286,1305,1928,1954]" pageId="34" pageNumber="35">E</emphasis>
. Dorsal view. 
<emphasis id="B903EAB4FFD5FF96FEA8F87FA613F81F" bold="true" box="[259,277,1963,1989]" pageId="34" pageNumber="35">F</emphasis>
. Left lateral view. 
<emphasis id="B903EAB4FFD5FF96FE5EF87FA50DF81F" bold="true" box="[501,523,1963,1989]" pageId="34" pageNumber="35">G</emphasis>
. Right lateral view. 
<emphasis id="B903EAB4FFD5FF96FD57F87FA414F81F" bold="true" box="[764,786,1963,1989]" pageId="34" pageNumber="35">H</emphasis>
. Left lateroventral view. 
<emphasis id="B903EAB4FFD5FF96FB95F87FA34FF81F" bold="true" box="[1086,1097,1963,1989]" pageId="34" pageNumber="35">I</emphasis>
. Right laterocaudal view. 
<emphasis id="B903EAB4FFD5FF96FF15F81AA7CAF832" bold="true" box="[190,204,1998,2024]" pageId="34" pageNumber="35">J</emphasis>
. Left laterocaudal view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF08662EFFD4FF97FF16F85CA38CF832" ID-DOI="http://doi.org/10.5281/zenodo.10559577" ID-Zenodo-Dep="10559577" httpUri="https://zenodo.org/record/10559577/files/figure.png" pageId="35" pageNumber="36" startId="35.[189,232,1928,1954]" targetBox="[242,1346,265,1888]" targetPageId="35">
<paragraph id="8BC836A6FFD4FF97FF16F85CA38CF832" blockId="35.[189,1399,1928,2025]" pageId="35" pageNumber="36">
<emphasis id="B903EAB4FFD4FF97FF16F85CA61BF878" bold="true" box="[189,285,1928,1954]" pageId="35" pageNumber="36">Fig. 19.</emphasis>
<taxonomicName id="4C774D25FFD4FF97FE83F85CA418F878" authority="(Melichar, 1906)" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[296,798,1928,1954]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="gravidus" status="gen. et comb. nov.">
<emphasis id="B903EAB4FFD4FF97FE83F85CA54CF878" box="[296,586,1928,1954]" italics="true" pageId="35" pageNumber="36">Melichergithus gravidus</emphasis>
(
<bibRefCitation id="EFE64B57FFD4FF97FDF6F85CA410F878" author="Melichar L." box="[605,790,1928,1954]" pageId="35" pageNumber="36" pagination="1 - 327" refId="ref35004" refString="Melichar L. 1906. Monographie der Issiden (Homoptera). Abhandlungen der K. K. Zoologischbotanischen Gesellschaft in Wien 3: 1 - 327." type="journal article" year="1906">Melichar, 1906</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFD4FF97FC82F85CA30DF878" box="[809,1035,1928,1954]" pageId="35" pageNumber="36" rank="species">gen. et comb. nov.</taxonomicNameLabel>
<emphasis id="B903EAB4FFD4FF97FBBDF85CA32DF878" bold="true" box="[1046,1067,1928,1954]" pageId="35" pageNumber="36">A</emphasis>
. Distribution map. 
<emphasis id="B903EAB4FFD4FF97FAB0F85CA229F878" bold="true" box="[1307,1327,1928,1954]" pageId="35" pageNumber="36">B</emphasis>
. Live specimen on fern, Xuan Lien Nature Reserve, 11 Jul. 2023. 
<emphasis id="B903EAB4FFD4FF97FC31F87FA4A9F81F" bold="true" box="[922,943,1963,1989]" pageId="35" pageNumber="36">C</emphasis>
. Live specimen on fern, Cuc Phuong National Park, 20 Jul. 2011. 
<emphasis id="B903EAB4FFD4FF97FDA7F81AA527F832" bold="true" box="[524,545,1998,2024]" pageId="35" pageNumber="36">D</emphasis>
. Habitat in Cuc Phuong National Park, 20 Jul. 2011.
</paragraph>
</caption>
<subSubSection id="C36D652DFFD3FF90FF16FE44A64EFD0C" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8BC836A6FFD3FF90FF16FE44A650FE70" blockId="36.[189,342,400,426]" box="[189,342,400,426]" pageId="36" pageNumber="37">
<heading id="D08081CAFFD3FF90FF16FE44A650FE70" bold="true" box="[189,342,400,426]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B903EAB4FFD3FF90FF16FE44A650FE70" bold="true" box="[189,342,400,426]" pageId="36" pageNumber="37">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFD3FF90FF16FE6BA379FE27" blockId="36.[189,1399,447,509]" pageId="36" pageNumber="37">
North 
<collectingCountry id="F3607636FFD3FF90FEA3FE6BA66DFE03" box="[264,363,447,473]" name="Vietnam" pageId="36" pageNumber="37">Vietnam</collectingCountry>
and northern Central 
<collectingCountry id="F3607636FFD3FF90FDC7FE6BA5D2FE03" box="[620,724,447,473]" name="Vietnam" pageId="36" pageNumber="37">Vietnam</collectingCountry>
: provinces of 
<collectingRegion id="49B3F844FFD3FF90FCD3FE6BA4C5FE03" box="[888,963,447,473]" country="Vietnam" name="Ha Noi" pageId="36" pageNumber="37">Hanoi</collectingRegion>
, 
<collectingRegion id="49B3F844FFD3FF90FC65FE14A336FE03" box="[974,1072,447,474]" country="Vietnam" name="Ha Tinh" pageId="36" pageNumber="37">Ha Tinh</collectingRegion>
, 
<collectingRegion id="49B3F844FFD3FF90FB97FE14A3B6FE03" box="[1084,1200,447,474]" country="Vietnam" name="Hoa Binh" pageId="36" pageNumber="37">Hoa Binh</collectingRegion>
, 
<collectingRegion id="49B3F844FFD3FF90FB17FE14A229FE03" box="[1212,1327,447,474]" country="Vietnam" name="Lang Son" pageId="36" pageNumber="37">Lang Son</collectingRegion>
, 
<collectingRegion id="49B3F844FFD3FF90FA97FE6BA7FFFE26" country="Vietnam" name="Ninh Binh" pageId="36" pageNumber="37">Ninh Binh</collectingRegion>
, 
<collectingRegion id="49B3F844FFD3FF90FEAEFE36A66DFE26" box="[261,363,482,508]" country="Vietnam" name="Phu Tho" pageId="36" pageNumber="37">Phu Tho</collectingRegion>
, 
<collectingRegion id="49B3F844FFD3FF90FEDCFE37A50DFE26" box="[375,523,482,509]" country="Vietnam" name="Quang Ninh" pageId="36" pageNumber="37">Quang Ninh</collectingRegion>
, 
<collectingRegion id="49B3F844FFD3FF90FDBEFE36A5B4FE27" box="[533,690,482,509]" country="Vietnam" name="Thai Nguyen" pageId="36" pageNumber="37">Thai Nguyen</collectingRegion>
, 
<collectingRegion id="49B3F844FFD3FF90FD16FE36A447FE27" box="[701,833,482,509]" country="Vietnam" name="Thanh Hoa" pageId="36" pageNumber="37">Thanh Hoa</collectingRegion>
and 
<collectingRegion id="49B3F844FFD3FF90FCD2FE36A4F2FE26" box="[889,1012,482,508]" country="Vietnam" name="Vinh Phuc" pageId="36" pageNumber="37">Vinh Phuc</collectingRegion>
(
<figureCitation id="134C2A23FFD3FF90FBAFFE36A374FE27" box="[1028,1138,482,509]" captionStart="Fig" captionStartId="35.[189,232,1928,1954]" captionTargetBox="[242,1346,265,1888]" captionTargetId="figure-12@35.[242,1346,265,1888]" captionTargetPageId="35" captionText="Fig. 19. Melichergithus gravidus (Melichar, 1906) gen. et comb. nov. A. Distribution map. B. Live specimen on fern, Xuan Lien Nature Reserve, 11 Jul. 2023. C. Live specimen on fern, Cuc Phuong National Park, 20 Jul. 2011. D. Habitat in Cuc Phuong National Park, 20 Jul. 2011." figureDoi="http://doi.org/10.5281/zenodo.10559577" httpUri="https://zenodo.org/record/10559577/files/figure.png" pageId="36" pageNumber="37">Fig. 19A</figureCitation>
).
</paragraph>
<paragraph id="8BC836A6FFD3FF90FF16FDF0A7FEFDE4" blockId="36.[189,248,548,574]" box="[189,248,548,574]" pageId="36" pageNumber="37">
<heading id="D08081CAFFD3FF90FF16FDF0A7FEFDE4" bold="true" box="[189,248,548,574]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B903EAB4FFD3FF90FF16FDF0A7FEFDE4" bold="true" box="[189,248,548,574]" pageId="36" pageNumber="37">Note</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFD3FF90FF16FD86A64EFD0C" blockId="36.[188,1399,594,727]" pageId="36" pageNumber="37">
The specimens from 
<collectingRegion id="49B3F844FFD3FF90FE1BFD86A513FDB6" box="[432,533,594,620]" country="China" name="Guangxi" pageId="36" pageNumber="37">Guangxi</collectingRegion>
recorded and illustrated in 
<bibRefCitation id="EFE64B57FFD3FF90FCF9FD86A31EFDB7" author="Che Y. &amp; Zhang Y. - L. &amp; Wang Y. - L." box="[850,1048,594,621]" pageId="36" pageNumber="37" pagination="611 - 627" refId="ref32906" refString="Che Y. - L, Zhang Y. - L. &amp; Wang Y. - L. 2007. Seven new species and one new record of Gergithus Stal (Hemiptera: Fulgoroidea: Issidae) from China. Proceedings of the Entomological Society of Washington 109 (3): 611 - 627." type="journal article" year="2007">
Che 
<emphasis id="B903EAB4FFD3FF90FC2DFD87A4BEFDB6" box="[902,952,594,620]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. (2007)
</bibRefCitation>
, 
<bibRefCitation id="EFE64B57FFD3FF90FB88FD86A3F3FDB7" author="Chen X. - S. &amp; Zhang Z. - G. &amp; Chang Z. - M." box="[1059,1269,594,621]" pageId="36" pageNumber="37" refId="ref32963" refString="Chen X. - S., Zhang Z. - G. &amp; Chang Z. - M. 2014. Issidae and Caliscelidae from China. Guizhou Science and Technology Publishing House, Guiyang." type="book" year="2014">
Chen 
<emphasis id="B903EAB4FFD3FF90FBCDFD87A39EFDB6" box="[1126,1176,594,620]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. (2014)
</bibRefCitation>
and 
<bibRefCitation id="EFE64B57FFD3FF90FA80FD86A657FD4A" author="Zhang Y. - L. &amp; Che Y. - L. &amp; Meng R. &amp; Wang Y. - L." pageId="36" pageNumber="37" refId="ref36146" refString="Zhang Y. - L., Che Y. - L., Meng R. &amp; Wang Y. - L. 2020. Hemiptera. Caliscelidae. Issidae. Insecta. Fauna Sinica. Vol. 70. Science Press, Beijing." type="book" year="2020">
Zhang 
<emphasis id="B903EAB4FFD3FF90FF17FDA2A7F6FD4A" box="[188,240,630,656]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. (2020)
</bibRefCitation>
, might represent an undescribed species based on the male terminalia illustration in 
<bibRefCitation id="EFE64B57FFD3FF90FA92FDA2A648FD69" author="Chen X. - S. &amp; Zhang Z. - G. &amp; Chang Z. - M." pageId="36" pageNumber="37" refId="ref32963" refString="Chen X. - S., Zhang Z. - G. &amp; Chang Z. - M. 2014. Issidae and Caliscelidae from China. Guizhou Science and Technology Publishing House, Guiyang." type="book" year="2014">
Chen 
<emphasis id="B903EAB4FFD3FF90FF15FD4EA7F6FD69" box="[190,240,665,691]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. (2014)
</bibRefCitation>
, so the presence of 
<taxonomicName id="4C774D25FFD3FF90FD99FD4EA5C6FD69" authorityName="Constant &amp; Pham" authorityYear="2024" baseAuthorityName="Melichar" baseAuthorityYear="1906" box="[562,704,665,691]" class="Insecta" family="Issidae" genus="Melichergithus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="gravidus" status="gen. et comb. nov.">
<emphasis id="B903EAB4FFD3FF90FD99FD4EA5C6FD69" box="[562,704,665,691]" italics="true" pageId="36" pageNumber="37">M. gravidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFD3FF90FD6CFD4DA49AFD69" box="[711,924,665,691]" pageId="36" pageNumber="37" rank="species">gen. et comb. nov.</taxonomicNameLabel>
in 
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(hence in 
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) is regarded as doubtful.
</paragraph>
</subSubSection>
</treatment>
</document>