119 lines
14 KiB
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119 lines
14 KiB
XML
<document id="D1FF51CC08245B922424AD45413AA96B" ID-DOI="10.11646/zootaxa.2186.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5311592" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629629438740" checkinUser="felipe" docAuthor="Jones, Joshua R. & Deitz, Lewis L." docDate="2009" docId="CF7A87E4FFCA894F7D9DA1B8B936FDAD" docLanguage="en" docName="zt02186p120.pdf" docOrigin="Zootaxa 2186 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2186.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hespenedrini Jones 2009, new tribe" docType="treatment" docVersion="3" lastPageNumber="30" masterDocId="3343FF9CFFD689527D0AA225BA23FC37" masterDocTitle="Phylogeny and systematics of the leafhopper subfamily Ledrinae (Hemiptera: Cicadellidae) 2186" masterLastPageNumber="120" masterPageNumber="1" pageNumber="29" updateTime="1699086546085" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0DC557FD1114F1617C662E227FF14B6D">Phylogeny and systematics of the leafhopper subfamily Ledrinae (Hemiptera: Cicadellidae) 2186</mods:title>
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<mods:namePart id="829FED0DB1C38E0FE17E26AA5391DD71">Jones, Joshua R.</mods:namePart>
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<mods:namePart id="7A2E6208090234BDB31B6800387D64EA">Deitz, Lewis L.</mods:namePart>
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<mods:title id="C3B02D64E4E2A47F033C47E1E65900C6">Zootaxa</mods:title>
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<mods:part id="323B9E11EC4EDB2D1876412239F3EF00">
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<mods:date id="5BB8EE032558E622E5525871D894C691">2009</mods:date>
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<mods:number id="16196415B366E4848CFC3D45E6EF5039">2009-08-06</mods:number>
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<mods:number id="45B66887692A68D487275DC9C06F2205">2186</mods:number>
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<mods:url id="31DAB95AD6DC1C3C8E43C2913EDF0094">https://biotaxa.org/Zootaxa/article/view/zootaxa.2186.1.1</mods:url>
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<mods:classification id="A9113842019BDCAF1BC33CD98E3C6BB6">journal article</mods:classification>
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<mods:identifier id="243C8597D690102A6A6B6010D7315B66" type="DOI">10.11646/zootaxa.2186.1.1</mods:identifier>
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<mods:identifier id="0E75BAF9CC23F0D525112384D986333B" type="ISSN">11755334</mods:identifier>
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<subSubSection id="0FC96579FFCA894E7D9DA1B8B878FF80" box="[151,603,925,951]" pageId="28" pageNumber="29" type="nomenclature">
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<paragraph id="476C36F2FFCA894E7D9DA1B8B878FF80" blockId="28.[151,603,925,951]" box="[151,603,925,951]" pageId="28" pageNumber="29">
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<emphasis id="75A7EAE0FFCA894E7D9DA1B8B878FF80" bold="true" box="[151,603,925,951]" pageId="28" pageNumber="29">
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Tribe
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<taxonomicName id="80D34D71FFCA894E7DE8A1B8BBFAFF80" ID-CoL="8JK9C" authority="Jones" authorityName="Jones" authorityYear="2009" box="[226,473,925,951]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Hespenedrini">Hespenedrini Jones</taxonomicName>
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,
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<taxonomicNameLabel id="6E94579BFFCA894E7CEFA1B8B878FF80" box="[485,603,925,951]" pageId="28" pageNumber="29" rank="tribe">new tribe</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0FC96579FFCA894E7D9DA1C2B871F837" box="[151,594,999,1024]" pageId="28" pageNumber="29" type="materials_examined">
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<paragraph id="476C36F2FFCA894E7D9DA1C2B871F837" blockId="28.[151,594,999,1024]" box="[151,594,999,1024]" pageId="28" pageNumber="29">
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<emphasis id="75A7EAE0FFCA894E7D9DA1C2B871F837" box="[151,594,999,1024]" italics="true" pageId="28" pageNumber="29">
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<typeStatus id="98688850FFCA894E7D9DA1C2BAEDF837" box="[151,206,999,1024]" pageId="28" pageNumber="29">Type</typeStatus>
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genus
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<taxonomicName id="80D34D71FFCA894E7C2BA1C2B871F837" authority="Kramer 1966" authorityName="Kramer" authorityYear="1966" box="[289,594,999,1024]" class="Insecta" family="Cicadellidae" genus="Hespenedra" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
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Hespenedra
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<bibRefCitation id="23424B03FFCA894E7CBFA1C2B871F837" author="Kramer, J. P." box="[437,594,999,1024]" pageId="28" pageNumber="29" pagination="469 - 502" refId="ref57713" refString="Kramer, J. P. (1966) A revision of the New World leafhoppers of the subfamily Ledrinae (Homoptera: Cicadellidae). Transactions of the American Entomological Society, 92, 469 - 502." type="journal article" year="1966">Kramer 1966</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="0FC96579FFCA894F7DCDA616BB94FCEF" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
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<paragraph id="476C36F2FFCA894F7DCDA616B8BCFC85" blockId="28.[151,1436,1075,2030]" lastBlockId="29.[151,1436,152,410]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
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<emphasis id="75A7EAE0FFCA894E7DCDA616BB7CF87A" bold="true" box="[199,351,1075,1101]" pageId="28" pageNumber="29">Description.</emphasis>
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Moderately large leafhoppers. Dorsum texture acinose-colliculate, rugose, punctate. Head spatulate, more or less lamellate toward anterolateral margins; crown parabolic, longer in females than in males, flat; longitudinal medial carina complete; marginal carina complete or incomplete, sometimes converging into crown texture laterally; ocelli closer to each other than to eyes, equidistant from hind and anterior margins in male, distance to anterior margin in female greater; surface with depressions and pits, more so in females; several deep pits arranged in a row, or two rows in the female, running from ocellus to point where longitudinal carina meets crown apex; anterior margin of pronotum slightly overlapping posterior margin of crown, straight, except laterally near the eyes, where it angles posterolaterally; pronotum essentially flat, with some depressions; mesoscutum and scutellum flat; forewings punctate, setose, sharply angled at second claval vein; claval area punctate, flat, and in same plane with mesoscutum and scutellum; remaining portion of wings angled downward; claval suture strongly depressed; veins raised, reticulate; face flattened, somewhat convex; frontoclypeus narrow, long, with apical extension; base with cibarial muscle attachment ridges visible laterally; row of pits running adjacent to the frontoclypeus from antennal bases to point where longitudinal medial carina on frontoclypeus meets crown apex, corresponding to pits on dorsal surface of crown; lora somewhat tumid, slightly angulate, genae narrow; proepisternum large, quadrate, situated proximate to lateral edge of pronotum; lateral carina of pronotum distinct; femora round, short, somewhat robust, with setae reduced; mesothoracic femur row II apex with triangular patch of scalelike yellowish setae; metathoracic tibia with two short macrosetae mounted on a narrow, prominent base; apical macrosetal formula 2 + 1; pro- and mesotibia intermediately flattened; metathoracic tibia quadrate in cross section, outer surface flat with carinate edges, setal rows reduced, row II with approximately six macrosetae, all bases cucullate; metathoracic tarsomere I intermediately long, plantar surface bearing numerous short stout white setae. Male genitalia: aedeagus short, somewhat broad throughout length; intermediately wide in distal view, with a ventroapical groove; connective long and narrow, medial section thin in dorsal view, flat in lateral view; subgenital plates shorter than pygofer, dorsoventrally flattened basally, laterally compressed distally; anal tube broad in dorsal view. Female genitalia: broadest medially, with two sclerotized dorsal teeth and numerous other less sclerotized dorsal teeth.
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</paragraph>
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<paragraph id="476C36F2FFCB894F7DCDA29BBB94FCEF" blockId="29.[151,1436,152,410]" box="[199,439,190,216]" pageId="29" pageNumber="30">
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<emphasis id="75A7EAE0FFCB894F7DCDA29BBB3DFCEF" bold="true" box="[199,286,190,216]" pageId="29" pageNumber="30">Range.</emphasis>
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Neotropical.
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</paragraph>
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</subSubSection>
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<subSubSection id="0FC96579FFCB894F7DCDA2C0BB91FCC8" box="[199,434,229,255]" pageId="29" pageNumber="30" type="biology_ecology">
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<paragraph id="476C36F2FFCB894F7DCDA2C0BB91FCC8" blockId="29.[151,1436,152,410]" box="[199,434,229,255]" pageId="29" pageNumber="30">
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<emphasis id="75A7EAE0FFCB894F7DCDA2C0BB13FCC8" bold="true" box="[199,304,229,255]" pageId="29" pageNumber="30">Ecology.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="0FC96579FFCB894F7DCDA32EB936FDAD" pageId="29" pageNumber="30" type="discussion">
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<paragraph id="476C36F2FFCB894F7DCDA32EB9C6FD44" blockId="29.[151,1436,152,410]" pageId="29" pageNumber="30">
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<emphasis id="75A7EAE0FFCB894F7DCDA32EBB63FD12" bold="true" box="[199,320,267,293]" pageId="29" pageNumber="30">Remarks.</emphasis>
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Besides
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<taxonomicName id="80D34D71FFCB894F7CA5A329B86EFD11" authorityName="Oman" authorityYear="1949" box="[431,589,268,294]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="tribe" tribe="Xerophloeini">Xerophloeini</taxonomicName>
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,
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<taxonomicName id="80D34D71FFCB894F7F51A329B8D4FD11" authorityName="Jones" authorityYear="2009" box="[603,759,268,294]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="tribe" tribe="Hespenedrini">Hespenedrini</taxonomicName>
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represent the only extant ledrine tribe distributed in the New World.
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<bibRefCitation id="23424B03FFCB894F7C38A317BBC5FD7B" author="Szwedo, J." box="[306,486,306,332]" pageId="29" pageNumber="30" pagination="153 - 163" refId="ref59765" refString="Szwedo, J. (2002) Studies on Xerophloeini leafhoppers with description of Pariacaca icanoensis gen. and sp. nov. from Argentina (Hemiptera: Cicadomorpha: Cicadellidae: Ledrinae). Genus, 13, 153 - 163." type="journal article" year="2002">Szwedo (2002)</bibRefCitation>
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postulated that
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<taxonomicName id="80D34D71FFCB894F7FA2A317B915FD7C" authorityName="Kramer" authorityYear="1966" box="[680,822,306,331]" class="Insecta" family="Cicadellidae" genus="Hespenedra" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="75A7EAE0FFCB894F7FA2A317B915FD7C" box="[680,822,306,331]" italics="true" pageId="29" pageNumber="30">Hespenedra</emphasis>
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</taxonomicName>
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belonged in
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<taxonomicName id="80D34D71FFCB894F7EDCA317BE57FD7B" authorityName="Oman" authorityYear="1949" box="[982,1140,306,332]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="tribe" tribe="Xerophloeini">Xerophloeini</taxonomicName>
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, but in this analysis it is shown to be clearly distinct within
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<taxonomicName id="80D34D71FFCB894F7F38A37CB8B9FD44" authorityName="Fairmaire" authorityYear="1855" box="[562,666,345,371]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subFamily" subFamily="Ledrinae">Ledrinae</taxonomicName>
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and much closer to
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<taxonomicName id="80D34D71FFCB894F7E8DA37CB9C2FD44" authorityName="Evans" authorityYear="1947" box="[903,993,345,371]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="tribe" tribe="Ledrini">Ledrini</taxonomicName>
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.
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</paragraph>
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<paragraph id="476C36F2FFCB894F7DCDA3A5B936FDAD" blockId="29.[151,1436,152,410]" box="[199,789,384,410]" pageId="29" pageNumber="30">
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This tribe is monotypic, and includes
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<taxonomicName id="80D34D71FFCB894F7F8BA3A5B92DFDAE" authorityName="Kramer" authorityYear="1966" box="[641,782,384,409]" class="Insecta" family="Cicadellidae" genus="Hespenedra" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="75A7EAE0FFCB894F7F8BA3A5B92DFDAE" box="[641,782,384,409]" italics="true" pageId="29" pageNumber="30">Hespenedra</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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