475 lines
67 KiB
XML
475 lines
67 KiB
XML
<document id="EA9AFDAB719F02B4F25763584A3FD1D3" ID-DOI="10.5252/geodiversitas2023v45a14" ID-ISSN="1638-9395" ID-Zenodo-Dep="8177407" ID-ZooBank="urn:lsid:zoobank.org:pub:B637BB5A-B41A-42F0-8236-00E040F8DD13" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1690198814183" checkinUser="felipe" docAuthor="Doriath-Döhler, Aimie, Hervet, Sophie & Béthoux, Olivier" docDate="2023" docId="CF4D017120279851FC4A5147FC42F86F" docLanguage="en" docName="Geodiversitas.45.14.401-407.pdf" docOrigin="Geodiversitas 45 (14)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Palaeophylolestes distinctus Doriath-Döhler & Hervet & Béthoux 2023, n. gen., n. sp." docType="treatment" docVersion="3" lastPageNumber="405" masterDocId="3374790920249857FFDA543BFFAFFF84" masterDocTitle="Palaeophylolestes distinctus n. gen., n. sp., a new malachite damselfly (Odonata: Zygoptera: Synlestidae) from the Paleocene Menat locality (France)" masterLastPageNumber="407" masterPageNumber="401" pageNumber="402" updateTime="1690225874833" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="297CB5057799123969DC24D4AA9CC526">Palaeophylolestes distinctus n. gen., n. sp., a new malachite damselfly (Odonata: Zygoptera: Synlestidae) from the Paleocene Menat locality (France)</mods:title>
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<mods:name id="43A2FF2753E58A0F6AA46BA8B9D5EA69" type="personal">
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<mods:namePart id="26554A48BB91AA896786F0D127325CD1">Doriath-Döhler, Aimie</mods:namePart>
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<mods:namePart id="9B31EDAB9554B281D3A19D3810EA85D0">Hervet, Sophie</mods:namePart>
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<mods:name id="90A31E8F9293A97B067AE9E7700118FC" type="personal">
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<mods:roleTerm id="9B672528E8A884136330815F8A251D7D">Author</mods:roleTerm>
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<mods:namePart id="014AA56E9873BFDC92A4A0514FA05BB8">Béthoux, Olivier</mods:namePart>
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<mods:title id="B8266D1C7BD3738FE6B4CCC29D19D8AD">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="54F0F5C8B55372FB024C2380D463B9B1">
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<mods:date id="9911DB37B2AB2E2BC9CCB65BE0B1103A">2023</mods:date>
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<mods:number id="AA4B0DEF47690DFC74F20E0894593AB9">2023-07-20</mods:number>
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<mods:number id="49C33D488AE9C30FAD20F4E213FADC90">45</mods:number>
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<mods:detail id="48B4DCA97275E668BA8C66A837BE5667" type="issue">
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<mods:number id="8EE2D02DE6F60420B7C18E0076027A1C">14</mods:number>
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<mods:identifier id="0E838EB8DA75C3AA52EC900D48E6CB27" type="DOI">10.5252/geodiversitas2023v45a14</mods:identifier>
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<treatment id="CF4D017120279851FC4A5147FC42F86F" LSID="urn:lsid:plazi:treatment:CF4D017120279851FC4A5147FC42F86F" httpUri="http://treatment.plazi.org/id/CF4D017120279851FC4A5147FC42F86F" lastPageId="6" lastPageNumber="405" pageId="3" pageNumber="402">
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<subSubSection id="0FFEE3EC20279854FC4A5147FAE2FA11" box="[912,1357,1403,1430]" pageId="3" pageNumber="402" type="nomenclature">
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<paragraph id="475BB06720279854FC4A5147FAE2FA11" blockId="3.[912,1357,1403,1461]" box="[912,1357,1403,1430]" pageId="3" pageNumber="402">
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<taxonomicName id="80E4CBE420279854FC4A5147FB1CFA12" authority="Doriath-Döhler & Hervet & Béthoux, 2023" authorityName="Doriath-Döhler & Hervet & Béthoux" authorityYear="2023" box="[912,1203,1404,1430]" class="Insecta" family="Synlestidae" genus="Palaeophylolestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="402" phylum="Arthropoda" rank="species" species="distinctus" status="n. gen., n. sp.">
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<emphasis id="75906C7520279854FC4A5147FB1CFA12" bold="true" box="[912,1203,1404,1430]" italics="true" pageId="3" pageNumber="402">Palaeophylolestes distinctus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6EA3D10E20279854FB605140FAE2FA11" box="[1210,1357,1403,1429]" pageId="3" pageNumber="402" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="0FFEE3EC20279854FBED51A1FA00FA6A" pageId="3" pageNumber="402" type="description">
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<paragraph id="475BB06720279854FBED51A1FB0AFA30" blockId="3.[912,1357,1403,1461]" box="[1079,1189,1434,1461]" pageId="3" pageNumber="402">
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(
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<figureCitation id="DFDFACE220279854FB9A51A1FB2FFA30" box="[1088,1152,1434,1460]" captionStart="FIG" captionStart-0="FIG" captionStart-1="FIG" captionStartId="4.[132,143,1592,1609]" captionStartId-0="4.[132,143,1592,1609]" captionStartId-1="5.[132,143,1299,1316]" captionTargetBox="[298,1366,155,1569]" captionTargetBox-0="[298,1366,155,1569]" captionTargetBox-1="[193,1399,215,1257]" captionTargetId="figure-200@4.[296,930,839,1380]" captionTargetId-0="figure-200@4.[296,930,839,1380]" captionTargetId-1="graphics-518@5.[236,1325,500,1043]" captionTargetPageId="4" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText="FIG. 1. — Palaeophylolestes distinctus n. gen., n. sp., holotype specimen (BDL 2002), overview; A, drawing; B, photograph (embedding matrix used as white reference). Scale bar: 5 mm." captionText-0="FIG. 1. — Palaeophylolestes distinctus n. gen., n. sp., holotype specimen (BDL 2002), overview; A, drawing; B, photograph (embedding matrix used as white reference). Scale bar: 5 mm." captionText-1="FIG. 2. — Palaeophylolestes distinctus n. gen., n. sp., holotype specimen (BDL 2002), drawings of wings, isolated (see Fig. 1 for their respective locations; faded vein portions reconstructed); A, B, first pair (located on the left side of the complete specimen); A, forewing; B, hindwing; C, D, second pair (located on the right side of the complete specimen); C, forewing; D, hindwing. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8179515" figureDoi-1="http://doi.org/10.5281/zenodo.8177411" httpUri="https://zenodo.org/record/8179515/files/figure.png" httpUri-1="https://zenodo.org/record/8177411/files/figure.png" pageId="3" pageNumber="402">Figs 1</figureCitation>
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;
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<figureCitation id="DFDFACE220279854FB5451A1FB33FA30" box="[1166,1180,1434,1460]" captionStart="FIG" captionStartId="5.[132,143,1299,1316]" captionTargetBox="[193,1399,215,1257]" captionTargetId="graphics-518@5.[236,1325,500,1043]" captionTargetPageId="5" captionText="FIG. 2. — Palaeophylolestes distinctus n. gen., n. sp., holotype specimen (BDL 2002), drawings of wings, isolated (see Fig. 1 for their respective locations; faded vein portions reconstructed); A, B, first pair (located on the left side of the complete specimen); A, forewing; B, hindwing; C, D, second pair (located on the right side of the complete specimen); C, forewing; D, hindwing. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8177411" httpUri="https://zenodo.org/record/8177411/files/figure.png" pageId="3" pageNumber="402">2</figureCitation>
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)
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</paragraph>
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<paragraph id="475BB06720279854FCF751E6FA00FA6A" blockId="3.[813,1455,1501,1518]" box="[813,1455,1501,1518]" pageId="3" pageNumber="402">
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urn:lsid:zoobank.org:act:
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<uuid id="33428AB220279854FBD051E6FA00FA6A" box="[1034,1455,1501,1518]" pageId="3" pageNumber="402">E1287128-20FA-4391-90E2-B1ABF30C00A6</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="0FFEE3EC20279854FCF7521BFBF5F9D5" pageId="3" pageNumber="402" type="materials_examined">
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<paragraph id="475BB06720279854FCF7521BFBF5F9D5" blockId="3.[813,1455,1567,1618]" pageId="3" pageNumber="402">
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<materialsCitation id="F78CBA3A20279854FCF7521BFBF5F9D5" collectionCode="BDL" country="France" county="Puy-de-Dome" location="Musee de Paleontologie de Menat" municipality="Menat" pageId="3" pageNumber="402" specimenCode="BDL 2002" specimenCount="1" typeStatus="holotype">
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<typeStatus id="985F0EC520279854FCF7521BFC35F9B2" box="[813,922,1568,1591]" pageId="3" pageNumber="402">HOLOTYPE</typeStatus>
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</smallCapsWord>
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. —
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<specimenCode id="1742181C20279854FC11521BFB92F9B3" box="[971,1085,1567,1591]" collectionCode="BDL" pageId="3" pageNumber="402">BDL 2002</specimenCode>
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,
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<location id="423BE6BC20279854FB935224FA06F9B3" LSID="urn:lsid:plazi:treatment:CF4D017120279851FC4A5147FC42F86F:423BE6BC20279854FB935224FA06F9B3" box="[1097,1449,1567,1591]" country="France" county="Puy-de-Dome" municipality="Menat" name="Musee de Paleontologie de Menat" pageId="3" pageNumber="402">Musée de Paléontologie de Menat</location>
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¸
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<collectingMunicipality id="A73F2A1D20279854FCF45201FCDFF9D5" box="[814,880,1594,1617]" pageId="3" pageNumber="402">Menat</collectingMunicipality>
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,
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<collectingCounty id="AE3AC8EB20279854FCA15201FBA7F9D6" box="[891,1032,1594,1618]" pageId="3" pageNumber="402">Puy-de-Dôme</collectingCounty>
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,
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<collectingCountry id="3FF3F0F720279854FBC85201FBF9F9D5" box="[1042,1110,1594,1617]" name="France" pageId="3" pageNumber="402">France</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="0FFEE3EC20279854FCF75250FB04F919" pageId="3" pageNumber="402" type="etymology">
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<paragraph id="475BB06720279854FCF75250FB04F919" blockId="3.[811,1455,1642,1693]" pageId="3" pageNumber="402">
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. — The species epithet refers to the length of MA, MP and CuA, distinctly long in the species.
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</paragraph>
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</subSubSection>
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<subSubSection id="0FFEE3EC20279854FCF7528EFB80F86F" pageId="3" pageNumber="402" type="materials_examined">
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<paragraph id="475BB06720279854FCF7528EFA1CF963" blockId="3.[813,1459,1717,1767]" pageId="3" pageNumber="402">
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<collectionCode id="21F528A220279854FCF7528EFCF2F94F" box="[813,861,1717,1740]" pageId="3" pageNumber="402">TYPE</collectionCode>
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<collectionCode id="21F528A220279854FCBC5281FC6EF94F" box="[870,961,1722,1739]" pageId="3" pageNumber="402">LOCALITY</collectionCode>
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<collectionCode id="21F528A220279854FC135281FC5CF94F" box="[969,1011,1722,1739]" pageId="3" pageNumber="402">AND</collectionCode>
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</smallCapsWord>
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<smallCapsWord id="41BD26BB20279854FC205281FB29F94F" baselines="1735" box="[1018,1158,1722,1739]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="stratigraphy" pageId="3" pageNumber="402">
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<collectionCode id="21F528A220279854FC205281FB29F94F" box="[1018,1158,1722,1739]" pageId="3" pageNumber="402">STRATIGRAPHY</collectionCode>
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</smallCapsWord>
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. — Bord du Lac des Grelins (
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<collectionCode id="21F528A220279854FCEF52F4FCC7F963" box="[821,872,1743,1767]" pageId="3" pageNumber="402">BDL</collectionCode>
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), Menat, Puy-de-Dôme,
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<collectingCountry id="3FF3F0F720279854FB9352EBFB26F963" box="[1097,1161,1744,1767]" name="France" pageId="3" pageNumber="402">France</collectingCountry>
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; middle Paleocene, Menat Basin.
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</paragraph>
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<paragraph id="475BB06720279854FCF1533BFB80F86F" blockId="3.[811,1457,1792,2027]" pageId="3" pageNumber="402">
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<smallCapsWord id="41BD26BB20279854FCF1533BFC1FF892" baselines="1810,1810" box="[811,944,1792,1816]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Commented" pageId="3" pageNumber="402">COMMENTED</smallCapsWord>
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<smallCapsWord id="41BD26BB20279854FC6C533EFBB3F892" baselines="1810" box="[950,1052,1797,1814]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="diagnosis" pageId="3" pageNumber="402">DIAGNOSIS</smallCapsWord>
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. — Wings: RP1+2 / RP3+4 fork closer to nodus than to arculus (within Lestoidea, apomorphic for
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<taxonomicName id="80E4CBE420279854FA815321FCE1F8C9" authorityName="Tillyard" authorityYear="1917" class="Insecta" family="Synlestidae" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="402" phylum="Arthropoda" rank="family">Synlestidae</taxonomicName>
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and
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<taxonomicName id="80E4CBE420279854FC5B530EFC5CF8C9" authorityName="Kennedy" authorityYear="1920" box="[897,1011,1845,1869]" class="Insecta" family="Perilestidae" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="402" phylum="Arthropoda" rank="family">Perilestidae</taxonomicName>
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);
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<collectionCode id="21F528A220279854FBDE530EFB81F8C9" box="[1028,1070,1845,1869]" country="Spain" lsid="urn:lsid:biocol.org:col:15682" name="Real Jardín Botánico" pageId="3" pageNumber="402" type="Herbarium">MA</collectionCode>
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||
,
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<collectionCode id="21F528A220279854FBE3530EFBF1F8C8" box="[1081,1118,1845,1868]" country="0" name="Mohonk Preserve, Inc." pageId="3" pageNumber="402">MP</collectionCode>
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and CuA reaching posterior wing margin very distally (unique to the species); forewing quadrangle with proximal edge about half length of posterior edge; cell below sub-discoidal (i.e. sub-quadrangular) cell well-developed, longer than sub-discoidal cell (plesiomorphy within
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<taxonomicName id="80E4CBE420279854FB6C53A4FAB3F833" box="[1206,1308,1951,1975]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="402" phylum="Tracheophyta" rank="subOrder" subOrder="Zygoptera">Zygoptera</taxonomicName>
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; as opposed to ‘cell reduced, with AA (or CuP+
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<collectionCode id="21F528A220279854FBBA5382FB2CF855" box="[1120,1155,1977,2001]" country="Kazakhstan" lsid="urn:lsid:biocol.org:col:13590" name="Ministry of Science, Academy of Sciences" pageId="3" pageNumber="402" type="Herbarium">AA</collectionCode>
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) diverging from posterior wing margin in distal position’).
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</paragraph>
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||
</subSubSection>
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||
<caption id="139BE0EF20209853FF5E5203FEDEF9E4" ID-DOI="http://doi.org/10.5281/zenodo.8179515" ID-Zenodo-Dep="8179515" httpUri="https://zenodo.org/record/8179515/files/figure.png" pageId="4" pageNumber="403" startId="4.[132,143,1592,1609]" targetBox="[298,1366,155,1569]" targetPageId="4" targetType="figure">
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<paragraph id="475BB06720209853FF5E5203FEDEF9E4" blockId="4.[132,1456,1592,1632]" pageId="4" pageNumber="403">
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<smallCapsWord id="41BD26BB20209853FF5E5203FF32F9CC" baselines="1605,1605" box="[132,157,1592,1609]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="4" pageNumber="403">FIG</smallCapsWord>
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||
. 1. —
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||
<taxonomicName id="80E4CBE420209853FF0D5203FE6CF9CD" authority="Doriath-Döhler & Hervet & Béthoux, 2023" authorityName="Doriath-Döhler & Hervet & Béthoux" authorityYear="2023" box="[215,451,1592,1609]" class="Insecta" family="Synlestidae" genus="Palaeophylolestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="403" phylum="Arthropoda" rank="species" species="distinctus" status="n. gen., n. sp.">
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<emphasis id="75906C7520209853FF0D5203FE6CF9CD" box="[215,451,1592,1609]" italics="true" pageId="4" pageNumber="403">Palaeophylolestes distinctus</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="6EA3D10E20209853FE105203FDECF9CD" box="[458,579,1592,1609]" pageId="4" pageNumber="403" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
, holotype specimen (BDL 2002), overview;
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||
<emphasis id="75906C7520209853FC755203FC13F9CD" bold="true" box="[943,956,1592,1609]" pageId="4" pageNumber="403">A</emphasis>
|
||
, drawing;
|
||
<emphasis id="75906C7520209853FBCF5203FB8DF9CD" bold="true" box="[1045,1058,1592,1609]" pageId="4" pageNumber="403">B</emphasis>
|
||
, photograph (embedding matrix used as white reference). Scale bar: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0FFEE3EC20209851FF5E52A8FC42F86F" lastPageId="6" lastPageNumber="405" pageId="4" pageNumber="403" type="description">
|
||
<paragraph id="475BB06720209853FF5E52A8FEBAF92F" blockId="4.[131,775,1683,1900]" box="[132,277,1683,1709]" pageId="4" pageNumber="403">
|
||
<smallCapsWord id="41BD26BB20209853FF5E52A8FEBAF92F" baselines="1702,1702" box="[132,277,1683,1709]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="4" pageNumber="403">DESCRIPTION</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="475BB06720209853FF5E5289FEB0F948" blockId="4.[131,775,1683,1900]" box="[132,287,1714,1740]" pageId="4" pageNumber="403">
|
||
<emphasis id="75906C7520209853FF5E5289FEB0F948" box="[132,287,1714,1740]" italics="true" pageId="4" pageNumber="403">General aspects</emphasis>
|
||
</paragraph>
|
||
<paragraph id="475BB06720209853FF5E52E9FDDCF8E8" blockId="4.[131,775,1683,1900]" pageId="4" pageNumber="403">Specimen preserving thoracic segments, fragments of a leg, 4 abdominal segments, and more or less complete wings, with pairs of one side largely overlapping; forewing of second pair (FW2) very damaged, creased along multiple folds, with a main disruption running along MA-MP area.</paragraph>
|
||
<paragraph id="475BB06720209853FF5E53A9FDC8F82F" blockId="4.[132,615,1937,1963]" box="[132,615,1937,1963]" pageId="4" pageNumber="403">
|
||
<heading id="1C13070B20209853FF5E53A9FDC8F82F" box="[132,615,1937,1963]" fontSize="11" level="2" pageId="4" pageNumber="403" reason="8">
|
||
<emphasis id="75906C7520209853FF5E53A9FDC8F82F" box="[132,615,1937,1963]" italics="true" pageId="4" pageNumber="403">Wing venation (essentially derived from HW1)</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="475BB06720209852FF5E5389FD77F868" blockId="4.[132,774,1969,2028]" lastBlockId="5.[131,777,1427,2028]" lastPageId="5" lastPageNumber="404" pageId="4" pageNumber="403">
|
||
Two antenodals in ScP-R area; at least 8 postnodals (as preserved in HW1), probably 11 or 12 originally in hindwing, and probably more in forewing; subnodus short, oblique; cross-veins between RA and RP1/RP1+2 aligned with postnodals near nodus, slightly misaligned with them near pterostigma; pterostigma extending over four cells (in the RA-RP1 area); quadrangle trapezoidal (as opposed to quadrangular) with proximal edge about ½ of posterior edge in forewing, less than ½ in hindwing (i.e. quadrangle slightly narrower and longer in hindwing); RP1+2 / RP3+4 fork located closer to nodus than to arculus; Irp
|
||
<subScript id="DB60B22220209853FB9F53A5FBCDF82B" attach="left" box="[1093,1122,1950,1967]" fontSize="7" pageId="4" pageNumber="403">1+2</subScript>
|
||
-rp
|
||
<subScript id="DB60B22220209853FB5853A5FB0FF82B" attach="left" box="[1154,1184,1950,1967]" fontSize="7" pageId="4" pageNumber="403">3+4</subScript>
|
||
starts opposite subnodus; RP1 / RP2 fork located proximally, just before postnodal 4; area between Irp
|
||
<subScript id="DB60B22220209853FBD253E5FBBDF86B" attach="right" box="[1032,1042,2014,2031]" fontSize="7" pageId="4" pageNumber="403">1</subScript>
|
||
-rp
|
||
<subScript id="DB60B22220209853FBE953E5FB92F86B" attach="left" box="[1075,1085,2014,2031]" fontSize="7" pageId="4" pageNumber="403">2</subScript>
|
||
and RP2 with 3 rows of cells at its broadest (probably forming one or two supplementary sectors); RP3+4-MA area broadening near posterior wing margin (broader than surrounding areas), with at least 3 rows of cells; MA very long, reaching posterior wing margin opposite beginning of pterostigma in hindwing (as observed in HW1), possibly longer in forewing (opposite pterostigma mid-length); MA-MP area with a single row of cells except in most distal part (preserved in hindwing, possibly present in forewing);
|
||
<collectionCode id="21F528A220219852FEFB52A8FEE4F929" box="[289,331,1683,1709]" country="0" name="Mohonk Preserve, Inc." pageId="5" pageNumber="404">MP</collectionCode>
|
||
slightly arched upwards at its point of divergence from
|
||
<collectionCode id="21F528A220219852FEED5289FECFF948" box="[311,352,1714,1740]" country="0" name="Mohonk Preserve, Inc." pageId="5" pageNumber="404">MP</collectionCode>
|
||
+CuA, simple and rectilinear, long; MPCuA area comparatively broad (broader than the MA-MP area), with a single row of cells; CuA long, mostly parallel to posterior wing margin; a single row of cells between CuA and posterior wing margin; cell located below sub-discoidal (i.e. sub-quadrangular) cell (i.e. anteriorly delimited by AA) well-developed, longer than sub-discoidal cell; in most areas cross-venation scalariform and forming quadrangular cells; cross-veins sigmoidal, reticulated and denser distal to pterostigma, reticulated near apex, and in RP3+4-MA area.
|
||
</paragraph>
|
||
<caption id="139BE0EF20219852FF5E5128FD53FAD6" ID-DOI="http://doi.org/10.5281/zenodo.8177411" ID-Zenodo-Dep="8177411" httpUri="https://zenodo.org/record/8177411/files/figure.png" pageId="5" pageNumber="404" startId="5.[132,143,1299,1316]" targetBox="[193,1399,215,1257]" targetPageId="5" targetType="figure">
|
||
<paragraph id="475BB06720219852FF5E5128FD53FAD6" blockId="5.[132,1455,1299,1362]" pageId="5" pageNumber="404">
|
||
<smallCapsWord id="41BD26BB20219852FF5E5128FF32FAA0" baselines="1312,1313" box="[132,157,1299,1316]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="5" pageNumber="404">FIG</smallCapsWord>
|
||
. 2. —
|
||
<taxonomicName id="80E4CBE420219852FF0E5128FE11FAA0" authority="Doriath-Döhler & Hervet & Béthoux, 2023" authorityName="Doriath-Döhler & Hervet & Béthoux" authorityYear="2023" box="[212,446,1299,1316]" class="Insecta" family="Synlestidae" genus="Palaeophylolestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="404" phylum="Arthropoda" rank="species" species="distinctus" status="n. gen., n. sp.">
|
||
<emphasis id="75906C7520219852FF0E5128FE11FAA0" box="[212,446,1299,1316]" italics="true" pageId="5" pageNumber="404">Palaeophylolestes distinctus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA3D10E20219852FE195128FD97FAA0" box="[451,568,1299,1316]" pageId="5" pageNumber="404" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
, holotype specimen (BDL 2002), drawings of wings, isolated (see Fig. 1 for their respective locations; faded vein portions reconstructed);
|
||
<emphasis id="75906C7520219852FEA25111FE2AFABF" bold="true" box="[376,389,1322,1339]" pageId="5" pageNumber="404">A</emphasis>
|
||
,
|
||
<emphasis id="75906C7520219852FE555111FE33FABF" bold="true" box="[399,412,1322,1339]" pageId="5" pageNumber="404">B</emphasis>
|
||
, first pair (located on the left side of the complete specimen);
|
||
<emphasis id="75906C7520219852FC465111FC06FABF" bold="true" box="[924,937,1322,1339]" pageId="5" pageNumber="404">A</emphasis>
|
||
, forewing;
|
||
<emphasis id="75906C7520219852FBDF5111FBBDFABF" bold="true" box="[1029,1042,1322,1339]" pageId="5" pageNumber="404">B</emphasis>
|
||
, hindwing;
|
||
<emphasis id="75906C7520219852FBA85111FB2FFABF" bold="true" box="[1138,1152,1322,1339]" pageId="5" pageNumber="404">C</emphasis>
|
||
,
|
||
<emphasis id="75906C7520219852FB505111FB37FABF" bold="true" box="[1162,1176,1322,1339]" pageId="5" pageNumber="404">D</emphasis>
|
||
, second pair (located on the right side of the complete specimen);
|
||
<emphasis id="75906C7520219852FE4E517AFE0DFAD6" bold="true" box="[404,418,1345,1362]" pageId="5" pageNumber="404">C</emphasis>
|
||
, forewing;
|
||
<emphasis id="75906C7520219852FE25517AFDA2FAD6" bold="true" box="[511,525,1345,1362]" pageId="5" pageNumber="404">D</emphasis>
|
||
, hindwing. Scale bar: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="475BB06720219852FCF751AFFC0AFA29" blockId="5.[813,933,1428,1453]" box="[813,933,1428,1453]" pageId="5" pageNumber="404">
|
||
<heading id="1C13070B20219852FCF751AFFC0AFA29" box="[813,933,1428,1453]" fontSize="11" level="2" pageId="5" pageNumber="404" reason="8">
|
||
<emphasis id="75906C7520219852FCF751AFFC0AFA29" box="[813,933,1428,1453]" italics="true" pageId="5" pageNumber="404">Dimensions</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="475BB06720219852FCF75188FB50F989" blockId="5.[812,1457,1459,1549]" pageId="5" pageNumber="404">
|
||
Forewing, length about
|
||
<quantity id="801C1D8220219852FBE25188FB0DFA49" box="[1080,1186,1459,1486]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="404" unit="mm" value="25.4">25.4 mm</quantity>
|
||
, maximum width about
|
||
<quantity id="801C1D8220219852FCF451E8FC2CFA69" box="[814,899,1491,1517]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="404" unit="mm" value="4.9">4.9 mm</quantity>
|
||
(both estimated from FW1); hindwing, length
|
||
<quantity id="801C1D8220219852FA5A51E8FCF4F989" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="5" pageNumber="404" unit="mm" value="23.6">23.6 mm</quantity>
|
||
, width
|
||
<quantity id="801C1D8220219852FC7651C9FBADF989" box="[940,1026,1522,1549]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="404" unit="mm" value="5.3">5.3 mm</quantity>
|
||
(deduced from HW1).
|
||
</paragraph>
|
||
<paragraph id="475BB06720219852FCF75208FC3EF9CF" blockId="5.[811,1456,1587,2028]" box="[813,913,1587,1613]" pageId="5" pageNumber="404">
|
||
<smallCapsWord id="41BD26BB20219852FCF75208FC3EF9CF" baselines="1606,1606" box="[813,913,1587,1613]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="5" pageNumber="404">REMARKS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="475BB06720219852FCF75269FA01F86F" blockId="5.[811,1456,1587,2028]" pageId="5" pageNumber="404">
|
||
Within Lestoidea (as delimited in
|
||
<bibRefCitation id="2375CD9620219852FB695269FADAF9E8" author="BYBEE S. M. & KALKMAN V. J. & ERICKSON R. J. & FRANDSEN P. B. & BREINHOLT J. W. & SUVOROV A. & DIJKSTRA K. - D. B. & CORDERO-RIVERA A. & SKEVINGTON J. H. & ABBOTT J. & SANCHEZ HERRERA M. & LEMMON A. R. & LEMMON E. M. & WARE J. L." box="[1203,1397,1618,1644]" pageId="5" pageNumber="404" pagination="107115" refId="ref3453" refString="BYBEE S. M., KALKMAN V. J., ERICKSON R. J., FRANDSEN P. B., BREINHOLT J. W., SUVOROV A., DIJKSTRA K. - D. B., CORDERO-RIVERA A., SKEVINGTON J. H., ABBOTT J., SANCHEZ HERRERA M., LEMMON A. R., LEMMON E. M. & WARE J. L. 2021. - Phylogeny and classification of Odonata using targeted genomics. Molecular Phylogenetics and Evolution 160: 107115. https: // doi. org / 10.1016 / j. ympev. 2021.107115" type="journal article" year="2021">
|
||
Bybee
|
||
<emphasis id="75906C7520219852FB275268FA9BF9E8" box="[1277,1332,1618,1644]" italics="true" pageId="5" pageNumber="404">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
; and see
|
||
<bibRefCitation id="2375CD9620219852FC895249FBA4F908" author="KOHLI M. & LETSCH H. & GREVE C. & BETHOUX O. & DEREGNAUCOURT I. & LIU S. & ZHOU X. & DONATH A. & MAYER C. & PODSIADLOWSKI L. & GUNKEL S. & MACHIDA R. & NIEHUIS O. & RUST J. & WAPPLER T. & YU X. & MISOF B. & WARE J." box="[851,1035,1650,1676]" pageId="5" pageNumber="404" pagination="103324" refId="ref4023" refString="KOHLI M., LETSCH H., GREVE C., BETHOUX O., DEREGNAUCOURT I., LIU S., ZHOU X., DONATH A., MAYER C., PODSIADLOWSKI L., GUNKEL S., MACHIDA R., NIEHUIS O., RUST J., WAPPLER T., YU X., MISOF B. & WARE J. 2021. - Evolutionary history and divergence times of Odonata (dragonflies and damselflies) revealed through transcriptomics. iScience 24: 103324. https: // doi. org / 10.1016 / j. isci. 2021.103324" type="journal article" year="2021">
|
||
Kohli
|
||
<emphasis id="75906C7520219852FC4D5248FC64F908" box="[919,971,1650,1676]" italics="true" pageId="5" pageNumber="404">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
), the new specimen can be assigned to the
|
||
<taxonomicName id="80E4CBE420219852FC8F52A9FC7EF928" authorityName="Tillyard" authorityYear="1917" box="[853,977,1682,1708]" class="Insecta" family="Synlestidae" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="404" phylum="Arthropoda" rank="family">Synlestidae</taxonomicName>
|
||
based on a particular combination of character states. The occurrence of the state ‘RP1+2 / RP3+4 fork located closer to nodus than to arculus’ is unique to this family indeed (see
|
||
<bibRefCitation id="2375CD9620219852FBEC52C9FA84F888" author="GARRISON R. W. & VON ELLENRIEDER N. & LOUTON J. A." box="[1078,1323,1778,1804]" pageId="5" pageNumber="404" refId="ref3699" refString="GARRISON R. W., VON ELLENRIEDER N. & LOUTON J. A. 2010. - Damselfly Genera of the New World. John Hopkins University Press, Baltimore, 490 p." type="book" year="2010">
|
||
Garrison
|
||
<emphasis id="75906C7520219852FB4552C8FB79F888" box="[1183,1238,1778,1804]" italics="true" pageId="5" pageNumber="404">et al.</emphasis>
|
||
[2010]
|
||
</bibRefCitation>
|
||
and
|
||
<figureCitation id="DFDFACE220219852FAB252C9FA07F888" box="[1384,1448,1778,1804]" captionStart="FIG" captionStartId="6.[132,143,1115,1132]" captionTargetBox="[142,1456,215,1079]" captionTargetId="graphics-832@6.[182,1400,521,784]" captionTargetPageId="6" captionText="FIG. 3. — Forewings of selected extant representatives of the Synlestidae Tillyard, 1917: A, Synlestes weyersii Selys Longchamps, 1868 (male; based on photograph by John Tann [Sydney, Australia], CC BY 2.0); B, Megalestes gyalsey Gyeltshen, Kalkman & Orr, 2017 (male; based on Payra et al. 2021: fig. 2F, approximate scale); C, Phylolestes ethelae Christiansen, 1947 (male; based on Garrison et al. 2010: fig. 527). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8177415" httpUri="https://zenodo.org/record/8177415/files/figure.png" pageId="5" pageNumber="404">Fig. 3</figureCitation>
|
||
; the RP1+2 / RP3+4 fork is located closer to the arculus in
|
||
<taxonomicName id="80E4CBE420219852FCF6530AFC29F8CF" authorityName="Calvert" authorityYear="1901" box="[812,902,1841,1867]" class="Insecta" family="Lestidae" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="404" phylum="Arthropoda" rank="family">Lestidae</taxonomicName>
|
||
, Menatlestidae and
|
||
<taxonomicName id="80E4CBE420219852FB8C530AFB6AF8CF" box="[1110,1221,1841,1867]" class="Insecta" family="Eolestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="404" phylum="Arthropoda" rank="family">Eolestidae</taxonomicName>
|
||
, very close or opposite to the nodus in
|
||
<taxonomicName id="80E4CBE420219852FC0F536AFB24F8EF" authorityName="Kennedy" authorityYear="1920" box="[981,1163,1873,1899]" class="Insecta" family="Hemiphlebiidae" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="404" phylum="Arthropoda" rank="family">Hemiphlebiidae</taxonomicName>
|
||
, and beyond it in
|
||
<taxonomicName id="80E4CBE420219852FA97536AFCF9F80F" authorityName="Kennedy" authorityYear="1920" class="Insecta" family="Perilestidae" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="404" phylum="Arthropoda" rank="family">Perilestidae</taxonomicName>
|
||
; exceptions occur within
|
||
<taxonomicName id="80E4CBE420219852FBAD534AFB59F80F" authorityName="Tillyard" authorityYear="1917" box="[1143,1270,1905,1931]" class="Insecta" family="Synlestidae" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="404" phylum="Arthropoda" rank="family">Synlestidae</taxonomicName>
|
||
, see
|
||
<figureCitation id="DFDFACE220219852FAF4534AFA2DF808" box="[1326,1410,1905,1932]" captionStart="FIG" captionStartId="6.[132,143,1115,1132]" captionTargetBox="[142,1456,215,1079]" captionTargetId="graphics-832@6.[182,1400,521,784]" captionTargetPageId="6" captionText="FIG. 3. — Forewings of selected extant representatives of the Synlestidae Tillyard, 1917: A, Synlestes weyersii Selys Longchamps, 1868 (male; based on photograph by John Tann [Sydney, Australia], CC BY 2.0); B, Megalestes gyalsey Gyeltshen, Kalkman & Orr, 2017 (male; based on Payra et al. 2021: fig. 2F, approximate scale); C, Phylolestes ethelae Christiansen, 1947 (male; based on Garrison et al. 2010: fig. 527). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8177415" httpUri="https://zenodo.org/record/8177415/files/figure.png" pageId="5" pageNumber="404">Fig. 3B</figureCitation>
|
||
; see
|
||
<bibRefCitation id="2375CD9620219852FCF453AAFC6DF82F" author="BRIDGES C. A." box="[814,962,1937,1963]" pageId="5" pageNumber="404" refId="ref3403" refString="BRIDGES C. A. 1994. - Catalogue of the Family-Group, Genus-Group and Species-Group Names of the Odonata of the World. 3 rd edition. The author, Urbana, 905 p. https: // doi. org / 10.5962 / bhl. title. 15291" type="book" year="1994">Bridges 1994</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2375CD9620219852FC0853AAFB53F82F" author="GREENWALT D. E. & BECHLY G." box="[978,1276,1937,1963]" pageId="5" pageNumber="404" pagination="138 - 156" refId="ref3735" refString="GREENWALT D. E. & BECHLY G. 2014. - A re-description of the fossil damselfly Eolestes syntheticus Cockerell, 1940 (Odonata: Zygoptera: Eolestidae n. fam.) with description of new taxa from the Eocene of North America. Zootaxa 3887: 138 - 156. https: // doi. org / 10.11646 / zootaxa. 3887.2.2" type="journal article" year="2014">Greenwalt & Bechly 2014</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2375CD9620219852FAD753AAFCC6F84F" author="NEL A. & JOUAULT C." pageId="5" pageNumber="404" pagination="631 - 648" refId="ref4231" refString="NEL A. & JOUAULT C. 2022. - The odonatan insects from the Paleocene of Menat, central France. Acta Palaeontologica Polonica 67: 631 - 648. https: // doi. org / 10.4202 / app. 00960.2021" type="journal article" year="2022">Nel & Jouault 2022</bibRefCitation>
|
||
). This assignment is also consistent with the position of the arculus being closer to the nodus than to the wing
|
||
</paragraph>
|
||
<paragraph id="475BB06720229851FF5454ECFDFDFF69" blockId="6.[142,1456,215,1079]" box="[142,594,215,240]" pageId="6" pageNumber="405">A ScP</paragraph>
|
||
<paragraph id="475BB06720229851FAAC55BFFAC9FE76" blockId="6.[142,1456,215,1079]" pageId="6" pageNumber="405">
|
||
RA RP1 1-rp2 B ScP CuA
|
||
<collectionCode id="21F528A220229851FBE055DFFBF5FE7C" box="[1082,1114,484,504]" country="0" name="Mohonk Preserve, Inc." pageId="6" pageNumber="405">MP</collectionCode>
|
||
MA RP3+4 Irp1+2-rp3+4
|
||
</paragraph>
|
||
<paragraph id="475BB06720229851FAA6569CFB07FC9B" blockId="6.[142,1456,215,1079]" pageId="6" pageNumber="405">
|
||
RA RP1 1-rp2 C ScP CuA
|
||
<collectionCode id="21F528A220229851FC01572DFC54FCAE" box="[987,1019,790,810]" country="0" name="Mohonk Preserve, Inc." pageId="6" pageNumber="405">MP</collectionCode>
|
||
MA RP3+4
|
||
</paragraph>
|
||
<paragraph id="475BB06720229851FAA3579CFB8CFBB7" blockId="6.[142,1456,215,1079]" pageId="6" pageNumber="405">
|
||
RA RP1 1-rp2 RP3+4 CuA
|
||
<collectionCode id="21F528A220229851FC095018FC5CFBB3" box="[979,1011,1059,1079]" country="0" name="Mohonk Preserve, Inc." pageId="6" pageNumber="405">MP</collectionCode>
|
||
MA
|
||
</paragraph>
|
||
<caption id="139BE0EF20229851FF5E5060FBC5FB1D" ID-DOI="http://doi.org/10.5281/zenodo.8177415" ID-Zenodo-Dep="8177415" httpUri="https://zenodo.org/record/8177415/files/figure.png" pageId="6" pageNumber="405" startId="6.[132,143,1115,1132]" targetBox="[142,1456,215,1079]" targetPageId="6" targetType="figure">
|
||
<paragraph id="475BB06720229851FF5E5060FBC5FB1D" blockId="6.[132,1455,1115,1177]" pageId="6" pageNumber="405">
|
||
<smallCapsWord id="41BD26BB20229851FF5E5060FF32FBEF" baselines="1128,1128" box="[132,157,1115,1132]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="6" pageNumber="405">FIG</smallCapsWord>
|
||
. 3. — Forewings of selected extant representatives of the
|
||
<taxonomicName id="80E4CBE420229851FD595060FCF4FBE8" authority="Tillyard, 1917" authorityName="Tillyard" authorityYear="1917" box="[643,859,1115,1132]" class="Insecta" family="Synlestidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">
|
||
Synlestidae
|
||
<bibRefCitation id="2375CD9620229851FD325060FCF4FBE8" author="TILLYARD R. J." box="[744,859,1115,1132]" pageId="6" pageNumber="405" refId="ref4960" refString="TILLYARD R. J. 1917. - The Biology of Dragonflies. Cambridge University Press, Cambridge, xi + 396 p. https: // doi. org / 10.5962 / bhl. title. 27401" type="book" year="1917">Tillyard, 1917</bibRefCitation>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="75906C7520229851FCB85060FCC0FBE8" bold="true" box="[866,879,1115,1132]" pageId="6" pageNumber="405">A</emphasis>
|
||
,
|
||
<taxonomicName id="80E4CBE420229851FCA35060FB46FBE8" authority="Selys Longchamps, 1868" authorityName="Selys Longchamps" authorityYear="1868" box="[889,1257,1115,1132]" class="Insecta" family="Synlestidae" genus="Synlestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="weyersii">
|
||
<emphasis id="75906C7520229851FCA35060FBBEFBE8" box="[889,1041,1115,1132]" italics="true" pageId="6" pageNumber="405">Synlestes weyersii</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FBCF5060FB46FBE8" author="SELYS LONGCHAMPS E. D." box="[1045,1257,1115,1132]" pageId="6" pageNumber="405" refId="ref4737" refString="SELYS LONGCHAMPS E. D. 1868. - Les nevropteres de Port Denison (in Assemblee mensuelle du 4 avril 1868). Annales de la Societe entomologique de Belgique (Comptes-rendus) 11: lxxvi-lxxvii. https: // www. biodiversitylibrary. org / page / 12563835" type="journal volume" year="1868">Selys Longchamps, 1868</bibRefCitation>
|
||
</taxonomicName>
|
||
(male; based on photograph by John Tann [Sydney, Australia], CC BY 2.0);
|
||
<emphasis id="75906C7520229851FDE6504AFDE6FB06" bold="true" box="[572,585,1137,1154]" pageId="6" pageNumber="405">B</emphasis>
|
||
,
|
||
<taxonomicName id="80E4CBE420229851FD8E504AFBA9FB06" authority="Gyeltshen, Kalkman & Orr, 2017" authorityName="Gyeltshen, Kalkman & Orr" authorityYear="2017" box="[596,1030,1137,1154]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="gyalsey">
|
||
<emphasis id="75906C7520229851FD8E504AFD59FB06" box="[596,758,1137,1154]" italics="true" pageId="6" pageNumber="405">Megalestes gyalsey</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FD20504AFBA9FB06" author="GYELTSHEN T. & KALKMAN V. & ORR A. G." box="[762,1030,1137,1154]" pageId="6" pageNumber="405" pagination="588 - 594" refId="ref3800" refString="GYELTSHEN T., KALKMAN V. & ORR A. G. 2017. - Honouring His Royal Highness the Crown Prince of Bhutan: Megalestes gyalsey (Odonata: Synlestidae). Zootaxa 4244: 588 - 594. https: // doi. org / 10.11646 / zootaxa. 4244.4.9" type="journal article" year="2017">Gyeltshen, Kalkman & Orr, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
(male; based on
|
||
<bibRefCitation id="2375CD9620229851FB4F504AFA8EFB06" author="PAYRA A. & DAWN P. & SUBRAMANIAN K. A. & DEEPAK C. K. & CHANDRA K. & TRIPATHY B." box="[1173,1313,1137,1154]" pageId="6" pageNumber="405" pagination="233 - 242" refId="ref4274" refString="PAYRA A., DAWN P., SUBRAMANIAN K. A., DEEPAK C. K., CHANDRA K. & TRIPATHY B. 2021. - New record of Megalestes gyalsey Gyeltshen, Kalkman & Orr, 2017 (Zygoptera: Synlestidae) from India, with first description of female and larva. Zootaxa 4938: 233 - 242. https: // doi. org / 10.11646 / zootaxa. 4938.2.4" type="journal article" year="2021">
|
||
Payra
|
||
<emphasis id="75906C7520229851FB10504AFB5CFB06" box="[1226,1267,1137,1154]" italics="true" pageId="6" pageNumber="405">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
: fig. 2F, approximate scale);
|
||
<emphasis id="75906C7520229851FF3450B3FF53FB1D" bold="true" box="[238,252,1160,1177]" pageId="6" pageNumber="405">C</emphasis>
|
||
,
|
||
<taxonomicName id="80E4CBE420229851FEDC50B3FDE8FB1D" authority="Christiansen, 1947" authorityName="Christiansen" authorityYear="1947" box="[262,583,1160,1177]" class="Insecta" family="Synlestidae" genus="Phylolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="ethelae">
|
||
<emphasis id="75906C7520229851FEDC50B3FE09FB1D" box="[262,422,1160,1177]" italics="true" pageId="6" pageNumber="405">Phylolestes ethelae</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FE7150B3FDE8FB1D" author="CHRISTIANSEN K. A." box="[427,583,1160,1177]" pageId="6" pageNumber="405" pagination="256 - 262" refId="ref3566" refString="CHRISTIANSEN K. A. 1947. - A new genus and species of damselfly from Southern Haiti (Odonata). Psyche 54: 256 - 262. https: // doi. org / 10.1155 / 1947 / 59418" type="journal article" year="1947">Christiansen, 1947</bibRefCitation>
|
||
</taxonomicName>
|
||
(male; based on
|
||
<bibRefCitation id="2375CD9620229851FD0C50B3FCD5FB1D" author="GARRISON R. W. & VON ELLENRIEDER N. & LOUTON J. A." box="[726,890,1160,1177]" pageId="6" pageNumber="405" refId="ref3699" refString="GARRISON R. W., VON ELLENRIEDER N. & LOUTON J. A. 2010. - Damselfly Genera of the New World. John Hopkins University Press, Baltimore, 490 p." type="book" year="2010">
|
||
Garrison
|
||
<emphasis id="75906C7520229851FCF950B3FCE3FB1D" box="[803,844,1160,1177]" italics="true" pageId="6" pageNumber="405">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
: fig. 527). Scale bars: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="475BB06720229851FF5E50C8FDEBFA49" blockId="6.[130,776,1267,2028]" pageId="6" pageNumber="405">
|
||
base (see
|
||
<bibRefCitation id="2375CD9620229851FF3650C8FE2BFA8A" author="TILLYARD R. J." box="[236,388,1267,1294]" pageId="6" pageNumber="405" refId="ref4960" refString="TILLYARD R. J. 1917. - The Biology of Dragonflies. Cambridge University Press, Cambridge, xi + 396 p. https: // doi. org / 10.5962 / bhl. title. 27401" type="book" year="1917">Tillyard 1917</bibRefCitation>
|
||
; more or less midway between the base and the nodus in
|
||
<taxonomicName id="80E4CBE420229851FE5B5128FE70FAA9" authorityName="Calvert" authorityYear="1901" box="[385,479,1299,1325]" class="Insecta" family="Lestidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Lestidae</taxonomicName>
|
||
; closer to the wing base in
|
||
<taxonomicName id="80E4CBE420229851FF5F5108FE94FAC9" authorityName="Kennedy" authorityYear="1920" box="[133,315,1331,1357]" class="Insecta" family="Hemiphlebiidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Hemiphlebiidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="80E4CBE420229851FE9E5108FE1BFAC9" box="[324,436,1331,1357]" class="Insecta" family="Eolestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Eolestidae</taxonomicName>
|
||
and Menatlestidae; conversely, very close to the nodus in
|
||
<taxonomicName id="80E4CBE420229851FE6B5168FD9BFAE9" authorityName="Kennedy" authorityYear="1920" box="[433,564,1363,1389]" class="Insecta" family="Perilestidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Perilestidae</taxonomicName>
|
||
), and the presence of intercalary veins in the area between Irp
|
||
<subScript id="DB60B22220229851FDBD51BBFDDEFA15" attach="left" box="[615,625,1408,1425]" fontSize="7" pageId="6" pageNumber="405">1</subScript>
|
||
-rp
|
||
<subScript id="DB60B22220229851FD4851BBFD33FA15" attach="left" box="[658,668,1408,1425]" fontSize="7" pageId="6" pageNumber="405">2</subScript>
|
||
and RP2 (absent in
|
||
<taxonomicName id="80E4CBE420229851FF2251A8FE04FA29" authorityName="Kennedy" authorityYear="1920" box="[248,427,1427,1453]" class="Insecta" family="Hemiphlebiidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Hemiphlebiidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="80E4CBE420229851FE3B51A8FDCDFA29" authorityName="Kennedy" authorityYear="1920" box="[481,610,1427,1453]" class="Insecta" family="Perilestidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Perilestidae</taxonomicName>
|
||
; also present in
|
||
<taxonomicName id="80E4CBE420229851FF595188FF4EFA49" authorityName="Calvert" authorityYear="1901" box="[131,225,1459,1485]" class="Insecta" family="Lestidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Lestidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="80E4CBE420229851FF375188FEF1FA49" box="[237,350,1459,1485]" class="Insecta" family="Eolestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Eolestidae</taxonomicName>
|
||
and Menatlestidae).
|
||
</paragraph>
|
||
<paragraph id="475BB06720229851FF4151E8FEF0F948" blockId="6.[130,776,1267,2028]" pageId="6" pageNumber="405">
|
||
It must be mentioned that, outside of Lestoidea, the
|
||
<taxonomicName id="80E4CBE420229851FF5E51C8FEF5F989" box="[132,346,1523,1549]" class="Insecta" family="Megapodagrionidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Megapodagrionidae</taxonomicName>
|
||
and the
|
||
<taxonomicName id="80E4CBE420229851FE6F51C8FDCFF989" box="[437,608,1523,1549]" class="Insecta" family="Coenagrionidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Coenagrionidae</taxonomicName>
|
||
also display the state ‘RP1+2 / RP3+4 fork located closer to nodus than to arculus’. However, assignment to the former can be excluded owing to the quadrangular shape of the quadrangle. As for the latter,
|
||
<taxonomicName id="80E4CBE420229851FF2F5249FE09F908" box="[245,422,1650,1676]" class="Insecta" family="Coenagrionidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Coenagrionidae</taxonomicName>
|
||
have a single row of cells in the area between Irp
|
||
<subScript id="DB60B22220229851FEE052A4FEEBF934" attach="right" box="[314,324,1695,1712]" fontSize="7" pageId="6" pageNumber="405">1</subScript>
|
||
-rp
|
||
<subScript id="DB60B22220229851FEBC52A4FEDFF934" attach="left" box="[358,368,1695,1712]" fontSize="7" pageId="6" pageNumber="405">2</subScript>
|
||
and RP2, while the new material has 3, as in
|
||
<taxonomicName id="80E4CBE420229851FF015289FEF4F948" authorityName="Tillyard" authorityYear="1917" box="[219,347,1714,1740]" class="Insecta" family="Synlestidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Synlestidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="475BB06720229851FF4152E9FC42F86F" blockId="6.[130,776,1267,2028]" lastBlockId="6.[811,1457,1267,2028]" pageId="6" pageNumber="405">
|
||
The new specimen differs from all known
|
||
<taxonomicName id="80E4CBE420229851FD5252E9FCA9F968" authorityName="Tillyard" authorityYear="1917" box="[648,774,1746,1772]" class="Insecta" family="Synlestidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Synlestidae</taxonomicName>
|
||
by the very long MA,
|
||
<collectionCode id="21F528A220229851FE5C52C9FE1FF888" box="[390,432,1778,1804]" country="0" name="Mohonk Preserve, Inc." pageId="6" pageNumber="405">MP</collectionCode>
|
||
and CuA. Also, the quadrangle, characterised by a comparatively long proximal edge, is atypical compared to known
|
||
<taxonomicName id="80E4CBE420229851FE31530AFDC4F8CF" authorityName="Tillyard" authorityYear="1917" box="[491,619,1841,1867]" class="Insecta" family="Synlestidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Synlestidae</taxonomicName>
|
||
. Additionally, unlike in
|
||
<taxonomicName id="80E4CBE420229851FF2F536AFDC1F8EF" authority="Selys Longchamps, 1868" authorityName="Selys Longchamps" authorityYear="1868" box="[245,622,1873,1899]" class="Insecta" family="Synlestidae" genus="Synlestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75906C7520229851FF2F536AFEE1F8EF" box="[245,334,1873,1899]" italics="true" pageId="6" pageNumber="405">Synlestes</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FE8C536AFDC1F8EF" author="SELYS LONGCHAMPS E. D." box="[342,622,1873,1899]" pageId="6" pageNumber="405" refId="ref4737" refString="SELYS LONGCHAMPS E. D. 1868. - Les nevropteres de Port Denison (in Assemblee mensuelle du 4 avril 1868). Annales de la Societe entomologique de Belgique (Comptes-rendus) 11: lxxvi-lxxvii. https: // www. biodiversitylibrary. org / page / 12563835" type="journal volume" year="1868">Selys Longchamps, 1868</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="DFDFACE220229851FD76536AFCAFF8E8" box="[684,768,1873,1900]" captionStart="FIG" captionStartId="6.[132,143,1115,1132]" captionTargetBox="[142,1456,215,1079]" captionTargetId="graphics-832@6.[182,1400,521,784]" captionTargetPageId="6" captionText="FIG. 3. — Forewings of selected extant representatives of the Synlestidae Tillyard, 1917: A, Synlestes weyersii Selys Longchamps, 1868 (male; based on photograph by John Tann [Sydney, Australia], CC BY 2.0); B, Megalestes gyalsey Gyeltshen, Kalkman & Orr, 2017 (male; based on Payra et al. 2021: fig. 2F, approximate scale); C, Phylolestes ethelae Christiansen, 1947 (male; based on Garrison et al. 2010: fig. 527). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8177415" httpUri="https://zenodo.org/record/8177415/files/figure.png" pageId="6" pageNumber="405">Fig. 3A</figureCitation>
|
||
) and a number of other extant genera, including
|
||
<taxonomicName id="80E4CBE420229851FD50534AFE1DF82F" authority="Selys Longchamps, 1862 b" authorityName="Selys Longchamps" authorityYear="1862" class="Insecta" family="Synlestidae" genus="Chlorolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75906C7520229851FD50534AFCAAF80F" box="[650,773,1905,1931]" italics="true" pageId="6" pageNumber="405">Chlorolestes</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FF5E53AAFE1DF82F" author="SELYS LONGCHAMPS E. D." box="[132,434,1937,1963]" pageId="6" pageNumber="405" pagination="5 - 44" refId="ref4688" refString="SELYS LONGCHAMPS E. D. 1862 b. - Synopsis des agrionines. 3 me Legion. - Podagrion. Bulletins de l'Academie royale de Belgique, serie 2, 14: 5 - 44. https: // www. biodiversitylibrary. org / page / 36014862" type="journal article" year="1862">Selys Longchamps, 1862b</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80E4CBE420229851FE1953AAFCAEF82F" authority="Barnard, 1937" authorityName="Barnard" authorityYear="1937" box="[451,769,1937,1963]" class="Insecta" family="Synlestidae" genus="Ecchlorolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75906C7520229851FE1953AAFDFCF82F" box="[451,595,1937,1963]" italics="true" pageId="6" pageNumber="405">Ecchlorolestes</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FD8053AAFCAEF82F" author="BARNARD K. H." box="[602,769,1937,1963]" pageId="6" pageNumber="405" pagination="169 - 260" refId="ref3259" refString="BARNARD K. H. 1937. - Notes on dragon-flies (Odonata) of the S. W. Cape with descriptions of the nymphs and of new species. Annals of the South African Museum 32: 169 - 260. https: // www. biodiversitylibrary. org / page / 40890966" type="journal article" year="1937">Barnard, 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80E4CBE420229851FF5D538AFE34F84F" authority="Kennedy, 1920" authorityName="Kennedy" authorityYear="1920" box="[135,411,1969,1995]" class="Insecta" family="Synlestidae" genus="Episynlestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75906C7520229851FF5D538AFF56F84F" box="[135,249,1969,1995]" italics="true" pageId="6" pageNumber="405">Episynlestes</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FF26538AFE34F84F" author="KENNEDY C. H." box="[252,411,1969,1995]" pageId="6" pageNumber="405" pagination="83 - 88" refId="ref3982" refString="KENNEDY C. H. 1920. - Forty-two hitherto unrecognized genera and subgenera of Zygoptera. Ohio Journal of Science 21: 83 - 88. https: // doi. org / 10.5962 / bhl. part. 14540" type="journal article" year="1920">Kennedy, 1920</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="80E4CBE420229851FE11538AFD75F84F" authority="Needham, 1930" authorityName="Needham" authorityYear="1930" box="[459,730,1969,1995]" class="Insecta" family="Synlestidae" genus="Sinolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75906C7520229851FE11538AFD87F84F" box="[459,552,1969,1995]" italics="true" pageId="6" pageNumber="405">Sinolestes</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FDEA538AFD75F84F" author="NEEDHAM J. G." box="[560,730,1969,1995]" pageId="6" pageNumber="405" pagination="1 - 344" refId="ref4193" refString="NEEDHAM J. G. 1930. - A manual of the dragonflies of China. A monographic study of the Chinese Odonata. Zoologica Sinica (A) 2: 1 - 344, 20 pls." type="journal article" year="1930">Needham, 1930</bibRefCitation>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="2375CD9620229851FF5853EAFEBAF86F" author="BRIDGES C. A." box="[130,277,2001,2027]" pageId="6" pageNumber="405" refId="ref3403" refString="BRIDGES C. A. 1994. - Catalogue of the Family-Group, Genus-Group and Species-Group Names of the Odonata of the World. 3 rd edition. The author, Urbana, 905 p. https: // doi. org / 10.5962 / bhl. title. 15291" type="book" year="1994">Bridges 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2375CD9620229851FEFC53EAFE54F86F" author="SIMAIKA J. P. & WARE J. L. & GARRISON R. W. & SAMWAYS M. J." box="[294,507,2001,2027]" pageId="6" pageNumber="405" pagination="15088" refId="ref4787" refString="SIMAIKA J. P., WARE J. L., GARRISON R. W. & SAMWAYS M. J. 2020. - Phylogeny of the Synlestidae (Odonata: Zygoptera), with an emphasis on Chlorolestes Selys and Ecchlorolestes Barnard. Scientific Reports 10: 15088. https: // doi. org / 10.1038 / s 41598 - 020 - 72001 - x" type="journal article" year="2020">
|
||
Simaika
|
||
<emphasis id="75906C7520229851FE5C53E9FE14F86F" box="[390,443,2001,2027]" italics="true" pageId="6" pageNumber="405">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
; and references therein) as well as some extinct genera such as
|
||
<taxonomicName id="80E4CBE420229851FB1F50C8FCCBFAAA" authority="Petrulevicius, 2018" authorityName="Petrulevicius" authorityYear="2018" class="Insecta" family="Synlestidae" genus="Madres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75906C7520229851FB1F50C8FABCFA89" box="[1221,1299,1267,1293]" italics="true" pageId="6" pageNumber="405">Madres</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FAC150C8FCCBFAAA" author="PETRULEVICIUS J. F." pageId="6" pageNumber="405" pagination="36 - 43" refId="ref4407" refString="PETRULEVICIUS J. F. 2018. - A new Malachite Damselfly (Synlestidae: Odonata) from the Eocene of Patagonia, Argentina. Life: The Excitement of Biology 6: 36 - 43. https: // doi. org / 10.9784 / LEB 6 (2) Petrulevicius. 01" type="journal article" year="2018">Petrulevičius, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="80E4CBE420229851FC795128FAA7FAAA" authority="Petrulevicius, 2015" authorityName="Petrulevicius" authorityYear="2015" box="[931,1288,1299,1326]" class="Insecta" family="Synlestidae" genus="Inacayalestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75906C7520229851FC795128FB86FAA9" box="[931,1065,1299,1325]" italics="true" pageId="6" pageNumber="405">Inacayalestes</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FBEB5128FAA7FAAA" author="PETRULEVICIUS J. F." box="[1073,1288,1299,1326]" pageId="6" pageNumber="405" pagination="287 - 294" refId="ref4351" refString="PETRULEVICIUS J. F. 2015. - A new Synlestidae damselfly (Insecta: Odonata: Zygoptera) from the early Eocene of Nahuel Huapi Este, Patagonia, Argentina. Arquivos Entomoloxicos 14: 287 - 294. http: // sedici. unlp. edu. ar / handle / 10915 / 99440" type="journal article" year="2015">Petrulevičius, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
(both Eocene,
|
||
<collectingCountry id="3FF3F0F720229851FCF45108FC0BFAC9" box="[814,932,1331,1357]" name="Argentina" pageId="6" pageNumber="405">Argentina</collectingCountry>
|
||
), the cell located below the sub-discoidal cell (colour-coded in light blue in
|
||
<figureCitation id="DFDFACE220229851FB5A5168FB6DFAE9" box="[1152,1218,1363,1389]" captionStart="FIG" captionStartId="5.[132,143,1299,1316]" captionTargetBox="[193,1399,215,1257]" captionTargetId="graphics-518@5.[236,1325,500,1043]" captionTargetPageId="5" captionText="FIG. 2. — Palaeophylolestes distinctus n. gen., n. sp., holotype specimen (BDL 2002), drawings of wings, isolated (see Fig. 1 for their respective locations; faded vein portions reconstructed); A, B, first pair (located on the left side of the complete specimen); A, forewing; B, hindwing; C, D, second pair (located on the right side of the complete specimen); C, forewing; D, hindwing. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8177411" httpUri="https://zenodo.org/record/8177411/files/figure.png" pageId="6" pageNumber="405">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="DFDFACE220229851FB0B5168FB70FAE9" box="[1233,1247,1363,1389]" captionStart="FIG" captionStartId="6.[132,143,1115,1132]" captionTargetBox="[142,1456,215,1079]" captionTargetId="graphics-832@6.[182,1400,521,784]" captionTargetPageId="6" captionText="FIG. 3. — Forewings of selected extant representatives of the Synlestidae Tillyard, 1917: A, Synlestes weyersii Selys Longchamps, 1868 (male; based on photograph by John Tann [Sydney, Australia], CC BY 2.0); B, Megalestes gyalsey Gyeltshen, Kalkman & Orr, 2017 (male; based on Payra et al. 2021: fig. 2F, approximate scale); C, Phylolestes ethelae Christiansen, 1947 (male; based on Garrison et al. 2010: fig. 527). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8177415" httpUri="https://zenodo.org/record/8177415/files/figure.png" pageId="6" pageNumber="405">3</figureCitation>
|
||
) is well-developed (i.e. AA diverges from the posterior margin in a basal position) in the new specimen. This state is shared with species of
|
||
<taxonomicName id="80E4CBE420229851FC915188FB4BFA49" authority="Selys Longchamps, 1862 a" authorityName="Selys Longchamps" authorityYear="1862" box="[843,1252,1459,1485]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75906C7520229851FC915188FC17FA49" box="[843,952,1459,1485]" italics="true" pageId="6" pageNumber="405">Megalestes</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FC185188FB4BFA49" author="SELYS LONGCHAMPS E. D." box="[962,1252,1459,1485]" pageId="6" pageNumber="405" pagination="288 - 338" refId="ref4614" refString="SELYS LONGCHAMPS E. D. 1862 a. - Synopsis des agrionines. Sous-ordre des Odonates, Fab. Famille 3 me. - Agrionidees. 2 me Sous-Famille - Agrionines. 2 me Division. Normostigmatees. 2 me Legion. - Lestes. Bulletins de l'Academie royale de Belgique, serie 2, 13: 288 - 338. https: // www. biodiversitylibrary. org / page / 36916985" type="journal article" year="1862">Selys Longchamps, 1862a</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="DFDFACE220229851FB2C5188FAE6FA49" box="[1270,1353,1459,1485]" captionStart="FIG" captionStartId="6.[132,143,1115,1132]" captionTargetBox="[142,1456,215,1079]" captionTargetId="graphics-832@6.[182,1400,521,784]" captionTargetPageId="6" captionText="FIG. 3. — Forewings of selected extant representatives of the Synlestidae Tillyard, 1917: A, Synlestes weyersii Selys Longchamps, 1868 (male; based on photograph by John Tann [Sydney, Australia], CC BY 2.0); B, Megalestes gyalsey Gyeltshen, Kalkman & Orr, 2017 (male; based on Payra et al. 2021: fig. 2F, approximate scale); C, Phylolestes ethelae Christiansen, 1947 (male; based on Garrison et al. 2010: fig. 527). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8177415" httpUri="https://zenodo.org/record/8177415/files/figure.png" pageId="6" pageNumber="405">Fig. 3B</figureCitation>
|
||
) and the monotypic genus
|
||
<taxonomicName id="80E4CBE420229851FC2851E8FA94FA69" authority="Christiansen, 1947" authorityName="Christiansen" authorityYear="1947" box="[1010,1339,1491,1517]" class="Insecta" family="Synlestidae" genus="Phylolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75906C7520229851FC2851E8FBCFFA69" box="[1010,1120,1491,1517]" italics="true" pageId="6" pageNumber="405">Phylolestes</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FBBD51E8FA94FA69" author="CHRISTIANSEN K. A." box="[1127,1339,1491,1517]" pageId="6" pageNumber="405" pagination="256 - 262" refId="ref3566" refString="CHRISTIANSEN K. A. 1947. - A new genus and species of damselfly from Southern Haiti (Odonata). Psyche 54: 256 - 262. https: // doi. org / 10.1155 / 1947 / 59418" type="journal article" year="1947">Christiansen, 1947</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="DFDFACE220229851FA9651E8FA0EFA69" box="[1356,1441,1491,1517]" captionStart="FIG" captionStartId="6.[132,143,1115,1132]" captionTargetBox="[142,1456,215,1079]" captionTargetId="graphics-832@6.[182,1400,521,784]" captionTargetPageId="6" captionText="FIG. 3. — Forewings of selected extant representatives of the Synlestidae Tillyard, 1917: A, Synlestes weyersii Selys Longchamps, 1868 (male; based on photograph by John Tann [Sydney, Australia], CC BY 2.0); B, Megalestes gyalsey Gyeltshen, Kalkman & Orr, 2017 (male; based on Payra et al. 2021: fig. 2F, approximate scale); C, Phylolestes ethelae Christiansen, 1947 (male; based on Garrison et al. 2010: fig. 527). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8177415" httpUri="https://zenodo.org/record/8177415/files/figure.png" pageId="6" pageNumber="405">Fig. 3C</figureCitation>
|
||
). It is likely a plesiomorphic condition. The new specimen sharply differs from
|
||
<taxonomicName id="80E4CBE420229851FBCE5229FB19F9A9" box="[1044,1206,1554,1581]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="75906C7520229851FBCE5229FB2EF9A8" box="[1044,1153,1554,1580]" italics="true" pageId="6" pageNumber="405">Megalestes</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
in the location of the RP1+2 / RP3+4 fork (unusually basal, for a
|
||
<taxonomicName id="80E4CBE420229851FACA5209FA20F9C8" authorityName="Tillyard" authorityYear="1917" box="[1296,1423,1586,1612]" class="Insecta" family="Synlestidae" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="family">Synlestidae</taxonomicName>
|
||
, in
|
||
<taxonomicName id="80E4CBE420229851FCF65269FC36F9E8" authorityName="Selys Longchamps" authorityYear="1862" box="[812,921,1618,1644]" class="Insecta" family="Synlestidae" genus="Megalestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75906C7520229851FCF65269FC36F9E8" box="[812,921,1618,1644]" italics="true" pageId="6" pageNumber="405">Megalestes</emphasis>
|
||
</taxonomicName>
|
||
). Finally, the new specimen is overly similar to
|
||
<taxonomicName id="80E4CBE420229851FCF45249FB10F908" authority="Christiansen, 1947" authorityName="Christiansen" authorityYear="1947" box="[814,1215,1650,1676]" class="Insecta" family="Synlestidae" genus="Phylolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="ethelae">
|
||
<emphasis id="75906C7520229851FCF45249FC48F908" box="[814,999,1650,1676]" italics="true" pageId="6" pageNumber="405">Phylolestes ethelae</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FC365249FB10F908" author="CHRISTIANSEN K. A." box="[1004,1215,1650,1676]" pageId="6" pageNumber="405" pagination="256 - 262" refId="ref3566" refString="CHRISTIANSEN K. A. 1947. - A new genus and species of damselfly from Southern Haiti (Odonata). Psyche 54: 256 - 262. https: // doi. org / 10.1155 / 1947 / 59418" type="journal article" year="1947">Christiansen, 1947</bibRefCitation>
|
||
</taxonomicName>
|
||
. However, in addition to traits highlighted above, RP3+4 is more rectilinear in the new specimen (distally, it remains parallel to the posterior wing margin for some distance in
|
||
<taxonomicName id="80E4CBE420229851FB3B52E9FA32F968" authorityName="Christiansen" authorityYear="1947" box="[1249,1437,1746,1772]" class="Insecta" family="Synlestidae" genus="Phylolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="ethelae">
|
||
<emphasis id="75906C7520229851FB3B52E9FA32F968" box="[1249,1437,1746,1772]" italics="true" pageId="6" pageNumber="405">Phylolestes ethelae</emphasis>
|
||
</taxonomicName>
|
||
). Finally, comparison with
|
||
<taxonomicName id="80E4CBE420229851FB9952C9FBC2F8A8" authority="Huang, Fu, Lian, Gao & Nel, 2022" authorityName="Huang, Fu, Lian, Gao & Nel" authorityYear="2022" class="Insecta" family="Synlestidae" genus="Cretaphylolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="cretacicus">
|
||
<emphasis id="75906C7520229851FB9952C9FAFBF888" box="[1091,1364,1778,1804]" italics="true" pageId="6" pageNumber="405">Cretaphylolestes cretacicus</emphasis>
|
||
<bibRefCitation id="2375CD9620229851FA8152C9FBC2F8A8" author="HUANG D. & FU Y. & LIAN X. & GAO J. & NEL A." pageId="6" pageNumber="405" pagination="105023" refId="ref3853" refString="HUANG D., FU Y., LIAN X., GAO J. & NEL A. 2022. - The oldest malachite damselfly (Odonata: Synlestidae) from the Lower Cretaceous of China. Cretaceous Research 129: 105023. https: // doi. org / 10.1016 / j. cretres. 2021.105023" type="journal article" year="2022">Huang, Fu, Lian, Gao & Nel, 2022</bibRefCitation>
|
||
</taxonomicName>
|
||
(Lower Cretaceous,
|
||
<collectingCountry id="3FF3F0F720229851FA855329FA09F8A8" box="[1375,1446,1810,1836]" name="China" pageId="6" pageNumber="405">China</collectingCountry>
|
||
) is made difficult by the fragmentary state of the available material. Moreover, the overlapping of elements belonging to fore- and hindwing jeopardizes the identification of critical elements, such as the quadrangle shape, in this species. Given the above, the erection of a new species and genus is justified.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |