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<document id="2394AF70737355B5A8C53CB5839DC846" ID-DOI="10.11646/zootaxa.4759.4.7" ID-GBIF-Dataset="9566bd4f-5391-4b65-ae39-9189f68d0f28" ID-ISSN="1175-5326" ID-Zenodo-Dep="3741094" ID-ZooBank="5E5A6906-E137-49BF-9270-B7DAF03DE57E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586154773837" checkinUser="plazi" docAuthor="Xin, Yafei, Yao, Zhiyuan &amp; Li, Shuqiang" docDate="2020" docId="CF2F9867E118CC058C84AF81FC0DF85D" docLanguage="en" docName="zootaxa.4759.4.7.pdf" docOrigin="Zootaxa 4759 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wanniyala batatota Xin &amp; Yao &amp; Li 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="570" masterDocId="3316E01FE11CCC018C13AD28FFF9FF92" masterDocTitle="Two new species of the spider genus Wanniyala Huber &amp; Benjamin, 2005 (Araneae Pholcidae) from Sri Lanka" masterLastPageNumber="574" masterPageNumber="566" pageNumber="570" updateTime="1698735634804" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="F628ABFFE91A037454344407BDB91EDF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3E5A8600852F72290AAEBC074593BB9D">
<mods:title id="56DA36C7B0CB916B8B548FB2E8836C0F">Two new species of the spider genus Wanniyala Huber &amp; Benjamin, 2005 (Araneae Pholcidae) from Sri Lanka</mods:title>
</mods:titleInfo>
<mods:name id="159DA6ABBE08FA93BAFCC39082D67FD5" type="personal">
<mods:role id="320FDFCDBBCF775D19FEA015109178B3">
<mods:roleTerm id="2D21952237C14D9AE9E579EA11A702E7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB23007E993CB86BE79362CD62307355">Xin, Yafei</mods:namePart>
</mods:name>
<mods:name id="82900C6E65A2D3B5C5AFE40D60886958" type="personal">
<mods:role id="BE2FB90BCB7362BD824E41ECDC6D5A5A">
<mods:roleTerm id="7442AD2446E447204A3AAD5FD370BD6A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5FA2EF6A0FF23198DB4B37F57057457D">Yao, Zhiyuan</mods:namePart>
</mods:name>
<mods:name id="1AF0D571473112463EB553E61C531918" type="personal">
<mods:role id="795250BB83BF126843372886B20F3E85">
<mods:roleTerm id="BB59E4177A336DBA569109A50C0883D4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A9FEB613F15F55E33CBAEE9F38B6024C">Li, Shuqiang</mods:namePart>
</mods:name>
<mods:typeOfResource id="34E567451F3D26B3AAA1DC129874FD0D">text</mods:typeOfResource>
<mods:relatedItem id="EC488DA7D56A9619F992D1920A264302" type="host">
<mods:titleInfo id="9236F8559A003704FF71F2BC15166C70">
<mods:title id="0A1B295DE6AD9A9E97675195AF8544B2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="20A8C5744D5D97A18DCEF60ADAE170F7">
<mods:date id="019213C32EE0C3A5714B7CF8CFA9ED13">2020</mods:date>
<mods:detail id="B3D10A230547F6A54288EF26DC6847B5" type="pubDate">
<mods:number id="A044CC6700F8C52D1AF33EFF6C519F96">2020-04-06</mods:number>
</mods:detail>
<mods:detail id="F9E9F7A6DCB339DBAA633209E364436B" type="volume">
<mods:number id="DA0E8C9359FEBB2CDAA6595F1F307EBC">4759</mods:number>
</mods:detail>
<mods:detail id="062ECE53CCABF6C9FF3D22A9748A8DDD" type="issue">
<mods:number id="A051F56F798ADFBBB6DBF960F3D3EBAF">4</mods:number>
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<mods:classification id="81D90BB317176EE5E345219AF6D1E652">journal article</mods:classification>
<mods:identifier id="3079FF10BB11E2CE5CBCA7653E8EB1F3" type="DOI">10.11646/zootaxa.4759.4.7</mods:identifier>
<mods:identifier id="2ED239638B2F9C55F3D77372B5F107B0" type="GBIF-Dataset">9566bd4f-5391-4b65-ae39-9189f68d0f28</mods:identifier>
<mods:identifier id="D6FDA6B86F7BAB67A1D7EFEF4FCEEAEA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8D525E53B7F0687796CED9ED05EFA145" type="Zenodo-Dep">3741094</mods:identifier>
<mods:identifier id="F13FAD588C565F64103C095A1ED2050B" type="ZooBank">5E5A6906-E137-49BF-9270-B7DAF03DE57E</mods:identifier>
</mods:mods>
<treatment id="CF2F9867E118CC058C84AF81FC0DF85D" ID-DOI="http://doi.org/10.5281/zenodo.3810535" ID-GBIF-Taxon="163437560" ID-Zenodo-Dep="3810535" LSID="urn:lsid:plazi:treatment:CF2F9867E118CC058C84AF81FC0DF85D" httpUri="http://treatment.plazi.org/id/CF2F9867E118CC058C84AF81FC0DF85D" lastPageNumber="570" pageId="4" pageNumber="570">
<subSubSection id="0F9C7AFAE118CC058C84AF81FD99FD56" box="[151,608,681,708]" pageId="4" pageNumber="570" type="nomenclature">
<paragraph id="47392971E118CC058C84AF81FD99FD56" blockId="4.[151,608,681,744]" box="[151,608,681,708]" pageId="4" pageNumber="570">
<heading id="1C719E1DE118CC058C84AF81FD99FD56" bold="true" box="[151,608,681,708]" fontSize="11" level="1" pageId="4" pageNumber="570" reason="1">
<emphasis id="75F2F563E118CC058C84AF81FD99FD56" bold="true" box="[151,608,681,708]" pageId="4" pageNumber="570">
<taxonomicName id="808652F2E118CC058C84AF81FE05FD56" ID-CoL="98ZQZ" authority="Yao &amp; Li" authorityName="Xin &amp; Yao &amp; Li" authorityYear="2020" box="[151,508,681,708]" class="Arachnida" family="Pholcidae" genus="Wanniyala" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="570" phylum="Arthropoda" rank="species" species="batatota" status="sp. nov.">
<emphasis id="75F2F563E118CC058C84AF81FE7BFD56" bold="true" box="[151,386,681,708]" italics="true" pageId="4" pageNumber="570">Wanniyala batatota</emphasis>
Yao &amp; Li
</taxonomicName>
<taxonomicNameLabel id="6EC14818E118CC058E10AF82FD99FD56" box="[515,608,682,708]" pageId="4" pageNumber="570" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0F9C7AFAE118CC058C84AFE6FF01FD7A" box="[151,248,718,744]" pageId="4" pageNumber="570" type="description">
<paragraph id="47392971E118CC058C84AFE6FF01FD7A" blockId="4.[151,608,681,744]" box="[151,248,718,744]" pageId="4" pageNumber="570">
<figureCitation id="DFBD35F4E118CC058C84AFE6FF01FD7A" box="[151,248,718,744]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1911,1936]" captionStartId-1="6.[151,250,1911,1936]" captionTargetBox-0="[218,1372,181,1872]" captionTargetBox-1="[226,1362,181,1872]" captionTargetId-0="figure@5.[214,1372,181,1885]" captionTargetId-1="figure@6.[226,1362,181,1885]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Wanniyala batatota Yao &amp; Li sp. nov., holotype male. A–B. Flipped right pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis on pedipalpal femur); C–D. Distal part of flipped right procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at rounded apophysis, arrow 2 points at blunt apophysis, arrow 3 points at medially slightly protruding rs2). Scale bars: 0.20 (A–B), 0.10 (C–D)." captionText-1="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi-0="http://doi.org/10.5281/zenodo.3741098" figureDoi-1="http://doi.org/10.5281/zenodo.3741100" httpUri-0="https://zenodo.org/record/3741098/files/figure.png" httpUri-1="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Figs 3–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0F9C7AFAE118CC058C84AE39FABFFCE0" pageId="4" pageNumber="570" type="materials_examined">
<paragraph id="47392971E118CC058C84AE39FABFFCE0" blockId="4.[151,1437,784,1999]" pageId="4" pageNumber="570">
<emphasis id="75F2F563E118CC058C84AE39FEBFFCB9" bold="true" box="[151,326,784,811]" pageId="4" pageNumber="570">Type material.</emphasis>
<materialsCitation id="F7EE232CE118CC058D5DAE39FD8EFCE0" ID-GBIF-Occurrence="2595809655" collectingDate="2014-09-28" collectorName="S. Kosala" country="Sri Lanka" county="Ratnapura District" elevation="268" latitude="6.8315835" location="Isthripura Cave" longLatPrecision="1" longitude="80.37428" municipality="Ekneligoda Town" pageId="4" pageNumber="570" specimenCount="1" stateProvince="Sabaragamuwa" typeStatus="holotype">
<emphasis id="75F2F563E118CC058D5DAE39FE3DFCB9" bold="true" box="[334,452,785,811]" pageId="4" pageNumber="570">
<typeStatus id="983D97D3E118CC058D5DAE39FE46FCB9" box="[334,447,785,811]" pageId="4" pageNumber="570" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (IZCAS-Ar 39838),
<location id="42597FAAE118CC058EE1AE38FC58FCB9" LSID="urn:lsid:plazi:treatment:CF2F9867E118CC058C84AF81FC0DF85D:42597FAAE118CC058EE1AE38FC58FCB9" box="[754,929,784,811]" country="Sri Lanka" county="Ratnapura District" latitude="6.8315835" longLatPrecision="1" longitude="80.37428" municipality="Ekneligoda Town" name="Isthripura Cave" pageId="4" pageNumber="570" stateProvince="Sabaragamuwa">Isthripura Cave</location>
(
<geoCoordinate id="22B24FB6E118CC058FA2AE38FBB1FCB9" box="[945,1096,784,811]" degrees="6" direction="north" minutes="49" orientation="latitude" pageId="4" pageNumber="570" precision="1" seconds="53.7" value="6.8315835">6°49’53.7’’N</geoCoordinate>
,
<geoCoordinate id="22B24FB6E118CC058846AE39FB0EFCB9" box="[1109,1271,785,811]" degrees="80" direction="east" minutes="22" orientation="longitude" pageId="4" pageNumber="570" precision="1" seconds="27.4" value="80.37428">80°22’27.4’’E</geoCoordinate>
,
<elevation id="CCABCE42E118CC058917AE38FF49FCDD" metricMagnitude="2" metricUnit="m" metricValue="2.68" pageId="4" pageNumber="570" unit="m" value="268.0">
elevation
<quantity id="807E8494E118CC058961AE38FF49FCDD" metricMagnitude="2" metricUnit="m" metricValue="2.68" pageId="4" pageNumber="570" unit="m" value="268.0">268 m</quantity>
</elevation>
),
<location id="42597FAAE118CC058CDBAE1DFE7AFCDC" LSID="urn:lsid:plazi:treatment:CF2F9867E118CC058C84AF81FC0DF85D:42597FAAE118CC058CDBAE1DFE7AFCDC" box="[200,387,820,847]" country="Sri Lanka" county="Ratnapura District" latitude="6.8315835" longLatPrecision="1" longitude="80.37428" municipality="Ekneligoda Town" name="Batatota Village" pageId="4" pageNumber="570" stateProvince="Sabaragamuwa">Batatota Village</location>
,
<location id="42597FAAE118CC058D83AE1CFD92FCDD" LSID="urn:lsid:plazi:treatment:CF2F9867E118CC058C84AF81FC0DF85D:42597FAAE118CC058D83AE1CFD92FCDD" box="[400,619,820,847]" country="Sri Lanka" county="Ratnapura District" latitude="6.8315835" longLatPrecision="1" longitude="80.37428" municipality="Ekneligoda Town" name="Adam's Peak Area" pageId="4" pageNumber="570" stateProvince="Sabaragamuwa">Adam’s Peak Area</location>
,
<collectingMunicipality id="A75DB30BE118CC058E69AE1CFCBFFCDD" box="[634,838,820,847]" pageId="4" pageNumber="570">Ekneligoda Town</collectingMunicipality>
,
<collectingCounty id="AE5851FDE118CC058F46AE1DFBD0FCDC" box="[853,1065,820,847]" pageId="4" pageNumber="570">Ratnapura District</collectingCounty>
,
<emphasis id="75F2F563E118CC05882BAE1CFA9EFCDD" bold="true" box="[1080,1383,820,847]" pageId="4" pageNumber="570">
<collectingRegion id="8542E793E118CC05882BAE1CFA9EFCDD" box="[1080,1383,820,847]" country="Sri Lanka" name="Sabaragamuwa" pageId="4" pageNumber="570">Sabaragamuwa Province</collectingRegion>
</emphasis>
,
<emphasis id="75F2F563E118CC05896BAE1CFF1CFCE1" bold="true" pageId="4" pageNumber="570">
<collectingCountry id="3F9169E1E118CC05896BAE1CFF1CFCE1" name="Sri Lanka" pageId="4" pageNumber="570">Sri Lanka</collectingCountry>
</emphasis>
,
<date id="33380FB1E118CC058CE0AE71FE29FCE1" box="[243,464,856,883]" pageId="4" pageNumber="570" value="2014-09-28">
<collectingDate id="237CF659E118CC058CE0AE71FE29FCE1" box="[243,464,856,883]" pageId="4" pageNumber="570" value="2014-09-28">28 September 2014</collectingDate>
</date>
,
<collectorName id="EA734CA7E118CC058DC8AE71FDBFFCE0" box="[475,582,856,883]" pageId="4" pageNumber="570">S. Kosala</collectorName>
leg.
</materialsCitation>
<materialsCitation id="F7EE232CE118CC058E6EAE71FABAFCE0" ID-GBIF-Occurrence="2595809658" box="[637,1347,856,883]" collectingDate="2014-09-28" collectorName="S. Kosala" country="Sri Lanka" county="Ratnapura District" elevation="268" latitude="6.8315835" location="Isthripura Cave" longLatPrecision="1" longitude="80.37428" municipality="Ekneligoda Town" pageId="4" pageNumber="570" specimenCount="1" specimenCount-female="1" stateProvince="Sabaragamuwa" typeStatus="paratype">
<emphasis id="75F2F563E118CC058E6EAE71FD0AFCE1" bold="true" box="[637,755,857,883]" pageId="4" pageNumber="570">
<typeStatus id="983D97D3E118CC058E6EAE71FD17FCE1" box="[637,750,857,883]" pageId="4" pageNumber="570" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="5180E2F8E118CC058EE9AE71FCA2FCE0" box="[762,859,856,883]" pageId="4" pageNumber="570" type="female">1 female</specimenCount>
(IZCAS-Ar 39839), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0F9C7AFAE118CC058CD4AE55FB85FC04" box="[199,1148,892,919]" pageId="4" pageNumber="570" type="etymology">
<paragraph id="47392971E118CC058CD4AE55FB85FC04" blockId="4.[151,1437,784,1999]" box="[199,1148,892,919]" pageId="4" pageNumber="570">
<emphasis id="75F2F563E118CC058CD4AE55FEB7FC05" bold="true" box="[199,334,893,919]" pageId="4" pageNumber="570">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="983D97D3E118CC058EA6AE55FD1CFC05" box="[693,741,893,919]" pageId="4" pageNumber="570">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="0F9C7AFAE118CC058CD4AE88FAC6FBD9" pageId="4" pageNumber="570" type="diagnosis">
<paragraph id="47392971E118CC058CD4AE88FAC6FBD9" blockId="4.[151,1437,784,1999]" pageId="4" pageNumber="570">
<emphasis id="75F2F563E118CC058CD4AE88FEBBFC29" bold="true" box="[199,322,928,955]" pageId="4" pageNumber="570">Diagnosis.</emphasis>
This species can be distinguished from all known congeners by combination of following characters: retrolateral sclerite 1 of procursus bifurcated into small rounded apophysis (arrow
<quantity id="807E8494E118CC058872AEEDFB72FC4C" box="[1121,1163,964,991]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="570" unit="in" value="1.0">1 in</quantity>
<figureCitation id="DFBD35F4E118CC058881AEECFB1EFC4D" box="[1170,1255,964,991]" captionStart="FIGURE 3" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[218,1372,181,1872]" captionTargetId="figure@5.[214,1372,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Wanniyala batatota Yao &amp; Li sp. nov., holotype male. A–B. Flipped right pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis on pedipalpal femur); C–D. Distal part of flipped right procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at rounded apophysis, arrow 2 points at blunt apophysis, arrow 3 points at medially slightly protruding rs2). Scale bars: 0.20 (A–B), 0.10 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3741098" httpUri="https://zenodo.org/record/3741098/files/figure.png" pageId="4" pageNumber="570">Fig. 3D</figureCitation>
) and large blunt apophysis (arrow
<quantity id="807E8494E118CC058D71AEC1FE77FB90" box="[354,398,1000,1027]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="570" unit="in" value="2.0">2 in</quantity>
<figureCitation id="DFBD35F4E118CC058D85AEC0FE16FB91" box="[406,495,1000,1027]" captionStart="FIGURE 3" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[218,1372,181,1872]" captionTargetId="figure@5.[214,1372,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Wanniyala batatota Yao &amp; Li sp. nov., holotype male. A–B. Flipped right pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis on pedipalpal femur); C–D. Distal part of flipped right procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at rounded apophysis, arrow 2 points at blunt apophysis, arrow 3 points at medially slightly protruding rs2). Scale bars: 0.20 (A–B), 0.10 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3741098" httpUri="https://zenodo.org/record/3741098/files/figure.png" pageId="4" pageNumber="570">Fig. 3D</figureCitation>
), retrolateral sclerite 2 of procursus not bifurcated and slightly protruding medially (arrow
<quantity id="807E8494E118CC058CFBA925FEEBFBB4" box="[232,274,1036,1063]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="570" unit="in" value="3.0">3 in</quantity>
<figureCitation id="DFBD35F4E118CC058D0BA924FE96FBB5" box="[280,367,1036,1063]" captionStart="FIGURE 3" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[218,1372,181,1872]" captionTargetId="figure@5.[214,1372,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Wanniyala batatota Yao &amp; Li sp. nov., holotype male. A–B. Flipped right pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis on pedipalpal femur); C–D. Distal part of flipped right procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at rounded apophysis, arrow 2 points at blunt apophysis, arrow 3 points at medially slightly protruding rs2). Scale bars: 0.20 (A–B), 0.10 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3741098" httpUri="https://zenodo.org/record/3741098/files/figure.png" pageId="4" pageNumber="570">Fig. 3D</figureCitation>
), epigynum strongly protruding and with distinct rounded median process process (arrow in
<figureCitation id="DFBD35F4E118CC058963A924FF43FBD9" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Fig. 4A</figureCitation>
), vulva with pair of large sclerites (arrow in
<figureCitation id="DFBD35F4E118CC058EBCA918FCFFFBD9" box="[687,774,1072,1099]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Fig. 4B</figureCitation>
), and vulval pore plates nearly n-shaped (
<figureCitation id="DFBD35F4E118CC0588CAA918FACBFBD9" box="[1241,1330,1072,1099]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Fig. 4B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0F9C7AFAE118CC058CD4A97CFB93F839" pageId="4" pageNumber="570" type="description">
<paragraph id="47392971E118CC058CD4A97CFA89F918" blockId="4.[151,1437,784,1999]" pageId="4" pageNumber="570">
<emphasis id="75F2F563E118CC058CD4A97CFE66FBFD" bold="true" box="[199,415,1108,1135]" pageId="4" pageNumber="570">Description. Male</emphasis>
(
<emphasis id="75F2F563E118CC058DA3A97DFDEEFBFD" bold="true" box="[432,535,1109,1135]" pageId="4" pageNumber="570">
<typeStatus id="983D97D3E118CC058DA3A97DFDEEFBFD" box="[432,535,1109,1135]" pageId="4" pageNumber="570" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS-Ar 39838): Total length 2.47 (2.60 with clypeus), carapace 0.77 long, 0.93 wide, opisthosoma 1.70 long, 0.90 wide. Leg I: 20.17 (4.85 + 0.38 + 5.10 + 7.9 + 1.94), leg II: 11.81 (3.15 + 0.34 + 2.88 + 4.29 + 1.15), leg III: 9.21 (2.45 + 0.32 + 2.27 + 3.35 + 0.82), leg IV: 12.4 (3.42 + 0.35 + 3.14 + 4.65 + 0.84); tibia I L/d: 56. Distance PME-PME 0.13, diameter PME 0.10, distance PME-ALE 0.04, AME absent. Sternum slightly wider than long (0.66/0.61). Habitus as in
<figureCitation id="DFBD35F4E118CC058F28A9CCFC78FB6D" box="[827,897,1252,1279]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Figs 4</figureCitation>
E–F. Carapace slightly yellow, with dark brown lateral margins and dark brown median stripe; ocular area brown; clypeus slightly yellow, with pair of brown bands; sternum dark brown but slightly yellow medially. Legs slightly brown, but slightly grey on distal parts of femora and tibiae, with darker rings on subdistal parts of femora and tibiae. Opisthosoma grey, with dark brown marks. Ocular area slightly elevated. Thoracic furrow shallow but distinct. Clypeus with pair of pointed apophyses (
<figureCitation id="DFBD35F4E118CC05891CA85CFA9DFA1D" box="[1295,1380,1396,1423]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Fig. 4E</figureCitation>
) and median hump (arrow in
<figureCitation id="DFBD35F4E118CC058DB7A8B0FE00FA21" box="[420,505,1432,1459]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Fig. 4F</figureCitation>
). Chelicerae as in
<figureCitation id="DFBD35F4E118CC058ED6A8B0FCE4FA21" box="[709,797,1432,1459]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Fig. 4D</figureCitation>
, with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses directed forwards, tips of apophyses distally directed downwards (distance between tips: 0.74). Pedipalps as in
<figureCitation id="DFBD35F4E118CC058DACA8C8FDF8FA69" box="[447,513,1504,1531]" captionStart="FIGURE 3" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[218,1372,181,1872]" captionTargetId="figure@5.[214,1372,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Wanniyala batatota Yao &amp; Li sp. nov., holotype male. A–B. Flipped right pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis on pedipalpal femur); C–D. Distal part of flipped right procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at rounded apophysis, arrow 2 points at blunt apophysis, arrow 3 points at medially slightly protruding rs2). Scale bars: 0.20 (A–B), 0.10 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3741098" httpUri="https://zenodo.org/record/3741098/files/figure.png" pageId="4" pageNumber="570">Figs 3</figureCitation>
A–B; coxa unmodified; trochanter with distinct apophyses; femur with small dorsal apophysis proximally (arrow in
<figureCitation id="DFBD35F4E118CC058DECAB2CFDAFF98D" box="[511,598,1540,1567]" captionStart="FIGURE 3" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[218,1372,181,1872]" captionTargetId="figure@5.[214,1372,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Wanniyala batatota Yao &amp; Li sp. nov., holotype male. A–B. Flipped right pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis on pedipalpal femur); C–D. Distal part of flipped right procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at rounded apophysis, arrow 2 points at blunt apophysis, arrow 3 points at medially slightly protruding rs2). Scale bars: 0.20 (A–B), 0.10 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3741098" httpUri="https://zenodo.org/record/3741098/files/figure.png" pageId="4" pageNumber="570">Fig. 3B</figureCitation>
); tibia very long (0.74); procursus simple proximally but complex distally, with three transparent processes and four sclerites (
<figureCitation id="DFBD35F4E118CC058EC6AB00FCE3F9D1" box="[725,794,1576,1603]" captionStart="FIGURE 3" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[218,1372,181,1872]" captionTargetId="figure@5.[214,1372,181,1885]" captionTargetPageId="5" captionText="FIGURE 3. Wanniyala batatota Yao &amp; Li sp. nov., holotype male. A–B. Flipped right pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at dorsal apophysis on pedipalpal femur); C–D. Distal part of flipped right procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at rounded apophysis, arrow 2 points at blunt apophysis, arrow 3 points at medially slightly protruding rs2). Scale bars: 0.20 (A–B), 0.10 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3741098" httpUri="https://zenodo.org/record/3741098/files/figure.png" pageId="4" pageNumber="570">Figs 3</figureCitation>
C–D); bulb with large proximal sclerite (
<figureCitation id="DFBD35F4E118CC0588F6AB00FAC4F9D1" box="[1253,1341,1576,1603]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Fig. 4C</figureCitation>
), simple membranous embolus, and short hooked apophysis. Retrolateral trichobothrium of tibia I at 10% proximally; legs with short vertical setae on metatarsi, without spines or curved setae; tarsus I with 24 distinct pseudosegments.
</paragraph>
<paragraph id="47392971E118CC058CD4ABBDFEFCF815" blockId="4.[151,1437,784,1999]" pageId="4" pageNumber="570">
<emphasis id="75F2F563E118CC058CD4ABBDFEE7F93D" bold="true" box="[199,286,1685,1711]" pageId="4" pageNumber="570">Female</emphasis>
(IZCAS-Ar 39839): Similar to male, habitus as in
<figureCitation id="DFBD35F4E118CC058F76ABBCFC52F93D" box="[869,939,1684,1711]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Figs 4</figureCitation>
G–H. Total length 2.04 (2.16 with clypeus), carapace 0.67 long, 0.89 wide, opisthosoma 1.37 long, 0.78 wide; leg I missing. Distance PME-PME 0.10, diameter PME 0.08, distance PME-ALE 0.02, AME absent. Sternum wider than long (0.78/0.68). Clypeus unmodified. Epigynum (
<figureCitation id="DFBD35F4E118CC058D0EAA28FE8FF889" box="[285,374,1792,1819]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Fig. 4A</figureCitation>
) strongly protruding, with rounded median process (arrow in
<figureCitation id="DFBD35F4E118CC05882AAA28FB6DF889" box="[1081,1172,1792,1819]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Fig. 4A</figureCitation>
); posterior plate apparently reduced to two lateral sclerites mostly hidden behind epigynal plate and barely visible in ventral view. Vulva with sclerotized anterior ‘valve’, pair of large sclerites (arrow in
<figureCitation id="DFBD35F4E118CC058F65AA60FC37F8F1" box="[886,974,1864,1891]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Fig. 4B</figureCitation>
), and pair of nearly n-shaped pore plates (
<figureCitation id="DFBD35F4E118CC058CB3AA44FF01F815" box="[160,248,1900,1927]" captionStart="FIGURE 4" captionStartId="6.[151,250,1911,1936]" captionTargetBox="[226,1362,181,1872]" captionTargetId="figure@6.[226,1362,181,1885]" captionTargetPageId="6" captionText="FIGURE 4. Wanniyala batatota Yao &amp; Li sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. Epigynum (ventral view, arrow points at rounded median process); B. Vulva, dorsal view, arrow points at large sclerite; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view, arrow points at hump; H. Ventral view). Scale bars: 0.10 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3741100" httpUri="https://zenodo.org/record/3741100/files/figure.png" pageId="4" pageNumber="570">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="47392971E118CC058CD4AAB9FB93F839" blockId="4.[151,1437,784,1999]" box="[199,1130,1936,1963]" pageId="4" pageNumber="570">
<emphasis id="75F2F563E118CC058CD4AAB9FE7CF839" bold="true" box="[199,389,1936,1963]" pageId="4" pageNumber="570">Natural history.</emphasis>
The species was found at the entrance (twilight) zone of the cave.
</paragraph>
</subSubSection>
<subSubSection id="0F9C7AFAE118CC058CD4AA9CFC0DF85D" box="[199,1012,1972,1999]" pageId="4" pageNumber="570" type="distribution">
<paragraph id="47392971E118CC058CD4AA9CFC0DF85D" blockId="4.[151,1437,784,1999]" box="[199,1012,1972,1999]" pageId="4" pageNumber="570">
<emphasis id="75F2F563E118CC058CD4AA9CFE98F85D" bold="true" box="[199,353,1972,1999]" pageId="4" pageNumber="570">Distribution.</emphasis>
<collectingCountry id="3F9169E1E118CC058D74AA9CFE2CF85C" box="[359,469,1972,1998]" name="Sri Lanka" pageId="4" pageNumber="570">Sri Lanka</collectingCountry>
(
<collectingRegion id="8542E793E118CC058DF7AA9CFD04F85C" box="[484,765,1972,1998]" country="Sri Lanka" name="Sabaragamuwa" pageId="4" pageNumber="570">Sabaragamuwa Province</collectingRegion>
,
<typeStatus id="983D97D3E118CC058F1AAA9DFCC0F85D" box="[777,825,1973,1999]" pageId="4" pageNumber="570">type</typeStatus>
locality;
<figureCitation id="DFBD35F4E118CC058FB0AA9CFC1FF85D" box="[931,998,1972,1999]" captionStart="FIGURE 5" captionStartId="7.[151,250,1835,1860]" captionTargetBox="[151,1435,448,1810]" captionTargetId="figure@7.[151,1435,448,1810]" captionTargetPageId="7" captionText="FIGURE 5. Distribution records of the species of Wanniyala from Sri Lanka. 1 Wanniyala agrabopath Huber &amp; Benjamin, 2005, 2 W. badulla Yao &amp; Li sp. nov., 3 W. batatota Yao &amp; Li sp. nov., 4 W. hakgala Huber &amp; Benjamin, 2005, 5 W. labugama Huber, 2019, 6 W. mapalena Huber, 2019, 7 W. mudita Huber, 2019, 8 W. ohiya Huber, 2019, 9 W. orientalis Huber, 2019, 10 W. upekkha Huber, 2019, 11 W. viharekele Huber, 2019. Locality coordinates for all known species are copied from the original publications (see Huber &amp; Benjamin 2005: 3312, 3317 and Huber 2019: 34–48)." figureDoi="http://doi.org/10.5281/zenodo.3741102" httpUri="https://zenodo.org/record/3741102/files/figure.png" pageId="4" pageNumber="570">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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