261 lines
31 KiB
XML
261 lines
31 KiB
XML
<document id="51167559211DDD58205C508D2C8F1AAE" ID-DOI="10.11646/phytotaxa.332.2.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="13722080" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725664193039" checkinUser="felipe" docAuthor="Cortés-Ceballos, Olga Lorena, Giraldo-Rodríguez, Andrés & Torres-Gonzalez, Alba Marina" docDate="2017" docId="03AA87CDFFAF927AFF0CFF13FDEE3BAB" docLanguage="en" docName="phytotaxa.332.2.5.pdf" docOrigin="Phytotaxa 332 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.332.2.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Besleria physaloides O. L. Cortes 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="185" masterDocId="FF93FFB5FFAE927EFF84FF8DFF903F02" masterDocTitle="Two new species of Besleria (Gesneriaceae) from the department of Valle del Cauca, Colombia" masterLastPageNumber="188" masterPageNumber="181" pageNumber="182" updateTime="1725675436918" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A103BFE1AD4C973124868EA6DE4B8D13">Two new species of Besleria (Gesneriaceae) from the department of Valle del Cauca, Colombia</mods:title>
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<mods:namePart id="C9AEB036CB9660911A9577196E2DE189">Cortés-Ceballos, Olga Lorena</mods:namePart>
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<mods:namePart id="C553160562B744D9D19B800A65DADAE9">Giraldo-Rodríguez, Andrés</mods:namePart>
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<mods:namePart id="385E90A4A47DDF637816446EC9639B7A">Torres-Gonzalez, Alba Marina</mods:namePart>
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<mods:title id="C3AE68ECD36336025E9C737D03C56952">Phytotaxa</mods:title>
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<mods:part id="A172A3400FA8F85321C31B5F553041C5">
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<mods:date id="CFA0BE70301013228D7EB50BF6FA6A02">2017</mods:date>
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<mods:number id="E2997C6972C51C9B2E4F14378571CDEF">2017-12-19</mods:number>
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<mods:number id="8092D58EBFBC27E69587058980A5CCF7">332</mods:number>
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<mods:url id="E289640E39AAA846925C7A66377E2EF3">http://dx.doi.org/10.11646/phytotaxa.332.2.5</mods:url>
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<treatment id="03AA87CDFFAF927AFF0CFF13FDEE3BAB" LSID="urn:lsid:plazi:treatment:03AA87CDFFAF927AFF0CFF13FDEE3BAB" httpUri="http://treatment.plazi.org/id/03AA87CDFFAF927AFF0CFF13FDEE3BAB" lastPageId="4" lastPageNumber="185" pageId="1" pageNumber="182">
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<subSubSection id="C3196550FFAF927FFF0CFF13FCAC3FBB" box="[136,828,158,185]" pageId="1" pageNumber="182" type="nomenclature">
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<paragraph id="8BBC36DBFFAF927FFF0CFF13FCAC3FBB" blockId="1.[136,828,158,185]" box="[136,828,158,185]" pageId="1" pageNumber="182">
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<heading id="D0F481B7FFAF927FFF0CFF13FCAC3FBB" box="[136,828,158,185]" fontSize="11" level="2" pageId="1" pageNumber="182" reason="3">
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<taxonomicName id="4C034D58FFAF927FFF0CFF13FE6A3FBB" ID-CoL="5WJXJ" authority="O.L.Cortes" authorityName="O. L. Cortes" authorityYear="2017" box="[136,506,158,185]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="182" phylum="Tracheophyta" rank="species" species="physaloides" status="sp. nov.">
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<emphasis id="B977EAC9FFAF927FFF0CFF13FEE43FBB" bold="true" box="[136,372,158,185]" italics="true" pageId="1" pageNumber="182">Besleria physaloides</emphasis>
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O.L.Cortés
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24457B2FFAF927FFD81FF2DFDCA3FBB" box="[517,602,160,185]" pageId="1" pageNumber="182" rank="species">
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<emphasis id="B977EAC9FFAF927FFD81FF2DFDC43FBB" box="[517,596,160,185]" italics="true" pageId="1" pageNumber="182">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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, (
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<figureCitation id="13382A5EFFAF927FFDF6FF2DFD2F3FBB" box="[626,703,160,185]" captionStart="FIGURE 1" captionStartId="1.[136,229,2021,2042]" captionTargetBox="[213,1335,458,1973]" captionTargetId="figure-115@1.[201,1386,433,1995]" captionTargetPageId="1" captionText="FIGURE 1. Besleria physaloides: A. habit; B. flower outer view; C. dorsal sepal D. lateral sepal; E ventral sepal; F. corolla (front view); G. corolla inner view; H gynoecium and disc annular. Drawn by Alba Marina Torres-Gonzalez." figureDoi="http://doi.org/10.5281/zenodo.13722082" httpUri="https://zenodo.org/record/13722082/files/figure.png" pageId="1" pageNumber="182">Figs. 1</figureCitation>
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,
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<figureCitation id="13382A5EFFAF927FFD49FF2DFC8A3FBB" box="[717,794,160,185]" captionStart="FIGURE 3" captionStartId="3.[136,229,1996,2017]" captionTargetBox="[240,1348,175,1971]" captionTargetId="figure-14@3.[240,1348,175,1971]" captionTargetPageId="3" captionText="FIGURE 3. Pollen micromorphology. A–D: Besleria physaloides. A. polar view; B. polar view (SEM); C. equatorial view (SEM); D. ornamentation (SEM). E–H: Besleria silverstoneana. E. polar view; F. polar view (SEM); G. equatorial view (SEM); H. ornamentation (SEM)." figureDoi="http://doi.org/10.5281/zenodo.13722086" httpUri="https://zenodo.org/record/13722086/files/figure.png" pageId="1" pageNumber="182">3 A–D</figureCitation>
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,
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<figureCitation id="13382A5EFFAF927FFCA1FF2DFCA43FBB" box="[805,820,160,185]" captionStart="FIGURE 4" captionStartId="5.[136,229,2056,2077]" captionTargetBox="[191,1408,526,2017]" captionTargetId="figure-163@5.[177,1410,515,2034]" captionTargetPageId="5" captionText="FIGURE 4. Geographical distribution of Besleria physaloides and Besleria silverstoneana in Valle del Cauca, Colombia." figureDoi="http://doi.org/10.5281/zenodo.13722088" httpUri="https://zenodo.org/record/13722088/files/figure.png" pageId="1" pageNumber="182">4</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3196550FFAF927AFF0CFF69FCBC3C67" lastPageId="4" lastPageNumber="185" pageId="1" pageNumber="182" type="materials_examined">
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<paragraph id="8BBC36DBFFAF927FFF0CFF69FEAD3E43" blockId="1.[136,1452,228,429]" pageId="1" pageNumber="182">
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<emphasis id="B977EAC9FFAF927FFF0CFF69FF2A3FFB" bold="true" box="[136,186,228,249]" pageId="1" pageNumber="182">Type</emphasis>
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:—
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<collectingCountry id="F314764BFFAF927FFF53FF69FECD3FFB" box="[215,349,228,249]" name="Colombia" pageId="1" pageNumber="182">COLOMBIA</collectingCountry>
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.
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<collectingRegion id="49C7F839FFAF927FFEE7FF69FE6B3FFB" box="[355,507,228,249]" country="Colombia" name="Valle del Cauca" pageId="1" pageNumber="182">Valle del Cauca</collectingRegion>
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: municipio Dagua, corregimiento San Bernardo, vía a Buenaventura, cordillera occidental, vertiente oriental,
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,
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<geoCoordinate id="EE37501CFFAF927FFE2CFE85FDA93E1F" box="[424,569,264,285]" degrees="76" direction="west" minutes="37" orientation="longitude" pageId="1" pageNumber="182" precision="1" seconds="27.8" value="-76.62439">76°37’27.8’’W</geoCoordinate>
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,
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,
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<date id="FFBD101BFFAF927FFD10FE85FC843E1F" box="[660,788,264,285]" pageId="1" pageNumber="182" value="2013-05-18">18 May 2013</date>
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,
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<emphasis id="B977EAC9FFAF927FFC99FE85FBB33E1F" box="[797,1059,264,285]" italics="true" pageId="1" pageNumber="182">
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<collectionCode id="ED12AE1EFFAF927FFC99FE85FCBF3E1F" box="[797,815,264,285]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="1" pageNumber="182" type="Herbarium">A</collectionCode>
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. Giraldo-Rodríguez 8021
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</emphasis>
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(
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<typeStatus id="54B88879FFAF927FFBAAFE85FB143E1F" box="[1070,1156,264,285]" pageId="1" pageNumber="182" type="holotype">holotype</typeStatus>
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<collectionCode id="ED12AE1EFFAF927FFB0DFE85FB5C3E1F" box="[1161,1228,264,285]" country="Colombia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15729" lsid="urn:lsid:biocol.org:col:15729" name="Universidad del Valle" pageId="1" pageNumber="182" type="Herbarium">CUVC</collectionCode>
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67092!,
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<typeStatus id="54B88879FFAF927FFAA4FE85FAFF3E1F" box="[1312,1391,264,285]" pageId="1" pageNumber="182" type="isotype">isotypes</typeStatus>
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<collectionCode id="ED12AE1EFFAF927FFAF7FE85FA383E1F" box="[1395,1448,264,285]" country="Colombia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15629" lsid="urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="1" pageNumber="182" type="Herbarium">COL</collectionCode>
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!
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<collectionCode id="ED12AE1EFFAF927FFF39FEA1FF663E43" box="[189,246,300,321]" country="Colombia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15677" lsid="urn:lsid:biocol.org:col:15677" name="Universidad de Antioquia" pageId="1" pageNumber="182" type="Herbarium">HUA</collectionCode>
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!
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<collectionCode id="ED12AE1EFFAF927FFF7BFEA1FEBD3E43" box="[255,301,300,321]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="1" pageNumber="182" type="Herbarium">MO</collectionCode>
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!).
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</paragraph>
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<paragraph id="8BBC36DBFFAF927FFF0CFEDDFC533EAF" blockId="1.[136,1452,228,429]" pageId="1" pageNumber="182">
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<emphasis id="B977EAC9FFAF927FFF0CFEDDFC533EAF" italics="true" pageId="1" pageNumber="182">
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This species differs from
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<taxonomicName id="4C034D58FFAF927FFEFFFEDDFDBF3E67" authorityName="Hanstein" authorityYear="1865" box="[379,559,336,357]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="182" phylum="Tracheophyta" rank="species" species="barbensis">Besleria barbensis</taxonomicName>
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in having broader, unequal sepals, which are only slightly serrate and usually purple at the base (sepals equal, serrate, and light green in
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<taxonomicName id="4C034D58FFAF927FFD02FEF9FC923E8B" authorityName="Hanstein" authorityYear="1865" box="[646,770,372,393]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="182" phylum="Tracheophyta" rank="species" species="barbensis">B. barbensis</taxonomicName>
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), shorter petioles, and elliptic to oblanceolate, entire leaf blades with fewer pairs of veins (blades elliptic and slightly serrate distally in
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<taxonomicName id="4C034D58FFAF927FFCB8FE15FC273EAF" authorityName="Hanstein" authorityYear="1865" box="[828,951,408,429]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="182" phylum="Tracheophyta" rank="species" species="barbensis">B. barbensis</taxonomicName>
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).
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</emphasis>
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</paragraph>
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<caption id="DF7C6653FFAF927FFF0CF868FB8B371C" ID-DOI="http://doi.org/10.5281/zenodo.13722082" ID-Zenodo-Dep="13722082" httpUri="https://zenodo.org/record/13722082/files/figure.png" pageId="1" pageNumber="182" startId="1.[136,229,2021,2042]" targetBox="[213,1335,458,1973]" targetPageId="1" targetType="figure">
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<paragraph id="8BBC36DBFFAF927FFF0CF868FB8B371C" blockId="1.[136,1451,2020,2078]" pageId="1" pageNumber="182">
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<emphasis id="B977EAC9FFAF927FFF0CF868FE5E38F8" bold="true" box="[136,462,2020,2042]" pageId="1" pageNumber="182">
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FIGURE 1.
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<taxonomicName id="4C034D58FFAF927FFE87F869FE5E38F8" authorityName="O. L. Cortes" authorityYear="2017" box="[259,462,2020,2042]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="182" phylum="Tracheophyta" rank="species" species="physaloides">
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<emphasis id="B977EAC9FFAF927FFE87F869FE5E38F8" bold="true" box="[259,462,2020,2042]" italics="true" pageId="1" pageNumber="182">Besleria physaloides</emphasis>
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</taxonomicName>
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</emphasis>
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: A. habit; B. flower outer view; C. dorsal sepal D. lateral sepal; E ventral sepal; F. corolla (front view); G. corolla inner view; H gynoecium and disc annular. Drawn by Alba
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<taxonomicName id="4C034D58FFAF927FFCADF784FB8B371C" authority="Torres-Gonzalez." authorityName="Torres-Gonzalez." box="[809,1051,2057,2078]" class="Magnoliopsida" family="Fabaceae" genus="Marina" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="182" phylum="Tracheophyta" rank="genus">Marina Torres-Gonzalez.</taxonomicName>
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</paragraph>
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||
</caption>
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||
<caption id="DF7C6653FFAC927CFF0CF866FC4F3726" ID-DOI="http://doi.org/10.5281/zenodo.13722084" ID-Zenodo-Dep="13722084" httpUri="https://zenodo.org/record/13722084/files/figure.png" pageId="2" pageNumber="183" startId="2.[136,229,2027,2048]" targetBox="[151,1434,190,2002]" targetPageId="2" targetType="figure">
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<paragraph id="8BBC36DBFFAC927CFF0CF866FC4F3726" blockId="2.[136,1452,2026,2084]" pageId="2" pageNumber="183">
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||
<emphasis id="B977EAC9FFAC927CFF0CF866FE7A3702" bold="true" box="[136,490,2026,2048]" pageId="2" pageNumber="183">
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FIGURE 2.
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<taxonomicName id="4C034D58FFAC927CFE80F867FE7A3702" authorityName="O. L. Cortes" authorityYear="2017" box="[260,490,2026,2048]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="183" phylum="Tracheophyta" rank="species" species="silverstoneana">
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||
<emphasis id="B977EAC9FFAC927CFE80F867FE7A3702" bold="true" box="[260,490,2026,2048]" italics="true" pageId="2" pageNumber="183">Besleria silverstoneana</emphasis>
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</taxonomicName>
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</emphasis>
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: A. habit; B. dorsal sepal; C. ventral sepal; D. lateral sepal; E. flower (front view); F. flower (lateral view); G. corolla inner view; H. gynoecium and disc annular. Drawn by Álvaro Vanegas.
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</paragraph>
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||
</caption>
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<caption id="DF7C6653FFAD927DFF0CF841FF40372B" ID-DOI="http://doi.org/10.5281/zenodo.13722086" ID-Zenodo-Dep="13722086" httpUri="https://zenodo.org/record/13722086/files/figure.png" pageId="3" pageNumber="184" startId="3.[136,229,1996,2017]" targetBox="[240,1348,175,1971]" targetPageId="3" targetType="figure">
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||
<paragraph id="8BBC36DBFFAD927DFF0CF841FF40372B" blockId="3.[136,1452,1995,2089]" pageId="3" pageNumber="184">
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||
<emphasis id="B977EAC9FFAD927DFF0CF841FF6F38E3" bold="true" box="[136,255,1996,2017]" pageId="3" pageNumber="184">FIGURE 3.</emphasis>
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Pollen micromorphology. A–D:
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<taxonomicName id="4C034D58FFAD927DFDC4F846FC9C38E3" authorityName="O. L. Cortes" authorityYear="2017" box="[576,780,1995,2017]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="184" phylum="Tracheophyta" rank="species" species="physaloides">
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||
<emphasis id="B977EAC9FFAD927DFDC4F846FC9C38E3" bold="true" box="[576,780,1995,2017]" italics="true" pageId="3" pageNumber="184">Besleria physaloides</emphasis>
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</taxonomicName>
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. A. polar view; B. polar view (SEM); C. equatorial view (SEM); D. ornamentation (SEM). E–H:
|
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<taxonomicName id="4C034D58FFAD927DFE25F862FD183707" authorityName="O. L. Cortes" authorityYear="2017" box="[417,648,2031,2053]" class="Magnoliopsida" family="Gesneriaceae" genus="Besleria" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="184" phylum="Tracheophyta" rank="species" species="silverstoneana">
|
||
<emphasis id="B977EAC9FFAD927DFE25F862FD183707" bold="true" box="[417,648,2031,2053]" italics="true" pageId="3" pageNumber="184">Besleria silverstoneana</emphasis>
|
||
</taxonomicName>
|
||
. E. polar view; F. polar view (SEM); G. equatorial view (SEM); H. ornamentation (SEM).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBC36DBFFAA927AFF0CFF2DFA913DFB" blockId="4.[136,1453,160,1193]" pageId="4" pageNumber="185">
|
||
Terrestrial shrub; stem erect,
|
||
<quantity id="4CFB9B3EFFAA927AFE50FF2DFDD63FBB" box="[468,582,160,185]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="4" pageNumber="185" unit="m" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 m</quantity>
|
||
tall, terete, hirsute toward apex; branches arising from base or
|
||
<quantity id="4CFB9B3EFFAA927AFA9DFF2DFAF03FBB" box="[1305,1376,160,185]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="185" unit="cm" value="10.0">10 cm</quantity>
|
||
above base,
|
||
<quantity id="4CFB9B3EFFAA927AFF43FF49FEBD3FDF" box="[199,301,196,221]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="3.0" metricValueMin="0.6" pageId="4" pageNumber="185" unit="cm" value="1.8" valueMax="3.0" valueMin="0.6">0.6–3 cm</quantity>
|
||
diam. Leaves opposite, decussate, members of each pair almost equal in length; petiole hirsute,
|
||
<quantity id="4CFB9B3EFFAA927AFAD4FF49FA3C3FDF" box="[1360,1452,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="8.0" metricValueMin="1.0" pageId="4" pageNumber="185" unit="mm" value="4.5" valueMax="8.0" valueMin="1.0">1–8 mm</quantity>
|
||
long; blade elliptic to oblanceolate, 4–12.6 ×
|
||
<quantity id="4CFB9B3EFFAA927AFDFFFF65FD6A3E03" box="[635,762,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="3.5" metricValueMin="1.2" pageId="4" pageNumber="185" unit="cm" value="2.35" valueMax="3.5" valueMin="1.2">1.2–3.5 cm</quantity>
|
||
, apex acuminate and mucronate, base attenuate, margin entire, 7–9 pairs of veins, pilose on adaxial surface, hirsute along midvein of abaxial surface. Inflorescence axillary, pedicels hirsute,
|
||
<quantity id="4CFB9B3EFFAA927AFF64FEBDFECA3E4B" box="[224,346,304,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="4" pageNumber="185" unit="cm" value="1.85" valueMax="2.5" valueMin="1.2">1.2–2.5 cm</quantity>
|
||
long. Flowers solitary or in pairs. Calyx with five free sepals that cover corolla tube loosely, unequal, ovate, with acuminate apex, very narrow near tip, margin slightly serrate, pale green with purple spots toward base of veins, purple color varies, from absent to very dark, hirsute on adaxial surface and pilose on abaxial surface, sepals of three sizes, dorsal sepal 1.5–2 ×
|
||
<quantity id="4CFB9B3EFFAA927AFE6FFE11FDF93EB7" box="[491,617,412,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="4" pageNumber="185" unit="cm" value="1.3" valueMax="1.5" valueMin="1.1">1.1–1.5 cm</quantity>
|
||
, two lateral sepals 1.3–2 ×
|
||
<quantity id="4CFB9B3EFFAA927AFC15FE11FC6A3EB7" box="[913,1018,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="4" pageNumber="185" unit="cm" value="0.8" valueMax="1.0" valueMin="0.6">0.6–1 cm</quantity>
|
||
, and two ventral sepals 1.3–2 ×
|
||
<quantity id="4CFB9B3EFFAA927AFADDFE11FF3D3EDB" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="4" pageNumber="185" unit="cm" value="0.65" valueMax="0.8" valueMin="0.5">0.5–0.8 cm</quantity>
|
||
. Corolla orange, tube cylindrical, at angle of 45° relative to stem, ventricose ventrally,
|
||
<quantity id="4CFB9B3EFFAA927AFB05FE4DFB7A3EDB" box="[1153,1258,448,473]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="4" pageNumber="185" unit="cm" value="1.8" valueMax="2.0" valueMin="1.6">1.6–2 cm</quantity>
|
||
long, constricted at throat and in middle part, tube
|
||
<quantity id="4CFB9B3EFFAA927AFD86FE69FDE33EFF" box="[514,627,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.0" metricValueMin="4.7" pageId="4" pageNumber="185" unit="mm" value="4.85" valueMax="5.0" valueMin="4.7">4.7–5 mm</quantity>
|
||
wide at throat,
|
||
<quantity id="4CFB9B3EFFAA927AFCA1FE69FC123EFF" box="[805,898,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="4" pageNumber="185" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
|
||
wide at the ventricose portion,
|
||
<quantity id="4CFB9B3EFFAA927AFB63FE69FAFD3EFF" box="[1255,1389,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="4.5" metricValueMin="4.3" pageId="4" pageNumber="185" unit="mm" value="4.4" valueMax="4.5" valueMin="4.3">4.3–4.5 mm</quantity>
|
||
wide at middle part,
|
||
<quantity id="4CFB9B3EFFAA927AFEB2FD85FE033D23" box="[310,403,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="4" pageNumber="185" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
|
||
wide at base, aperture
|
||
<quantity id="4CFB9B3EFFAA927AFD19FD85FC9F3D23" box="[669,783,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.0" metricValueMin="3.8" pageId="4" pageNumber="185" unit="mm" value="3.9" valueMax="4.0" valueMin="3.8">3.8–4 mm</quantity>
|
||
wide, glabrous outside, with lobes villous outside, internal surface with glandular trichomes
|
||
<quantity id="4CFB9B3EFFAA927AFE73FDA1FDF63D47" box="[503,614,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="4" pageNumber="185" unit="mm" value="1.9" valueMax="2.0" valueMin="1.8">1.8–2 mm</quantity>
|
||
long, becoming sparse toward base, limb with five unequal lobes, which are orbicular, erect or spreading, villous on the outer and inner surface, margin slightly ciliate and slightly revolute, three lobes two lateral and one ventral 5.9–6 ×
|
||
<quantity id="4CFB9B3EFFAA927AFDD9FDF9FD5E3D8F" box="[605,718,628,653]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" metricValueMax="8.0" metricValueMin="7.8" pageId="4" pageNumber="185" unit="mm" value="7.9" valueMax="8.0" valueMin="7.8">7.8–8 mm</quantity>
|
||
each, two dorsal lobes 4.9–5 ×
|
||
<quantity id="4CFB9B3EFFAA927AFBB3FDF9FB3C3D8F" box="[1079,1196,628,653]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="6.0" metricValueMin="5.8" pageId="4" pageNumber="185" unit="mm" value="5.9" valueMax="6.0" valueMin="5.8">5.8–6 mm</quantity>
|
||
. Stamens didynamous, staminode one, dorsal, filaments curved to spiral, adnate to corolla base,
|
||
<quantity id="4CFB9B3EFFAA927AFC3DFD15FBBA3DB3" box="[953,1066,664,689]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.0" metricValueMin="3.8" pageId="4" pageNumber="185" unit="mm" value="3.9" valueMax="4.0" valueMin="3.8">3.8–4 mm</quantity>
|
||
long, glabrous, anthers connivent. Disc annular, complete, glabrous, white. Ovary glabrous, pale green, 3.2–3.3 ×
|
||
<quantity id="4CFB9B3EFFAA927AFB85FD31FBE63DD7" box="[1025,1142,700,725]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="4" pageNumber="185" unit="mm" value="2.95" valueMax="3.0" valueMin="2.9">2.9–3 mm</quantity>
|
||
; style
|
||
<quantity id="4CFB9B3EFFAA927AFB3AFD31FA8A3DD7" box="[1214,1306,700,725]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="4" pageNumber="185" unit="mm" value="7.5" valueMax="9.0" valueMin="6.0">6–9 mm</quantity>
|
||
long, pilose, trichomes thin, short and translucent, 1–3-celled; stigma bilobed. Fruit a fleshy berry, white at maturity.
|
||
</paragraph>
|
||
<paragraph id="8BBC36DBFFAA927AFF39FC89FCBC3C67" blockId="4.[136,1453,160,1193]" pageId="4" pageNumber="185">
|
||
<emphasis id="B977EAC9FFAA927AFF39FC89FED13C1F" bold="true" box="[189,321,772,797]" pageId="4" pageNumber="185">Palynology</emphasis>
|
||
—Pollen grains triporate, subspheroidal, (17–)18.3(–20) × (18–)20.9(–21.7) μm,
|
||
<collectionCode id="ED12AE1EFFAA927AFB6DFC89FB683C1F" box="[1257,1272,772,797]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="185" type="Herbarium">P</collectionCode>
|
||
/
|
||
<collectionCode id="ED12AE1EFFAA927AFA84FC89FA813C1F" box="[1280,1297,772,797]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="4" pageNumber="185" type="Herbarium">E</collectionCode>
|
||
ratio approx. 0.87, radially symmetrical, isopolar, circular in polar view. Exine 1.1 μm thick. Pores 1.9 × 2.1 μm, apocolpe 16.8–17 μm long. Exine with perforate ornamentation (
|
||
<figureCitation id="13382A5EFFAA927AFD16FCC1FC8F3C67" box="[658,799,844,869]" captionStart="FIGURE 3" captionStartId="3.[136,229,1996,2017]" captionTargetBox="[240,1348,175,1971]" captionTargetId="figure-14@3.[240,1348,175,1971]" captionTargetPageId="3" captionText="FIGURE 3. Pollen micromorphology. A–D: Besleria physaloides. A. polar view; B. polar view (SEM); C. equatorial view (SEM); D. ornamentation (SEM). E–H: Besleria silverstoneana. E. polar view; F. polar view (SEM); G. equatorial view (SEM); H. ornamentation (SEM)." figureDoi="http://doi.org/10.5281/zenodo.13722086" httpUri="https://zenodo.org/record/13722086/files/figure.png" pageId="4" pageNumber="185">Figs. 3 A–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3196550FFAA927AFF39FCFDFE473B63" pageId="4" pageNumber="185" type="distribution">
|
||
<paragraph id="8BBC36DBFFAA927AFF39FCFDFD5A3CF7" blockId="4.[136,1453,160,1193]" pageId="4" pageNumber="185">
|
||
<emphasis id="B977EAC9FFAA927AFF39FCFDFDF23C8B" bold="true" box="[189,610,880,905]" pageId="4" pageNumber="185">Distribution, habitat and phenology</emphasis>
|
||
:—Known only from the
|
||
<typeStatus id="54B88879FFAA927AFCF1FCFDFC353C8B" box="[885,933,880,905]" pageId="4" pageNumber="185">type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="F314764BFFAA927AFB8EFCFDFBEB3C8B" box="[1034,1147,880,905]" name="Colombia" pageId="4" pageNumber="185">Colombia</collectingCountry>
|
||
,
|
||
<collectingRegion id="49C7F839FFAA927AFB00FCFDFAA13C8B" box="[1156,1329,880,905]" country="Colombia" name="Valle del Cauca" pageId="4" pageNumber="185">Valle del Cauca</collectingRegion>
|
||
, municipio Dagua, corregimiento San Bernardo (
|
||
<figureCitation id="13382A5EFFAA927AFDA9FC19FDE23CAF" box="[557,626,916,941]" captionStart="FIGURE 4" captionStartId="5.[136,229,2056,2077]" captionTargetBox="[191,1408,526,2017]" captionTargetId="figure-163@5.[177,1410,515,2034]" captionTargetPageId="5" captionText="FIGURE 4. Geographical distribution of Besleria physaloides and Besleria silverstoneana in Valle del Cauca, Colombia." figureDoi="http://doi.org/10.5281/zenodo.13722088" httpUri="https://zenodo.org/record/13722088/files/figure.png" pageId="4" pageNumber="185">Fig. 4</figureCitation>
|
||
). Small patches of Andean forest of secondary succession, surrounded by constructions of rural housing. It grows on edge of the forest. Flowering from March to May, fruiting from April to June, according to the sampling done for this study.
|
||
</paragraph>
|
||
<paragraph id="8BBC36DBFFAA927AFF39FB8DFE473B63" blockId="4.[136,1453,160,1193]" pageId="4" pageNumber="185">
|
||
<emphasis id="B977EAC9FFAA927AFF39FB8DFEBA3B1B" bold="true" box="[189,298,1024,1049]" pageId="4" pageNumber="185">
|
||
<typeStatus id="54B88879FFAA927AFF39FB8DFEBA3B1B" box="[189,298,1024,1049]" pageId="4" pageNumber="185" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
:—
|
||
<collectingCountry id="F314764BFFAA927AFECFFB8DFE773B1B" box="[331,487,1024,1049]" name="Colombia" pageId="4" pageNumber="185">COLOMBIA</collectingCountry>
|
||
.
|
||
<collectingRegion id="49C7F839FFAA927AFE75FB8DFD353B1B" box="[497,677,1024,1049]" country="Colombia" name="Valle del Cauca" pageId="4" pageNumber="185">Valle del Cauca</collectingRegion>
|
||
: municipio Dagua, corregimiento San Bernardo, vía a Buenaventura, cordillera occidental, vertiente oriental,
|
||
<geoCoordinate id="EE37501CFFAA927AFDDFFBA9FC903B3F" box="[603,768,1060,1085]" degrees="03" direction="north" minutes="30" orientation="latitude" pageId="4" pageNumber="185" precision="1" seconds="52.0" value="3.5144444">03°30’52.0’’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE37501CFFAA927AFC94FBA9FC283B3F" box="[784,952,1060,1085]" degrees="76" direction="west" minutes="37" orientation="longitude" pageId="4" pageNumber="185" precision="1" seconds="27.8" value="-76.62439">76°37’27.8’’W</geoCoordinate>
|
||
,
|
||
<quantity id="4CFB9B3EFFAA927AFC4CFBA9FBB43B3F" box="[968,1060,1060,1085]" metricMagnitude="3" metricUnit="m" metricValue="1.9919999999999998" pageId="4" pageNumber="185" unit="m" value="1992.0">1992 m</quantity>
|
||
,
|
||
<date id="FFBD101BFFAA927AFBB1FBA9FB4E3B3F" box="[1077,1246,1060,1085]" pageId="4" pageNumber="185" value="2016-03-02">2 March 2016</date>
|
||
,
|
||
<emphasis id="B977EAC9FFAA927AFB69FBA9FA3B3B3F" box="[1261,1451,1060,1085]" italics="true" pageId="4" pageNumber="185">O.L. Cortés 256</emphasis>
|
||
(CUVC 67091! PSO! HUQ!).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3196550FFAA927AFF39FBE1FDEE3BAB" pageId="4" pageNumber="185" type="etymology">
|
||
<paragraph id="8BBC36DBFFAA927AFF39FBE1FDEE3BAB" blockId="4.[136,1453,160,1193]" pageId="4" pageNumber="185">
|
||
<emphasis id="B977EAC9FFAA927AFF39FBE1FEAE3B87" bold="true" box="[189,318,1132,1157]" pageId="4" pageNumber="185">Etymology</emphasis>
|
||
:—The specific epithet refers to the form of the sepals, which are similar to the sepals of the genus
|
||
<taxonomicName id="4C034D58FFAA927AFF0CFB1DFE4F3BAB" authority="Linnaeus (1753: 182)" authorityName="Linnaeus" authorityPageNumber="182" authorityYear="1753" box="[136,479,1168,1193]" class="Magnoliopsida" family="Solanaceae" genus="Physalis" kingdom="Plantae" order="Solanales" pageId="4" pageNumber="185" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B977EAC9FFAA927AFF0CFB1DFF773BAB" box="[136,231,1168,1193]" italics="true" pageId="4" pageNumber="185">Physalis</emphasis>
|
||
Linnaeus (1753: 182)
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C034D58FFAA927AFE6AFB1DFDE33BAB" box="[494,627,1168,1193]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="4" pageNumber="185" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |