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<document id="2C6E5EA22D29630F1485D7D45D554E32" ID-CLB-Dataset="304964" ID-DOI="10.5852/ejt.2024.967.2699" ID-GBIF-Dataset="0647b761-1496-4dcd-abb6-495b67abe85f" ID-ISSN="2118-9773" ID-Zenodo-Dep="13990819" ID-ZooBank="C08B8027-50CC-417E-BCD4-5183B9FF6738" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1729849508920" checkinUser="plazi" docAuthor="Zonstein, Sergei L." docDate="2024" docId="03A8B723551AFF8AFD8DE622FAC4C9B3" docLanguage="en" docName="EJT.2024.967.1-185.pdf" docOrigin="European Journal of Taxonomy 967" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2699/12459" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola Zonstein 1987" docType="key" docVersion="8" lastPageNumber="49" masterDocId="FF91CF5B552AFFBAFFEFE72AFFB2CE5B" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia" masterLastPageNumber="185" masterPageNumber="1" pageNumber="49" updateTime="1730721133903" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4C6D1E557B1961A18279469B7C9BF3AA">A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia</mods:title>
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<mods:namePart id="47745F2E544789DE18349B9618AD3761">Zonstein, Sergei L.</mods:namePart>
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<mods:affiliation id="AB3586AE577D6115650DF0426333C86A">Steinhardt Museum of Natural History, Tel-Aviv University, 69978 Tel-Aviv, Israel</mods:affiliation>
<mods:nameIdentifier id="D496E0D86C1FD2B1367860C99C7F8DBC" type="email">znn@tauex.tau.ac.il</mods:nameIdentifier>
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<mods:title id="90C60988193A069B429284966EB905E1">European Journal of Taxonomy</mods:title>
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<mods:date id="EBDA060590166275DD13526C25010B71">2024</mods:date>
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<subSubSection id="C31B55BE551AFF8AFD8DE622FC63CF78" box="[610,977,264,291]" pageId="48" pageNumber="49" type="nomenclature">
<paragraph id="8BBE0635551AFF8AFD8DE622FC63CF78" blockId="48.[610,977,264,291]" box="[610,977,264,291]" pageId="48" pageNumber="49">
<heading id="D0F6B159551AFF8AFD8DE622FC63CF78" box="[610,977,264,291]" centered="true" fontSize="11" level="2" pageId="48" pageNumber="49" reason="2">
<emphasis id="B975DA27551AFF8AFD8DE622FC98CF78" bold="true" box="[610,810,264,291]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="4C017DB6551AFF8AFD8DE622FD6ECF78" ID-CoL="77BJ" authority="Zonstein, 1987" authorityName="Zonstein" authorityYear="1987" box="[610,732,264,291]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">Raveniola</taxonomicName>
diluta
</emphasis>
species group
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE551AFF8AFF52E665FF48CCCA" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="8BBE0635551AFF8AFF52E665FE7ECF31" blockId="48.[189,460,335,362]" box="[189,460,335,362]" pageId="48" pageNumber="49">
<heading id="D0F6B159551AFF8AFF52E665FE7ECF31" bold="true" box="[189,460,335,362]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B975DA27551AFF8AFF52E665FE7ECF31" bold="true" box="[189,460,335,362]" pageId="48" pageNumber="49">Diagnostic characters</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE0635551AFF8AFF52E6A9FF48CCCA" blockId="48.[189,1398,387,657]" pageId="48" pageNumber="49">
Maxillae with a few cuspules confined to probasal maxillary edge (
<figureCitation id="133A1AB0551AFF8AFC55E6A9FBEFCFC6" box="[954,1117,387,413]" captionStart="Figs 211–219" captionStartId="130.[189,241,1847,1873]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-85@130.[189,1398,265,1816]" captionTargetPageId="130" captionText="Figs 211–219. Raveniola spp., ♂♂, holotypes (except 215, 217), sternum, labium and maxillae, ventral views. 211. R. insolita sp. nov. (ZISP). 212. R. ornata sp. nov. (SMNH). 213. R. ornatula sp. nov. (SMNH). 214. R. pamira sp. nov. (SMNH). 215. R. pamira, paratype (SMNH). 216. R. diluta sp. nov. (SMNH). 217. R. fedotovi (Charitonov, 1946), ♂ from Hojakurgan (SMNH). 218. R. pallens sp. nov. (SMNH). 219. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990880" httpUri="https://zenodo.org/record/13990880/files/figure.png" pageId="48" pageNumber="49">Figs 216–219</figureCitation>
,
<figureCitation id="133A1AB0551AFF8AFB88E6A9FB60CFC6" box="[1127,1234,387,413]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="48" pageNumber="49">243–246</figureCitation>
). PMS absent; apical segment of PLS triangular or shortly digitiform (
<figureCitation id="133A1AB0551AFF8AFCA6E686FC43CF9C" box="[841,1009,428,455]" captionStart-0="Figs 584–592" captionStart-1="Figs 593–601" captionStartId-0="163.[189,241,1821,1847]" captionStartId-1="164.[189,241,1759,1785]" captionTargetBox-0="[189,1398,265,1789]" captionTargetBox-1="[189,1399,265,1718]" captionTargetId-0="figure-92@163.[189,1398,265,1789]" captionTargetId-1="figure-74@164.[189,1399,265,1718]" captionTargetPageId-1="164" captionText-0="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." captionText-1="Figs 593–601. Raveniola spp., ♂♂, holotypes (593, 595, 597–598, 601) and ♀♀ paratypes (594, 596, 599–600). Spinnerets, lateral (593,596–597), ventral (595,598–599, 601) and posterior (594, 600) views. 593–594. R. pallens sp. nov. (SMNH). 595–596. R. zyuzini sp. nov. (SMNH). 597–600. R. ferghanensis (Zonstein, 1984) (ZISP). 601. R. kirgizica sp. nov. (SMNH). Strictly reduced PMS indicated by white arrows. Scale bars: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14018408" figureDoi-1="http://doi.org/10.5281/zenodo.14018411" httpUri-0="https://zenodo.org/record/14018408/files/figure.png" httpUri-1="https://zenodo.org/record/14018411/files/figure.png" pageId="48" pageNumber="49">Figs 589–596</figureCitation>
). Males: tibiae and metatarsi I–II without modified hairs (
<figureCitation id="133A1AB0551AFF8AFE38E6FEFDCECFB4" box="[471,636,468,495]" captionStart-0="Figs 265–273" captionStart-1="Figs 274–281" captionStartId-0="136.[189,241,1765,1791]" captionStartId-1="137.[189,241,1743,1769]" captionTargetBox-0="[189,1398,265,1731]" captionTargetBox-1="[189,1398,265,1701]" captionTargetId-0="figure-88@136.[189,1398,265,1731]" captionTargetId-1="figure-78@137.[189,1398,265,1703]" captionTargetPageId-0="136" captionTargetPageId-1="137" captionText-0="Figs 265–273. Raveniola spp., ♂♂, holotypes (except 272), tibia and metatarsus I, retrolateral views. 265. R. dolosa sp. nov. (SMNH). 266. R. hirta sp. nov. (SMNH). 267. R. ignobilis sp. nov. (SMNH). 268. R. insolita sp. nov. (ZISP). 269. R. ornata sp. nov. (SMNH). 270. R. ornatula sp. nov. (SMNH). 271. R. pamira sp. nov. (SMNH). 272. R. pamira, paratype (SMNH). 273. R. diluta sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." captionText-1="Figs 274–281. Raveniola spp., ♂♂, holotypes (except 275, 279), tibia and metatarsus I, retrolateral views. 274. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 275. R. fedotovi, ♂ from Hojakurgan (SMNH). 276. R. pallens sp. nov. (SMNH). 277. R. zyuzini sp. nov. (SMNH). 278. R. ferghanensis (Zonstein, 1984) (ZISP). 279. R. ferghanensis, paratype (SMNH). 280. R. kirgizica sp. nov. (SMNH). 281. R. kopetdaghensis (Fet, 1984) (ZISP). Scale bars: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990895" figureDoi-1="http://doi.org/10.5281/zenodo.13990897" httpUri-0="https://zenodo.org/record/13990895/files/figure.png" httpUri-1="https://zenodo.org/record/13990897/files/figure.png" pageId="48" pageNumber="49">Figs 273–277</figureCitation>
,
<figureCitation id="133A1AB0551AFF8AFD67E6FFFD0ACFB4" box="[648,696,469,495]" captionStart="Figs 290–309" captionStartId="139.[189,241,1678,1704]" captionTargetBox="[189,1398,265,1651]" captionTargetId="figure-180@139.[189,1398,265,1651]" captionTargetPageId="139" captionText="Figs 290–309. Raveniola spp., ♂♂, holotypes (except 290, 294, 300, 302, 308), metatarsus I, ventral views. 290. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 291. R. alajensis sp. nov. (SMNH). 292. R. concolor Zonstein, 2000 (MCSN AR 18). 293. R. cucullata sp. nov. (SMNH). 294. R. cucullata, paratype from Shahob (SMNH).295. R. dolosa sp. nov.(SMNH). 296. R. hirtasp. nov. (SMNH). 297. R. ignobilis sp. nov. (SMNH). 298. R. ornata sp. nov. (SMNH). 299. R. ornatula sp. nov. (SMNH). 300. R. ornatula, paratype (SMNH).301. R. pamirasp. nov. (SMNH). 302. R. pamira, paratype (SMNH). 303. R. diluta sp. nov. (SMNH). 304. R. kirgizica sp. nov. (SMNH). 305. R. kopetdaghensis (Fet, 1984) (ZISP). 306. R. mikhailovi Zonstein, 2021 (SMNH). 307. R. nenilini sp. nov. (SMNH). 308. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 309. R. vulpina sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990903" httpUri="https://zenodo.org/record/13990903/files/figure.png" pageId="48" pageNumber="49">303</figureCitation>
); cymbium moderately short (as in
<figureCitation id="133A1AB0551AFF8AFBB6E6FEFB4DCFB4" box="[1113,1279,468,495]" captionStart="Figs 364–378" captionStartId="144.[189,241,1714,1740]" captionTargetBox="[189,1398,271,1690]" captionTargetId="figure-142@144.[189,1398,265,1690]" captionTargetPageId="144" captionText="Figs 364–378. Raveniola spp., ♂♂, holotypes (except 366, 369, 376, 378), palpal tibia, cymbium and copulatory bulb, retrolateral (364–377) and retroventral (378) views. 364. R. pamira sp. nov. (SMNH). 365. R. diluta sp. nov. (SMNH). 366. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 367. R. pallens sp. nov. (SMNH). 368. R. zyuzini sp. nov. (SMNH). 369. R. zyuzini, paratype (SMNH). 370. R. ferghanensis (Zonstein, 1984) (ZISP). 371. R. kirgizica sp. nov. (SMNH).372. R. kopetdaghensis (Fet, 1984) (ZISP). 373. R. mikhailovi Zonstein, 2021 (SMNH). 374. R. nenilini sp. nov. (SMNH). 375. R. ovchinnikovi sp. nov. (SMNH). 376. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 377. R. vulpina sp. nov. (SMNH). 378. R. vulpina, paratype from Tereksai (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990915" httpUri="https://zenodo.org/record/13990915/files/figure.png" pageId="48" pageNumber="49">Figs 365–369</figureCitation>
); embolus bent and screwed subapically, with or without subapical keel (
<figureCitation id="133A1AB0551AFF8AFC59E6D7FBD3CC4C" box="[950,1121,509,535]" captionStart="Figs 429–438" captionStartId="150.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1818]" captionTargetId="figure-75@150.[189,1398,265,1818]" captionTargetPageId="150" captionText="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990927" httpUri="https://zenodo.org/record/13990927/files/figure.png" pageId="48" pageNumber="49">Figs 429–438</figureCitation>
). Females: each paired spermatheca Ⱶ-shaped, mostly low and moderately wide, with wide base and inner branch incorporated into the entire cone-shaped trunk, and with short and club-like or fusiform lateral diverticulum (
<figureCitation id="133A1AB0551AFF8AFAEDE564FF5FCCCA" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="48" pageNumber="49">Figs 526– 533</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE551AFF8AFF52E597FAFACD50" pageId="48" pageNumber="49" type="taxon_list">
<paragraph id="8BBE0635551AFF8AFF52E597FE3ECC8C" blockId="48.[189,396,701,727]" box="[189,396,701,727]" pageId="48" pageNumber="49">
<heading id="D0F6B159551AFF8AFF52E597FE3ECC8C" bold="true" box="[189,396,701,727]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B975DA27551AFF8AFF52E597FE3ECC8C" bold="true" box="[189,396,701,727]" pageId="48" pageNumber="49">Species included</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE0635551AFF8AFF52E5DBFAFACD50" blockId="48.[189,1352,753,779]" box="[189,1352,753,779]" pageId="48" pageNumber="49">
<taxonomicName id="4C017DB6551AFF8AFF52E5DBFECDCD50" authorityName="Zonstein" authorityYear="2024" box="[189,383,753,779]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="diluta" status="sp. nov.">
<emphasis id="B975DA27551AFF8AFF52E5DBFECDCD50" box="[189,383,753,779]" italics="true" pageId="48" pageNumber="49">Raveniola diluta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C551AFF8AFE68E5DBFE56CD50" box="[391,484,753,779]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C017DB6551AFF8AFE1FE5D8FCEACD50" ID-CoL="4RMTR" authority="(Charitonov, 1946)" baseAuthorityName="Charitonov" baseAuthorityYear="1946" box="[496,856,753,779]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="fedotovi">
<emphasis id="B975DA27551AFF8AFE1FE5D8FDDDCD50" box="[496,623,753,779]" italics="true" pageId="48" pageNumber="49">R. fedotovi</emphasis>
(
<bibRefCitation id="EF907BC4551AFF8AFD91E5DBFCFDCD50" author="Charitonov" box="[638,847,753,779]" firstAuthor="Charitonov" pageId="48" pageNumber="49" pagination="19 - 32" refId="ref77775" refString="Charitonov D. E. 1946. New forms of spiders of the USSR. Izvestija Estedvenno-Nauchnogo Instituta pri Molotovskom Gosudarstvennom Universitete imeni M. Gor'kogo 12: 19 - 32. [In Russian and Latin.]" type="journal article" year="1946">Charitonov, 1946</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C017DB6551AFF8AFC8BE5D8FC6BCD50" authorityName="Zonstein" authorityYear="2024" box="[868,985,753,779]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="pallens" status="sp. nov.">
<emphasis id="B975DA27551AFF8AFC8BE5D8FC6BCD50" box="[868,985,753,779]" italics="true" pageId="48" pageNumber="49">R. pallens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C551AFF8AFC0EE5DBFB89CD50" box="[993,1083,753,779]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C017DB6551AFF8AFB9BE5D8FB57CD50" authorityName="Zonstein" authorityYear="2024" box="[1140,1253,754,779]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="zyuzini" status="sp. nov.">
<emphasis id="B975DA27551AFF8AFB9BE5D8FB57CD50" box="[1140,1253,754,779]" italics="true" pageId="48" pageNumber="49">R. zyuzini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C551AFF8AFB02E5DBFAFACD50" box="[1261,1352,753,779]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE551AFF8AFF52E412FAC4C9B3" pageId="48" pageNumber="49" type="key">
<paragraph id="8BBE0635551AFF8AFF52E412FD3ACD09" blockId="48.[189,648,823,850]" box="[189,648,823,850]" pageId="48" pageNumber="49">
<heading id="D0F6B159551AFF8AFF52E412FD3ACD09" bold="true" box="[189,648,823,850]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B975DA27551AFF8AFF52E412FD3ACD09" bold="true" box="[189,648,823,850]" pageId="48" pageNumber="49">
Key to the species of the
<taxonomicName id="4C017DB6551AFF8AFE1EE41DFD85CD09" authorityName="Zonstein" authorityYear="2024" box="[497,567,823,850]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="diluta">
<emphasis id="B975DA27551AFF8AFE1EE41DFD85CD09" bold="true" box="[497,567,823,850]" italics="true" pageId="48" pageNumber="49">diluta</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE0635551AFF8AFF05E446FE87CDDD" blockId="48.[189,1399,876,983]" box="[234,309,876,902]" pageId="48" pageNumber="49">
<emphasis id="B975DA27551AFF8AFF05E446FE87CDDD" bold="true" box="[234,309,876,902]" pageId="48" pageNumber="49">Males</emphasis>
</paragraph>
<paragraph id="8BBE0635551AFF8AFF52E4BEFAC5CDF5" blockId="48.[189,1399,876,983]" box="[189,1399,916,943]" pageId="48" pageNumber="49">
1. Embolus with well-defined and raised triangular keel (
<figureCitation id="133A1AB0551AFF8AFC87E4BEFBBECDF5" box="[872,1036,916,942]" captionStart="Figs 429–438" captionStartId="150.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1818]" captionTargetId="figure-75@150.[189,1398,265,1818]" captionTargetPageId="150" captionText="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990927" httpUri="https://zenodo.org/record/13990927/files/figure.png" pageId="48" pageNumber="49">Figs 429–433</figureCitation>
)............................................. 2
</paragraph>
<paragraph id="8BBE0635551AFF8AFF2EE497FAC5CD8C" blockId="48.[189,1399,876,983]" box="[193,1399,956,983]" pageId="48" pageNumber="49">
– Embolic keel rudimentary or absent (
<figureCitation id="133A1AB0551AFF8AFD71E496FCF0CD8C" box="[670,834,956,983]" captionStart="Figs 429–438" captionStartId="150.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1818]" captionTargetId="figure-75@150.[189,1398,265,1818]" captionTargetPageId="150" captionText="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990927" httpUri="https://zenodo.org/record/13990927/files/figure.png" pageId="48" pageNumber="49">Figs 434–438</figureCitation>
) ........................................................................ 3
</paragraph>
<paragraph id="8BBE0635551AFF8AFF52E329FAC5CA1D" blockId="48.[189,1399,1027,1175]" pageId="48" pageNumber="49">
2. Copulatory bulb elongate; proximal part of embolus much longer than its apical part; pointed triangular keel less acute and forms an angle&gt; 60° (
<figureCitation id="133A1AB0551AFF8AFCA3E306FC42CA1D" box="[844,1008,1068,1094]" captionStart="Figs 429–438" captionStartId="150.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1818]" captionTargetId="figure-75@150.[189,1398,265,1818]" captionTargetPageId="150" captionText="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990927" httpUri="https://zenodo.org/record/13990927/files/figure.png" pageId="48" pageNumber="49">Figs 429–431</figureCitation>
)........................
<taxonomicName id="4C017DB6551AFF8AFB42E306FAA7CA1D" authorityName="Zonstein" authorityYear="2024" box="[1197,1301,1067,1094]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="diluta" status="sp. nov.">
<emphasis id="B975DA27551AFF8AFB42E306FAA7CA1D" bold="true" box="[1197,1301,1067,1094]" italics="true" pageId="48" pageNumber="49">R. diluta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C551AFF8AFAF3E306FAC5CA1D" box="[1308,1399,1068,1094]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BBE0635551AFF8AFF2EE37FFAC4CACC" blockId="48.[189,1399,1027,1175]" pageId="48" pageNumber="49">
– Copulatory bulb short; proximal and apical parts of embolus subequal in length; short and acute triangular keel forms an angle &lt;45° (
<figureCitation id="133A1AB0551AFF8AFD4DE357FCF4CACC" box="[674,838,1149,1175]" captionStart="Figs 429–438" captionStartId="150.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1818]" captionTargetId="figure-75@150.[189,1398,265,1818]" captionTargetPageId="150" captionText="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990927" httpUri="https://zenodo.org/record/13990927/files/figure.png" pageId="48" pageNumber="49">Figs 432–433</figureCitation>
).........................
<taxonomicName id="4C017DB6551AFF8AFBE1E357FAC4CACC" ID-CoL="4RMTR" authority="(Charitonov, 1946)" baseAuthorityName="Charitonov" baseAuthorityYear="1946" box="[1038,1398,1148,1175]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="fedotovi">
<emphasis id="B975DA27551AFF8AFBE1E357FB22CACC" bold="true" box="[1038,1168,1148,1175]" italics="true" pageId="48" pageNumber="49">R. fedotovi</emphasis>
(
<bibRefCitation id="EF907BC4551AFF8AFB70E357FADCCACC" author="Charitonov" box="[1183,1390,1149,1175]" firstAuthor="Charitonov" pageId="48" pageNumber="49" pagination="19 - 32" refId="ref77775" refString="Charitonov D. E. 1946. New forms of spiders of the USSR. Izvestija Estedvenno-Nauchnogo Instituta pri Molotovskom Gosudarstvennom Universitete imeni M. Gor'kogo 12: 19 - 32. [In Russian and Latin.]" type="journal article" year="1946">Charitonov, 1946</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BBE0635551AFF8AFF52E3E9FAC5CB5D" blockId="48.[189,1399,1219,1367]" pageId="48" pageNumber="49">
3. Metatarsus I weakly curved (
<figureCitation id="133A1AB0551AFF8AFDAEE3E9FD1BCA86" box="[577,681,1219,1245]" captionStart="Figs 274–281" captionStartId="137.[189,241,1743,1769]" captionTargetBox="[189,1398,265,1701]" captionTargetId="figure-78@137.[189,1398,265,1703]" captionTargetPageId="137" captionText="Figs 274–281. Raveniola spp., ♂♂, holotypes (except 275, 279), tibia and metatarsus I, retrolateral views. 274. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 275. R. fedotovi, ♂ from Hojakurgan (SMNH). 276. R. pallens sp. nov. (SMNH). 277. R. zyuzini sp. nov. (SMNH). 278. R. ferghanensis (Zonstein, 1984) (ZISP). 279. R. ferghanensis, paratype (SMNH). 280. R. kirgizica sp. nov. (SMNH). 281. R. kopetdaghensis (Fet, 1984) (ZISP). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990897" httpUri="https://zenodo.org/record/13990897/files/figure.png" pageId="48" pageNumber="49">Fig. 276</figureCitation>
); scopula on tarsi and metatarsi I–II sparser. Copulatory bulb shorter; proximal part of embolus more tapering (
<figureCitation id="133A1AB0551AFF8AFCDFE3C6FC66CB5C" box="[816,980,1260,1287]" captionStart="Figs 429–438" captionStartId="150.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1818]" captionTargetId="figure-75@150.[189,1398,265,1818]" captionTargetPageId="150" captionText="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990927" httpUri="https://zenodo.org/record/13990927/files/figure.png" pageId="48" pageNumber="49">Figs 434–435</figureCitation>
)..........................
<taxonomicName id="4C017DB6551AFF8AFB73E3C6FAA7CB5D" authorityName="Zonstein" authorityYear="2024" box="[1180,1301,1259,1286]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="pallens" status="sp. nov.">
<emphasis id="B975DA27551AFF8AFB73E3C6FAA7CB5D" bold="true" box="[1180,1301,1259,1286]" italics="true" pageId="48" pageNumber="49">R. pallens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C551AFF8AFAF3E3C6FAC5CB5D" box="[1308,1399,1260,1286]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BBE0635551AFF8AFF2EE23FFAC5CB0C" blockId="48.[189,1399,1219,1367]" pageId="48" pageNumber="49">
– Metatarsus I strongly curved (
<figureCitation id="133A1AB0551AFF8AFDA4E23EFD03CB75" box="[587,689,1300,1326]" captionStart="Figs 274–281" captionStartId="137.[189,241,1743,1769]" captionTargetBox="[189,1398,265,1701]" captionTargetId="figure-78@137.[189,1398,265,1703]" captionTargetPageId="137" captionText="Figs 274–281. Raveniola spp., ♂♂, holotypes (except 275, 279), tibia and metatarsus I, retrolateral views. 274. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 275. R. fedotovi, ♂ from Hojakurgan (SMNH). 276. R. pallens sp. nov. (SMNH). 277. R. zyuzini sp. nov. (SMNH). 278. R. ferghanensis (Zonstein, 1984) (ZISP). 279. R. ferghanensis, paratype (SMNH). 280. R. kirgizica sp. nov. (SMNH). 281. R. kopetdaghensis (Fet, 1984) (ZISP). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990897" httpUri="https://zenodo.org/record/13990897/files/figure.png" pageId="48" pageNumber="49">Fig. 277</figureCitation>
); scopula on tarsi and metatarsi I–II denser. Copulatory bulb longer; proximal part of embolus less tapering (
<figureCitation id="133A1AB0551AFF8AFCF7E217FC0ECB0C" box="[792,956,1341,1367]" captionStart="Figs 429–438" captionStartId="150.[189,241,1855,1881]" captionTargetBox="[189,1398,265,1818]" captionTargetId="figure-75@150.[189,1398,265,1818]" captionTargetPageId="150" captionText="Figs 429–438. Raveniola spp., ♂♂, holotypes (except 432–433, 437–438), details of copulatoty bulb, ventral (429, 435, 438), retroventral (430, 434), retrolateral (431, 436–437), proventral (432) and prolateral (433) views. 429–431. R. diluta sp. nov. (SMNH). 432–433. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 434–435. R. pallens sp. nov. (SMNH). 436. R. zyuzini sp. nov. (SMNH). 437– 438. R. zyuzini, paratype (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990927" httpUri="https://zenodo.org/record/13990927/files/figure.png" pageId="48" pageNumber="49">Figs 436–438</figureCitation>
).............................
<taxonomicName id="4C017DB6551AFF8AFB71E217FAA7CB0C" authorityName="Zonstein" authorityYear="2024" box="[1182,1301,1341,1367]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="zyuzini" status="sp. nov.">
<emphasis id="B975DA27551AFF8AFB71E217FAA7CB0C" bold="true" box="[1182,1301,1341,1367]" italics="true" pageId="48" pageNumber="49">R. zyuzini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C551AFF8AFAF3E217FAC5CB0C" box="[1308,1399,1341,1367]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BBE0635551AFF8AFF05E2AEFEE3CBC5" blockId="48.[189,1399,1412,1600]" box="[234,337,1412,1438]" pageId="48" pageNumber="49">
<emphasis id="B975DA27551AFF8AFF05E2AEFEE3CBC5" bold="true" box="[234,337,1412,1438]" pageId="48" pageNumber="49">Females</emphasis>
</paragraph>
<key id="FF905104551AFF8AFF52E286FAC4C9B3" pageId="48" pageNumber="49">
<keyStep id="30F51E90551AFF8AFF52E286FAC5C81B" pageId="48" pageNumber="49">
<paragraph id="8BBE0635551AFF8AFF52E286FAC5CBB4" blockId="48.[189,1399,1412,1600]" pageId="48" pageNumber="49">
<keyLead id="30F0A500551AFF8AFF52E286FAC5CBB4" pageId="48" pageNumber="49">
1. Carapace, palps and legs intensely brownish orange (
<figureCitation id="133A1AB0551AFF8AFCB3E286FC05CB9D" box="[860,951,1452,1478]" captionStart="Figs 10–18" captionStartId="108.[189,241,1826,1852]" captionTargetBox="[189,1398,265,1785]" captionTargetId="figure-83@108.[189,1398,265,1785]" captionTargetPageId="108" captionText="Figs 10–18. Raveniola spp., ♂♂, holotypes (except 14, 16), habitus, dorsal views. 10. R. insolita sp. nov. (ZISP). 11. R. ornata sp. nov. (SMNH). 12. R. ornatula sp. nov. (SMNH). 13. R. pamira sp. nov. (SMNH). 14. R. pamira sp. nov., paratype (SMNH). 15. R. diluta sp. nov. (SMNH). 16. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 17. R. pallens sp. nov. (SMNH). 18. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990826" httpUri="https://zenodo.org/record/13990826/files/figure.png" pageId="48" pageNumber="49">Figs 18</figureCitation>
,
<figureCitation id="133A1AB0551AFF8AFC2AE286FC57CB9C" box="[965,997,1452,1479]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="48" pageNumber="49">53</figureCitation>
,
<figureCitation id="133A1AB0551AFF8AFC1EE286FBF2CB9D" box="[1009,1088,1452,1478]" captionStart="Figs 69–75" captionStartId="115.[189,241,1831,1857]" captionTargetBox="[191,1400,265,1794]" captionTargetId="figure-66@115.[191,1400,265,1794]" captionTargetPageId="115" captionText="Figs 69–75. Raveniola spp., ♀♀ (69, 71–72, 74) and ♂♂ (70, 73, 75), live spiders; predominantly dorsal (69, 71–72, 74–75) and dorsolateral (70, 73) views. 69–70. R. cucullata sp. nov. (SMNH). 71. R. dolosa sp. nov. (SMNH).72. R. ornatasp. nov. (SMNH). 73–74. R. zyuzinisp. nov. (SMNH). 75. R. ferghanensis (Zonstein, 1984) (SMNH)." figureDoi="http://doi.org/10.5281/zenodo.13990842" httpUri="https://zenodo.org/record/13990842/files/figure.png" pageId="48" pageNumber="49">73–74</figureCitation>
). Spermathecal trunks and fundus of lateral diverticulum weakly sclerotized (
<figureCitation id="133A1AB0551AFF8AFCD5E2FEFC6CCBB4" box="[826,990,1492,1519]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="48" pageNumber="49">Figs 532–533</figureCitation>
) ........................
<taxonomicName id="4C017DB6551AFF8AFB71E2FFFAA7CBB5" authorityName="Zonstein" authorityYear="2024" box="[1182,1301,1492,1519]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="zyuzini" status="sp. nov.">
<emphasis id="B975DA27551AFF8AFB71E2FFFAA7CBB5" bold="true" box="[1182,1301,1492,1519]" italics="true" pageId="48" pageNumber="49">R. zyuzini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C551AFF8AFAF3E2FFFAC5CBB4" box="[1308,1399,1493,1519]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBE0635551AFF8AFF2EE2D4FAC5C81B" blockId="48.[189,1399,1412,1600]" pageId="48" pageNumber="49">
<keyLead id="30F0A500551AFF8AFF2EE2D4FAC5C81B" pageId="48" pageNumber="49">
– Carapace, palps and legs pale brownish yellow or pale brownish orange (
<figureCitation id="133A1AB0551AFF8AFBD8E2D7FB72C843" box="[1079,1216,1533,1560]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="48" pageNumber="49">Figs 50–52</figureCitation>
). Spermathecae differ in their structure (
<figureCitation id="133A1AB0551AFF8AFDECE10FFD14C81B" box="[515,678,1573,1600]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="48" pageNumber="49">Figs 526–531</figureCitation>
).............................................................................................. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E90551AFF8AFF52E146FAC5C972" pageId="48" pageNumber="49">
<paragraph id="8BBE0635551AFF8AFF52E146FAC5C8F4" blockId="48.[189,1399,1644,1833]" pageId="48" pageNumber="49">
<keyLead id="30F0A500551AFF8AFF52E146FAC5C8F4" pageId="48" pageNumber="49">
2. Interdistance AME–AME slightly wider than diameter of AME (
<figureCitation id="133A1AB0551AFF8AFC07E146FB22C8DD" box="[1000,1168,1644,1670]" captionStart="Figs 172–183" captionStartId="126.[189,241,1749,1775]" captionTargetBox="[189,1399,265,1701]" captionTargetId="figure-107@126.[189,1399,265,1701]" captionTargetPageId="126" captionText="Figs 172–183. Raveniola spp., ♀♀, paratypes (except 176, 181, 183), clypeus and eye tubercle, dorsal views. 172. R. dolosa sp. nov. (SMNH). 173–174. R. hirta sp. nov. (SMNH). 175. R. ignobilis sp. nov. (SMNH). 176. R. karategensis sp. nov., holotype (SMNH). 177. R. karategensis, paratype (SMNH). 178. R. ornata sp. nov., from Sanglok (SMNH). 179. R. ornatula sp. nov. (SMNH). 180. R. pamira sp. nov. (SMNH). 181. R. sororcula sp. nov., holotype (SMNH). 182. R. diluta sp. nov. (SMNH). 183. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990872" httpUri="https://zenodo.org/record/13990872/files/figure.png" pageId="48" pageNumber="49">Figs 182–183</figureCitation>
). Palpal tarsus with entire scopula. Cone-shaped spermathecal trunk clearly longer than wide (
<figureCitation id="133A1AB0551AFF8AFBBEE1BFFB47C8F4" box="[1105,1269,1684,1711]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="48" pageNumber="49">Figs 526–529</figureCitation>
) ............. 3
</keyLead>
</paragraph>
<paragraph id="8BBE0635551AFF8AFF2EE194FAC5C972" blockId="48.[189,1399,1644,1833]" pageId="48" pageNumber="49">
<keyLead id="30F0A500551AFF8AFF2EE194FAC5C972" pageId="48" pageNumber="49">
– Interdistance AME–AME considerably wider than diameter of AME (
<figureCitation id="133A1AB0551AFF8AFBBDE197FB72C88C" box="[1106,1216,1725,1751]" captionStart="Figs 184–195" captionStartId="127.[189,241,1720,1746]" captionTargetBox="[189,1399,266,1676]" captionTargetId="figure-110@127.[189,1399,265,1676]" captionTargetPageId="127" captionText="Figs 184–195. Raveniola spp., ♀♀, paratypes (except 191–193), clypeus and eye tubercle, dorsal views. 184. R. pallens sp. nov. (SMNH). 185. R. zyuzini sp. nov. (SMNH). 186. R. ferghanensis (Zonstein, 1984) (SMNH). 187. R. kirgizica sp. nov. (SMNH). 188–189. R. mikhailovi Zonstein, 2021, from Aflatun (ZMMU) and Sary-Chelek Reserve (SMNH), respectively. 190. R. nenilini sp. nov. (SMNH). 191. R. tarabaevi sp. nov., holotype (SMNH). 192–193. R. virgata (Simon, 1891), ♀♀ from Arslanbob (SMNH) and Gava (SMF), respectively. 194–195. R. vulpina sp. nov., from Sary-Chelek Reserve and Tereksai, respectively (both SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990874" httpUri="https://zenodo.org/record/13990874/files/figure.png" pageId="48" pageNumber="49">Fig. 184</figureCitation>
). Palpal tarsus with widely divided scopula. Dome-shaped spermathecal trunk about as wide as long (
<figureCitation id="133A1AB0551AFF8AFB11E1CCFEA5C972" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="48" pageNumber="49">Figs 530– 531</figureCitation>
).........................................................................................................................
<taxonomicName id="4C017DB6551AFF8AFB73E024FAA7C972" authorityName="Zonstein" authorityYear="2024" box="[1180,1301,1806,1833]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="pallens" status="sp. nov.">
<emphasis id="B975DA27551AFF8AFB73E024FAA7C972" bold="true" box="[1180,1301,1806,1833]" italics="true" pageId="48" pageNumber="49">R. pallens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C551AFF8AFAF3E025FAC5C972" box="[1308,1399,1807,1833]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E90551AFF8AFF52E07FFAC4C9B3" pageId="48" pageNumber="49">
<paragraph id="8BBE0635551AFF8AFF52E07FFAC5C9C3" blockId="48.[189,1399,1877,2025]" pageId="48" pageNumber="49">
<keyLead id="30F0A500551AFF8AFF52E07FFAC5C9C3" pageId="48" pageNumber="49">
3. Patellae I–II with spines. Scopula on tarsi II and III narrowly and widely divided, respectively. Spermathecal trunk shorter and more tapering toward apex (
<figureCitation id="133A1AB0551AFF8AFC43E057FBE2C9C3" box="[940,1104,1917,1944]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="48" pageNumber="49">Figs 526–527</figureCitation>
)...........
<taxonomicName id="4C017DB6551AFF8AFB42E054FAA7C9C3" authorityName="Zonstein" authorityYear="2024" box="[1197,1301,1917,1944]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="diluta" status="sp. nov.">
<emphasis id="B975DA27551AFF8AFB42E054FAA7C9C3" bold="true" box="[1197,1301,1917,1944]" italics="true" pageId="48" pageNumber="49">R. diluta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C551AFF8AFAF3E054FAC5C9C3" box="[1308,1399,1918,1944]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBE0635551AFF8AFF2EE08CFAC4C9B3" blockId="48.[189,1399,1877,2025]" pageId="48" pageNumber="49">
<keyLead id="30F0A500551AFF8AFF2EE08CFAC4C9B3" pageId="48" pageNumber="49">
– Patellae I–II aspinose.Scopula on tarsi II and III widely divided and absent, respectively.Spermathecal trunk longer and less tapering toward apex (
<figureCitation id="133A1AB0551AFF8AFD04E0E4FC3DC9B2" box="[747,911,1998,2025]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="48" pageNumber="49">Figs 528–529</figureCitation>
) ...............
<taxonomicName id="4C017DB6551AFF8AFBE1E0E5FAC4C9B3" ID-CoL="4RMTR" authority="(Charitonov, 1946)" baseAuthorityName="Charitonov" baseAuthorityYear="1946" box="[1038,1398,1998,2025]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="fedotovi">
<emphasis id="B975DA27551AFF8AFBE1E0E5FB22C9B2" bold="true" box="[1038,1168,1998,2025]" italics="true" pageId="48" pageNumber="49">R. fedotovi</emphasis>
(
<bibRefCitation id="EF907BC4551AFF8AFB70E0E4FADCC9B3" author="Charitonov" box="[1183,1390,1998,2024]" firstAuthor="Charitonov" pageId="48" pageNumber="49" pagination="19 - 32" refId="ref77775" refString="Charitonov D. E. 1946. New forms of spiders of the USSR. Izvestija Estedvenno-Nauchnogo Instituta pri Molotovskom Gosudarstvennom Universitete imeni M. Gor'kogo 12: 19 - 32. [In Russian and Latin.]" type="journal article" year="1946">Charitonov, 1946</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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