140 lines
17 KiB
XML
140 lines
17 KiB
XML
<document id="4A9CC5C7E8474C7324C316337FD6F14C" ID-DOI="10.5281/zenodo.12626496" ID-ISSN="2064-2474" ID-Zenodo-Dep="12626496" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638059390968" checkinUser="felipe" docAuthor="Papp, LÁsZLÓ" docDate="2014" docId="2D64A424FFD5FFA8C369FA9FCA50BC5B" docLanguage="en" docName="ActaZoolAcadSciH.60.2.109-155.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 60 (2)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Ceroptera MACQUART 1835" docType="treatment" docVersion="2" lastPageNumber="111" masterDocId="D15DDC5CFFD4FFAAC13CFFF1C943B961" masterDocTitle="A Review Of The Old World Species Of Ceroptera Macquart, 1835 (Diptera, Sphaeroceridae)" masterLastPageNumber="155" masterPageNumber="109" pageNumber="110" updateTime="1719930603981" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
|
||
<mods:mods id="607F8FE243DAAE5953DF56DDA12DADD9" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="4A8D3235F9FFDCDDBE8D0FB42EA84945">
|
||
<mods:title id="D4AE7BDB588F2720701F84FE6978105B">A Review Of The Old World Species Of Ceroptera Macquart, 1835 (Diptera, Sphaeroceridae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="7E3A87D9B30731E8DF1BDA4E5424F72F" type="personal">
|
||
<mods:role id="BEC8C752EC2ABA3CA829C2F342E7DC21">
|
||
<mods:roleTerm id="D4C7A7815891F1089CD88337561FD4FF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B86068DA6D6D35BBA6B19360F52580CB">Papp, LÁsZLÓ</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="DB684DD7070A0905B1A19AA9FC535FED">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F55CC16832F34C278AC045C1352BE3E3" type="host">
|
||
<mods:titleInfo id="38B1760051BD446D62E5B171DBB93EAD">
|
||
<mods:title id="F4F6C7B914CE65D46EA8BF7068C6F98F">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="D787DD0B54BA9AF3EFAAB0A5946D01DD">
|
||
<mods:date id="B2DF1B0BFCF69B9ACD56CD3C891729F4">2014</mods:date>
|
||
<mods:detail id="F2B783375208E6CFC56FB8628B10139B" type="pubDate">
|
||
<mods:number id="A9897A31A776200F270F4A91ED19184F">2014-04-04</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="722D480B3310E308FCFF72A406B80B5F" type="volume">
|
||
<mods:number id="36915BEE0B609C288687AB3F052FA0C0">60</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4AD7FE44875697B6E016072DDC56DAFF" type="issue">
|
||
<mods:number id="3250A5B425824459C767B4C750B421CB">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="D2229495D96D993878FF1D8510AD89E7" unit="page">
|
||
<mods:start id="CB8FBDA81DE1AB21BF317449168D668C">109</mods:start>
|
||
<mods:end id="3625303B7337BF5A6D019A0C19C67389">155</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="F85E750600F9960FA989BDF9A9BFC6A2">journal article</mods:classification>
|
||
<mods:identifier id="8B675667EA663979A2C3031DA8E93E02" type="DOI">10.5281/zenodo.12626496</mods:identifier>
|
||
<mods:identifier id="0D1487CF85BD5A3B78E0C334431050C6" type="ISSN">2064-2474</mods:identifier>
|
||
<mods:identifier id="214D817B0E3E248D7582BB4B261AFDEA" type="Zenodo-Dep">12626496</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="2D64A424FFD5FFA8C369FA9FCA50BC5B" LSID="urn:lsid:plazi:treatment:2D64A424FFD5FFA8C369FA9FCA50BC5B" httpUri="http://treatment.plazi.org/id/2D64A424FFD5FFA8C369FA9FCA50BC5B" lastPageId="2" lastPageNumber="111" pageId="1" pageNumber="110">
|
||
<subSubSection id="EDD746B9FFD5FFABC369FA9FCD47BCE9" box="[597,1028,1389,1416]" pageId="1" pageNumber="110" type="nomenclature">
|
||
<paragraph id="A5721532FFD5FFABC369FA9FCD47BCE9" blockId="1.[597,1028,1389,1416]" box="[597,1028,1389,1416]" pageId="1" pageNumber="110">
|
||
<heading id="FE3AA25EFFD5FFABC369FA9FCD47BCE9" box="[597,1028,1389,1416]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="110" reason="2">
|
||
<taxonomicName id="62CD6EB1FFD5FFABC369FA9FCD47BCE9" ID-CoL="3L7Y" authority="MACQUART, 1835" authorityName="MACQUART" authorityYear="1835" box="[597,1028,1389,1416]" class="Insecta" family="Sphaeroceridae" genus="Ceroptera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="97B9C920FFD5FFABC369FA9FCA40BCE9" box="[597,771,1390,1416]" italics="true" pageId="1" pageNumber="110">CEROPTERA</emphasis>
|
||
MACQUART, 1835
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EDD746B9FFD5FFABC06CFA57CBCDBCDA" box="[336,654,1445,1467]" pageId="1" pageNumber="110" type="reference_group">
|
||
<paragraph id="A5721532FFD5FFABC06CFA57CBCDBCDA" blockId="1.[336,895,1445,1496]" box="[336,654,1445,1467]" pageId="1" pageNumber="110">
|
||
<taxonomicName id="62CD6EB1FFD5FFABC06CFA57CBC9BCDA" ID-CoL="3L7Y" authority="Macquart, 1835: 564" authorityName="MACQUART" authorityPageNumber="564" authorityYear="1835" box="[336,650,1445,1467]" class="Insecta" family="Sphaeroceridae" genus="Ceroptera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="97B9C920FFD5FFABC06CFA57C8EDBCDA" box="[336,430,1446,1467]" italics="true" pageId="1" pageNumber="110">Ceroptera</emphasis>
|
||
Macquart, 1835: 564
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EDD746B9FFD5FFA8C06CFA32CA50BC5B" lastPageId="2" lastPageNumber="111" pageId="1" pageNumber="110" type="materials_examined">
|
||
<paragraph id="A5721532FFD5FFABC06CFA32CA3CBCB9" blockId="1.[336,895,1445,1496]" box="[336,895,1474,1496]" pageId="1" pageNumber="110">
|
||
<typeStatus id="7A76AB90FFD5FFABC06CFA32C8C7BCB9" box="[336,388,1475,1496]" pageId="1" pageNumber="110">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="62CD6EB1FFD5FFABC0DEFA33CA6FBCB6" authority="Meigen, 1830" authorityName="Meigen" authorityYear="1830" box="[482,812,1474,1496]" class="Insecta" family="Sphaeroceridae" genus="Borborus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="rufitarsis">
|
||
<emphasis id="97B9C920FFD5FFABC0DEFA33CBD9BCB6" box="[482,666,1474,1495]" italics="true" pageId="1" pageNumber="110">Borborus rufitarsis</emphasis>
|
||
Meigen, 1830
|
||
</taxonomicName>
|
||
(mon.).
|
||
</paragraph>
|
||
<paragraph id="A5721532FFD5FFABC0B1FA0CC8CFBF58" blockId="1.[336,1289,1533,1761]" pageId="1" pageNumber="110">
|
||
The genus was re-described by
|
||
<bibRefCitation id="C15C68C3FFD5FFABC22DFA0CCAD7BF76" author="PAPP" box="[785,916,1533,1559]" pageId="1" pageNumber="110" pagination="371 - 385" refId="ref15875" refString="PAPP, L. (1977) A contribution to the knowledge of species of the subfamily Ceropterinae (Diptera: Sphaeroceridae). ActaZoologicaAcademiaeScientiarumHungaricae 23: 371 - 385." type="journal article" year="1977">PAPP (1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="C15C68C3FFD5FFABC2A3FA0CCA9FBF76" author="PAPP" box="[927,988,1533,1559]" pageId="1" pageNumber="110" pagination="47 - 209" refId="ref15978" refString="PAPP, L. (2008) NewgeneraoftheOldWorldLimosininae (Diptera, Sphaeroceridae). Acta Zoologica Academiae Scientiarum Hungaricae 54 (Suppl.): 47 - 209." type="journal article" year="2008">2008</bibRefCitation>
|
||
). Here I give a diagnosis only.
|
||
</paragraph>
|
||
<paragraph id="A5721532FFD5FFA8C0B1F9B0CBC9BDD5" blockId="1.[336,1289,1533,1761]" lastBlockId="2.[298,1251,405,1339]" lastPageId="2" lastPageNumber="111" pageId="1" pageNumber="110">
|
||
Headcomparativelylargeandlongwithnumerousinterfrontalpairsof setae. Lunulesmall, withoutsetae (inoppositionto
|
||
<taxonomicName id="62CD6EB1FFD5FFABC2F2F993CD15BF1D" authorityName="Richards" authorityYear="1953" box="[974,1110,1634,1660]" class="Insecta" family="Sphaeroceridae" genus="Ceropterella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="97B9C920FFD5FFABC2F2F993CD15BF1D" box="[974,1110,1634,1660]" italics="true" pageId="1" pageNumber="110">Ceropterella</emphasis>
|
||
</taxonomicName>
|
||
). Orbitalsetulaecontinuedventrallyin 1 ormorerowsonparafacialia, aswellasmostlyalso subocularsetulaepresent (Fig. 98). Thenumberofdorsocentralsetaedefine speciesgroups: therearealsopresuturaldorsocentrals (
|
||
<taxonomicName id="62CD6EB1FFD5FFABC2C1F936CDC6BF80" authorityName="Meigen" authorityYear="1830" box="[1021,1157,1735,1761]" class="Insecta" family="Sphaeroceridae" genus="Borborus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="rufitarsis">
|
||
<emphasis id="97B9C920FFD5FFABC2C1F936CDC6BF80" box="[1021,1157,1735,1761]" italics="true" pageId="1" pageNumber="110">C. rufitarsis</emphasis>
|
||
</taxonomicName>
|
||
group), or there are 0 + 2 large pairs (
|
||
<taxonomicName id="62CD6EB1FFD6FFA8C352FE67CB91B8CE" baseAuthorityName="Duda" baseAuthorityYear="1925" box="[622,722,406,432]" class="Insecta" family="Sphaeroceridae" genus="Ceroptera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="crispa">
|
||
<emphasis id="97B9C920FFD6FFA8C352FE67CB91B8CE" box="[622,722,406,432]" italics="true" pageId="2" pageNumber="111">C. crispa</emphasis>
|
||
</taxonomicName>
|
||
group), or, there is only 1 prescutellar dorsocentralpair (
|
||
<emphasis id="97B9C920FFD6FFA8C0D9FE46CB9BB8B0" box="[485,728,439,465]" italics="true" pageId="2" pageNumber="111">
|
||
<taxonomicName id="62CD6EB1FFD6FFA8C0D9FE46CB1BB8B1" box="[485,600,439,465]" class="Insecta" family="Sphaeroceridae" genus="Ceroptera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="armata">C. armata</taxonomicName>
|
||
,
|
||
<taxonomicName id="62CD6EB1FFD6FFA8C358FE46CB9BB8B0" box="[612,728,439,465]" class="Insecta" family="Sphaeroceridae" genus="Ceroptera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="globosa">C. globosa</taxonomicName>
|
||
</emphasis>
|
||
, etc.). Theprescutellarpairofacrostichals isusuallystrong. 1 or 2 pairsofkatepisternals. Thereisnogeneralfeaturein wing venation: veins
|
||
<collectionCode id="C3DC8DF7FFD6FFA8C30FFE0BCB05BB75" box="[563,582,506,532]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="111">R</collectionCode>
|
||
<subScript id="39491777FFD6FFA8C379FDFDCB1DBB7B" attach="left" box="[581,606,524,538]" fontSize="6" pageId="2" pageNumber="111">2+3</subScript>
|
||
and
|
||
<collectionCode id="C3DC8DF7FFD6FFA8C3A7FE0BCBEDBB75" box="[667,686,506,532]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="111">R</collectionCode>
|
||
<subScript id="39491777FFD6FFA8C392FDFDCB85BB7B" attach="left" box="[686,710,524,538]" fontSize="6" pageId="2" pageNumber="111">4+5</subScript>
|
||
are long (
|
||
<figureCitation id="3DF609B7FFD6FFA8C27DFE0BCAD5BB75" box="[833,918,506,532]" captionStart="Fig" captionStartId="40.[468,504,929,951]" captionTargetBox="[369,1181,405,911]" captionTargetId="figure-38@40.[369,1182,405,912]" captionTargetPageId="40" captionText="Fig. 97. Habitus of Ceroptera crispa (Duda), lectotype male." figureDoi="http://doi.org/10.5281/zenodo.12626520" httpUri="https://zenodo.org/record/12626520/files/figure.png" pageId="2" pageNumber="111">Fig. 97</figureCitation>
|
||
), or shorter and upcurving (
|
||
<figureCitation id="3DF609B7FFD6FFA8C00FFDEAC8CFBB57" box="[307,396,539,566]" captionStart="Fig" captionStartId="40.[761,797,1666,1688]" captionTargetBox="[761,1249,982,1638]" captionTargetId="figure-61@40.[761,1250,982,1638]" captionTargetPageId="40" captionText="Fig. 99. Habitus of Ceroptera nigra sp. n. para- type male." figureDoi="http://doi.org/10.5281/zenodo.12626524" httpUri="https://zenodo.org/record/12626524/files/figure.png" pageId="2" pageNumber="111">Fig. 99</figureCitation>
|
||
). Ahighvarietyisalsopresentinthearmatureoflegs, particularly so for the mid tibia. There are species with numerous setae on mid tibia (
|
||
<taxonomicName id="62CD6EB1FFD6FFA8C5F5FDCFC82DBB19" baseAuthorityName="Duda" baseAuthorityYear="1925" class="Insecta" family="Sphaeroceridae" genus="Ceroptera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="crispa">
|
||
<emphasis id="97B9C920FFD6FFA8C5F5FDCFC82DBB19" italics="true" pageId="2" pageNumber="111">C. crispa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="3DF609B7FFD6FFA8C040FDAEC892BB18" box="[380,465,607,633]" captionStart="Fig" captionStartId="40.[468,504,929,951]" captionTargetBox="[369,1181,405,911]" captionTargetId="figure-38@40.[369,1182,405,912]" captionTargetPageId="40" captionText="Fig. 97. Habitus of Ceroptera crispa (Duda), lectotype male." figureDoi="http://doi.org/10.5281/zenodo.12626520" httpUri="https://zenodo.org/record/12626520/files/figure.png" pageId="2" pageNumber="111">Fig. 97</figureCitation>
|
||
), or there is only an anterodorsal seta at 3/4 and a strong dorsal setaatc. 4/5 (e.g.
|
||
<taxonomicName id="62CD6EB1FFD6FFA8C33EFD70CB3BBBFB" box="[514,632,641,667]" class="Insecta" family="Sphaeroceridae" genus="Ceroptera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="inermis">
|
||
<emphasis id="97B9C920FFD6FFA8C33EFD70CB3BBBFB" box="[514,632,641,667]" italics="true" pageId="2" pageNumber="111">C. inermis</emphasis>
|
||
</taxonomicName>
|
||
). Clawsusuallylargebuttherearespecies, where claws are reduced to thin setae (
|
||
<figureCitation id="3DF609B7FFD6FFA8C381FD53CA52BBDC" box="[701,785,674,701]" captionStart="Figs 64–70" captionStartId="29.[336,381,1637,1659]" captionTargetBox="[339,1286,549,1601]" captionTargetId="figure-78@29.[336,1288,545,1602]" captionTargetPageId="29" captionText="Figs 64–70. Ceroptera inermis sp. n., male: 64 = hind 5th tarsomere, ventral view, 65 = sternite 5, ventralview, 66 = cerciandsurstyliwithsubepandrialsclerite (covered), caudalview, 67 = hypandrium, ventral view, 68 = postgonite with medial part of hypandrium, lateral view, 69 = leftsurstylus, lateralview, 70 = phallusandphallapodeme, lateralview (e: empodium," figureDoi="http://doi.org/10.5281/zenodo.12626514" httpUri="https://zenodo.org/record/12626514/files/figure.png" pageId="2" pageNumber="111">Fig. 64</figureCitation>
|
||
) and the function of claws was taken overbythelateralapicalprojectionsoftheterminaltarsomere. Abdominal tergamostlyreducednotreachinglateralmarginofabdomen. Sternitesare comparativelylargeinmostofthespecies. Sternite 5 withoutappendages butwithnumerousfineandbluntorthickandpointedsetaecaudally. Epandrium more or less asymmetrical. There are less generalities in the form of the surstylus, than it was thought before (cf.
|
||
<bibRefCitation id="C15C68C3FFD6FFA8C213FC9DCA95BAE7" author="PAPP" box="[815,982,875,902]" pageId="2" pageNumber="111" pagination="47 - 209" refId="ref15978" refString="PAPP, L. (2008) NewgeneraoftheOldWorldLimosininae (Diptera, Sphaeroceridae). Acta Zoologica Academiae Scientiarum Hungaricae 54 (Suppl.): 47 - 209." type="journal article" year="2008">PAPP 2008: 51</bibRefCitation>
|
||
). I am quite sure that the groundplan of the
|
||
<taxonomicName id="62CD6EB1FFD6FFA8C377FC7FCBFABAC9" authorityName="MACQUART" authorityYear="1835" box="[587,697,910,936]" class="Insecta" family="Sphaeroceridae" genus="Ceroptera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="97B9C920FFD6FFA8C377FC7FCBFABAC9" box="[587,697,910,936]" italics="true" pageId="2" pageNumber="111">Ceroptera</emphasis>
|
||
</taxonomicName>
|
||
male genitalia is with two lobes, which are connected on their base only (see that of
|
||
<taxonomicName id="62CD6EB1FFD6FFA8C209FC5ECA82BAA8" authorityName="Meigen" authorityYear="1830" box="[821,961,943,969]" class="Insecta" family="Sphaeroceridae" genus="Borborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="rufitarsis">
|
||
<emphasis id="97B9C920FFD6FFA8C209FC5ECA82BAA8" box="[821,961,943,969]" italics="true" pageId="2" pageNumber="111">C. rufitarsis</emphasis>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="7A76AB90FFD6FFA8C53EFC5ECD7BBAA8" box="[1026,1080,943,969]" pageId="2" pageNumber="111">type</typeStatus>
|
||
species). The twolobesmaybefusedtovariousextent, andasamostapomorphousstate, thesurstylusisaslendersinglescleritewithacompletereductionoftheother lobe. Distiphallusisusuallyslender (Ithinkitofasynapomorphy), basiphal- lusiswithadistincttoverylargeepiphallus, or, withoutanyepiphallus. Femalepostabdomenusuallyshort, always 3 spermathecaebuttheirshapeis variablewithinthegenus. ActuallyIhavenotfoundmoregeneraldiagnostic featuresinfemaleabdomen.
|
||
</paragraph>
|
||
<paragraph id="A5721532FFD6FFA8C05BFB4DCA50BC5B" blockId="2.[298,1251,405,1339]" pageId="2" pageNumber="111">
|
||
Incontrasttothehighvariabilityinmorphology, thelife-habitsofthe
|
||
<taxonomicName id="62CD6EB1FFD6FFA8C016FB2FC8DBBD99" authorityName="MACQUART" authorityYear="1835" box="[298,408,1246,1272]" class="Insecta" family="Sphaeroceridae" genus="Ceroptera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="97B9C920FFD6FFA8C016FB2FC8DBBD99" box="[298,408,1246,1272]" italics="true" pageId="2" pageNumber="111">Ceroptera</emphasis>
|
||
</taxonomicName>
|
||
speciesaresimilartoeachother: alltheyarecommensalistsofscara- baeid beetles; the female lays eggs into the dung balls of the beetles, so the larvaedevelopprotectedunderground.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |