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<document id="72DFE76079645713420CC012C045070E" ID-DOI="10.1649/0010-065X-69.2.297" ID-Zenodo-Dep="5332194" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630170744189" checkinUser="diego" docAuthor="Arce-Pérez, Roberto &amp; Novelo-Gutiérrez, Rodolfo" docDate="2015" docId="03D3F676FFADFF88FF6EFEAC3912F94B" docLanguage="en" docName="0010-065X-69.2.297.pdf" docOrigin="The Coleopterists Bulletin 69 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-69.2.297" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Neoclypeodytes moroni Arce-Perez and Novelo-Gutierrez 2015, new species" docType="treatment" docVersion="3" lastPageNumber="300" masterDocId="FFEA8E0EFFAFFF8BFFADFFB03A3FFF8A" masterDocTitle="Neoclypeodytes moroniArce-Pérez and Novelo-Gutiérrez, New Species (Coleoptera: Dytiscidae: Hydroporinae: Bidessini) from Mexico, with Notes on the Distribution ofNeoclypeodytes haroldiMiller" masterLastPageNumber="300" masterPageNumber="297" pageNumber="297" updateTime="1699172944065" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9A1CC6B4675078518C6760C21D9C1199">Neoclypeodytes moroniArce-Pérez and Novelo-Gutiérrez, New Species (Coleoptera: Dytiscidae: Hydroporinae: Bidessini) from Mexico, with Notes on the Distribution ofNeoclypeodytes haroldiMiller</mods:title>
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<mods:namePart id="3F4E1450AC26D69E268842116AD038B8">Arce-Pérez, Roberto</mods:namePart>
</mods:name>
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<mods:namePart id="C5ADA542F3E7D26851AC64536FB0B3A3">Novelo-Gutiérrez, Rodolfo</mods:namePart>
</mods:name>
<mods:typeOfResource id="186BEF1B908C26031F84CEF4B427293D">text</mods:typeOfResource>
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<mods:title id="D6D4A9D3496F3B1485830EAE63056ED1">The Coleopterists Bulletin</mods:title>
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<mods:date id="1BFDBE9B7CA81399C3A26E9D0800009E">2015</mods:date>
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<mods:number id="E4FDD3D218DB2BA97513EC2EE5A5A6A9">2015-06-30</mods:number>
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<mods:number id="5A7A3F09D0B614912A6FAE1C9BAE8428">69</mods:number>
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<mods:classification id="0544304E14D350FEF5DBB765E9F3751A">journal article</mods:classification>
<mods:identifier id="7726FAE67F15EE186339C8B548B1FECA" type="DOI">10.1649/0010-065X-69.2.297</mods:identifier>
<mods:identifier id="A745CD57AF15BDC8FC96AC9FF209203F" type="Zenodo-Dep">5332194</mods:identifier>
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<treatment id="03D3F676FFADFF88FF6EFEAC3912F94B" LSID="urn:lsid:plazi:treatment:03D3F676FFADFF88FF6EFEAC3912F94B" httpUri="http://treatment.plazi.org/id/03D3F676FFADFF88FF6EFEAC3912F94B" lastPageId="3" pageId="2">
<subSubSection id="C36014EBFFADFF89FF6EFEAC3818FEC6" pageId="2" type="nomenclature">
<paragraph id="8BC54760FFADFF89FF6EFEAC3818FEC6" blockId="2.[195,594,284,359]" pageId="2">
<heading id="D08DF00CFFADFF89FF6EFEAC3818FEC6" centered="true" fontSize="10" level="2" pageId="2" reason="2">
<emphasis id="B90E9B72FFADFF89FF6EFEAC3818FEC6" bold="true" pageId="2">
<taxonomicName id="4C7A3CE3FFADFF89FF6EFEAC3B9BFEC6" ID-CoL="8LZW8" authority="Arce-Perez and Novelo-Gutierrez" authorityName="Arce-Perez and Novelo-Gutierrez" authorityYear="2015" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="moroni" status="sp. nov.">
<emphasis id="B90E9B72FFADFF89FF6EFEAC3B98FEBB" bold="true" box="[195,423,284,305]" italics="true" pageId="2">Neoclypeodytes moroni</emphasis>
Arce-Pérez and Novelo-Gutiérrez
</taxonomicName>
,
<taxonomicNameLabel id="A23D2609FFADFF89FE1CFE873818FEC6" box="[433,551,311,332]" pageId="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C36014EBFFADFF89FEF8FEE13BFFFEEC" box="[341,448,337,359]" pageId="2" type="description">
<paragraph id="8BC54760FFADFF89FEF8FEE13BFFFEEC" blockId="2.[195,594,284,359]" box="[341,448,337,359]" pageId="2">
(
<figureCitation id="13415BE5FFADFF89FEF1FEE13B86FEEC" box="[348,441,337,359]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="2.[175,215,1686,1705]" captionStartId-1="3.[197,238,738,757]" captionTargetBox-0="[153,1167,1075,1663]" captionTargetBox-1="[173,1193,193,715]" captionTargetId-0="figure-452@2.[151,1171,1075,1680]" captionTargetId-1="figure-513@3.[173,1193,193,715]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs. 13. Neoclypeodytes moroni. 1) Habitus, dorsal view; 2) Habitus, ventral view; 3) Head showing the shape of clypeus and frontal punctation, dorsal view." captionText-1="Figs. 46. Male genitalia of Neoclypeodytes moroni. 4) Median lobe of aedeagus, lateral view; 5) Median lobe of aedeagus, ventral view; 6) Paramere, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.5332198" figureDoi-1="http://doi.org/10.5281/zenodo.5332202" httpUri-0="https://zenodo.org/record/5332198/files/figure.png" httpUri-1="https://zenodo.org/record/5332202/files/figure.png" pageId="2">Figs. 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36014EBFFADFF89FF02FECC3BF5FD3F" pageId="2" type="materials_examined">
<paragraph id="8BC54760FFADFF89FF02FECC3BF5FD3F" blockId="2.[151,639,380,1012]" pageId="2">
<emphasis id="B90E9B72FFADFF89FF02FECC3B05FE1B" bold="true" box="[175,314,380,401]" pageId="2">Type Material.</emphasis>
<materialsCitation id="3B124D3DFFADFF89FEEBFECC3840FE4C" collectingDate="1997-07-30" collectionCode="R" collectorName="Arce" country="Mexico" elevation="850" location="Laguna Miradores" municipality="Municipality of Emiliano Zapata" pageId="2" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<typeStatus id="54C1F9C2FFADFF89FEEBFECC3BA5FE1B" box="[326,410,380,401]" pageId="2" type="holotype">Holotype</typeStatus>
male, labeled:
<collectingCountry id="F36D07F0FFADFF89FDB3FECC3845FE1B" box="[542,634,380,401]" name="Mexico" pageId="2">MÉXICO</collectingCountry>
,
<collectingRegion id="49BE8982FFADFF89FF3AFE273AD3FE26" box="[151,236,407,428]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="2">Veracruz</collectingRegion>
,
<collectingMunicipality id="6BA1DD1AFFADFF89FF5BFE273812FE26" box="[246,557,407,428]" pageId="2">Municipality of Emiliano Zapata</collectingMunicipality>
,
<location id="8EA511BBFFADFF89FD95FE273AC2FE4C" LSID="urn:lsid:plazi:treatment:03D3F676FFADFF88FF6EFEAC3912F94B:8EA511BBFFADFF89FD95FE273AC2FE4C" country="Mexico" municipality="Municipality of Emiliano Zapata" name="Laguna Miradores" pageId="2" stateProvince="Veracruz de Ignacio de la Llave">Laguna Miradores</location>
,
<quantity id="4C82EA85FFADFF89FEA0FE013B6CFE4C" box="[269,339,433,454]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="2" unit="m" value="850.0">
<elevation id="0057A053FFADFF89FEA0FE013B6CFE4C" box="[269,339,433,454]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="2" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<date id="FFC461A0FFADFF89FECEFE013BD3FE4C" box="[355,492,433,454]" pageId="2" value="1997-07-30">
<collectingDate id="EF809848FFADFF89FECEFE013BD3FE4C" box="[355,492,433,454]" pageId="2" value="1997-07-30">30. vii. 1997</collectingDate>
</date>
,
<collectionCode id="ED6BDFA5FFADFF89FE50FE01382FFE4C" box="[509,528,433,454]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2">R</collectionCode>
.
<collectorName id="268F22B6FFADFF89FDB2FE013871FE4C" box="[543,590,433,454]" pageId="2">Arce</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B124D3DFFADFF89FF3AFE7C3AFCFE71" collectingDate="1997-07-30" collectionCode="R" collectorName="Arce" country="Mexico" elevation="850" location="Laguna Miradores" municipality="Municipality of Emiliano Zapata" pageId="2" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<typeStatus id="54C1F9C2FFADFF89FF3AFE7C3ACBFE6B" box="[151,244,460,481]" pageId="2" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D7C8CE9FFADFF89FEACFE7C3B70FE6B" box="[257,335,460,481]" pageId="2" type="male">3 males</specimenCount>
,
<specimenCount id="9D7C8CE9FFADFF89FEF6FE7C3B85FE6B" box="[347,442,460,481]" pageId="2" type="female">2 females</specimenCount>
, same data as holotype
</materialsCitation>
;
<materialsCitation id="3B124D3DFFADFF89FF7FFE563B7DFD9C" collectingDate="1997-05-02" collectionCode="R" collectorName="Arce" country="Mexico" elevation="850" location="Laguna Miradores" municipality="Municipality of Emiliano Zapata" pageId="2" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<specimenCount id="9D7C8CE9FFADFF89FF7FFE563B25FE71" box="[210,282,486,507]" pageId="2" type="male">1 male</specimenCount>
,
<specimenCount id="9D7C8CE9FFADFF89FE87FE563BB2FE71" box="[298,397,486,507]" pageId="2" type="female">4 females</specimenCount>
, same data as holotype except
<date id="FFC461A0FFADFF89FF70FDB13B7DFD9C" box="[221,322,513,534]" pageId="2" value="1997-05-02">
<collectingDate id="EF809848FFADFF89FF70FDB13B7DFD9C" box="[221,322,513,534]" pageId="2" value="1997-05-02">2. v. 1997</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B124D3DFFADFF89FEE3FDB13B6DFDBA" collectingDate="1997-05-18" collectionCode="R" collectorName="Arce" country="Mexico" elevation="850" location="Laguna Miradores" municipality="Municipality of Emiliano Zapata" pageId="2" specimenCount="3" specimenCount-male="3" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<specimenCount id="9D7C8CE9FFADFF89FEE3FDB13BA3FD9C" box="[334,412,513,534]" pageId="2" type="male">3 males</specimenCount>
, same data as holotype except
<date id="FFC461A0FFADFF89FF73FDAB3B6DFDBA" box="[222,338,539,560]" pageId="2" value="1997-05-18">
<collectingDate id="EF809848FFADFF89FF73FDAB3B6DFDBA" box="[222,338,539,560]" pageId="2" value="1997-05-18">18. v. 1997</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B124D3DFFADFF89FEF2FDAB3BB6FDC1" collectingDate="1997-10-30" collectionCode="R" collectorName="Arce" country="Mexico" elevation="850" location="Laguna Miradores" municipality="Municipality of Emiliano Zapata" pageId="2" specimenCount="1" specimenCount-female="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<specimenCount id="9D7C8CE9FFADFF89FEF2FDAB3B89FDBA" box="[351,438,539,560]" pageId="2" type="female">1 female</specimenCount>
, same data as holotype except
<date id="FFC461A0FFADFF89FEBEFD863BB6FDC1" box="[275,393,566,587]" pageId="2" value="1997-10-30">
<collectingDate id="EF809848FFADFF89FEBEFD863BB6FDC1" box="[275,393,566,587]" pageId="2" value="1997-10-30">30. x. 1997</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B124D3DFFADFF89FE3AFD863BA4FDEC" collectingDate="1998-01-11" collectionCode="R" collectorName="Arce" country="Mexico" elevation="850" location="Laguna Miradores" municipality="Municipality of Emiliano Zapata" pageId="2" specimenCount="1" specimenCount-female="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<specimenCount id="9D7C8CE9FFADFF89FE3AFD863BCFFDC1" box="[407,496,566,587]" pageId="2" type="female">1 female</specimenCount>
, same data as holotype except
<date id="FFC461A0FFADFF89FE9EFDE13BA4FDEC" box="[307,411,593,614]" pageId="2" value="1998-01-11">
<collectingDate id="EF809848FFADFF89FE9EFDE13BA4FDEC" box="[307,411,593,614]" pageId="2" value="1998-01-11">11. i. 1998</collectingDate>
</date>
</materialsCitation>
, all
<collectionCode id="ED6BDFA5FFADFF89FE69FDE13BE8FDEC" box="[452,471,593,614]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2">R</collectionCode>
.
<collectorName id="268F22B6FFADFF89FE4DFDE13832FDEC" box="[480,525,593,614]" pageId="2">Arce</collectorName>
leg;
<materialsCitation id="3B124D3DFFADFF89FD93FDE13B78FD3F" collectingDate="2010-09-30" collectorName="Colonia San Pedro" country="Mexico" location="Municipality of Coatepec" pageId="2" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<specimenCount id="9D7C8CE9FFADFF89FD93FDE13842FDEC" box="[574,637,593,614]" pageId="2" type="male">1 male</specimenCount>
labeled:
<collectingCountry id="F36D07F0FFADFF89FF58FDDB3B64FD0A" box="[245,347,619,640]" name="Mexico" pageId="2">MÉXICO</collectingCountry>
,
<collectingRegion id="49BE8982FFADFF89FEC7FDDB3BF6FD0A" box="[362,457,619,640]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="2">Veracruz</collectingRegion>
,
<location id="8EA511BBFFADFF89FE75FDDB3ACFFD11" LSID="urn:lsid:plazi:treatment:03D3F676FFADFF88FF6EFEAC3912F94B:8EA511BBFFADFF89FE75FDDB3ACFFD11" country="Mexico" name="Municipality of Coatepec" pageId="2" stateProvince="Veracruz de Ignacio de la Llave">Municipality of Coatepec</location>
, Coatepec,
<collectorName id="268F22B6FFADFF89FEF3FD36382CFD11" box="[350,531,646,667]" pageId="2">Colonia San Pedro</collectorName>
, pool aside road,
<date id="FFC461A0FFADFF89FF62FD103B78FD3F" box="[207,327,672,693]" pageId="2" value="2010-09-30">
<collectingDate id="EF809848FFADFF89FF62FD103B78FD3F" box="[207,327,672,693]" pageId="2" value="2010-09-30">30. ix. 2010</collectingDate>
</date>
</materialsCitation>
,
<collectionCode id="ED6BDFA5FFADFF89FEFEFD103B59FD3F" box="[339,358,672,693]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2">R</collectionCode>
.
<collectorName id="268F22B6FFADFF89FEDCFD103BA1FD3F" box="[369,414,672,693]" pageId="2">Arce</collectorName>
leg.
</paragraph>
</subSubSection>
<subSubSection id="C36014EBFFADFF88FF02FD0B3B24FA67" lastPageId="3" pageId="2" type="description">
<paragraph id="8BC54760FFADFF88FF02FD0B3825FAE2" blockId="2.[151,639,380,1012]" lastBlockId="3.[173,660,858,1730]" lastPageId="3" pageId="2">
<emphasis id="B90E9B72FFADFF89FF02FD0B3B18FD5A" bold="true" box="[175,295,699,720]" pageId="2">Description.</emphasis>
<typeStatus id="54C1F9C2FFADFF89FE99FD0B3BB3FD5A" box="[308,396,699,720]" pageId="2" type="holotype">Holotype</typeStatus>
. Male [In square brackets are male
<typeStatus id="54C1F9C2FFADFF89FF47FD653B07FD60" box="[234,312,725,746]" pageId="2" type="paratype">paratype</typeStatus>
variations,
<emphasis id="B90E9B72FFADFF89FE0CFD653B92FD61" box="[417,429,725,747]" italics="true" pageId="2">n</emphasis>
= 6]. Body (
<figureCitation id="13415BE5FFADFF89FD8CFD653849FD60" box="[545,630,725,747]" captionStart="Figs" captionStartId="2.[175,215,1686,1705]" captionTargetBox="[153,1167,1075,1663]" captionTargetId="figure-452@2.[151,1171,1075,1680]" captionTargetPageId="2" captionText="Figs. 13. Neoclypeodytes moroni. 1) Habitus, dorsal view; 2) Habitus, ventral view; 3) Head showing the shape of clypeus and frontal punctation, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5332198" httpUri="https://zenodo.org/record/5332198/files/figure.png" pageId="2">Figs. 12</figureCitation>
) oval, elongate, convex dorsally, in dorsal view with margins of pronotum and elytra continuous. Total length (from apex of clypeus to apex of elytra)
<quantity id="4C82EA85FFADFF89FF3AFCF03AD4FCDF" box="[151,235,832,853]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="2" unit="mm" value="2.25">2.25 mm</quantity>
[range 2.152.30, mean = 2.20], maximum width
<quantity id="4C82EA85FFADFF89FF75FCEA3B0EFCE5" box="[216,305,858,879]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" unit="mm" value="1.2">1.20 mm</quantity>
[range 1.151.25, mean = 1.18]. Integument dark brown [yellowish].
<emphasis id="B90E9B72FFADFF89FE50FCC53805FC00" bold="true" box="[509,570,885,906]" pageId="2">Head:</emphasis>
Wider than long (
<figureCitation id="13415BE5FFADFF89FEA5FC3F3B78FC2E" box="[264,327,911,932]" captionStart="Figs" captionStartId="2.[175,215,1686,1705]" captionTargetBox="[153,1167,1075,1663]" captionTargetId="figure-452@2.[151,1171,1075,1680]" captionTargetPageId="2" captionText="Figs. 13. Neoclypeodytes moroni. 1) Habitus, dorsal view; 2) Habitus, ventral view; 3) Head showing the shape of clypeus and frontal punctation, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5332198" httpUri="https://zenodo.org/record/5332198/files/figure.png" pageId="2">Fig. 3</figureCitation>
); dark brown; compound eyes silvery; antennae, palpi, and legs yellow [reddish]; pronotum and elytra yellow with irregular, dark brown spot; clypeus elongate, trapezoid, fissured, with margins raised and apex slightly rounded and smooth; compound eyes large, lateral; frontal region with 2 rows of fine punctures bearing short, golden setae, 1 of these rows along internal margin of eye, other starting on frons without entering clypeus; coarse punctures on frons; 1 stria behind eyes; antenna 11-segmented, scape long, pedicel globose, first 8 antennomeres short, subtriangular, last antennomere long, acuminate; maxillary palp 4-segmented, 2
<superScript id="7C0FEA28FFADFF89FC91FE1C3971FE30" attach="left" box="[828,846,428,442]" fontSize="6" pageId="2">nd</superScript>
and 3
<superScript id="7C0FEA28FFADFF89FC25FE1C39A8FE30" attach="left" box="[904,919,428,442]" fontSize="6" pageId="2">rd</superScript>
segments slightly globose, 4
<superScript id="7C0FEA28FFADFF89FD14FE7638F8FE5E" attach="left" box="[697,711,454,468]" fontSize="6" pageId="2">th</superScript>
acuminate and as long as 2 preceding together; labial palp short, 3-segmented, first 2 segments globose, apical segment acuminate, bifid at apex.
<emphasis id="B90E9B72FFADFF89FD00FDAB3926FDBA" bold="true" box="[685,793,539,560]" pageId="2">Pronotum:</emphasis>
Convex; light yellow; with longitudinal, medial, dark brown band extending laterally on posterior half of pronotum as inverted “T”; shallow, sinuate striae extending over 0.45 of posterior part of pronotum; punctation coarser than that on head, more dense in interstrial area.
<emphasis id="B90E9B72FFADFF89FC65FD103E32FD3F" bold="true" box="[968,1037,672,693]" pageId="2">Elytra:</emphasis>
Convex; light yellow; without sutural stria; most of surface with irregular pattern of dark spots; punctation coarser than that on pronotum; punctures separated by 12 times diameter of each puncture and distributed in parallel lines along length of elytron; striae shallow, short, occupying 0.20 basal length of elytra and slightly sinuate; with parallel lines of punctures bearing sparse, delicate, golden setae.
<emphasis id="B90E9B72FFADFF89FB9EFCC53EACFC00" bold="true" box="[1075,1171,885,906]" pageId="2">Sternum:</emphasis>
Prosternum yellow; convex; prosternal process reddish yellow, elongated, convex, apically lanceolate and fissured at middle, with a longitudinal sulcus. Metaventrite and metacoxae (
<figureCitation id="13415BE5FFADFF89FC7CFC6F3E31FC7E" box="[977,1038,991,1012]" captionStart="Figs" captionStartId="2.[175,215,1686,1705]" captionTargetBox="[153,1167,1075,1663]" captionTargetId="figure-452@2.[151,1171,1075,1680]" captionTargetPageId="2" captionText="Figs. 13. Neoclypeodytes moroni. 1) Habitus, dorsal view; 2) Habitus, ventral view; 3) Head showing the shape of clypeus and frontal punctation, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5332198" httpUri="https://zenodo.org/record/5332198/files/figure.png" pageId="2">Fig. 2</figureCitation>
) with sparse, rough, moderately coarse punctation; length of metacoxa (ML1) along metacoxal suture
<quantity id="4C82EA85FFACFF88FD91FCC538ADFC00" box="[572,658,885,906]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="3" unit="mm" value="0.31">0.31 mm</quantity>
[
<quantity id="4C82EA85FFACFF88FF1FFC3F3B78FC2E" box="[178,327,911,933]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="3" unit="mm" value="0.315" valueMax="0.32" valueMin="0.31">0.310.32 mm</quantity>
]; maximum width of medial portion of metacoxa (MW1) from metacoxal suture and perpendicular to metacoxal line
<quantity id="4C82EA85FFACFF88FD9BFC7438ACFC53" box="[566,659,964,985]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="3" unit="mm" value="0.08">0.08 mm</quantity>
[
<quantity id="4C82EA85FFACFF88FF1FFC6F3B6FFC7E" box="[178,336,990,1012]" metricMagnitude="-5" metricUnit="m" metricValue="8.499999999999998" metricValueMax="9.0" metricValueMin="8.0" pageId="3" unit="mm" value="0.08499999999999999" valueMax="0.09" valueMin="0.08">0.080.09 mm</quantity>
], ratio ML1/MW1 3.65 [3.55 3.87].
<emphasis id="B90E9B72FFACFF88FF46FC493B6BFB84" bold="true" box="[235,340,1017,1038]" pageId="3">Abdomen:</emphasis>
Convex; with 6 visible ventrites; 2
<superScript id="7C0FEA28FFACFF88FF17FBBF3AF2FB97" attach="left" box="[186,205,1039,1053]" fontSize="6" pageId="3">nd</superScript>
ventrite as large as following 4 ventrites together; ventrites 12 with coarse punctures on margins and medial portion; ventrites 45 bearing medial row of long, golden setae; ventrite 6 (anal plate) with dense, coarse punctation and minute, golden setae.
<emphasis id="B90E9B72FFACFF88FE87FB293BB4FB24" bold="true" box="[298,395,1177,1198]" pageId="3">Genitalia:</emphasis>
Median lobe (
<figureCitation id="13415BE5FFACFF88FDBDFB293878FB24" box="[528,583,1177,1198]" captionStart="Figs" captionStartId="3.[197,238,738,757]" captionTargetBox="[173,1193,193,715]" captionTargetId="figure-513@3.[173,1193,193,715]" captionTargetPageId="3" captionText="Figs. 46. Male genitalia of Neoclypeodytes moroni. 4) Median lobe of aedeagus, lateral view; 5) Median lobe of aedeagus, ventral view; 6) Paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5332202" httpUri="https://zenodo.org/record/5332202/files/figure.png" pageId="3">Fig. 4</figureCitation>
) shorter than parameres, slender and curved, in lateral view with medial portion not expanded, convergent to rounded apex; in ventral view (
<figureCitation id="13415BE5FFACFF88FE47FB593819FB74" box="[490,550,1257,1278]" captionStart="Figs" captionStartId="3.[197,238,738,757]" captionTargetBox="[173,1193,193,715]" captionTargetId="figure-513@3.[173,1193,193,715]" captionTargetPageId="3" captionText="Figs. 46. Male genitalia of Neoclypeodytes moroni. 4) Median lobe of aedeagus, lateral view; 5) Median lobe of aedeagus, ventral view; 6) Paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5332202" httpUri="https://zenodo.org/record/5332202/files/figure.png" pageId="3">Fig. 5</figureCitation>
) wide and sinuate, concave in basal third, convex at middle, convergent in apical third, ending in rounded point; parameres (
<figureCitation id="13415BE5FFACFF88FEB0FA883B66FAC7" box="[285,345,1336,1357]" captionStart="Figs" captionStartId="3.[197,238,738,757]" captionTargetBox="[173,1193,193,715]" captionTargetId="figure-513@3.[173,1193,193,715]" captionTargetPageId="3" captionText="Figs. 46. Male genitalia of Neoclypeodytes moroni. 4) Median lobe of aedeagus, lateral view; 5) Median lobe of aedeagus, ventral view; 6) Paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5332202" httpUri="https://zenodo.org/record/5332202/files/figure.png" pageId="3">Fig. 6</figureCitation>
) narrow, with apical 0.43 elongated and slender, with setae at apex.
</paragraph>
<caption id="DF0517E8FFADFF89FF02F9263815F94B" ID-DOI="http://doi.org/10.5281/zenodo.5332198" ID-Zenodo-Dep="5332198" httpUri="https://zenodo.org/record/5332198/files/figure.png" pageId="2" startId="2.[175,215,1686,1705]" targetBox="[153,1167,1075,1663]" targetPageId="2">
<paragraph id="8BC54760FFADFF89FF02F9263815F94B" blockId="2.[151,1172,1686,1729]" pageId="2">
<emphasis id="B90E9B72FFADFF89FF02F9263B34F923" bold="true" box="[175,267,1686,1705]" pageId="2">Figs. 13.</emphasis>
<taxonomicName id="4C7A3CE3FFADFF89FE8DF9263BD8F923" authorityName="Arce-Perez and Novelo-Gutierrez" authorityYear="2015" box="[288,487,1686,1705]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="moroni">
<emphasis id="B90E9B72FFADFF89FE8DF9263BD8F923" box="[288,487,1686,1705]" italics="true" pageId="2">Neoclypeodytes moroni</emphasis>
</taxonomicName>
.
<emphasis id="B90E9B72FFADFF89FE59F9263BC0F923" bold="true" box="[500,511,1686,1705]" pageId="2">1</emphasis>
) Habitus, dorsal view;
<emphasis id="B90E9B72FFADFF89FD67F92638EAF923" bold="true" box="[714,725,1686,1705]" pageId="2">2</emphasis>
) Habitus, ventral view;
<emphasis id="B90E9B72FFADFF89FC05F926398CF923" bold="true" box="[936,947,1686,1705]" pageId="2">3</emphasis>
) Head showing the shape of clypeus and frontal punctation, dorsal view.
</paragraph>
</caption>
<caption id="DF0517E8FFACFF88FF68FD52384BFC87" ID-DOI="http://doi.org/10.5281/zenodo.5332202" ID-Zenodo-Dep="5332202" httpUri="https://zenodo.org/record/5332202/files/figure.png" pageId="3" startId="3.[197,238,738,757]" targetBox="[173,1193,193,715]" targetPageId="3">
<paragraph id="8BC54760FFACFF88FF68FD52384BFC87" blockId="3.[173,1193,738,781]" pageId="3">
<emphasis id="B90E9B72FFACFF88FF68FD523B1DFD7F" bold="true" box="[197,290,738,757]" pageId="3">Figs. 46.</emphasis>
Male genitalia of
<taxonomicName id="4C7A3CE3FFACFF88FE78FD5238A1FD7F" authorityName="Arce-Perez and Novelo-Gutierrez" authorityYear="2015" box="[469,670,738,757]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="moroni">
<emphasis id="B90E9B72FFACFF88FE78FD5238A1FD7F" box="[469,670,738,757]" italics="true" pageId="3">Neoclypeodytes moroni</emphasis>
</taxonomicName>
.
<emphasis id="B90E9B72FFACFF88FD06FD523889FD7F" bold="true" box="[683,694,738,757]" pageId="3">4</emphasis>
) Median lobe of aedeagus, lateral view;
<emphasis id="B90E9B72FFACFF88FB8FFD523E12FD7F" bold="true" box="[1058,1069,738,757]" pageId="3">5</emphasis>
) Median lobe of aedeagus, ventral view;
<emphasis id="B90E9B72FFACFF88FE3BFD4A3B9EFC87" bold="true" box="[406,417,762,781]" pageId="3">6</emphasis>
) Paramere, lateral view.
</paragraph>
</caption>
<paragraph id="8BC54760FFACFF88FF68FADD3B24FA67" blockId="3.[173,660,858,1730]" pageId="3">
<emphasis id="B90E9B72FFACFF88FF68FADD3B2AFA08" bold="true" box="[197,277,1389,1410]" pageId="3">Female.</emphasis>
Similar to male except: Body slightly more robust, total length 2.152.35 (mean = 2.25,
<emphasis id="B90E9B72FFACFF88FF00FA123A86FA32" box="[173,185,1442,1464]" italics="true" pageId="3">n</emphasis>
= 7), maximum width 1.151.25 (mean = 1.22); margin of clypeus convex, not raised at apex, dorsal surface flat.
</paragraph>
</subSubSection>
<subSubSection id="C36014EBFFACFF88FF68FA4239FCFB42" pageId="3" type="diagnosis">
<paragraph id="8BC54760FFACFF88FF68FA4239FCFB42" blockId="3.[173,660,858,1730]" lastBlockId="3.[707,1194,857,1729]" pageId="3">
<emphasis id="B90E9B72FFACFF88FF68FA423B12F98D" bold="true" box="[197,301,1522,1543]" pageId="3">Diagnosis.</emphasis>
By the shape of the clypeus and the large metacoxal ratio, we consider
<taxonomicName id="4C7A3CE3FFACFF88FE55F9BC3868F9A8" authorityName="Arce-Perez and Novelo-Gutierrez" authorityYear="2015" box="[504,599,1548,1570]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="moroni">
<emphasis id="B90E9B72FFACFF88FE55F9BC3868F9A8" box="[504,599,1548,1570]" italics="true" pageId="3">N. moroni</emphasis>
</taxonomicName>
likely to be closely related to
<taxonomicName id="4C7A3CE3FFACFF88FE31F9973AD7F9DD" authority="Miller" authorityName="Miller" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="anasinus">
<emphasis id="B90E9B72FFACFF88FE31F99738ACF9B7" box="[412,659,1575,1597]" italics="true" pageId="3">Neoclypeodytes anasinus</emphasis>
Miller
</taxonomicName>
from
<collectingCountry id="F36D07F0FFACFF88FE8AF9F23B4CF9DD" box="[295,371,1602,1623]" name="Jamaica" pageId="3">Jamaica</collectingCountry>
(Greater Antilles). Both species share the unique characteristic of a projecting, trapezoidal clypeus (
<figureCitation id="13415BE5FFACFF88FED2F9C73B80F906" box="[383,447,1655,1676]" captionStart="Figs" captionStartId="2.[175,215,1686,1705]" captionTargetBox="[153,1167,1075,1663]" captionTargetId="figure-452@2.[151,1171,1075,1680]" captionTargetPageId="2" captionText="Figs. 13. Neoclypeodytes moroni. 1) Habitus, dorsal view; 2) Habitus, ventral view; 3) Head showing the shape of clypeus and frontal punctation, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5332198" httpUri="https://zenodo.org/record/5332198/files/figure.png" pageId="3">Fig. 2</figureCitation>
), which is unlike all other species of
<taxonomicName id="4C7A3CE3FFACFF88FEFAF9213BCEF92D" authorityName="Young" authorityYear="1967" box="[343,497,1681,1703]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90E9B72FFACFF88FEFAF9213BCEF92D" box="[343,497,1681,1703]" italics="true" pageId="3">Neoclypeodytes</emphasis>
</taxonomicName>
known to date. The main differences between
<taxonomicName id="4C7A3CE3FFACFF88FE57F91C385CF948" authorityName="Arce-Perez and Novelo-Gutierrez" authorityYear="2015" box="[506,611,1708,1730]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="moroni">
<emphasis id="B90E9B72FFACFF88FE57F91C385CF948" box="[506,611,1708,1730]" italics="true" pageId="3">N. moroni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7A3CE3FFACFF88FD6EFCE93901FCE5" box="[707,830,857,879]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="anasinus">
<emphasis id="B90E9B72FFACFF88FD6EFCE93901FCE5" box="[707,830,857,879]" italics="true" pageId="3">N. anasinus</emphasis>
</taxonomicName>
are (characters of
<taxonomicName id="4C7A3CE3FFACFF88FBA3FCE93EB6FCE5" box="[1038,1161,857,879]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="anasinus">
<emphasis id="B90E9B72FFACFF88FBA3FCE93EB6FCE5" box="[1038,1161,857,879]" italics="true" pageId="3">N. anasinus</emphasis>
</taxonomicName>
in parenthesis): Total length of body from anterior margin of clypeus to apex of elytra
<quantity id="4C82EA85FFACFF88FBFAFC3F3E96FC2E" box="[1111,1193,911,932]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" unit="mm" value="2.5">2.5 mm</quantity>
(
<quantity id="4C82EA85FFACFF88FD64FC1A3919FC35" box="[713,806,938,959]" metricMagnitude="-3" metricUnit="m" metricValue="2.13" pageId="3" unit="mm" value="2.13">2.13 mm</quantity>
), greatest width dorsally across widest point of body
<quantity id="4C82EA85FFACFF88FCCDFC743982FC53" box="[864,957,964,985]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" unit="mm" value="1.2">1.20 mm</quantity>
(
<quantity id="4C82EA85FFACFF88FC7CFC743E0CFC53" box="[977,1075,964,985]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="3" unit="mm" value="1.16">1.16 mm</quantity>
), length of basal pronotal stria from base to discernible apex 0.45 (0.52), length of basal elytral stria from base to discernible apex 0.20 (0.16), ratio ML1/MW1 3.553.87 (5.50); frontal region of head with two rows of fine punctures (without rows of fine punctures); elytra without sutural stria (with sutural stria); male genitalia with median lobe strongly convergent in apical third, ending in a rounded tip (not strongly convergent in apical third).
</paragraph>
</subSubSection>
<subSubSection id="C36014EBFFACFF88FD76FB7E3E7CFB77" pageId="3" type="materials_examined">
<paragraph id="8BC54760FFACFF88FD76FB7E3E7CFB77" blockId="3.[707,1194,857,1729]" pageId="3">
<materialsCitation id="3B124D3DFFACFF88FD76FB7E3E01FB77" country="Mexico" location="Laguna Miradores" municipality="Municipality of Emiliano Zapata" pageId="3" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<emphasis id="B90E9B72FFACFF88FD76FB7E3955FB69" bold="true" box="[731,874,1230,1251]" pageId="3">
<typeStatus id="54C1F9C2FFACFF88FD76FB7E3934FB69" box="[731,779,1230,1251]" pageId="3">Type</typeStatus>
Locality.
</emphasis>
<collectingCountry id="F36D07F0FFACFF88FCD5FB7E39FCFB69" box="[888,963,1230,1251]" name="Mexico" pageId="3">Mexico</collectingCountry>
,
<collectingRegion id="49BE8982FFACFF88FC60FB7E3E1CFB69" box="[973,1059,1230,1251]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3">Veracruz</collectingRegion>
,
<collectingMunicipality id="6BA1DD1AFFACFF88FB82FB7E39BFFB77" pageId="3">Municipality of Emiliano Zapata</collectingMunicipality>
,
<location id="8EA511BBFFACFF88FC26FB583E01FB77" LSID="urn:lsid:plazi:treatment:03D3F676FFADFF88FF6EFEAC3912F94B:8EA511BBFFACFF88FC26FB583E01FB77" box="[907,1086,1256,1277]" country="Mexico" municipality="Municipality of Emiliano Zapata" name="Laguna Miradores" pageId="3" stateProvince="Veracruz de Ignacio de la Llave">Laguna Miradores</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36014EBFFACFF88FD76FAB33912F94B" pageId="3" type="etymology">
<paragraph id="8BC54760FFACFF88FD76FAB33E66FAC7" blockId="3.[707,1194,857,1729]" pageId="3">
<emphasis id="B90E9B72FFACFF88FD76FAB33972FA92" bold="true" box="[731,845,1283,1304]" pageId="3">Etymology.</emphasis>
We gladly dedicate this species to Dr. Miguel Angel Morón Ríos for his contribution to the study of the
<taxonomicName id="4C7A3CE3FFACFF88FCD3FA8839D9FAC7" box="[894,998,1336,1357]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="3" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
of
<collectingRegion id="49BE8982FFACFF88FBA4FA883E6AFAC7" box="[1033,1109,1336,1357]" country="Mexico" name="Mexico" pageId="3">Mexico</collectingRegion>
.
</paragraph>
<paragraph id="8BC54760FFACFF88FD76FAE33912F94B" blockId="3.[707,1194,857,1729]" pageId="3">
<emphasis id="B90E9B72FFACFF88FD76FAE339AEFAE2" bold="true" box="[731,913,1363,1384]" pageId="3">New State Records.</emphasis>
Hitherto, only
<taxonomicName id="4C7A3CE3FFACFF88FBB0FAE239AEFA08" authority="Sharp" authorityName="Sharp" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="quadrinotatus">
<emphasis id="B90E9B72FFACFF88FBB0FAE23971FA09" italics="true" pageId="3">Neoclypeodytes quadrinotatus</emphasis>
Sharp
</taxonomicName>
had been recorded for the state of
<collectingRegion id="49BE8982FFACFF88FCA2FA38395EFA17" box="[783,865,1416,1437]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3">Veracruz</collectingRegion>
(Córdoba). Herein, two more species are incorporated into the fauna of this Mexican state:
<taxonomicName id="4C7A3CE3FFACFF88FD57FA0D3967FA59" authorityName="Arce-Perez and Novelo-Gutierrez" authorityYear="2015" box="[762,856,1469,1491]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="moroni">
<emphasis id="B90E9B72FFACFF88FD57FA0D3967FA59" box="[762,856,1469,1491]" italics="true" pageId="3">N. moroni</emphasis>
</taxonomicName>
, here described from the Municipalities of Emiliano Zapata and Coatepec; and the first record of
<taxonomicName id="4C7A3CE3FFACFF88FCCAFA423E96F98D" authority="Miller" authorityName="Miller" box="[871,1193,1522,1544]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="haroldi">
<emphasis id="B90E9B72FFACFF88FCCAFA423E62F982" box="[871,1117,1522,1544]" italics="true" pageId="3">Neoclypeodytes haroldi</emphasis>
Miller
</taxonomicName>
(
<specimenCount id="9D7C8CE9FFACFF88FD61F9BD3927F9A8" box="[716,792,1549,1570]" pageId="3" type="male">1 male</specimenCount>
,
<specimenCount id="9D7C8CE9FFACFF88FC84F9BD39ABF9A8" box="[809,916,1549,1570]" pageId="3" type="female">2 females</specimenCount>
, labeled: Municipality of Coatepec, Coatepec, Río Ahuacatlán,
<geoCoordinate id="EE4E21A7FFACFF88FB86F9973E9AF9B6" box="[1067,1189,1575,1597]" degrees="19" direction="north" minutes="28" orientation="latitude" pageId="3" precision="15" seconds="20" value="19.472223">19°2820″ N</geoCoordinate>
;
<geoCoordinate id="EE4E21A7FFACFF88FD6EF9F2397FF9DD" box="[707,832,1601,1623]" degrees="96" direction="west" minutes="59" orientation="longitude" pageId="3" precision="15" seconds="05" value="-96.98472">96°5905″ W</geoCoordinate>
,
<quantity id="4C82EA85FFACFF88FCE6F9F239AAF9DD" box="[843,917,1602,1623]" metricMagnitude="3" metricUnit="m" metricValue="1.369" pageId="3" unit="m" value="1369.0">1369 m</quantity>
asl, pool close to a cascade,
<date id="FFC461A0FFACFF88FD6EF9EC3905F9FB" box="[707,826,1628,1649]" pageId="3" value="2010-09-30">30. ix. 2010</date>
, R. Arce leg), which was only known from
<collectingRegion id="49BE8982FFACFF88FD55F9C73975F906" box="[760,842,1655,1676]" country="Mexico" name="Durango" pageId="3">Durango</collectingRegion>
and
<collectingRegion id="49BE8982FFACFF88FCD0F9C739F9F906" box="[893,966,1655,1676]" country="Mexico" name="Sinaloa" pageId="3">Sinaloa</collectingRegion>
. This extends considerably its distributional range to the southeastern part of
<collectingCountry id="F36D07F0FFACFF88FD72F91C3916F94B" box="[735,809,1708,1729]" name="Mexico" pageId="3">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>