179 lines
15 KiB
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179 lines
15 KiB
XML
<document ID-DOI="10.5733/afin.053.0119" ID-ISSN="2305-2562" ID-Zenodo-Dep="7917198" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683666793957" checkinUser="felipe" docAuthor="Wagner, Rüdiger & Stuckenberg, Brian R." docDate="2012" docId="03DE879F2336FFE662FAFCA0C6E48605" docLanguage="en" docName="AfrInvertebr.53.1.355-367.pdf" docOrigin="African Invertebrates 53 (1)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.053.0119" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Nemopalpus velteni Wagner & Stuckenberg 2012, sp. n." docType="treatment" docVersion="2" lastPageNumber="358" masterDocId="FFE7FFE72335FFE2635DFFF8C435842D" masterDocTitle="New fossil and extant species of Nemopalpus Macquart (Diptera: Psychodidae: Bruchomyiinae)" masterLastPageNumber="355" masterPageNumber="355" pageNumber="357" updateTime="1683694472035" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New fossil and extant species of Nemopalpus Macquart (Diptera: Psychodidae: Bruchomyiinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wagner, Rüdiger</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Stuckenberg, Brian R.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2012-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>53</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>355</mods:start>
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<mods:end>355</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.053.0119</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5733/afin.053.0119</mods:identifier>
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<mods:identifier type="ISSN">2305-2562</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7917198</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03DE879F2336FFE662FAFCA0C6E48605" httpUri="http://treatment.plazi.org/id/03DE879F2336FFE662FAFCA0C6E48605" lastPageId="4" lastPageNumber="358" pageId="3" pageNumber="357">
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<subSubSection box="[423,824,856,882]" pageId="3" pageNumber="357" type="nomenclature">
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<paragraph blockId="3.[144,1105,856,1674]" box="[423,824,856,882]" pageId="3" pageNumber="357">
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<heading box="[423,824,856,882]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="357" reason="2">
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<taxonomicName ID-CoL="8CYTF" authority="Wagner" authorityName="Wagner & Stuckenberg" authorityYear="2012" box="[423,753,856,882]" class="Insecta" family="Psychodidae" genus="Nemopalpus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="357" phylum="Arthropoda" rank="species" species="velteni" status="sp. n.">
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<emphasis bold="true" box="[423,658,856,882]" pageId="3" pageNumber="357">Nemopalpus velteni</emphasis>
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Wagner
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</taxonomicName>
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,
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<taxonomicNameLabel box="[764,824,856,882]" pageId="3" pageNumber="357" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="358" pageId="3" pageNumber="357" type="description">
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<paragraph blockId="3.[144,1105,856,1674]" box="[575,672,896,922]" pageId="3" pageNumber="357">
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<figureCitation box="[575,672,896,922]" captionStart="Figs 1–4" captionStartId="2.[181,220,1582,1603]" captionTargetBox="[202,1125,976,1556]" captionTargetId="figure-371@2.[181,1141,972,1560]" captionTargetPageId="2" captionText="Figs 1–4. New fossil species of Nemopalpus: (1–3) N. velteni Wagner, sp. n., male: (1) habitus in amber; (2) head with mouthparts and basal antennomeres; (3) terminalia, dorsal view; (4) N. inexpectatus:DJQHU, VS. Q., PDOH WHUPLQDOLD, ODWHUDO YLHZ. 6RXUFHV RI ¿JXUHV:)LJV Ɨ± (5:);)LJ. 4 (%6 ZLWK additions by RW). Scale bars: Fig. 1 = 0.5 mm; Figs 2, 3 = 0.1 mm; Fig. 4 = scale unknown." figureDoi="http://doi.org/10.5281/zenodo.7917200" httpUri="https://zenodo.org/record/7917200/files/figure.png" pageId="3" pageNumber="357">Figs 1–3</figureCitation>
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</paragraph>
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<paragraph blockId="3.[144,1105,856,1674]" pageId="3" pageNumber="357">
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<emphasis bold="true" box="[144,1103,934,962]" pageId="3" pageNumber="357">(W\PRORJ\: 7KH VSHFL¿F QDPH LV GHGLFDWHG WR -üUJHQ 9HOWHQ (,GVWHLQ, *HUPDQ\), ZKR</emphasis>
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kindly provided the inclusion for study.
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</paragraph>
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<paragraph blockId="3.[144,1105,856,1674]" box="[144,283,1008,1034]" pageId="3" pageNumber="357">Description:</paragraph>
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<paragraph blockId="3.[144,1105,856,1674]" box="[144,208,1048,1074]" pageId="3" pageNumber="357">
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<emphasis box="[144,202,1050,1074]" italics="true" pageId="3" pageNumber="357">Male</emphasis>
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.
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</paragraph>
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<paragraph blockId="3.[144,1105,856,1674]" box="[144,826,1088,1114]" pageId="3" pageNumber="357">
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<emphasis box="[144,306,1090,1114]" italics="true" pageId="3" pageNumber="357">Measurements</emphasis>
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: Overall length
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<quantity box="[489,578,1088,1114]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="357" unit="mm" value="2.1">2.1 mm</quantity>
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. Wing length
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<quantity box="[733,822,1088,1114]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="357" unit="mm" value="2.2">2.2 mm</quantity>
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.
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</paragraph>
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<paragraph blockId="3.[144,1105,856,1674]" pageId="3" pageNumber="357">
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<emphasis box="[144,205,1130,1154]" italics="true" pageId="3" pageNumber="357">Head</emphasis>
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(
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<figureCitation box="[221,291,1128,1154]" captionStart="Figs 1–4" captionStartId="2.[181,220,1582,1603]" captionTargetBox="[202,1125,976,1556]" captionTargetId="figure-371@2.[181,1141,972,1560]" captionTargetPageId="2" captionText="Figs 1–4. New fossil species of Nemopalpus: (1–3) N. velteni Wagner, sp. n., male: (1) habitus in amber; (2) head with mouthparts and basal antennomeres; (3) terminalia, dorsal view; (4) N. inexpectatus:DJQHU, VS. Q., PDOH WHUPLQDOLD, ODWHUDO YLHZ. 6RXUFHV RI ¿JXUHV:)LJV Ɨ± (5:);)LJ. 4 (%6 ZLWK additions by RW). Scale bars: Fig. 1 = 0.5 mm; Figs 2, 3 = 0.1 mm; Fig. 4 = scale unknown." figureDoi="http://doi.org/10.5281/zenodo.7917200" httpUri="https://zenodo.org/record/7917200/files/figure.png" pageId="3" pageNumber="357">Fig. 2</figureCitation>
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): Typical of the genus, with circular eyes, no eye bridge. Mouthparts of male non-functional. Relative lengths of four clearly visible palpomeres: 20–15–25–25; overall length of palpus ~
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<quantity box="[432,538,1192,1219]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="357" unit="mm" value="0.56">0.56 mm</quantity>
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. Antenna with spherical scape and pedicel, and 14
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<emphasis bold="true" pageId="3" pageNumber="357">HORQJDWH WXEXODU ÀDJHOORPHUHV FORWKHG LQ ORQJ VHWDH. 'XH WR GHQVH YHVWLWXUH WKH SUHVHQFH RI DVFRLGV RQ ÀDJHOORPHUHV FRXOG QRW EH GHWHFWHG. 5HODWLYH OHQJWKV RI DQWHQQRPHUHV:</emphasis>
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8–8–26–21–22–22–20–17–16–16–15–14 –12–10–10–10; antennal length
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.6749999999999998" metricValueMax="1.7" metricValueMin="1.65" pageId="3" pageNumber="357" unit="mm" value="1.6749999999999998" valueMax="1.7" valueMin="1.65">1.65– 1.70mm</quantity>
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. Measurements of palpus and antennal segments are approximate as articles are not arranged in a single plane.
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</paragraph>
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<paragraph blockId="3.[144,1105,856,1674]" pageId="3" pageNumber="357">
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<emphasis box="[144,225,1394,1418]" italics="true" pageId="3" pageNumber="357">Thorax</emphasis>
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.
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<emphasis bold="true" box="[240,1103,1390,1419]" pageId="3" pageNumber="357">+DOWHUH DQG OHJV ODFNLQJ VSHFL¿F IHDWXUHV.:LQJV GLI¿FXOW WR UHFRQVWUXFW GXH</emphasis>
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to high degree of translucency, partly folded, with one wing tip missing. Wing
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<emphasis box="[1020,1046,1426,1450]" italics="true" pageId="3" pageNumber="357">ca</emphasis>
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2.5× longer than wide;
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<emphasis box="[356,382,1458,1482]" italics="true" pageId="3" pageNumber="357">Sc</emphasis>
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ending free in wing area at
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<emphasis box="[716,742,1458,1482]" italics="true" pageId="3" pageNumber="357">ca</emphasis>
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0.25 wing length;
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<emphasis box="[966,991,1458,1489]" italics="true" pageId="3" pageNumber="357">
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R
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<subScript attach="left" box="[983,991,1475,1489]" fontSize="6" pageId="3" pageNumber="357">1</subScript>
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</emphasis>
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elongate; fork
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<emphasis box="[198,243,1490,1521]" italics="true" pageId="3" pageNumber="357">R 2+3</emphasis>
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/
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<emphasis box="[252,296,1490,1521]" italics="true" pageId="3" pageNumber="357">R 4+5</emphasis>
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positioned basally; forks
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<emphasis box="[588,613,1490,1521]" italics="true" pageId="3" pageNumber="357">R 2</emphasis>
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+
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<emphasis box="[629,654,1490,1521]" italics="true" pageId="3" pageNumber="357">R 3</emphasis>
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and
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<emphasis box="[710,735,1490,1521]" italics="true" pageId="3" pageNumber="357">R 4</emphasis>
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+
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<emphasis box="[751,776,1490,1521]" italics="true" pageId="3" pageNumber="357">R 5</emphasis>
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at
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<emphasis box="[812,838,1490,1514]" italics="true" pageId="3" pageNumber="357">ca</emphasis>
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basal third of wing; tip positioned between
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<emphasis box="[368,393,1522,1553]" italics="true" pageId="3" pageNumber="357">R 4</emphasis>
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and
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<emphasis box="[447,472,1522,1553]" italics="true" pageId="3" pageNumber="357">R 5</emphasis>
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; fork
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<emphasis box="[541,592,1522,1553]" italics="true" pageId="3" pageNumber="357">M 1+2</emphasis>
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(most distal of forks) positioned at
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<emphasis box="[992,1018,1522,1546]" italics="true" pageId="3" pageNumber="357">ca</emphasis>
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middle of wing;
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<emphasis box="[246,277,1554,1585]" italics="true" pageId="3" pageNumber="357">M 3</emphasis>
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arising basally from
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<emphasis box="[518,568,1554,1585]" italics="true" pageId="3" pageNumber="357">M 1+2</emphasis>
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stem; short cross-vein positioned between
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<emphasis box="[1053,1103,1554,1585]" italics="true" pageId="3" pageNumber="357">M 1+2</emphasis>
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and
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<emphasis box="[192,223,1586,1617]" italics="true" pageId="3" pageNumber="357">M 3</emphasis>
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at short distance distally from that fork;
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<emphasis box="[689,746,1586,1617]" italics="true" pageId="3" pageNumber="357">CuA 1</emphasis>
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arising near point where cross-vein joins
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<emphasis box="[263,295,1618,1649]" italics="true" pageId="3" pageNumber="357">M 3</emphasis>
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;
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<emphasis box="[311,369,1618,1649]" italics="true" pageId="3" pageNumber="357">CuA 2</emphasis>
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moderately short, not reaching posterior wing margin; anal vein indistinguishable.
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</paragraph>
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<paragraph blockId="4.[181,1143,203,382]" pageId="4" pageNumber="358">
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<emphasis box="[181,287,206,230]" italics="true" pageId="4" pageNumber="358">Abdomen</emphasis>
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: With 8 pregenital segments, plus terminalia (
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<figureCitation box="[797,896,203,229]" captionStart="Figs 1–4" captionStartId="2.[181,220,1582,1603]" captionTargetBox="[202,1125,976,1556]" captionTargetId="figure-371@2.[181,1141,972,1560]" captionTargetPageId="2" captionText="Figs 1–4. New fossil species of Nemopalpus: (1–3) N. velteni Wagner, sp. n., male: (1) habitus in amber; (2) head with mouthparts and basal antennomeres; (3) terminalia, dorsal view; (4) N. inexpectatus:DJQHU, VS. Q., PDOH WHUPLQDOLD, ODWHUDO YLHZ. 6RXUFHV RI ¿JXUHV:)LJV Ɨ± (5:);)LJ. 4 (%6 ZLWK additions by RW). Scale bars: Fig. 1 = 0.5 mm; Figs 2, 3 = 0.1 mm; Fig. 4 = scale unknown." figureDoi="http://doi.org/10.5281/zenodo.7917200" httpUri="https://zenodo.org/record/7917200/files/figure.png" pageId="4" pageNumber="358">Figs 1, 3</figureCitation>
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). Terminalia inverted, segmentation forming torsion unclear. Terminalia relatively simple, gonocoxite pipeshaped, virtually straight, shorter than slightly bent gonostylus.Tergum 9 sub-rectangular,
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<emphasis bold="true" box="[181,1096,298,327]" pageId="4" pageNumber="358">ZLWK ÀHVK\, VHWRVH FHUFL; WHUJXP ƗƟ FRQLFDO. $HGHDJXV HORQJDWH WKLQ, WXEH‒VKDSHG.</emphasis>
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</paragraph>
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<paragraph blockId="4.[181,1143,203,382]" box="[181,1143,337,361]" pageId="4" pageNumber="358">
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<emphasis bold="true" box="[181,1143,337,361]" pageId="4" pageNumber="358">+RORW\SH: ƃ 0<$10$5: %XUPHVH DPEHU; &UHWDFHRXV, $OELDQ (SULYDWH FROOHFWLRQ RI -. 9HOWHQ,,GVWHLQ,</emphasis>
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</paragraph>
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<paragraph blockId="4.[181,1143,203,382]" lastBlockId="4.[181,1142,399,552]" pageId="4" pageNumber="358">
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<collectingCountry box="[181,270,362,383]" name="Germany" pageId="4" pageNumber="358">Germany</collectingCountry>
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). Notes:
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<taxonomicName authority="Wagner & Stuckenberg, 2012" authorityName="Wagner & Stuckenberg" authorityYear="2012" box="[264,489,400,424]" class="Insecta" family="Psychodidae" genus="Nemopalpus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="358" phylum="Arthropoda" rank="species" species="velteni" status="sp. n.">
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<emphasis box="[264,489,400,424]" italics="true" pageId="4" pageNumber="358">Nemopalpus velteni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[498,560,399,425]" pageId="4" pageNumber="358" rank="species">sp. n.</taxonomicNameLabel>
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represents the oldest known fossil
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<taxonomicName box="[969,1142,398,424]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="358" phylum="Arthropoda" rank="subFamily" subFamily="Bruchomyiinae">Bruchomyiinae</taxonomicName>
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and is distinguished by its overall smaller size and in the shape of the terminalia. The structure of the terminalia is much simpler than the geologically younger Baltic amber species
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<taxonomicName authority="Wagner & Stuckenberg, 2012" authorityName="Wagner & Stuckenberg" authorityYear="2012" box="[269,441,495,521]" class="Insecta" family="Psychodidae" genus="Nemopalpus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="358" phylum="Arthropoda" rank="species" species="inexpectatus" status="sp. n.">
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<emphasis box="[269,288,496,520]" italics="true" pageId="4" pageNumber="358">N</emphasis>
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.
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<emphasis box="[301,441,496,520]" italics="true" pageId="4" pageNumber="358">inexpectatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[448,508,495,521]" pageId="4" pageNumber="358" rank="species">sp. n.</taxonomicNameLabel>
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, described below. The specimen record greatly increases the known distribution of fossil
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<taxonomicName box="[540,716,526,552]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="358" phylum="Arthropoda" rank="subFamily" subFamily="Bruchomyiinae">Bruchomyiinae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |