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<document id="0AAC29260970B91AE1E19A03E55B3658" ID-DOI="10.5281/zenodo.179977" ID-GBIF-Dataset="8a222f0b-c42b-456b-ba16-ff056e4ee6bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="179977" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460024914887" checkinUser="plazi" docAuthor="Mueses-Cisneros, Jonh Jairo" docDate="2007" docId="0386BD2A4466B51E7DF6662663B6FE42" docLanguage="en" docName="zt01662p059.pdf" docOrigin="Zootaxa 1662" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Rhaebo lynchi Mueses-Cisneros, 2007, sp. nov." docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="FFBFC5524467B51B7D616779662FFFB0" masterDocTitle="A new species of Rhaebo (Anura: Bufonidae) from the Cordillera Occidental of Colombia" masterLastPageNumber="59" masterPageNumber="53" pageNumber="54" updateTime="1698233827927" updateUser="plazi">
<mods:mods id="75C20E4C4A2A43BA390455C02AACC21C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="107DC28CB6037360ED924655CE969C5C">
<mods:title id="D6C8B1070E504FD449B54353B55CBB7A">A new species of Rhaebo (Anura: Bufonidae) from the Cordillera Occidental of Colombia</mods:title>
</mods:titleInfo>
<mods:name id="7ED18F92E08C55F9931657A7A85E7A27" type="personal">
<mods:role id="7D90A6071CDAD974DDD07F4CB3EBAD31">
<mods:roleTerm id="8C52F10FA334BD84AD23EE3CAA892BC1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3018D92C24AB8720D67314B457B88C5A">Mueses-Cisneros, Jonh Jairo</mods:namePart>
</mods:name>
<mods:typeOfResource id="44F8BF1139652CD0703260DA63881F13">text</mods:typeOfResource>
<mods:relatedItem id="F8C0EEA1374A72D353CAD25C077EFBC5" type="host">
<mods:titleInfo id="F18AA22B2876147FDEB1D0957C99513E">
<mods:title id="96109FA73BF275F82C275849798200B7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8EE1A2102E4C1B9CF65182182F2CF705">
<mods:date id="A0CA31307304AED0AABA9720F173A352">2007</mods:date>
<mods:detail id="F210C2655AF0EA090361657E20806160" type="volume">
<mods:number id="A55AFD3B0EFF6032A2DB4A9FF6FEAB92">1662</mods:number>
</mods:detail>
<mods:extent id="7657C6E4FA18B5D9C741759F986D6F3B" unit="page">
<mods:start id="AD8F47ED27DD8CB824DF380F1A15EC01">53</mods:start>
<mods:end id="51E877B431A5148A5EE7D7F1486B1D4A">59</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F7B6D9798DEDEE47619039C499EECAFC">journal article</mods:classification>
<mods:identifier id="EE89FF187A13D125167AE5E217C5050E" type="DOI">10.5281/zenodo.179977</mods:identifier>
<mods:identifier id="D7F715C4411E2B08892A609903B0FBAC" type="GBIF-Dataset">8a222f0b-c42b-456b-ba16-ff056e4ee6bd</mods:identifier>
<mods:identifier id="E305394F55DFC83F96FE88B7AA06DBF7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="01178AD091BD68B3B2F6B4761DDC9797" type="Zenodo-Dep">179977</mods:identifier>
</mods:mods>
<treatment id="0386BD2A4466B51E7DF6662663B6FE42" ID-DOI="http://doi.org/10.5281/zenodo.6243973" ID-GBIF-Taxon="119382034" ID-Zenodo-Dep="6243973" LSID="urn:lsid:plazi:treatment:0386BD2A4466B51E7DF6662663B6FE42" httpUri="http://treatment.plazi.org/id/0386BD2A4466B51E7DF6662663B6FE42" lastPageId="5" lastPageNumber="58" pageId="1" pageNumber="54">
<subSubSection id="C3355FB74466B51A7DF6662667C2FE2C" pageId="1" pageNumber="54" type="nomenclature">
<paragraph id="8B900C3C4466B51A7DF666266784FEC9" blockId="1.[151,493,351,412]" box="[151,427,351,377]" pageId="1" pageNumber="54">
<heading id="D0D8BB504466B51A7DF666266784FEC9" bold="true" box="[151,427,351,377]" fontSize="11" level="1" pageId="1" pageNumber="54" reason="1">
<emphasis id="B95BD02E4466B51A7DF666266784FEC9" bold="true" box="[151,427,351,377]" pageId="1" pageNumber="54">
<taxonomicName id="4C2F77BF4466B51A7DF666266769FEC9" ID-CoL="4RY97" box="[151,326,351,377]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="lynchi" status="sp. nov.">
<emphasis id="B95BD02E4466B51A7DF666266769FEC9" bold="true" box="[151,326,351,377]" italics="true" pageId="1" pageNumber="54">Rhaebo lynchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2686D554466B51A7C2C66266784FEC9" box="[333,427,351,377]" pageId="1" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B900C3C4466B51A7DF666FB67C2FE2C" blockId="1.[151,493,351,412]" box="[151,493,386,412]" pageId="1" pageNumber="54">
(
<figureCitation id="131410B94466B51A7DFE66FB66D1FE2C" box="[159,254,386,412]" captionStart="FIGURE 1" captionStartId="1.[151,255,1647,1671]" captionTargetBox="[174,1415,759,1614]" captionTargetId="figure@1.[164,1422,751,1620]" captionTargetPageId="1" captionText="FIGURE 1. Rhaebo lynchi sp. nov.: dorsal (A) and ventral (B) views of holotype, ICN 1161, adult female SVL 37.5 mm. Photos by Jorge Contreras." httpUri="https://zenodo.org/record/179978/files/figure.png" pageId="1" pageNumber="54">Figure 1</figureCitation>
A–B, 3A–C, 4A–B)
</paragraph>
</subSubSection>
<subSubSection id="C3355FB74466B51A7DF666AB673BFDD4" pageId="1" pageNumber="54" type="materials_examined">
<paragraph id="8B900C3C4466B51A7DF666AB655EFDA4" blockId="1.[151,1437,466,692]" pageId="1" pageNumber="54">
<emphasis id="B95BD02E4466B51A7DF666AB673FFE5C" bold="true" box="[151,272,466,492]" pageId="1" pageNumber="54">
<typeStatus id="5494B29E4466B51A7DF666AB6724FE5C" box="[151,267,466,492]" pageId="1" pageNumber="54" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED3E94F94466B51A7C7866AB6764FE5C" box="[281,331,466,492]" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="1" pageNumber="54">ICN</collectionCode>
1161, an adult female collected between
<date id="FF912AFC4466B51A7E5F66AB658DFE5C" box="[830,930,466,492]" pageId="1" pageNumber="54" value="1928-04">28 April</date>
and 0
<date id="FF912AFC4466B51A7E8C66AB6250FE5C" box="[1005,1151,466,492]" pageId="1" pageNumber="54" value="1972-05-04">4 May 1972</date>
by Pedro M. Ruiz-Carranza, Gustavo Lozano, F. Flórez and José Vicente Rodriguez.
</paragraph>
<paragraph id="8B900C3C4466B51A7DA6655B673BFDD4" blockId="1.[151,1437,466,692]" pageId="1" pageNumber="54">
<emphasis id="B95BD02E4466B51A7DA6655B6741FD8C" bold="true" box="[199,366,546,572]" pageId="1" pageNumber="54">
<typeStatus id="5494B29E4466B51A7DA6655B672BFD8C" box="[199,260,546,572]" pageId="1" pageNumber="54">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3384CAC4466B51A7C18655B6436FD8C" box="[377,537,546,572]" name="Colombia" pageId="1" pageNumber="54">COLOMBIA</collectingCountry>
, Antioquia, Urrao, boundaries with Caicedo, Vereda “El Chuscal”,
<quantity id="4CD7A1D94466B51A7822655B63B7FD8C" box="[1347,1432,546,572]" metricMagnitude="3" metricUnit="m" metricValue="2.585" pageId="1" pageNumber="54" unit="m" value="2585.0">2585m</quantity>
. (
<figureCitation id="131410B94466B51A7DFE6533672AFDD4" box="[159,261,586,612]" captionStart="FIGURE 2" captionStartId="2.[151,255,1768,1792]" captionTargetBox="[387,1177,721,1705]" captionTargetId="figure@2.[368,1219,700,1744]" captionTargetPageId="2" captionText="FIGURE 2. Type locality (squares) of Rhaebo lynchi sp. nov. The shaded areas represent elevations above 2000 meters." httpUri="https://zenodo.org/record/179979/files/figure.png" pageId="1" pageNumber="54">Figure 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3355FB74466B51A7DA6650B6590F854" pageId="1" pageNumber="54" type="diagnosis">
<paragraph id="8B900C3C4466B51A7DA6650B63B3FD04" blockId="1.[151,1437,466,692]" pageId="1" pageNumber="54">
<emphasis id="B95BD02E4466B51A7DA6650B6768FD3C" bold="true" box="[199,327,626,652]" pageId="1" pageNumber="54">Diagnosis.</emphasis>
A species of
<taxonomicName id="4C2F77BF4466B51A7C91650B6465FD3B" box="[496,586,626,651]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="1" pageNumber="54" phylum="Chordata" rank="genus">
<emphasis id="B95BD02E4466B51A7C91650B6465FD3B" box="[496,586,626,651]" italics="true" pageId="1" pageNumber="54">Rhaebo</emphasis>
</taxonomicName>
characterized by having a row of very prominent, subacuminate tarsal tubercles on the outer edge (
<figureCitation id="131410B94466B51A7CBE65E36410FD04" box="[479,575,666,692]" captionStart="FIGURE 1" captionStartId="1.[151,255,1647,1671]" captionTargetBox="[174,1415,759,1614]" captionTargetId="figure@1.[164,1422,751,1620]" captionTargetPageId="1" captionText="FIGURE 1. Rhaebo lynchi sp. nov.: dorsal (A) and ventral (B) views of holotype, ICN 1161, adult female SVL 37.5 mm. Photos by Jorge Contreras." httpUri="https://zenodo.org/record/179978/files/figure.png" pageId="1" pageNumber="54">Figure 1</figureCitation>
A, 3A–B) and very prominent, subacuminate ulnar tubercles (
<figureCitation id="131410B94466B51A787265E36357FD04" box="[1299,1400,666,692]" captionStart="FIGURE 3" captionStartId="3.[151,255,1750,1774]" captionTargetBox="[430,1098,910,1697]" captionTargetId="figure@3.[430,1156,900,1727]" captionTargetPageId="3" captionText="FIGURE 3. Tarsal region and metatarsal fold (A), Foot (B) and Hand (C) of holotype of Rhaebo lynchi sp. nov. Scales equal 2 mm." httpUri="https://zenodo.org/record/179980/files/figure.png" pageId="1" pageNumber="54">Figure 3</figureCitation>
C).
</paragraph>
<caption id="DF505CB44466B51A7DF6611667C2F919" httpUri="https://zenodo.org/record/179978/files/figure.png" pageId="1" pageNumber="54" targetBox="[174,1415,759,1614]" targetPageId="1">
<paragraph id="8B900C3C4466B51A7DF6611667C2F919" blockId="1.[151,1436,1647,1705]" pageId="1" pageNumber="54">
<emphasis id="B95BD02E4466B51A7DF661166734F937" bold="true" box="[151,283,1647,1671]" pageId="1" pageNumber="54">FIGURE 1.</emphasis>
<taxonomicName id="4C2F77BF4466B51A7C4561096790F937" box="[292,447,1648,1671]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="lynchi" status="sp. nov.">
<emphasis id="B95BD02E4466B51A7C4561096790F937" box="[292,447,1648,1671]" italics="true" pageId="1" pageNumber="54">Rhaebo lynchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2686D554466B51A7CA96116640FF937" box="[456,544,1647,1671]" pageId="1" pageNumber="54" rank="species">
<emphasis id="B95BD02E4466B51A7CA96116640FF937" bold="true" box="[456,544,1647,1671]" pageId="1" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
: dorsal (
<emphasis id="B95BD02E4466B51A7FE3611664B0F937" bold="true" box="[642,671,1647,1671]" pageId="1" pageNumber="54">A)</emphasis>
and ventral (
<emphasis id="B95BD02E4466B51A7E536116656BF937" bold="true" box="[818,836,1647,1671]" pageId="1" pageNumber="54">B</emphasis>
) views of holotype, ICN 1161, adult female SVL 37.5 mm. Photos by Jorge Contreras.
</paragraph>
</caption>
<paragraph id="8B900C3C4466B51A7DA6607B6590F854" blockId="1.[151,1436,1794,2020]" pageId="1" pageNumber="54">
<emphasis id="B95BD02E4466B51A7DA6607B647CF8AC" bold="true" box="[199,595,1794,1820]" pageId="1" pageNumber="54">Comparison with other species.</emphasis>
<taxonomicName id="4C2F77BF4466B51A7F3D607B6529F8AB" box="[604,774,1794,1819]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4466B51A7F3D607B6529F8AB" box="[604,774,1794,1819]" italics="true" pageId="1" pageNumber="54">Rhaebo lynchi</emphasis>
</taxonomicName>
also differs from any other known
<taxonomicName id="4C2F77BF4466B51A79CF607B6327F8AB" box="[1198,1288,1794,1819]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="1" pageNumber="54" phylum="Chordata" rank="genus">
<emphasis id="B95BD02E4466B51A79CF607B6327F8AB" box="[1198,1288,1794,1819]" italics="true" pageId="1" pageNumber="54">Rhaebo</emphasis>
</taxonomicName>
by having a smaller size (
<quantity id="4CD7A1D94466B51A7C5F60536780F8F4" box="[318,431,1834,1860]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="1" pageNumber="54" unit="mm" value="37.5">37.5 mm</quantity>
SVL in the adult female) versus
<quantity id="4CD7A1D94466B51A7E3E6053621EF8F4" box="[863,1073,1834,1860]" metricMagnitude="-1" metricUnit="m" metricValue="2.125" metricValueMax="2.3449999999999998" metricValueMin="1.905" pageId="1" pageNumber="54" unit="mm" value="212.5" valueMax="234.5" valueMin="190.5">190.5–234.5 mm</quantity>
. SVL in adult females of
<taxonomicName id="4C2F77BF4466B51A78E260536721F8DB" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="blombergi">
<emphasis id="B95BD02E4466B51A78E260536721F8DB" italics="true" pageId="1" pageNumber="54">R. blombergi</emphasis>
</taxonomicName>
;
<quantity id="4CD7A1D94466B51A7C7F602B67E3F8DC" box="[286,460,1874,1900]" metricMagnitude="-2" metricUnit="m" metricValue="8.194999999999999" metricValueMax="9.23" metricValueMin="7.159999999999999" pageId="1" pageNumber="54" unit="mm" value="81.94999999999999" valueMax="92.3" valueMin="71.6">71.6–92.3 mm</quantity>
. SVL in adult females of
<taxonomicName id="4C2F77BF4466B51A7F98602B62B0F8DC" authority="Hoogmoed 1989" authorityName="Hoogmoed" authorityYear="1989" box="[761,1183,1874,1900]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="caerulestictus">
<emphasis id="B95BD02E4466B51A7F98602B6593F8DB" box="[761,956,1874,1899]" italics="true" pageId="1" pageNumber="54">R. caerulestictus</emphasis>
(
<bibRefCitation id="EFBE71CD4466B51A7EAA602B62B9F8DC" author="Hoogmoed" box="[971,1174,1874,1900]" pageId="1" pageNumber="59" refString="Hoogmoed, M. S. (1989) On the identity of some toads of the genus Bufo from Ecuador, with additional remarks on Andinophryne colomai Hoogmoed, 1985 (Amphibia: Anura: Bufonidae). Zoologische Verhandelingen, 250,1 - 32." type="book chapter" year="1989">Hoogmoed 1989</bibRefCitation>
)
</taxonomicName>
; 58.8–69.0 mm. SVL in adult females of
<taxonomicName id="4C2F77BF4466B51A7C1660036409F823" box="[375,550,1914,1939]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="glaberrimus">
<emphasis id="B95BD02E4466B51A7C1660036409F823" box="[375,550,1914,1939]" italics="true" pageId="1" pageNumber="54">R. glaberrimus</emphasis>
</taxonomicName>
;
<quantity id="4CD7A1D94466B51A7F546003652EF824" box="[565,769,1914,1940]" metricMagnitude="-1" metricUnit="m" metricValue="1.341" metricValueMax="1.469" metricValueMin="1.2129999999999999" pageId="1" pageNumber="54" unit="mm" value="134.1" valueMax="146.9" valueMin="121.3">121.3–146.9 mm</quantity>
. SVL in adult males of
<taxonomicName id="4C2F77BF4466B51A7972600362BEF823" box="[1043,1169,1914,1939]" class="Amphibia" family="Bufonidae" genus="Bufo" kingdom="Animalia" order="Anura" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="guttatus">
<emphasis id="B95BD02E4466B51A7972600362BEF823" box="[1043,1169,1914,1939]" italics="true" pageId="1" pageNumber="54">R. guttatus</emphasis>
</taxonomicName>
;
<quantity id="4CD7A1D94466B51A79C160036375F824" box="[1184,1370,1914,1940]" metricMagnitude="-2" metricUnit="m" metricValue="9.45" metricValueMax="11.129999999999999" metricValueMin="7.77" pageId="1" pageNumber="54" unit="mm" value="94.5" valueMax="111.3" valueMin="77.7">77.7–111.3 mm</quantity>
. SVL in adult females of
<taxonomicName id="4C2F77BF4466B51A7C1960DB6401F80B" box="[376,558,1954,1979]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="haematiticus">
<emphasis id="B95BD02E4466B51A7C1960DB6401F80B" box="[376,558,1954,1979]" italics="true" pageId="1" pageNumber="54">R. haematiticus</emphasis>
</taxonomicName>
[although
<bibRefCitation id="EFBE71CD4466B51A7FCD60DB6579F80C" author="Savage" box="[684,854,1954,1980]" pageId="1" pageNumber="59" refString="Savage, J. M. (2002) The Amphibians and Reptiles of Costa Rica: A Herpetofauna Between Two Continents, Between Two Seas. The University of Chicago Press, Chicago, 954 pp." type="book" year="2002">Savage (2002)</bibRefCitation>
reported adult females between
<quantity id="4CD7A1D94466B51A79B360DB6379F80C" box="[1234,1366,1954,1980]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="1" pageNumber="54" unit="mm" value="65.0" valueMax="80.0" valueMin="50.0">50–80 mm</quantity>
]; 64.4 and
<quantity id="4CD7A1D94466B51A7DA960B3671AF854" box="[200,309,1994,2020]" metricMagnitude="-2" metricUnit="m" metricValue="6.65" pageId="1" pageNumber="54" unit="mm" value="66.5">66.5 mm</quantity>
. SVL in adult females of
<taxonomicName id="4C2F77BF4466B51A7F0160B36594F854" authority="Hoogmoed 1977" authorityName="Hoogmoed" authorityYear="1977" box="[608,955,1994,2020]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="nasicus">
<emphasis id="B95BD02E4466B51A7F0160B364F6F853" box="[608,729,1994,2019]" italics="true" pageId="1" pageNumber="54">R. nasicus</emphasis>
(
<bibRefCitation id="EFBE71CD4466B51A7F8960B3659DF854" author="Hoogmoed" box="[744,946,1994,2020]" pageId="1" pageNumber="59" refString="Hoogmoed, M. S. (1977) On the presence of Bufo nasicus Werner in Guiana, with a redescription of the species on the basis of recently collected material. Zoologische Mededelingen, 51 (16), 265 - 275." type="journal article" year="1977">Hoogmoed 1977</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3355FB74465B51E7DA667E16374FECA" lastPageId="5" lastPageNumber="58" pageId="2" pageNumber="55" type="description">
<paragraph id="8B900C3C4465B5197DA667E162FDFD22" blockId="2.[151,1436,152,658]" pageId="2" pageNumber="55">
<taxonomicName id="4C2F77BF4465B5197DA667E16798FF01" box="[199,439,152,177]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="glaberrimus">
<emphasis id="B95BD02E4465B5197DA667E16798FF01" box="[199,439,152,177]" italics="true" pageId="2" pageNumber="55">Rhaebo glaberrimus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F77BF4465B5197C9067E16445FF01" box="[497,618,152,177]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="nasicus">
<emphasis id="B95BD02E4465B5197C9067E16445FF01" box="[497,618,152,177]" italics="true" pageId="2" pageNumber="55">R. nasicus</emphasis>
</taxonomicName>
, like
<taxonomicName id="4C2F77BF4465B5197FCD67E1653AFF01" box="[684,789,152,177]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197FCD67E1653AFF01" box="[684,789,152,177]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
, have ulnar and tarsal tubercles, but these are very low and round in
<taxonomicName id="4C2F77BF4465B5197C5767B967C9FF69" box="[310,486,192,217]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="glaberrimus">
<emphasis id="B95BD02E4465B5197C5767B967C9FF69" box="[310,486,192,217]" italics="true" pageId="2" pageNumber="55">R. glaberrimus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F77BF4465B5197F4367B964B3FF69" box="[546,668,192,217]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="nasicus">
<emphasis id="B95BD02E4465B5197F4367B964B3FF69" box="[546,668,192,217]" italics="true" pageId="2" pageNumber="55">R. nasicus</emphasis>
</taxonomicName>
(very prominent and subacuminate in
<taxonomicName id="4C2F77BF4465B519790667B962FFFF69" box="[1127,1232,192,217]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B519790667B962FFFF69" box="[1127,1232,192,217]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
);
<taxonomicName id="4C2F77BF4465B519798A67B963B3FF69" box="[1259,1436,192,217]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="glaberrimus">
<emphasis id="B95BD02E4465B519798A67B963B3FF69" box="[1259,1436,192,217]" italics="true" pageId="2" pageNumber="55">R. glaberrimus</emphasis>
</taxonomicName>
also has a short inner tarsal fold (absent in
<taxonomicName id="4C2F77BF4465B5197FFF67916528FEB1" box="[670,775,232,257]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197FFF67916528FEB1" box="[670,775,232,257]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
),
<taxonomicName id="4C2F77BF4465B5197E43679165FCFEB1" box="[802,979,232,257]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="glaberrimus">
<emphasis id="B95BD02E4465B5197E43679165FCFEB1" box="[802,979,232,257]" italics="true" pageId="2" pageNumber="55">R. glaberrimus</emphasis>
</taxonomicName>
lacks metacarpal and metatarsal folds (present in
<taxonomicName id="4C2F77BF4465B5197C7A666967ACFE99" box="[283,387,272,297]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197C7A666967ACFE99" box="[283,387,272,297]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
) and the cloacal opening is at lower level of thighs or ventral in
<taxonomicName id="4C2F77BF4465B519791A66696304FE99" box="[1147,1323,272,297]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="glaberrimus">
<emphasis id="B95BD02E4465B519791A66696304FE99" box="[1147,1323,272,297]" italics="true" pageId="2" pageNumber="55">R. glaberrimus</emphasis>
</taxonomicName>
(at upper level of thighs in
<taxonomicName id="4C2F77BF4465B5197C06664167FFFEE1" box="[359,464,312,337]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197C06664167FFFEE1" box="[359,464,312,337]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
).
<taxonomicName id="4C2F77BF4465B5197C886641647DFEE1" box="[489,594,312,337]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197C886641647DFEE1" box="[489,594,312,337]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
, like
<taxonomicName id="4C2F77BF4465B5197FF466416520FEE1" box="[661,783,312,337]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="nasicus">
<emphasis id="B95BD02E4465B5197FF466416520FEE1" box="[661,783,312,337]" italics="true" pageId="2" pageNumber="55">R. nasicus</emphasis>
</taxonomicName>
, has prominent metatarsal and metacarpal folds (Plates 1–3;
<bibRefCitation id="EFBE71CD4465B5197DB2661967B3FECA" author="Hoogmoed" box="[211,412,352,378]" pageId="2" pageNumber="59" refString="Hoogmoed, M. S. (1977) On the presence of Bufo nasicus Werner in Guiana, with a redescription of the species on the basis of recently collected material. Zoologische Mededelingen, 51 (16), 265 - 275." type="journal article" year="1977">Hoogmoed 1977</bibRefCitation>
), but
<taxonomicName id="4C2F77BF4465B5197C8166196476FEC9" box="[480,601,352,377]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="nasicus">
<emphasis id="B95BD02E4465B5197C8166196476FEC9" box="[480,601,352,377]" italics="true" pageId="2" pageNumber="55">R. nasicus</emphasis>
</taxonomicName>
differs also from
<taxonomicName id="4C2F77BF4465B5197E4E661965B7FEC9" box="[815,920,352,377]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197E4E661965B7FEC9" box="[815,920,352,377]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
in having a prominent inner tarsal fold and the head with low but prominent bony ridges (absent in
<taxonomicName id="4C2F77BF4465B5197E2666F1659DFE11" box="[839,946,392,417]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197E2666F1659DFE11" box="[839,946,392,417]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
).
<taxonomicName id="4C2F77BF4465B5197EAF66F16246FE11" box="[974,1129,392,417]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="blombergi">
<emphasis id="B95BD02E4465B5197EAF66F16246FE11" box="[974,1129,392,417]" italics="true" pageId="2" pageNumber="55">R. blombergi</emphasis>
</taxonomicName>
has low and round ulnar tubercles (very prominent and subacuminate in
<taxonomicName id="4C2F77BF4465B5197FBF66C96564FE79" box="[734,843,432,457]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197FBF66C96564FE79" box="[734,843,432,457]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
),
<taxonomicName id="4C2F77BF4465B5197E0666C9622BFE79" box="[871,1028,432,457]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="blombergi">
<emphasis id="B95BD02E4465B5197E0666C9622BFE79" box="[871,1028,432,457]" italics="true" pageId="2" pageNumber="55">R. blombergi</emphasis>
</taxonomicName>
lacks outer tarsal tubercles (very prominent and subacuminate in
<taxonomicName id="4C2F77BF4465B5197F4A66A164B5FE41" box="[555,666,472,497]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197F4A66A164B5FE41" box="[555,666,472,497]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
),
<taxonomicName id="4C2F77BF4465B5197FD866A16577FE41" box="[697,856,472,497]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="blombergi">
<emphasis id="B95BD02E4465B5197FD866A16577FE41" box="[697,856,472,497]" italics="true" pageId="2" pageNumber="55">R. blombergi</emphasis>
</taxonomicName>
has long limbs (short limbs in
<taxonomicName id="4C2F77BF4465B519799566A1634CFE41" box="[1268,1379,472,497]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B519799566A1634CFE41" box="[1268,1379,472,497]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
),
<taxonomicName id="4C2F77BF4465B51978E266A16720FDA9" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="blombergi">
<emphasis id="B95BD02E4465B51978E266A16720FDA9" italics="true" pageId="2" pageNumber="55">R. blombergi</emphasis>
</taxonomicName>
lacks metacarpal and metatarsal folds (present in
<taxonomicName id="4C2F77BF4465B5197E0A657965F9FDA9" box="[875,982,512,537]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197E0A657965F9FDA9" box="[875,982,512,537]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
) and
<taxonomicName id="4C2F77BF4465B519797D65796298FDA9" box="[1052,1207,512,537]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="blombergi">
<emphasis id="B95BD02E4465B519797D65796298FDA9" box="[1052,1207,512,537]" italics="true" pageId="2" pageNumber="55">R. blombergi</emphasis>
</taxonomicName>
has an inner tarsal fold (absent in
<taxonomicName id="4C2F77BF4465B5197C2A6551679BFDF1" box="[331,436,552,577]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197C2A6551679BFDF1" box="[331,436,552,577]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
).
<taxonomicName id="4C2F77BF4465B5197CAC6551648FFDF1" box="[461,672,552,577]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="caeruleostictus">
<emphasis id="B95BD02E4465B5197CAC6551648FFDF1" box="[461,672,552,577]" italics="true" pageId="2" pageNumber="55">R. caeruleostictus</emphasis>
</taxonomicName>
has tarsal tubercles, although these are not very prominent as in
<taxonomicName id="4C2F77BF4465B5197DF66529672DFDD9" box="[151,258,592,617]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197DF66529672DFDD9" box="[151,258,592,617]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2F77BF4465B5197C73652967C9FDD9" box="[274,486,592,617]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="caeruleostictus">
<emphasis id="B95BD02E4465B5197C73652967C9FDD9" box="[274,486,592,617]" italics="true" pageId="2" pageNumber="55">R. caeruleostictus</emphasis>
</taxonomicName>
lacks ulnar tubercles (present in
<taxonomicName id="4C2F77BF4465B5197E19652965CCFDD9" box="[888,995,592,617]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197E19652965CCFDD9" box="[888,995,592,617]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
),
<taxonomicName id="4C2F77BF4465B5197E9F652962FDFDD9" box="[1022,1234,592,617]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="caeruleostictus">
<emphasis id="B95BD02E4465B5197E9F652962FDFDD9" box="[1022,1234,592,617]" italics="true" pageId="2" pageNumber="55">R. caeruleostictus</emphasis>
</taxonomicName>
have long limbs (short limbs in
<taxonomicName id="4C2F77BF4465B5197C286501679DFD21" box="[329,434,632,657]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197C286501679DFD21" box="[329,434,632,657]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
) and
<taxonomicName id="4C2F77BF4465B5197C95650164E9FD21" box="[500,710,632,657]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="caeruleostictus">
<emphasis id="B95BD02E4465B5197C95650164E9FD21" box="[500,710,632,657]" italics="true" pageId="2" pageNumber="55">R. caeruleostictus</emphasis>
</taxonomicName>
has an inner tarsal fold (absent in
<taxonomicName id="4C2F77BF4465B5197938650162EDFD21" box="[1113,1218,632,657]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi">
<emphasis id="B95BD02E4465B5197938650162EDFD21" box="[1113,1218,632,657]" italics="true" pageId="2" pageNumber="55">R. lynchi</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF505CB44465B5197DF6619163B2F94F" box="[151,1437,1768,1792]" httpUri="https://zenodo.org/record/179979/files/figure.png" pageId="2" pageNumber="55" targetBox="[387,1177,721,1705]" targetPageId="2">
<paragraph id="8B900C3C4465B5197DF6619163B2F94F" blockId="2.[151,1437,1768,1792]" box="[151,1437,1768,1792]" pageId="2" pageNumber="55">
<emphasis id="B95BD02E4465B5197DF661916736F8B0" bold="true" box="[151,281,1768,1792]" pageId="2" pageNumber="55">FIGURE 2.</emphasis>
Type locality (squares) of
<taxonomicName id="4C2F77BF4465B5197F56619164FFF94F" box="[567,720,1768,1791]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="lynchi" status="sp. nov.">
<emphasis id="B95BD02E4465B5197F56619164FFF94F" box="[567,720,1768,1791]" italics="true" pageId="2" pageNumber="55">Rhaebo lynchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2686D554465B5197FB661916503F8B0" box="[727,812,1768,1792]" pageId="2" pageNumber="55" rank="species">
<emphasis id="B95BD02E4465B5197FB661916503F8B0" bold="true" box="[727,812,1768,1792]" pageId="2" pageNumber="55">sp. nov.</emphasis>
</taxonomicNameLabel>
The shaded areas represent elevations above 2000 meters.
</paragraph>
</caption>
<paragraph id="8B900C3C4465B5187DA6602C65FFFCEA" blockId="2.[151,1436,1877,2023]" lastBlockId="3.[151,1437,152,858]" lastPageId="3" lastPageNumber="56" pageId="2" pageNumber="55">
<emphasis id="B95BD02E4465B5197DA6602C642FF8DF" bold="true" box="[199,512,1877,1903]" pageId="2" pageNumber="55">
Description of
<typeStatus id="5494B29E4465B5197CE6602C67D4F8DF" box="[391,507,1877,1903]" pageId="2" pageNumber="55" type="holotype">Holotype</typeStatus>
.
</emphasis>
Toad of small size, female
<quantity id="4CD7A1D94465B5197E3B602C6594F8DF" box="[858,955,1877,1903]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="2" pageNumber="55" unit="mm" value="37.5">37.5mm</quantity>
SVL. Head narrower than body width, slightly longer than wide. Width of head at corners of jaws slightly wider than at level of tympanum. HW 98.4% of the head length and 33.3% of SVL. Head length 33.9% of the SVL. Head subacuminate in dorsal view (
<figureCitation id="131410B94465B5197DBE60B4676BF857" box="[223,324,1997,2023]" captionStart="FIGURE 4" captionStartId="4.[151,255,1814,1838]" captionTargetBox="[451,1147,805,1742]" captionTargetId="figure@4.[385,1202,774,1793]" captionTargetPageId="4" captionText="FIGURE 4. Dorsal view (A) and lateral profile (B) of head of holotype of Rhaebo lynchi sp. nov. Scales equal 2 mm." httpUri="https://zenodo.org/record/179981/files/figure.png" pageId="2" pageNumber="55">Figure 4</figureCitation>
A); snout acuminate dorsally, weakly truncate in lateral profile (
<figureCitation id="131410B94465B519794E60B462BAF857" box="[1071,1173,1997,2023]" captionStart="FIGURE 4" captionStartId="4.[151,255,1814,1838]" captionTargetBox="[451,1147,805,1742]" captionTargetId="figure@4.[385,1202,774,1793]" captionTargetPageId="4" captionText="FIGURE 4. Dorsal view (A) and lateral profile (B) of head of holotype of Rhaebo lynchi sp. nov. Scales equal 2 mm." httpUri="https://zenodo.org/record/179981/files/figure.png" pageId="2" pageNumber="55">Figure 4</figureCitation>
B). Tip of snout with a fleshy vertical ridge. Distance between nostril and tip of snout 73.3% of distance between nostril and eye. Nostrils situated posterior to the anterior rim of the mouth, below
<emphasis id="B95BD02E4464B5187EC067B96249FF69" box="[929,1126,192,217]" italics="true" pageId="3" pageNumber="56">canthus rostralis</emphasis>
in a slightly swollen area. Nostrils oblique, oval, directed laterally. Distance between nostrils 130.0% of that between nostril and eye. Internarial area concave. Top of head up to anterior corner of eyelid weakly concave. Interorbital and occipital region flat, with some low and subacuminate tubercles extending up to inter-parotoid region and back. Upper eyelid narrower than interorbital area. Upper eyelid width 64.4 of IOD. Upper eyelid with some low and rounded tubercles in central area. Inner and outer rim of upper eyelid not delineated by tubercles. Outer rim of upper eyelid slightly projected beyond eye. Cephalic crests absent.
<emphasis id="B95BD02E4464B5187ED166C96253FE79" box="[944,1148,432,457]" italics="true" pageId="3" pageNumber="56">Canthus rostralis</emphasis>
straight and fleshy, projected over loreal region. Loreal region concave, with some, small and thin spines extending up to middle part of flanks. Spines on infra-parotoid and infra-tympanic regions much more prominent and denser than those on loreal region. Lips not flared. Eyes with horizontally oval pupil. Preorbital ridge absent. Tympanum large, rounded, 63.6% of eye length. Anterior near 4/5 (or all) of tympanum visible. Supratympanic fleshy ridge present. Parotoids distinct, oval, rounded anteriorly and pointed posteriorly. Parotoids longer than wide, parotoid width 40.2% of parotoid length and 34.5% of inter-parotoid distance. Parotoids with some pores. Outer margin of parotoids without indentations. Inter-parotoidal region with two very low and weakly evident crests. Skin of back smooth, bearing some low tubercles in sacral region and round warts on postsacral region. Interrupted lateral oblique row of five to six subacuminate tubercles, from parotoids to groin. Ventral skin finely tuberculate, with some minute spines mainly on chest and belly.
</paragraph>
<caption id="DF505CB44464B5187DF661AF6733F8A0" httpUri="https://zenodo.org/record/179980/files/figure.png" pageId="3" pageNumber="56" targetBox="[430,1098,910,1697]" targetPageId="3">
<paragraph id="8B900C3C4464B5187DF661AF6733F8A0" blockId="3.[151,1435,1750,1808]" pageId="3" pageNumber="56">
<emphasis id="B95BD02E4464B5187DF661AF6736F95E" bold="true" box="[151,281,1750,1774]" pageId="3" pageNumber="56">FIGURE 3.</emphasis>
Tarsal region and metatarsal fold (
<emphasis id="B95BD02E4464B5187FF361AF648AF95E" bold="true" box="[658,677,1750,1774]" pageId="3" pageNumber="56">A</emphasis>
), Foot (
<emphasis id="B95BD02E4464B5187F9C61AF6520F95E" bold="true" box="[765,783,1750,1774]" pageId="3" pageNumber="56">B</emphasis>
) and Hand (
<emphasis id="B95BD02E4464B5187EF661AF6585F95E" bold="true" box="[919,938,1750,1774]" pageId="3" pageNumber="56">C</emphasis>
) of holotype of
<taxonomicName id="4C2F77BF4464B518793B61AE62DCF95E" box="[1114,1267,1751,1774]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="lynchi" status="sp. nov.">
<emphasis id="B95BD02E4464B518793B61AE62DCF95E" box="[1114,1267,1751,1774]" italics="true" pageId="3" pageNumber="56">Rhaebo lynchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2686D554464B518799A61AF637EF95E" box="[1275,1361,1750,1774]" pageId="3" pageNumber="56" rank="species">
<emphasis id="B95BD02E4464B518799A61AF637EF95E" bold="true" box="[1275,1361,1750,1774]" pageId="3" pageNumber="56">sp. nov.</emphasis>
</taxonomicNameLabel>
Scales equal 2 mm.
</paragraph>
</caption>
<paragraph id="8B900C3C4464B51F7DA6601F6250FE9A" blockId="3.[151,1436,1894,2003]" lastBlockId="4.[151,1437,152,740]" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
Forelimbs slender and short, with numerous spines and some rounded and low tubercles on upper surface and flanks. Hand length 72.4% of SVL. Fingers basally webbed. Hand-web formula
<date id="FF912AFC4464B51879EE60E862E9F81B" box="[1167,1222,1937,1963]" pageId="3" pageNumber="56">
<emphasis id="B95BD02E4464B51879EE60E862B5F81B" bold="true" box="[1167,1178,1937,1963]" pageId="3" pageNumber="56">I</emphasis>
2–2
</date>
-
<date id="FF912AFC4464B51879AD60E862DEF81B" box="[1228,1265,1937,1963]" pageId="3" pageNumber="56">
<emphasis id="B95BD02E4464B51879AD60E862CCF81B" bold="true" box="[1228,1251,1937,1963]" pageId="3" pageNumber="56">II</emphasis>
2
</date>
-–3
<date id="FF912AFC4464B518787560E8636CF81B" box="[1300,1347,1937,1963]" pageId="3" pageNumber="56">
<emphasis id="B95BD02E4464B518787560E86319F81B" bold="true" box="[1300,1334,1937,1963]" pageId="3" pageNumber="56">III</emphasis>
3
</date>
–2.5
<emphasis id="B95BD02E4464B518781960E863BAF81B" bold="true" box="[1400,1429,1937,1963]" pageId="3" pageNumber="56">IV</emphasis>
. Fingers with slender, fleshy and non bulbous tips; basally broadened. Finger I slightly shorter than II. Supernumerary palmar tubercles not evident. Subarticular tubercles rounded and low. Palmar and thenar tubercles weakly evident, palmar tubercle large, rounded, 1.1 times the size of oval thenar tubercle. A row of six to seven subacuminate ulnar tubercles and three to four low tubercles along inner edge. Ulnar fold absent. Metacarpal fold present, extending from metatarsus to nearly half of length finger IV.
</paragraph>
<paragraph id="8B900C3C4463B51F7DAF664166C5FDDC" blockId="4.[151,1437,152,740]" pageId="4" pageNumber="57">
<collectingCountry id="F3384CAC4463B51F7DAF66416726FEE2" box="[206,265,312,338]" name="India" pageId="4" pageNumber="57">Hind</collectingCountry>
limbs short. Upper surface with some low tubercles extending up the thighs. Outer edge of tibia with four to six prominent and rounded tubercles and some spines. Inner tarsal fold absent. Tarsus bearing a row of five to six subacuminate tubercles on outer edge and a row of two to four low tubercles on inner edge. Foot length and tibia length 35.7% and 32.3% of SVL. Toes with slender, fleshy, with non bulbous tips; basally broadened or not. Toes basally webbed. Foot-web formula
<date id="FF912AFC4463B51F7EC566A365F3FE44" box="[932,988,474,500]" pageId="4" pageNumber="57">
<emphasis id="B95BD02E4463B51F7EC566A36580FE44" bold="true" box="[932,943,474,500]" pageId="4" pageNumber="57">I</emphasis>
1–2
</date>
-
<date id="FF912AFC4463B51F7E8066A3620CFE44" box="[993,1059,474,500]" pageId="4" pageNumber="57">
<emphasis id="B95BD02E4463B51F7E8066A365D7FE44" bold="true" box="[993,1016,474,500]" pageId="4" pageNumber="57">II</emphasis>
1–3
</date>
-
<date id="FF912AFC4463B51F794866A36258FE44" box="[1065,1143,474,500]" pageId="4" pageNumber="57">
<emphasis id="B95BD02E4463B51F794866A36264FE44" bold="true" box="[1065,1099,474,500]" pageId="4" pageNumber="57">III</emphasis>
2–3
</date>
+
<emphasis id="B95BD02E4463B51F79E166A3628EFE44" bold="true" box="[1152,1185,474,500]" pageId="4" pageNumber="57">IV</emphasis>
3.5–
<date id="FF912AFC4463B51F79B466A362D4FE44" box="[1237,1275,474,500]" pageId="4" pageNumber="57">
2
<emphasis id="B95BD02E4463B51F798266A362DBFE44" bold="true" box="[1251,1268,474,500]" pageId="4" pageNumber="57">V</emphasis>
.
</date>
Supernumerary plantar tubercles weakly evident. Metatarsal inner tubercle oval, elongated, 1.5 times size of rounded outer. Metatarsal fold present, extending from metatarsus to near tip of toe V.
<taxonomicName id="4C2F77BF4463B51F793D655263B3FDF4" box="[1116,1436,555,580]" class="Bivalvia" family="Mytilidae" genus="Musculus" kingdom="Animalia" order="Mytiloida" pageId="4" pageNumber="57" phylum="Mollusca" rank="subSpecies" species="adductor" subSpecies="longus">
<emphasis id="B95BD02E4463B51F793D655263B3FDF4" box="[1116,1436,555,580]" italics="true" pageId="4" pageNumber="57">Musculus adductor longus</emphasis>
</taxonomicName>
absent.
</paragraph>
<paragraph id="8B900C3C4463B51F7DAF65036211FD54" blockId="4.[151,1437,152,740]" pageId="4" pageNumber="57">Tongue oval, not notched posteriorly, 2/3 of its extension adherent to floor of mouth. Choanae rounded, not concealed by palatal shelf of maxillary arch. Cloacal opening at upper level of thighs. Supracloacal region with two rounded or prominent tubercles and some small tubercles on the sides.</paragraph>
<caption id="DF505CB44463B51F7DF6606F63AEF89E" box="[151,1409,1814,1838]" httpUri="https://zenodo.org/record/179981/files/figure.png" pageId="4" pageNumber="57" targetBox="[451,1147,805,1742]" targetPageId="4">
<paragraph id="8B900C3C4463B51F7DF6606F63AEF89E" blockId="4.[151,1409,1814,1838]" box="[151,1409,1814,1838]" pageId="4" pageNumber="57">
<emphasis id="B95BD02E4463B51F7DF6606F6736F89E" bold="true" box="[151,281,1814,1838]" pageId="4" pageNumber="57">FIGURE 4.</emphasis>
Dorsal view (
<emphasis id="B95BD02E4463B51F7CD0606F67EBF89E" bold="true" box="[433,452,1814,1838]" pageId="4" pageNumber="57">A</emphasis>
) and lateral profile (
<emphasis id="B95BD02E4463B51F7FC0606F649CF89E" bold="true" box="[673,691,1814,1838]" pageId="4" pageNumber="57">B</emphasis>
) of head of holotype of
<taxonomicName id="4C2F77BF4463B51F7ED4606E6261F89E" box="[949,1102,1815,1838]" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="lynchi" status="sp. nov.">
<emphasis id="B95BD02E4463B51F7ED4606E6261F89E" box="[949,1102,1815,1838]" italics="true" pageId="4" pageNumber="57">Rhaebo lynchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2686D554463B51F7934606F6284F89E" box="[1109,1195,1814,1838]" pageId="4" pageNumber="57" rank="species">
<emphasis id="B95BD02E4463B51F7934606F6284F89E" bold="true" box="[1109,1195,1814,1838]" pageId="4" pageNumber="57">sp. nov.</emphasis>
</taxonomicNameLabel>
Scales equal 2 mm.
</paragraph>
</caption>
<paragraph id="8B900C3C4463B51F7DA660F065A2F844" blockId="4.[151,1436,1929,2036]" pageId="4" pageNumber="57">
<emphasis id="B95BD02E4463B51F7DA660F06480F813" bold="true" box="[199,687,1929,1955]" pageId="4" pageNumber="57">Color in preservative (Ethanol 70%).</emphasis>
Specimen bicolored, dorsally pale brown; sacral warts dark brown (
<figureCitation id="131410B94463B51F7D9360CB677CF87C" box="[242,339,1970,1996]" captionStart="FIGURE 1" captionStartId="1.[151,255,1647,1671]" captionTargetBox="[174,1415,759,1614]" captionTargetId="figure@1.[164,1422,751,1620]" captionTargetPageId="1" captionText="FIGURE 1. Rhaebo lynchi sp. nov.: dorsal (A) and ventral (B) views of holotype, ICN 1161, adult female SVL 37.5 mm. Photos by Jorge Contreras." httpUri="https://zenodo.org/record/179978/files/figure.png" pageId="4" pageNumber="57">Figure 1</figureCitation>
A); flanks dark brown, separated from the dorsum by a delicate cream line (
<figureCitation id="131410B94463B51F79B760CB6312F87C" box="[1238,1341,1970,1996]" captionStart="FIGURE 4" captionStartId="4.[151,255,1814,1838]" captionTargetBox="[451,1147,805,1742]" captionTargetId="figure@4.[385,1202,774,1793]" captionTargetPageId="4" captionText="FIGURE 4. Dorsal view (A) and lateral profile (B) of head of holotype of Rhaebo lynchi sp. nov. Scales equal 2 mm." httpUri="https://zenodo.org/record/179981/files/figure.png" pageId="4" pageNumber="57">Figure 4</figureCitation>
B); ventrally pale brown with reticulated dark brown stains (
<figureCitation id="131410B94463B51F7E6560A36549F844" box="[772,870,2010,2036]" captionStart="FIGURE 1" captionStartId="1.[151,255,1647,1671]" captionTargetBox="[174,1415,759,1614]" captionTargetId="figure@1.[164,1422,751,1620]" captionTargetPageId="1" captionText="FIGURE 1. Rhaebo lynchi sp. nov.: dorsal (A) and ventral (B) views of holotype, ICN 1161, adult female SVL 37.5 mm. Photos by Jorge Contreras." httpUri="https://zenodo.org/record/179978/files/figure.png" pageId="4" pageNumber="57">Figure 1</figureCitation>
B).
</paragraph>
<paragraph id="8B900C3C4462B51E7DA667EE64C3FE9A" blockId="5.[151,1437,151,498]" pageId="5" pageNumber="58">
<emphasis id="B95BD02E4462B51E7DA667EE64FAFF01" bold="true" box="[199,725,151,177]" pageId="5" pageNumber="58">
Measurements in millimeters of
<typeStatus id="5494B29E4462B51E7F3D67EE64FFFF01" box="[604,720,151,177]" pageId="5" pageNumber="58" type="holotype">Holotype</typeStatus>
.
</emphasis>
SVL 37.5; HW 12.5; head length 12.7; IOD 5.9; upper eyelid width 3.8; internarial distance 3.9; parotoid length 9.7; parotoid width 3.9; inter-parotoid distance 11.3; eye-parotoid distance 1.8; eye diameter 3.3; tympanum 2.1; eye-nare distance 3.0; nare-snout distance 2.2; tibia length 12.1; foot length 13.4; hand length 9.2.
</paragraph>
<paragraph id="8B900C3C4462B51E7DA6664E6374FECA" blockId="5.[151,1437,151,498]" pageId="5" pageNumber="58">
<emphasis id="B95BD02E4462B51E7DA6664E67A0FEE1" bold="true" box="[199,399,311,337]" pageId="5" pageNumber="58">Natural history.</emphasis>
The
<typeStatus id="5494B29E4462B51E7CAE6641641AFEE2" box="[463,565,312,338]" pageId="5" pageNumber="58" type="holotype">holotype</typeStatus>
was captured in primary forest, under logs during a rainy season (Fieldnotes of the amphibian collection of ICN). The female has a mass of pale eggs and convoluted oviducts.
</paragraph>
</subSubSection>
<subSubSection id="C3355FB74462B51E7DA666FE63B6FE42" pageId="5" pageNumber="58" type="etymology">
<paragraph id="8B900C3C4462B51E7DA666FE63B6FE42" blockId="5.[151,1437,151,498]" pageId="5" pageNumber="58">
<emphasis id="B95BD02E4462B51E7DA666FE677BFE11" bold="true" box="[199,340,391,417]" pageId="5" pageNumber="58">Etymology.</emphasis>
This toad is dedicated to my professor Dr. John Douglas Lynch, in acknowledgment for his great help to my formation, for his unconditional support during my residence in the Amphibian Laboratory of ICN, and in recognition and admiration to his enormous contributions to the study of Colombian amphibians.
</paragraph>
</subSubSection>
</treatment>
</document>
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