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<mods:title id="6C66694E63279C62AC39A9C62D2FC20C">Description of a model tardigrade Paramacrobiotus metropolitanus sp. nov (Eutardigrada) from Japan with a summary of its life history, reproduction and genomics</mods:title>
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<mods:namePart id="6295EB87F6EB762E61AD84A64521BF36">Matsumoto, Midori</mods:namePart>
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Species 
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<emphasis id="3DA6FE104135FFC1FF65F966B9EA30E0" bold="true" box="[250,649,1736,1763]" italics="true" pageId="4" pageNumber="96">Paramacrobiotus metropolitanus</emphasis>
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Sugiura, Matsumoto &amp; Kunieda, 2022
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(
<figureCitation id="97E93E874135FFC1FF00F942BA673104" box="[159,260,1772,1799]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="5.[151,250,1913,1937]" captionStartId-1="6.[151,250,1101,1125]" captionStartId-2="7.[151,250,1718,1742]" captionStartId-3="8.[151,250,1442,1466]" captionStartId-4="10.[151,250,1735,1759]" captionStartId-5="11.[151,250,1943,1967]" captionStartId-6="12.[151,250,1665,1689]" captionTargetBox-0="[151,1435,417,1888]" captionTargetBox-1="[151,1436,633,1077]" captionTargetBox-2="[151,1435,273,1695]" captionTargetBox-3="[151,1436,1065,1419]" captionTargetBox-4="[151,1435,181,1711]" captionTargetBox-5="[248,1337,181,1918]" captionTargetBox-6="[184,1394,222,1613]" captionTargetId-0="figure-92@5.[151,1436,417,1889]" captionTargetId-1="figure-235@6.[151,1436,633,1077]" captionTargetId-2="figure-55@7.[151,1436,273,1695]" captionTargetId-3="figure-451@8.[151,1436,1065,1419]" captionTargetId-4="figure-18@10.[151,1435,181,1711]" captionTargetId-5="figure-18@11.[248,1339,181,1920]" captionTargetId-6="figure-18@12.[151,1436,198,1641]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionText-0="FIGURE 1. Morphology of Paramacrobiotus metropolitanus sp. nov. Ventral image under LM of Paramacrobiotus metropolitanus sp. nov. (A, holotype), under SEM (B). Lateral image under SEM (C). Scale bars = μm." captionText-1="FIGURE 2. Oral morphology of Paramacrobiotus metropolitanus sp. nov. Observation of mouth opening (A), ventral (B) and dorsal (C) of oral cavity armature. Macroplacoids (D) and microplacoid (E). A is taken with SEM, others are with LM. Black-filled flat arrowhead: the first band of teeth; white-filled flat arrowhead: the second band of teeth; black-filled indent arrowhead: the third band of teeth; white-filled indent arrowhead: subterminal constriction; white-filled arrows: microplacoid. Ve: ventral, Do: dorsal side. Scale bars = μm." captionText-2="FIGURE 3. Morphology of inner and external surface of legs I–III SEM images of inner and external surface of legs I–III. A panel with labelled LM is photo under LM. Asterisks indicate the cuticular bulges. Scale bars = μm." captionText-3="FIGURE 4. Morphology of leg IV The leg IV observed with LM (A) and SEM (B, C). white-filled indent arrowhead and white dashed line indicate the granulation pattern. White-filled arrows indicate the accessory points. Scale bars = μm." captionText-4="FIGURE 5. Dorsal cuticle apodemes (muscle attachments/cribriform areas)" captionText-5="FIGURE 6. Egg characters of Paramacrobiotus metropolitanus sp. nov. Egg surface structures observed with phase-contrast microscopy (A, B) and SEM (C, D, E). The structure of a process is shown in D. E shows smooth areoles on the egg surface. Scale bars = μm." captionText-6="FIGURE 7A. Phylogenetic position of the new species based on the concatenated 18S+28S+ITS-2+COI dataset. Values at nodes indicate Bayesian posterior probabilities. Scale bars = substitutions per position." figureDoi-0="http://doi.org/10.5281/zenodo.6531342" figureDoi-1="http://doi.org/10.5281/zenodo.6531344" figureDoi-2="http://doi.org/10.5281/zenodo.6531346" figureDoi-3="http://doi.org/10.5281/zenodo.6531350" figureDoi-4="http://doi.org/10.5281/zenodo.6531352" figureDoi-5="http://doi.org/10.5281/zenodo.6531354" figureDoi-6="http://doi.org/10.5281/zenodo.6531356" httpUri-0="https://zenodo.org/record/6531342/files/figure.png" httpUri-1="https://zenodo.org/record/6531344/files/figure.png" httpUri-2="https://zenodo.org/record/6531346/files/figure.png" httpUri-3="https://zenodo.org/record/6531350/files/figure.png" httpUri-4="https://zenodo.org/record/6531352/files/figure.png" httpUri-5="https://zenodo.org/record/6531354/files/figure.png" httpUri-6="https://zenodo.org/record/6531356/files/figure.png" pageId="4" pageNumber="96">Figs 1–7</figureCitation>
, 
<tableCitation id="425017B94135FFC1FE90F942BAE93104" box="[271,394,1772,1799]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStartId-0="9.[151,239,512,536]" captionStartId-1="15.[151,239,151,175]" captionTargetPageId-0="9" captionTargetPageId-1="15" captionText-0="TABLE 2. Morphometrics of Paramacrobiotus metropolitanus sp. nov. animals." captionText-1="TABLE 3. Morphometrics of Paramacrobiotus metropolitanus sp. nov. eggs." httpUri-0="http://table.plazi.org/id/5BAD728A4138FFCCFF08FDAEBF70341B" httpUri-1="http://table.plazi.org/id/5BAD728A413EFFCAFF08FF39B88836AC" pageId="4" pageNumber="96" tableUuid-0="5BAD728A4138FFCCFF08FDAEBF70341B" tableUuid-1="5BAD728A413EFFCAFF08FF39B88836AC">Tables 2–3</tableCitation>
)
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<taxonomicName id="C8D259814135FFC1FF08F89FBA0C314B" box="[151,367,1841,1865]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="96" phylum="Tardigrada" rank="species" species="undetermined">
<emphasis id="3DA6FE104135FFC1FF08F89FBA24314A" box="[151,327,1841,1865]" italics="true" pageId="4" pageNumber="96">Paramacrobiotus</emphasis>
sp.
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TYO in: 
<bibRefCitation id="6B435FF34135FFC1FE44F89FB9DA314A" author="Sugiura, K. &amp; Minato, H. &amp; Suzuki, A. C. &amp; Arakawa, K. &amp; Kunieda, T. &amp; Matsumoto, M." box="[475,697,1841,1865]" pageId="4" pageNumber="96" pagination="120 - 127" refId="ref11087" refString="Sugiura, K., Minato, H., Suzuki, A. C., Arakawa, K., Kunieda, T. &amp; Matsumoto, M. (2019) Comparison of sexual reproductive behaviors in two species of Macrobiotidae (Tardigrada: Eutardigrada). Zoological Science, 36 (2), 120 - 127. https: // doi. org / 10.2108 / zs 180103" type="journal article" year="2019">
Sugiura 
<emphasis id="3DA6FE104135FFC1FDACF89CB907314A" box="[563,612,1841,1865]" italics="true" pageId="4" pageNumber="96">et al</emphasis>
. (2019)
</bibRefCitation>
, 
<bibRefCitation id="6B435FF34135FFC1FD57F89FB8EA314A" author="Hara, Y. &amp; Shibahara, R. &amp; Kondo, K. &amp; Abe, W. &amp; Kunieda T." box="[712,905,1841,1865]" pageId="4" pageNumber="96" pagination="7" refId="ref8751" refString="Hara, Y., Shibahara, R., Kondo, K., Abe, W. &amp; Kunieda T. (2021) Parallel evolution of trehalose production machinery in anhydrobiotic animals via recurrent gene loss and horizontal transfer. Open Biology, 11, 7. https: // doi. org / 10.1098 / rsob. 200413" type="journal article" year="2021">
Hara 
<emphasis id="3DA6FE104135FFC1FC9CF89CB850314A" box="[771,819,1841,1865]" italics="true" pageId="4" pageNumber="96">et al</emphasis>
. (2021)
</bibRefCitation>
, 
<bibRefCitation id="6B435FF34135FFC1FC08F89FBFB2314A" author="Sugiura, K. &amp; Matsumoto, M." box="[919,1233,1841,1865]" pageId="4" pageNumber="96" pagination="42 - 48" refId="ref11221" refString="Sugiura, K. &amp; Matsumoto, M. (2021) Spermatozoa morphology changes during reproduction and first observation of acrosomal contact in two dioecious species of Macrobiotidae (Tardigrada: Eutardigrada). Zygote, 29, 42 - 48. https: // doi. org / 10.1017 / S 0967199420000490" type="journal article" year="2021">Sugiura &amp; Matsumoto (2021)</bibRefCitation>
, and 
<bibRefCitation id="6B435FF34135FFC1FA92F89FBA6E316B" author="Sugiura, K. &amp; Shiba, K. &amp; Inaba, K. &amp; Matsumoto, M." pageId="4" pageNumber="96" pagination="8" refId="ref11275" refString="Sugiura, K., Shiba, K., Inaba, K. &amp; Matsumoto, M. (2022) Morphological differences in tardigrade spermatozoa induce variation in gamete motility. BMC Zoology, 7, 8. https: // doi. org / 10.1186 / s 40850 - 022 - 00109 - w" type="journal article" year="2022">
Sugiura 
<emphasis id="3DA6FE104135FFC1FAF9F89CBEF5314A" box="[1382,1430,1841,1865]" italics="true" pageId="4" pageNumber="96">et al</emphasis>
. (2022)
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.
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<emphasis id="3DA6FE104135FFC1FF08F838BA0831B3" box="[151,363,1942,1968]" italics="true" pageId="4" pageNumber="96">Material examined</emphasis>
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Forty animals (NMST-Tg-281–310, NHMD-918038–918047, 
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: NMST-Tg-299) and 
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(NMST-Tg-311–333, NHMD-918048–918057) were examined under light microscopes and were deposited in 
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and 
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, 
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addition, over 30 animals and eggs were observed with the scanning electron microscope and stored in the laboratory of 
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<emphasis id="3DA6FE104134FFC0FF08FE89BA783743" box="[151,283,295,320]" italics="true" pageId="5" pageNumber="97">Description</emphasis>
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<paragraph id="0F6D22024134FFC3FF58FEE5BAEF3462" blockId="5.[151,1436,295,393]" lastBlockId="6.[151,1437,151,609]" lastPageId="6" lastPageNumber="98" pageId="5" pageNumber="97">
Measurements and statistics of animals are given in 
<tableCitation id="425017B94134FFC0FCB0FEE5B8EB3766" box="[815,904,331,357]" captionStart="TABLE 2" captionStartId="9.[151,239,512,536]" captionTargetPageId="9" captionText="TABLE 2. Morphometrics of Paramacrobiotus metropolitanus sp. nov. animals." httpUri="http://table.plazi.org/id/5BAD728A4138FFCCFF08FDAEBF70341B" pageId="5" pageNumber="97" tableUuid="5BAD728A4138FFCCFF08FDAEBF70341B">Table 2</tableCitation>
. The 
<typeStatus id="D0699CA04134FFC0FC52FEE5BF563766" box="[973,1077,331,357]" pageId="5" pageNumber="97" type="holotype">Holotype</typeStatus>
is illustrated in 
<figureCitation id="97E93E874134FFC0FB65FEE5BEFF3766" box="[1274,1436,331,357]" captionStart="FIGURE 1" captionStartId="5.[151,250,1913,1937]" captionTargetBox="[151,1435,417,1888]" captionTargetId="figure-92@5.[151,1436,417,1889]" captionTargetPageId="5" captionText="FIGURE 1. Morphology of Paramacrobiotus metropolitanus sp. nov. Ventral image under LM of Paramacrobiotus metropolitanus sp. nov. (A, holotype), under SEM (B). Lateral image under SEM (C). Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531342" httpUri="https://zenodo.org/record/6531342/files/figure.png" pageId="5" pageNumber="97">Fig. 1A and a</figureCitation>
<typeStatus id="D0699CA04134FFC0FF08FEC1BB94378A" box="[151,247,367,393]" pageId="5" pageNumber="97" type="paratype">paratype</typeStatus>
observed with 
<collectionCode id="69C3BAC74134FFC0FE36FEC1BA80378A" box="[425,483,367,393]" pageId="5" pageNumber="97">SEM</collectionCode>
in 
<figureCitation id="97E93E874134FFC0FD96FEC1B9E4378A" box="[521,647,367,393]" captionStart="FIGURE 1" captionStartId="5.[151,250,1913,1937]" captionTargetBox="[151,1435,417,1888]" captionTargetId="figure-92@5.[151,1436,417,1889]" captionTargetPageId="5" captionText="FIGURE 1. Morphology of Paramacrobiotus metropolitanus sp. nov. Ventral image under LM of Paramacrobiotus metropolitanus sp. nov. (A, holotype), under SEM (B). Lateral image under SEM (C). Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531342" httpUri="https://zenodo.org/record/6531342/files/figure.png" pageId="5" pageNumber="97">Figs 1B–C</figureCitation>
. Adult 
<taxonomicName id="C8D259814134FFC0FD45FEC1B8C2378A" authority="Sugiura &amp; Matsumoto &amp; Kunieda, 2022" authorityName="Sugiura &amp; Matsumoto &amp; Kunieda" authorityYear="2022" box="[730,929,367,393]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="97" phylum="Tardigrada" rank="species" species="metropolitanus" status="sp. nov.">
<emphasis id="3DA6FE104134FFC0FD45FEC1B8C2378A" box="[730,929,367,393]" italics="true" pageId="5" pageNumber="97">P. metropolitanus</emphasis>
</taxonomicName>
<emphasis id="3DA6FE104134FFC0FC36FEC1BF67378A" bold="true" box="[937,1028,367,393]" pageId="5" pageNumber="97">
<taxonomicNameLabel id="2695436B4134FFC0FC36FEC1BF67378A" box="[937,1028,367,393]" pageId="5" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens whitish under dark-field illumination, the juvenile transparent (data not shown). Eyes absent in live animals. Mouth anteroventral (
<figureCitation id="97E93E874137FFC3FAF5FF39BBBD36D6" captionStart="FIGURE 2" captionStartId="6.[151,250,1101,1125]" captionTargetBox="[151,1436,633,1077]" captionTargetId="figure-235@6.[151,1436,633,1077]" captionTargetPageId="6" captionText="FIGURE 2. Oral morphology of Paramacrobiotus metropolitanus sp. nov. Observation of mouth opening (A), ventral (B) and dorsal (C) of oral cavity armature. Macroplacoids (D) and microplacoid (E). A is taken with SEM, others are with LM. Black-filled flat arrowhead: the first band of teeth; white-filled flat arrowhead: the second band of teeth; black-filled indent arrowhead: the third band of teeth; white-filled indent arrowhead: subterminal constriction; white-filled arrows: microplacoid. Ve: ventral, Do: dorsal side. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531344" httpUri="https://zenodo.org/record/6531344/files/figure.png" pageId="6" pageNumber="98">Figs 2A–C</figureCitation>
). Buccopharyngeal apparatus the 
<taxonomicName id="C8D259814137FFC3FDFAFF15B99036D6" authorityName="C. A. S. Schultze" authorityYear="1834" box="[613,755,187,213]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="98" phylum="Tardigrada" rank="genus">
<emphasis id="3DA6FE104137FFC3FDFAFF15B99036D6" box="[613,755,187,213]" italics="true" pageId="6" pageNumber="98">Macrobiotus</emphasis>
</taxonomicName>
type, with ten peribuccal lamellae around the mouth (
<figureCitation id="97E93E874137FFC3FAF0FF15BBDE36FA" captionStart="FIGURE 2" captionStartId="6.[151,250,1101,1125]" captionTargetBox="[151,1436,633,1077]" captionTargetId="figure-235@6.[151,1436,633,1077]" captionTargetPageId="6" captionText="FIGURE 2. Oral morphology of Paramacrobiotus metropolitanus sp. nov. Observation of mouth opening (A), ventral (B) and dorsal (C) of oral cavity armature. Macroplacoids (D) and microplacoid (E). A is taken with SEM, others are with LM. Black-filled flat arrowhead: the first band of teeth; white-filled flat arrowhead: the second band of teeth; black-filled indent arrowhead: the third band of teeth; white-filled indent arrowhead: subterminal constriction; white-filled arrows: microplacoid. Ve: ventral, Do: dorsal side. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531344" httpUri="https://zenodo.org/record/6531344/files/figure.png" pageId="6" pageNumber="98">Fig. 2A</figureCitation>
). Numerous conical teeth present, forming the first band of teeth posterior to the peribuccal lamellae. Conical teeth present at just behind the ring fold, forming the second band, and ridges parallel to the main axis of the buccal tube as the third band (
<figureCitation id="97E93E874137FFC3FE01FE89B97C3742" box="[414,543,295,321]" captionStart="FIGURE 2" captionStartId="6.[151,250,1101,1125]" captionTargetBox="[151,1436,633,1077]" captionTargetId="figure-235@6.[151,1436,633,1077]" captionTargetPageId="6" captionText="FIGURE 2. Oral morphology of Paramacrobiotus metropolitanus sp. nov. Observation of mouth opening (A), ventral (B) and dorsal (C) of oral cavity armature. Macroplacoids (D) and microplacoid (E). A is taken with SEM, others are with LM. Black-filled flat arrowhead: the first band of teeth; white-filled flat arrowhead: the second band of teeth; black-filled indent arrowhead: the third band of teeth; white-filled indent arrowhead: subterminal constriction; white-filled arrows: microplacoid. Ve: ventral, Do: dorsal side. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531344" httpUri="https://zenodo.org/record/6531344/files/figure.png" pageId="6" pageNumber="98">Figs 2A–C</figureCitation>
). Approximately 20–30 small and sharp ridges on the second band, and two to four larger ridges on the third band (
<figureCitation id="97E93E874137FFC3FDADFEE5B9CD3766" box="[562,686,331,357]" captionStart="FIGURE 2" captionStartId="6.[151,250,1101,1125]" captionTargetBox="[151,1436,633,1077]" captionTargetId="figure-235@6.[151,1436,633,1077]" captionTargetPageId="6" captionText="FIGURE 2. Oral morphology of Paramacrobiotus metropolitanus sp. nov. Observation of mouth opening (A), ventral (B) and dorsal (C) of oral cavity armature. Macroplacoids (D) and microplacoid (E). A is taken with SEM, others are with LM. Black-filled flat arrowhead: the first band of teeth; white-filled flat arrowhead: the second band of teeth; black-filled indent arrowhead: the third band of teeth; white-filled indent arrowhead: subterminal constriction; white-filled arrows: microplacoid. Ve: ventral, Do: dorsal side. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531344" httpUri="https://zenodo.org/record/6531344/files/figure.png" pageId="6" pageNumber="98">Fig. 2A–C</figureCitation>
). Three macroplacoids present in the pharynx, the length sequence 2&lt;1&lt;3 (
<figureCitation id="97E93E874137FFC3FF70FEC1BA2A378A" box="[239,329,367,393]" captionStart="FIGURE 2" captionStartId="6.[151,250,1101,1125]" captionTargetBox="[151,1436,633,1077]" captionTargetId="figure-235@6.[151,1436,633,1077]" captionTargetPageId="6" captionText="FIGURE 2. Oral morphology of Paramacrobiotus metropolitanus sp. nov. Observation of mouth opening (A), ventral (B) and dorsal (C) of oral cavity armature. Macroplacoids (D) and microplacoid (E). A is taken with SEM, others are with LM. Black-filled flat arrowhead: the first band of teeth; white-filled flat arrowhead: the second band of teeth; black-filled indent arrowhead: the third band of teeth; white-filled indent arrowhead: subterminal constriction; white-filled arrows: microplacoid. Ve: ventral, Do: dorsal side. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531344" httpUri="https://zenodo.org/record/6531344/files/figure.png" pageId="6" pageNumber="98">Fig. 2D</figureCitation>
). The third macroplacoid with a subterminal construction. Microplacoid present (
<figureCitation id="97E93E874137FFC3FB50FEC1BE45378A" box="[1231,1318,367,393]" captionStart="FIGURE 2" captionStartId="6.[151,250,1101,1125]" captionTargetBox="[151,1436,633,1077]" captionTargetId="figure-235@6.[151,1436,633,1077]" captionTargetPageId="6" captionText="FIGURE 2. Oral morphology of Paramacrobiotus metropolitanus sp. nov. Observation of mouth opening (A), ventral (B) and dorsal (C) of oral cavity armature. Macroplacoids (D) and microplacoid (E). A is taken with SEM, others are with LM. Black-filled flat arrowhead: the first band of teeth; white-filled flat arrowhead: the second band of teeth; black-filled indent arrowhead: the third band of teeth; white-filled indent arrowhead: subterminal constriction; white-filled arrows: microplacoid. Ve: ventral, Do: dorsal side. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531344" httpUri="https://zenodo.org/record/6531344/files/figure.png" pageId="6" pageNumber="98">Fig. 2E</figureCitation>
). Claws of the 
<taxonomicName id="C8D259814137FFC3FF5EFE3DBA4837AE" authorityName="C. A. S. Schultze" authorityYear="1834" box="[193,299,403,429]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="98" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="3DA6FE104137FFC3FF5EFE3DBA4837AE" box="[193,299,403,429]" italics="true" pageId="6" pageNumber="98">hufelandi</emphasis>
</taxonomicName>
type (Y-shaped, 
<figureCitation id="97E93E874137FFC3FE74FE3DB93237AE" box="[491,593,403,429]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="7.[151,250,1718,1742]" captionStartId-1="8.[151,250,1442,1466]" captionTargetBox-0="[151,1435,273,1695]" captionTargetBox-1="[151,1436,1065,1419]" captionTargetId-0="figure-55@7.[151,1436,273,1695]" captionTargetId-1="figure-451@8.[151,1436,1065,1419]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 3. Morphology of inner and external surface of legs I–III SEM images of inner and external surface of legs I–III. A panel with labelled LM is photo under LM. Asterisks indicate the cuticular bulges. Scale bars = μm." captionText-1="FIGURE 4. Morphology of leg IV The leg IV observed with LM (A) and SEM (B, C). white-filled indent arrowhead and white dashed line indicate the granulation pattern. White-filled arrows indicate the accessory points. Scale bars = μm." figureDoi-0="http://doi.org/10.5281/zenodo.6531346" figureDoi-1="http://doi.org/10.5281/zenodo.6531350" httpUri-0="https://zenodo.org/record/6531346/files/figure.png" httpUri-1="https://zenodo.org/record/6531350/files/figure.png" pageId="6" pageNumber="98">Figs 3–4</figureCitation>
). Smooth cuticle with a cuticular bulge on the inner side of legs I–III (
<figureCitation id="97E93E874137FFC3FAF0FE3DBBC537D2" captionStart="FIGURE 3" captionStartId="7.[151,250,1718,1742]" captionTargetBox="[151,1435,273,1695]" captionTargetId="figure-55@7.[151,1436,273,1695]" captionTargetPageId="7" captionText="FIGURE 3. Morphology of inner and external surface of legs I–III SEM images of inner and external surface of legs I–III. A panel with labelled LM is photo under LM. Asterisks indicate the cuticular bulges. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531346" httpUri="https://zenodo.org/record/6531346/files/figure.png" pageId="6" pageNumber="98">Fig. 3</figureCitation>
), whereas granulation on the inner surface of leg IV visible under both 
<collectionCode id="69C3BAC74137FFC3FC77FE19BF4037D2" box="[1000,1059,439,465]" country="Poland" httpUri="http://grbio.org/cool/f3v1-41fg" name="Polish Collection of Microorganisms" pageId="6" pageNumber="98">PCM</collectionCode>
and 
<collectionCode id="69C3BAC74137FFC3FBC0FE19BFFA37D2" box="[1119,1177,439,465]" pageId="6" pageNumber="98">SEM</collectionCode>
(
<figureCitation id="97E93E874137FFC3FB35FE19BE4F37D2" box="[1194,1324,439,465]" captionStart="FIGURE 4" captionStartId="8.[151,250,1442,1466]" captionTargetBox="[151,1436,1065,1419]" captionTargetId="figure-451@8.[151,1436,1065,1419]" captionTargetPageId="8" captionText="FIGURE 4. Morphology of leg IV The leg IV observed with LM (A) and SEM (B, C). white-filled indent arrowhead and white dashed line indicate the granulation pattern. White-filled arrows indicate the accessory points. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531350" httpUri="https://zenodo.org/record/6531350/files/figure.png" pageId="6" pageNumber="98">Figs 4A–B</figureCitation>
). Primary branches with two distinct accessory points (
<figureCitation id="97E93E874137FFC3FD0AFE75B99337F6" box="[661,752,475,501]" captionStart="FIGURE 4" captionStartId="8.[151,250,1442,1466]" captionTargetBox="[151,1436,1065,1419]" captionTargetId="figure-451@8.[151,1436,1065,1419]" captionTargetPageId="8" captionText="FIGURE 4. Morphology of leg IV The leg IV observed with LM (A) and SEM (B, C). white-filled indent arrowhead and white dashed line indicate the granulation pattern. White-filled arrows indicate the accessory points. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531350" httpUri="https://zenodo.org/record/6531350/files/figure.png" pageId="6" pageNumber="98">Fig. 4C</figureCitation>
). A long common tract with a stalk connects the claw to the lunule. Lunules smooth (
<figureCitation id="97E93E874137FFC3FE2BFE51B96C341A" box="[436,527,511,537]" captionStart="FIGURE 4" captionStartId="8.[151,250,1442,1466]" captionTargetBox="[151,1436,1065,1419]" captionTargetId="figure-451@8.[151,1436,1065,1419]" captionTargetPageId="8" captionText="FIGURE 4. Morphology of leg IV The leg IV observed with LM (A) and SEM (B, C). white-filled indent arrowhead and white dashed line indicate the granulation pattern. White-filled arrows indicate the accessory points. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531350" httpUri="https://zenodo.org/record/6531350/files/figure.png" pageId="6" pageNumber="98">Fig. 4C</figureCitation>
). Smooth body cuticle without pores, sculpturing or granulation (
<figureCitation id="97E93E874137FFC3FB64FE51BE27341A" box="[1275,1348,511,537]" captionStart="FIGURE 1" captionStartId="5.[151,250,1913,1937]" captionTargetBox="[151,1435,417,1888]" captionTargetId="figure-92@5.[151,1436,417,1889]" captionTargetPageId="5" captionText="FIGURE 1. Morphology of Paramacrobiotus metropolitanus sp. nov. Ventral image under LM of Paramacrobiotus metropolitanus sp. nov. (A, holotype), under SEM (B). Lateral image under SEM (C). Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531342" httpUri="https://zenodo.org/record/6531342/files/figure.png" pageId="6" pageNumber="98">Figs 1</figureCitation>
and 
<figureCitation id="97E93E874137FFC3FAE1FE50BEED341A" box="[1406,1422,510,537]" captionStart="FIGURE 5" captionStartId="10.[151,250,1735,1759]" captionTargetBox="[151,1435,181,1711]" captionTargetId="figure-18@10.[151,1435,181,1711]" captionTargetPageId="10" captionText="FIGURE 5. Dorsal cuticle apodemes (muscle attachments/cribriform areas)" figureDoi="http://doi.org/10.5281/zenodo.6531352" httpUri="https://zenodo.org/record/6531352/files/figure.png" pageId="6" pageNumber="98">5</figureCitation>
). Muscle attachments (apodemes/cribriform areas) clearly visible on the cuticle, with two holes, each with densely packed pores (
<figureCitation id="97E93E874137FFC3FEA6FDE9BA1D3462" box="[313,382,582,609]" captionStart="FIGURE 5" captionStartId="10.[151,250,1735,1759]" captionTargetBox="[151,1435,181,1711]" captionTargetId="figure-18@10.[151,1435,181,1711]" captionTargetPageId="10" captionText="FIGURE 5. Dorsal cuticle apodemes (muscle attachments/cribriform areas)" figureDoi="http://doi.org/10.5281/zenodo.6531352" httpUri="https://zenodo.org/record/6531352/files/figure.png" pageId="6" pageNumber="98">Fig. 5</figureCitation>
).
</paragraph>
<caption id="5BAD728A4134FFC0FF08F8D7BA1331DA" ID-DOI="http://doi.org/10.5281/zenodo.6531342" ID-Zenodo-Dep="6531342" httpUri="https://zenodo.org/record/6531342/files/figure.png" pageId="5" pageNumber="97" startId="5.[151,250,1913,1937]" targetBox="[151,1435,417,1888]" targetPageId="5">
<paragraph id="0F6D22024134FFC0FF08F8D7B8133192" blockId="5.[151,1436,1912,2009]" box="[151,880,1912,1937]" pageId="5" pageNumber="97">
<emphasis id="3DA6FE104134FFC0FF08F8D7B8133192" bold="true" box="[151,880,1912,1937]" pageId="5" pageNumber="97">
FIGURE 1. Morphology of 
<taxonomicName id="C8D259814134FFC0FE5EF8D6B87A3193" authority="Sugiura &amp; Matsumoto &amp; Kunieda, 2022" authorityName="Sugiura &amp; Matsumoto &amp; Kunieda" authorityYear="2022" box="[449,793,1912,1936]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="97" phylum="Tardigrada" rank="species" species="metropolitanus" status="sp. nov.">
<emphasis id="3DA6FE104134FFC0FE5EF8D6B87A3193" bold="true" box="[449,793,1912,1936]" italics="true" pageId="5" pageNumber="97">Paramacrobiotus metropolitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2695436B4134FFC0FC80F8D7B8133192" box="[799,880,1913,1937]" pageId="5" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="0F6D22024134FFC0FF08F833BA1331DA" blockId="5.[151,1436,1912,2009]" pageId="5" pageNumber="97">
Ventral image under LM of 
<taxonomicName id="C8D259814134FFC0FE2FF833B99C31B6" authority="Sugiura &amp; Matsumoto &amp; Kunieda, 2022" authorityName="Sugiura &amp; Matsumoto &amp; Kunieda" authorityYear="2022" box="[432,767,1949,1973]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="97" phylum="Tardigrada" rank="species" species="metropolitanus" status="sp. nov.">
<emphasis id="3DA6FE104134FFC0FE2FF833B99C31B6" box="[432,767,1949,1973]" italics="true" pageId="5" pageNumber="97">Paramacrobiotus metropolitanus</emphasis>
</taxonomicName>
<emphasis id="3DA6FE104134FFC0FC9AF833B83631B6" bold="true" box="[773,853,1949,1973]" pageId="5" pageNumber="97">
<taxonomicNameLabel id="2695436B4134FFC0FC9AF833B83631B6" box="[773,853,1949,1973]" pageId="5" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A, holotype), under SEM (B). Lateral image under SEM (C). Scale bars = μm.
</paragraph>
</caption>
<caption id="5BAD728A4137FFC3FF08FBE3BF2A32F6" ID-DOI="http://doi.org/10.5281/zenodo.6531344" ID-Zenodo-Dep="6531344" httpUri="https://zenodo.org/record/6531344/files/figure.png" pageId="6" pageNumber="98" startId="6.[151,250,1101,1125]" targetBox="[151,1436,633,1077]" targetPageId="6">
<paragraph id="0F6D22024137FFC3FF08FBE3B8C53266" blockId="6.[151,1437,1100,1269]" box="[151,934,1100,1125]" pageId="6" pageNumber="98">
<emphasis id="3DA6FE104137FFC3FF08FBE3B8C53266" bold="true" box="[151,934,1100,1125]" pageId="6" pageNumber="98">
FIGURE 2. Oral morphology of 
<taxonomicName id="C8D259814137FFC3FE68FBE2B8333267" authority="Sugiura &amp; Matsumoto &amp; Kunieda, 2022" authorityName="Sugiura &amp; Matsumoto &amp; Kunieda" authorityYear="2022" box="[503,848,1100,1124]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="98" phylum="Tardigrada" rank="species" species="metropolitanus" status="sp. nov.">
<emphasis id="3DA6FE104137FFC3FE68FBE2B8333267" bold="true" box="[503,848,1100,1124]" italics="true" pageId="6" pageNumber="98">Paramacrobiotus metropolitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2695436B4137FFC3FCC9FBE3B8C53266" box="[854,934,1101,1125]" pageId="6" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="0F6D22024137FFC3FF08FBDFBF2A32F6" blockId="6.[151,1437,1100,1269]" pageId="6" pageNumber="98">Observation of mouth opening (A), ventral (B) and dorsal (C) of oral cavity armature. Macroplacoids (D) and microplacoid (E). A is taken with SEM, others are with LM. Black-filled flat arrowhead: the first band of teeth; white-filled flat arrowhead: the second band of teeth; black-filled indent arrowhead: the third band of teeth; white-filled indent arrowhead: subterminal constriction; white-filled arrows: microplacoid. Ve: ventral, Do: dorsal side. Scale bars = μm.</paragraph>
</caption>
<paragraph id="0F6D22024137FFC3FF58FA8ABA3833CE" blockId="6.[151,1437,1315,1485]" pageId="6" pageNumber="98">
Eggs laid freely, yellow yolk seen internally and transparent conical processes on the surface present in 
<collectionCode id="69C3BAC74137FFC3FAFEFA8DBEFF333E" box="[1377,1436,1315,1341]" country="Poland" httpUri="http://grbio.org/cool/f3v1-41fg" name="Polish Collection of Microorganisms" pageId="6" pageNumber="98">PCM</collectionCode>
(
<figureCitation id="97E93E874137FFC3FF00FAE9BA7C3361" box="[159,287,1351,1378]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[248,1337,181,1918]" captionTargetId="figure-18@11.[248,1339,181,1920]" captionTargetPageId="11" captionText="FIGURE 6. Egg characters of Paramacrobiotus metropolitanus sp. nov. Egg surface structures observed with phase-contrast microscopy (A, B) and SEM (C, D, E). The structure of a process is shown in D. E shows smooth areoles on the egg surface. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531354" httpUri="https://zenodo.org/record/6531354/files/figure.png" pageId="6" pageNumber="98">Figs 6A–B</figureCitation>
). Reticulated pattern with annual rings present on the surface of the processes, with slightly rough tops (
<figureCitation id="97E93E874137FFC3FF00FAC5BA423385" box="[159,289,1387,1414]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[248,1337,181,1918]" captionTargetId="figure-18@11.[248,1339,181,1920]" captionTargetPageId="11" captionText="FIGURE 6. Egg characters of Paramacrobiotus metropolitanus sp. nov. Egg surface structures observed with phase-contrast microscopy (A, B) and SEM (C, D, E). The structure of a process is shown in D. E shows smooth areoles on the egg surface. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531354" httpUri="https://zenodo.org/record/6531354/files/figure.png" pageId="6" pageNumber="98">Figs 6C–D</figureCitation>
). Approximately 10 areolae around each process on the surface of the egg (
<figureCitation id="97E93E874137FFC3FBE2FAC5BFB43385" box="[1149,1239,1387,1414]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[248,1337,181,1918]" captionTargetId="figure-18@11.[248,1339,181,1920]" captionTargetPageId="11" captionText="FIGURE 6. Egg characters of Paramacrobiotus metropolitanus sp. nov. Egg surface structures observed with phase-contrast microscopy (A, B) and SEM (C, D, E). The structure of a process is shown in D. E shows smooth areoles on the egg surface. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531354" httpUri="https://zenodo.org/record/6531354/files/figure.png" pageId="6" pageNumber="98">Fig. 6B</figureCitation>
). Smooth surface of areoles (
<figureCitation id="97E93E874137FFC3FE87FA21BA1233A9" box="[280,369,1423,1450]" captionStart="FIGURE 6" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[248,1337,181,1918]" captionTargetId="figure-18@11.[248,1339,181,1920]" captionTargetPageId="11" captionText="FIGURE 6. Egg characters of Paramacrobiotus metropolitanus sp. nov. Egg surface structures observed with phase-contrast microscopy (A, B) and SEM (C, D, E). The structure of a process is shown in D. E shows smooth areoles on the egg surface. Scale bars = μm." figureDoi="http://doi.org/10.5281/zenodo.6531354" httpUri="https://zenodo.org/record/6531354/files/figure.png" pageId="6" pageNumber="98">Fig. 6E</figureCitation>
). Morphometric data are available in 
<tableCitation id="425017B94137FFC3FCBEFA21B81B33AA" box="[801,888,1423,1449]" captionStart="TABLE 3" captionStartId="15.[151,239,151,175]" captionTargetPageId="15" captionText="TABLE 3. Morphometrics of Paramacrobiotus metropolitanus sp. nov. eggs." httpUri="http://table.plazi.org/id/5BAD728A413EFFCAFF08FF39B88836AC" pageId="6" pageNumber="98" tableUuid="5BAD728A413EFFCAFF08FF39B88836AC">Table 3</tableCitation>
. All morphometric data are listed in Supporting Materials 
<collectionCode id="69C3BAC74137FFC3FE97FA1DBA5533CE" box="[264,310,1459,1485]" country="Malaysia" name="Sarawak Museum" pageId="6" pageNumber="98">SM</collectionCode>
.01.
</paragraph>
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<emphasis id="3DA6FE104137FFC3FF08FA52BA443016" box="[151,295,1531,1557]" italics="true" pageId="6" pageNumber="98">
<typeStatus id="D0699CA04137FFC3FF08FA52BBA83016" box="[151,203,1532,1557]" pageId="6" pageNumber="98">Type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="0F6D22024137FFC3FF58F9B1BA053082" blockId="6.[151,1436,1531,1666]" pageId="6" pageNumber="98">
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, 
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; 
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asl: 
<collectingCountry id="77C562924137FFC3FD3EF98EB9873039" box="[673,740,1568,1594]" name="Japan" pageId="6" pageNumber="98">Japan</collectingCountry>
, Honshu mainland, Kanto region, 
<collectingRegion id="CD16ECE04137FFC3FBE5F9B1BFA7303A" box="[1146,1220,1567,1593]" country="Japan" name="Tokyo" pageId="6" pageNumber="98">Tokyo</collectingRegion>
, Nishi-Tokyo city, Tozenji temple; urban cemetery; bamboo leaf litter. The locality described in 
<bibRefCitation id="6B435FF34137FFC3FC68F9EDBFBD305E" author="Sugiura, K. &amp; Minato, H. &amp; Suzuki, A. C. &amp; Arakawa, K. &amp; Kunieda, T. &amp; Matsumoto, M." box="[1015,1246,1603,1630]" pageId="6" pageNumber="98" pagination="120 - 127" refId="ref11087" refString="Sugiura, K., Minato, H., Suzuki, A. C., Arakawa, K., Kunieda, T. &amp; Matsumoto, M. (2019) Comparison of sexual reproductive behaviors in two species of Macrobiotidae (Tardigrada: Eutardigrada). Zoological Science, 36 (2), 120 - 127. https: // doi. org / 10.2108 / zs 180103" type="journal article" year="2019">
Sugiura 
<emphasis id="3DA6FE104137FFC3FBCAF9EABFE5305E" box="[1109,1158,1603,1629]" italics="true" pageId="6" pageNumber="98">et al</emphasis>
. (2019)
</bibRefCitation>
was corrected in 
<bibRefCitation id="6B435FF34137FFC3FF08F9C6BA023082" author="Hara, Y. &amp; Shibahara, R. &amp; Kondo, K. &amp; Abe, W. &amp; Kunieda T." box="[151,353,1639,1666]" pageId="6" pageNumber="98" pagination="7" refId="ref8751" refString="Hara, Y., Shibahara, R., Kondo, K., Abe, W. &amp; Kunieda T. (2021) Parallel evolution of trehalose production machinery in anhydrobiotic animals via recurrent gene loss and horizontal transfer. Open Biology, 11, 7. https: // doi. org / 10.1098 / rsob. 200413" type="journal article" year="2021">
Hara 
<emphasis id="3DA6FE104137FFC3FF4AF9C6BA653082" box="[213,262,1639,1665]" italics="true" pageId="6" pageNumber="98">et al</emphasis>
. (2021)
</bibRefCitation>
.
</paragraph>
<paragraph id="0F6D22024137FFC3FF08F901BA6C30CA" blockId="6.[151,1436,1711,1810]" box="[151,271,1711,1737]" pageId="6" pageNumber="98">
<emphasis id="3DA6FE104137FFC3FF08F901BA6C30CA" box="[151,271,1711,1737]" italics="true" pageId="6" pageNumber="98">Etymology</emphasis>
</paragraph>
<paragraph id="0F6D22024137FFC3FF58F97DB9953112" blockId="6.[151,1436,1711,1810]" pageId="6" pageNumber="98">
<materialsCitation id="BFBA285F4137FFC3FF58F97DB9913112" ID-GBIF-Occurrence="3765284302" location="Metropolitan Area" pageId="6" pageNumber="98" specimenCount="1" stateProvince="Tokyo">
The species name of “ 
<taxonomicName id="C8D259814137FFC3FE48F97DB9E330EE" authorityName="Sugiura &amp; Matsumoto &amp; Kunieda" authorityYear="2022" box="[471,640,1747,1773]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="98" phylum="Tardigrada" rank="species" species="metropolitanus">
<emphasis id="3DA6FE104137FFC3FE48F97DB9E330EE" box="[471,640,1747,1773]" italics="true" pageId="6" pageNumber="98">metropolitanus</emphasis>
</taxonomicName>
” refers to the 
<collectingRegion id="CD16ECE04137FFC3FCAAF97DB81F30EE" box="[821,892,1747,1773]" country="Japan" name="Tokyo" pageId="6" pageNumber="98">Tokyo</collectingRegion>
<location id="0A0D74D94137FFC3FC16F97DBF3F30EE" LSID="urn:lsid:plazi:treatment:877B93144135FFCFFF08F966B953314F:0A0D74D94137FFC3FC16F97DBF3F30EE" box="[905,1116,1747,1773]" name="Metropolitan Area" pageId="6" pageNumber="98" stateProvince="Tokyo">Metropolitan Area</location>
(
<collectingRegion id="CD16ECE04137FFC3FBEEF97DBFD930EE" box="[1137,1210,1747,1773]" country="Japan" name="Tokyo" pageId="6" pageNumber="98">Tokyo</collectingRegion>
Cho-mei mushi in Japanese), the region where the species was collected
</materialsCitation>
.
</paragraph>
<paragraph id="0F6D22024137FFC3FF08F891BA59315A" blockId="6.[151,1436,1855,2026]" box="[151,314,1855,1881]" pageId="6" pageNumber="98">
<emphasis id="3DA6FE104137FFC3FF08F891BA59315A" box="[151,314,1855,1881]" italics="true" pageId="6" pageNumber="98">Sequence data</emphasis>
</paragraph>
<paragraph id="0F6D22024137FFC2FF58F8CAB85A36F9" blockId="6.[151,1436,1855,2026]" lastBlockId="7.[151,1436,151,250]" lastPageId="7" lastPageNumber="99" pageId="6" pageNumber="98">
<materialsCitation id="BFBA285F4137FFC2FF58F8CAB85636F9" ID-GBIF-Occurrence="3765284303" accessionNumber="LC637243, LC649795, LC649797, LC649794, LC637242, LC649796" collectionCode="COI" lastPageId="7" lastPageNumber="99" pageId="6" pageNumber="98" specimenCount="1">
We confirmed 100% matches between the sequence of PCR-amplicon and the sequence retrieved from the published genome of the species for 18S rDNA, 28S rDNA and ITS-2 sequences. As for COI sequence, one mismatched base was found between the sequence of 658 bp PCR-amplicon and the sequence retrieved from the genome. The sequences were deposited in GenBank with IDs: 
<accessionNumber id="1081BFE14137FFC3FCFFF861B8BB31EA" box="[864,984,1999,2025]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC637243" pageId="6" pageNumber="98">LC637243</accessionNumber>
(18S rDNA), 
<accessionNumber id="1081BFE14137FFC3FBE4F861BF9031E9" box="[1147,1267,1999,2026]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC649795" pageId="6" pageNumber="98">LC649795</accessionNumber>
(28S rDNA of the PCR amplicon using primer-set of 28S_Eutar_F and 28SR0090), 
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(28S rDNA of the PCR amplicon using primer-set of 28SF0001 and 28SR0990), 
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(ITS-2), 
<accessionNumber id="1081BFE14136FFC2FC00FF15BF7536D6" box="[927,1046,187,213]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC637242" pageId="7" pageNumber="99">LC637242</accessionNumber>
(
<collectionCode id="69C3BAC74136FFC2FBB7FF15BF3936D6" box="[1064,1114,187,213]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="7" pageNumber="99" type="Herbarium">COI</collectionCode>
sequence deduced from the genome), and 
<accessionNumber id="1081BFE14136FFC2FEA8FF71BACD36FA" box="[311,430,223,249]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/LC649796" pageId="7" pageNumber="99">LC649796</accessionNumber>
(
<collectionCode id="69C3BAC74136FFC2FE22FF71BA8C36FA" box="[445,495,223,249]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="7" pageNumber="99" type="Herbarium">COI</collectionCode>
sequence of PCR-amplicon)
</materialsCitation>
.
</paragraph>
<caption id="5BAD728A4136FFC2FF08F918BA933114" ID-DOI="http://doi.org/10.5281/zenodo.6531346" ID-Zenodo-Dep="6531346" httpUri="https://zenodo.org/record/6531346/files/figure.png" pageId="7" pageNumber="99" startId="7.[151,250,1718,1742]" targetBox="[151,1435,273,1695]" targetPageId="7">
<paragraph id="0F6D22024136FFC2FF08F918B80030CD" blockId="7.[151,1436,1718,1815]" box="[151,867,1718,1742]" pageId="7" pageNumber="99">
<emphasis id="3DA6FE104136FFC2FF08F918B80030CD" bold="true" box="[151,867,1718,1742]" pageId="7" pageNumber="99">FIGURE 3. Morphology of inner and external surface of legs I–III</emphasis>
</paragraph>
<paragraph id="0F6D22024136FFC2FF08F974BA933114" blockId="7.[151,1436,1718,1815]" pageId="7" pageNumber="99">SEM images of inner and external surface of legs I–III. A panel with labelled LM is photo under LM. Asterisks indicate the cuticular bulges. Scale bars = μm.</paragraph>
</caption>
<paragraph id="0F6D22024136FFC2FF08F8EAB969315D" blockId="7.[151,1436,1860,2031]" box="[151,522,1860,1886]" pageId="7" pageNumber="99">
<emphasis id="3DA6FE104136FFC2FF08F8EAB969315D" box="[151,522,1860,1886]" italics="true" pageId="7" pageNumber="99">Phenotypic differential diagnosis</emphasis>
</paragraph>
<paragraph id="0F6D22024136FFC2FF58F8C6B92631C8" blockId="7.[151,1436,1860,2031]" pageId="7" pageNumber="99">
The new species 
<taxonomicName id="C8D259814136FFC2FE0CF8C7B9383181" authority="Sugiura &amp; Matsumoto &amp; Kunieda, 2022" authorityName="Sugiura &amp; Matsumoto &amp; Kunieda" authorityYear="2022" box="[403,603,1896,1922]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="99" phylum="Tardigrada" rank="species" species="metropolitanus" status="sp. nov.">
<emphasis id="3DA6FE104136FFC2FE0CF8C7B9383181" box="[403,603,1896,1922]" italics="true" pageId="7" pageNumber="99">P. metropolitanus</emphasis>
</taxonomicName>
<emphasis id="3DA6FE104136FFC2FDF9F8C7B9A03180" bold="true" box="[614,707,1897,1923]" pageId="7" pageNumber="99">
<taxonomicNameLabel id="2695436B4136FFC2FDF9F8C7B9A03180" box="[614,707,1897,1923]" pageId="7" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with microplacoid, smooth lunules, and its egg without pored surface on the areolae is similar to the following seven species but differs specifically from (data from original descriptions of the compared species):
</paragraph>
</subSubSection>
<subSubSection id="47C871894136FFCDFF58F87BBB9A3211" lastPageId="8" lastPageNumber="100" pageId="7" pageNumber="99" type="reference_group">
<paragraph id="0F6D22024136FFCDFF58F87BBB9A36D5" blockId="7.[151,1436,1860,2031]" lastBlockId="8.[151,1437,151,1042]" lastPageId="8" lastPageNumber="100" pageId="7" pageNumber="99">
• 
<taxonomicName id="C8D259814136FFC2FF68F87BB81931EC" ID-CoL="340270aa-a889-4581-a627-f37151c96a54" authority="Pilato, Binda, Napolitano &amp; Moncada 2001" authorityName="Pilato, Binda, Napolitano &amp; Moncada" authorityYear="2001" box="[247,890,2004,2031]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="99" phylum="Tardigrada" rank="species" species="danielae">
<emphasis id="3DA6FE104136FFC2FF68F87BBA1431EC" bold="true" box="[247,375,2004,2031]" italics="true" pageId="7" pageNumber="99">P. danielae</emphasis>
<bibRefCitation id="6B435FF34136FFC2FE1EF87AB81931EC" author="Pilato, G. &amp; Binda, M. G. &amp; Napolitano, A. &amp; Moncada E." box="[385,890,2004,2031]" pageId="7" pageNumber="99" pagination="223 - 231" refId="ref10152" refString="Pilato, G., Binda, M. G., Napolitano, A. &amp; Moncada E. (2001) Notes on South American tardigrades with the description of two new species: Pseudechiniscus spinerectus and Macrobiotus danielae. Tropical Zoology, 14, 2, 223 - 231. https: // doi. org / 10.1080 / 03946975.2001.10531154" type="journal article" year="2001">Pilato, Binda, Napolitano &amp; Moncada 2001</bibRefCitation>
</taxonomicName>
, known from 
<collectingCountry id="77C562924136FFC2FB82F87ABF1931ED" box="[1053,1146,2004,2030]" name="Ecuador" pageId="7" pageNumber="99">Ecuador</collectingCountry>
and 
<collectingCountry id="77C562924136FFC2FB28F87BBF8E31EC" box="[1207,1261,2005,2031]" name="Peru" pageId="7" pageNumber="99">Peru</collectingCountry>
, by the lack of eyes, and the lack of body granulation, larger body length (200–390 μm in 
<emphasis id="3DA6FE104139FFCDFC73FF39BFEB36B3" box="[1004,1160,151,177]" italics="true" pageId="8" pageNumber="100">
<taxonomicName id="C8D259814139FFCDFC73FF39BF0B36B2" ID-CoL="340270aa-a889-4581-a627-f37151c96a54" authorityName="Pilato, Binda, Napolitano &amp; Moncada" authorityYear="2001" box="[1004,1128,151,177]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="danielae">P. danielae</taxonomicName>
vs
</emphasis>
393–711 μm in the new species).
</paragraph>
<paragraph id="0F6D22024139FFCDFF58FF71BE61371E" blockId="8.[151,1437,151,1042]" pageId="8" pageNumber="100">
• 
<taxonomicName id="C8D259814139FFCDFF68FF71BADD36FA" ID-CoL="985a6c07-4e54-4122-9af1-a5743157121a" authorityName="Kaczmarek, Mioduchowska, Poprawa &amp; Roszkowska" authorityYear="2020" box="[247,446,222,249]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="experimentalis">
<emphasis id="3DA6FE104139FFCDFF68FF71BADD36FA" bold="true" box="[247,446,222,249]" italics="true" pageId="8" pageNumber="100">P. experimentalis</emphasis>
</taxonomicName>
, known only from 
<collectingCountry id="77C562924139FFCDFD05FF71B84036FA" box="[666,803,223,249]" name="Madagascar" pageId="8" pageNumber="100">Madagascar</collectingCountry>
, by the lack of wrinkled surface on the egg areolae, the presence of cuticular bulge on inner surface of claws I–III and granulation on inner surface of leg IV.
</paragraph>
<paragraph id="0F6D22024139FFCDFF58FE89BE3C37AD" blockId="8.[151,1437,151,1042]" pageId="8" pageNumber="100">
• 
<taxonomicName id="C8D259814139FFCDFF68FE89B8063742" ID-CoL="df59e194-bf07-449e-ad11-fba764d8731e" authority="Kaczmarek, Michalczyk &amp; Diduszko, 2005" authorityName="Kaczmarek, Michalczyk &amp; Diduszko" authorityYear="2005" box="[247,869,294,321]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="garynahi">
<emphasis id="3DA6FE104139FFCDFF68FE89BA193742" bold="true" box="[247,378,294,321]" italics="true" pageId="8" pageNumber="100">P. garynahi</emphasis>
Kaczmarek, 
<bibRefCitation id="6B435FF34139FFCDFD93FE89B8063742" author="Michalczyk, L. &amp; Diduszko, D." box="[524,869,294,321]" pageId="8" pageNumber="100" pagination="35 - 45" refId="ref8813" refString="Kaczmarek. L., Michalczyk, L. &amp; Diduszko, D. (2005) Some tardigrades from Siberia (Russia, Baikal region) with a description of Macrobiotus garynahi sp. nov. (Eutardigrada: Macrobiotidae: richtersi group). Zootaxa, 1053, 35 - 45. http: // doi. org / 10.11646 / zootaxa. 1053.1.3" type="journal article" year="2005">Michalczyk &amp; Diduszko, 2005</bibRefCitation>
</taxonomicName>
, known only from 
<collectingCountry id="77C562924139FFCDFBA9FE89BFE03742" box="[1078,1155,295,321]" name="Russia" pageId="8" pageNumber="100">Russia</collectingCountry>
, by the lack of oval pores on the cuticle, the lack of granulation in the first three pairs of legs, the lack of a cap-like structure on the top of the egg process, smaller egg bare diameter (96.0–132.0 μm in 
<emphasis id="3DA6FE104139FFCDFCB9FEC1B8A6378B" box="[806,965,367,393]" italics="true" pageId="8" pageNumber="100">
<taxonomicName id="C8D259814139FFCDFCB9FEC1B8C4378A" ID-CoL="df59e194-bf07-449e-ad11-fba764d8731e" authorityName="Kaczmarek, Michalczyk &amp; Diduszko" authorityYear="2005" box="[806,935,367,393]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="garynahi">P. garynahi</taxonomicName>
vs
</emphasis>
58.1–90.1 μm in the new species), and by a smaller egg diameter with processes (142.0–180.0 μm in 
<emphasis id="3DA6FE104139FFCDFCB3FE3DB8A837AF" box="[812,971,403,429]" italics="true" pageId="8" pageNumber="100">
<taxonomicName id="C8D259814139FFCDFCB3FE3DB8CE37AE" ID-CoL="df59e194-bf07-449e-ad11-fba764d8731e" authorityName="Kaczmarek, Michalczyk &amp; Diduszko" authorityYear="2005" box="[812,941,403,429]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="garynahi">P. garynahi</taxonomicName>
vs
</emphasis>
81.7–112.1 μm in the new species).
</paragraph>
<paragraph id="0F6D22024139FFCDFF58FE19B85F37F6" blockId="8.[151,1437,151,1042]" pageId="8" pageNumber="100">
• 
<taxonomicName id="C8D259814139FFCDFF68FE19B9B537D2" ID-CoL="1424212f-6bba-4d74-8109-fdfb5dc5bc2f" authority="Pilato, Bind &amp; Lisi, 2006" authorityName="Pilato, Bind &amp; Lisi" authorityYear="2006" box="[247,726,438,465]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="hapukuensis">
<emphasis id="3DA6FE104139FFCDFF68FE19BAC437D2" bold="true" box="[247,423,438,465]" italics="true" pageId="8" pageNumber="100">P. hapukuensis</emphasis>
Pilato, Bind &amp; Lisi, 2006
</taxonomicName>
, known only from 
<collectingCountry id="77C562924139FFCDFC2CFE19BF2E37D2" box="[947,1101,439,465]" name="New Zealand" pageId="8" pageNumber="100">New Zealand</collectingCountry>
, by the lack of finger-shaped apices of egg processes, and the presence of leg granulation.
</paragraph>
<paragraph id="0F6D22024139FFCDFF58FE51BAFB34A9" blockId="8.[151,1437,151,1042]" pageId="8" pageNumber="100">
• 
<taxonomicName id="C8D259814139FFCDFF68FE51B9A4341A" ID-CoL="4c0368fe-19da-4b72-b8c3-d173b7259bef" authority="Pilato, Claxton &amp; Binda, 1989" authorityName="Pilato, Claxton &amp; Binda" authorityYear="1989" box="[247,711,511,537]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="peteri">
<emphasis id="3DA6FE104139FFCDFF68FE51BA34341A" bold="true" box="[247,343,511,537]" italics="true" pageId="8" pageNumber="100">P. peteri</emphasis>
<bibRefCitation id="6B435FF34139FFCDFEFDFE51B9A4341A" author="Pilato, G. &amp; Claxton, S. &amp; Binda, M. G." box="[354,711,511,537]" pageId="8" pageNumber="100" pagination="43 - 48" refId="ref10104" refString="Pilato, G., Claxton, S. &amp; Binda, M. G. (1989) Tardigrades from Australia. III. Echiniscus marcusi and Macrobiotus peteri, new species of tardigrades from New South Wales. Animalia, 16, 43 - 48." type="journal article" year="1989">Pilato, Claxton &amp; Binda, 1989</bibRefCitation>
</taxonomicName>
, known only from 
<collectingCountry id="77C562924139FFCDFC38FE51BF6D341A" box="[935,1038,511,537]" name="Australia" pageId="8" pageNumber="100">Australia</collectingCountry>
and 
<collectingCountry id="77C562924139FFCDFBD4FE51BFDB341A" box="[1099,1208,511,537]" name="Indonesia" pageId="8" pageNumber="100">Indonesia</collectingCountry>
(
<bibRefCitation id="6B435FF34139FFCDFB56FE51BEF4341A" author="Pilato, G. &amp; Claxton, S. &amp; Binda, M. G." box="[1225,1431,511,537]" pageId="8" pageNumber="100" pagination="43 - 48" refId="ref10104" refString="Pilato, G., Claxton, S. &amp; Binda, M. G. (1989) Tardigrades from Australia. III. Echiniscus marcusi and Macrobiotus peteri, new species of tardigrades from New South Wales. Animalia, 16, 43 - 48." type="journal article" year="1989">
Pilato 
<emphasis id="3DA6FE104139FFCDFA88FE51BE31341A" box="[1303,1362,511,537]" italics="true" pageId="8" pageNumber="100">et al.</emphasis>
1989
</bibRefCitation>
; 
<bibRefCitation id="6B435FF34139FFCDFF08FD8DBAC3343E" author="Pilato, G. &amp; Binda M. G." box="[151,416,547,573]" pageId="8" pageNumber="100" pagination="209 - 218" refId="ref10073" refString="Pilato, G. &amp; Binda M. G. (1990) Notizie sui tardigradi muscicoli di Bali (Indonesia). Animalia, 17, 209 - 218." type="journal article" year="1990">Pilato and Binda, 1990</bibRefCitation>
; 
<bibRefCitation id="6B435FF34139FFCDFE32FD8DB90A343E" author="Mackness, B. S." box="[429,617,547,573]" pageId="8" pageNumber="100" pagination="1 - 5" refId="ref9583" refString="Mackness, B. S. (2003) A review of nematophagy by terrestrial tardigrades with new observations on some Australian species. Proceedings of the Royal Society of Victoria, 115, 1 - 5." type="journal article" year="2003">Mackness, 2003</bibRefCitation>
), by the lack of a cuticular thickening near the lunules I–III, the lack of subdivided apices of egg processes, a lower number of processes on the egg circumference (
<quantity id="C82A8FE74139FFCDFB2EFDE9BE753462" box="[1201,1302,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="8" pageNumber="100" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
<emphasis id="3DA6FE104139FFCDFA80FDE9BEFF3463" box="[1311,1436,583,608]" italics="true" pageId="8" pageNumber="100">
<taxonomicName id="C8D259814139FFCDFA80FDE9BE1F3463" ID-CoL="4c0368fe-19da-4b72-b8c3-d173b7259bef" authorityName="Pilato, Claxton &amp; Binda" authorityYear="1989" box="[1311,1404,583,608]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="peteri">P. peteri</taxonomicName>
vs
</emphasis>
<quantity id="C82A8FE74139FFCDFF08FDC4BB993486" box="[151,250,618,645]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.81" metricValueMin="2.54" pageId="8" pageNumber="100" unit="in" value="12.5" valueMax="15.0" valueMin="10.0">10–15 in</quantity>
the new species), and by a higher number of areolae around egg processes (
<quantity id="C82A8FE74139FFCDFBDCFDC5BFE83486" box="[1091,1163,619,645]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="8" pageNumber="100" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">6–7 in</quantity>
<taxonomicName id="C8D259814139FFCDFB0EFDC5BF8E3487" ID-CoL="4c0368fe-19da-4b72-b8c3-d173b7259bef" authorityName="Pilato, Claxton &amp; Binda" authorityYear="1989" box="[1169,1261,619,644]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="peteri">
<emphasis id="3DA6FE104139FFCDFB0EFDC5BF8E3487" box="[1169,1261,619,644]" italics="true" pageId="8" pageNumber="100">P. peteri</emphasis>
</taxonomicName>
, approximately 
<quantity id="C82A8FE74139FFCDFF08FD21BBB334AA" box="[151,208,655,681]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="100" unit="in" value="10.0">10 in</quantity>
the new species).
</paragraph>
<paragraph id="0F6D22024139FFCDFF58FD1DB9993581" blockId="8.[151,1437,151,1042]" pageId="8" pageNumber="100">
• 
<taxonomicName id="C8D259814139FFCDFF68FD1DB9E034CE" ID-CoL="7d67af7e-17c8-4507-a424-a226a1feb594" authority="Claps &amp; Rossi, 1997" authorityName="Claps &amp; Rossi" authorityYear="1997" box="[247,643,690,717]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="rioplatensis">
<emphasis id="3DA6FE104139FFCDFF68FD1DBAF434CE" bold="true" box="[247,407,690,717]" italics="true" pageId="8" pageNumber="100">P. rioplatensis</emphasis>
<bibRefCitation id="6B435FF34139FFCDFE02FD1DB9E034CE" author="Claps, M. C. &amp; Rossi, G. C." box="[413,643,691,717]" pageId="8" pageNumber="100" pagination="17 - 22" refId="ref8180" refString="Claps, M. C. &amp; Rossi, G. C. (1997) Tardigrados de Uruguay, com descripcion de dos nuevas especies (Echiniscidae, Macrobiotidae). Iheringia Serie Zoologia, 83, 17 - 22." type="journal article" year="1997">Claps &amp; Rossi, 1997</bibRefCitation>
</taxonomicName>
, known only from 
<collectingCountry id="77C562924139FFCDFCCEFD1DB8D734CE" box="[849,948,691,717]" name="Uruguay" pageId="8" pageNumber="100">Uruguay</collectingCountry>
, by the lack of eyespots, a larger body (227– 372 μm in 
<emphasis id="3DA6FE104139FFCDFE8AFD79BAB534F3" box="[277,470,727,753]" italics="true" pageId="8" pageNumber="100">
<taxonomicName id="C8D259814139FFCDFE8AFD79BAD534F2" ID-CoL="7d67af7e-17c8-4507-a424-a226a1feb594" authorityName="Claps &amp; Rossi" authorityYear="1997" box="[277,438,727,753]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="rioplatensis">P. rioplatensis</taxonomicName>
vs
</emphasis>
392.5–710.8 μm in the new species), a longer buccal tube (38 μm in 
<taxonomicName id="C8D259814139FFCDFB6BFD79BEF634F2" ID-CoL="7d67af7e-17c8-4507-a424-a226a1feb594" authorityName="Claps &amp; Rossi" authorityYear="1997" box="[1268,1429,727,753]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="rioplatensis">
<emphasis id="3DA6FE104139FFCDFB6BFD79BEF634F2" box="[1268,1429,727,753]" italics="true" pageId="8" pageNumber="100">P. rioplatensis</emphasis>
</taxonomicName>
, over 40–58.4 μm in the new species), a larger egg full diameter (68–79 μm in 
<emphasis id="3DA6FE104139FFCDFB91FD55BFAE3517" box="[1038,1229,763,789]" italics="true" pageId="8" pageNumber="100">
<taxonomicName id="C8D259814139FFCDFB91FD55BFCD3516" ID-CoL="7d67af7e-17c8-4507-a424-a226a1feb594" authorityName="Claps &amp; Rossi" authorityYear="1997" box="[1038,1198,763,789]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="rioplatensis">P. rioplatensis</taxonomicName>
vs
</emphasis>
81.7–112.1 μm in the new species), the lack of the a filament-shaped apex of egg processes, higher egg processes (4.6 μm height in 
<emphasis id="3DA6FE104139FFCDFF08FCEDBA34355F" box="[151,343,835,861]" italics="true" pageId="8" pageNumber="100">
<taxonomicName id="C8D259814139FFCDFF08FCEDBA5B355E" ID-CoL="7d67af7e-17c8-4507-a424-a226a1feb594" authorityName="Claps &amp; Rossi" authorityYear="1997" box="[151,312,835,861]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="rioplatensis">P. rioplatensis</taxonomicName>
vs
</emphasis>
over 7.4 μm in the new species), and by a lower number of processes on the egg circumference (
<quantity id="C82A8FE74139FFCDFF00FCC9BA673582" box="[159,260,871,897]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" metricValueMax="4.826" metricValueMin="4.318" pageId="8" pageNumber="100" unit="in" value="18.0" valueMax="19.0" valueMin="17.0">17–19 in</quantity>
<emphasis id="3DA6FE104139FFCDFE95FCC9BAAB3583" box="[266,456,871,897]" italics="true" pageId="8" pageNumber="100">
<taxonomicName id="C8D259814139FFCDFE95FCC9BAC93582" ID-CoL="7d67af7e-17c8-4507-a424-a226a1feb594" authorityName="Claps &amp; Rossi" authorityYear="1997" box="[266,426,871,897]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="rioplatensis">P. rioplatensis</taxonomicName>
vs
</emphasis>
<quantity id="C82A8FE74139FFCDFE50FCC8B9513582" box="[463,562,870,897]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.81" metricValueMin="2.54" pageId="8" pageNumber="100" unit="in" value="12.5" valueMax="15.0" valueMin="10.0">10–15 in</quantity>
the new species).
</paragraph>
<paragraph id="0F6D22024139FFCDFF58FC25BB9A3211" blockId="8.[151,1437,151,1042]" pageId="8" pageNumber="100">
• 
<taxonomicName id="C8D259814139FFCDFF68FC25B92435A6" ID-CoL="eae3347c-4d41-4f5b-bf97-69ed172f9539" authority="Binda &amp; Pilato, 2001" authorityName="Binda &amp; Pilato" authorityYear="2001" box="[247,583,907,933]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="savai">
<emphasis id="3DA6FE104139FFCDFF68FC25BA2D35A6" bold="true" box="[247,334,907,933]" italics="true" pageId="8" pageNumber="100">P. savai</emphasis>
<bibRefCitation id="6B435FF34139FFCDFECAFC25B92435A6" author="Pilato, G. &amp; Binda, M. G. &amp; Napolitano, A. &amp; Moncada E." box="[341,583,907,933]" pageId="8" pageNumber="100" pagination="223 - 231" refId="ref10152" refString="Pilato, G., Binda, M. G., Napolitano, A. &amp; Moncada E. (2001) Notes on South American tardigrades with the description of two new species: Pseudechiniscus spinerectus and Macrobiotus danielae. Tropical Zoology, 14, 2, 223 - 231. https: // doi. org / 10.1080 / 03946975.2001.10531154" type="journal article" year="2001">Binda &amp; Pilato, 2001</bibRefCitation>
</taxonomicName>
, known from only 
<collectingCountry id="77C562924139FFCDFC82FC25B8EE35A6" box="[797,909,907,933]" name="Sri Lanka" pageId="8" pageNumber="100">Sri Lanka</collectingCountry>
, by the lack of eyespots, the lack of a cuticular thickening near the lunules of the first three legs, a different egg process shapes (blunt in 
<emphasis id="3DA6FE104139FFCDFB37FC01BE4135CB" box="[1192,1314,943,968]" italics="true" pageId="8" pageNumber="100">
<taxonomicName id="C8D259814139FFCDFB37FC01BE6235CB" ID-CoL="eae3347c-4d41-4f5b-bf97-69ed172f9539" authorityName="Binda &amp; Pilato" authorityYear="2001" box="[1192,1281,943,968]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="savai">P. savai</taxonomicName>
vs
</emphasis>
conical in the new species), and by a slightly smaller egg full diameter (115 μm in 
<emphasis id="3DA6FE104139FFCDFC77FC7DBF0135EF" box="[1000,1122,979,1004]" italics="true" pageId="8" pageNumber="100">
<taxonomicName id="C8D259814139FFCDFC77FC7DBF2235EF" ID-CoL="eae3347c-4d41-4f5b-bf97-69ed172f9539" authorityName="Binda &amp; Pilato" authorityYear="2001" box="[1000,1089,979,1004]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="savai">P. savai</taxonomicName>
vs
</emphasis>
81.7–112.1 μm in the new species).
</paragraph>
</subSubSection>
<caption id="5BAD728A4139FFCDFF08FA0CB8F23000" ID-DOI="http://doi.org/10.5281/zenodo.6531350" ID-Zenodo-Dep="6531350" httpUri="https://zenodo.org/record/6531350/files/figure.png" pageId="8" pageNumber="100" startId="8.[151,250,1442,1466]" targetBox="[151,1436,1065,1419]" targetPageId="8">
<paragraph id="0F6D22024139FFCDFF08FA0CB96133B8" blockId="8.[151,1436,1442,1539]" box="[151,514,1442,1467]" pageId="8" pageNumber="100">
<emphasis id="3DA6FE104139FFCDFF08FA0CB96133B8" bold="true" box="[151,514,1442,1467]" pageId="8" pageNumber="100">FIGURE 4. Morphology of leg IV</emphasis>
</paragraph>
<paragraph id="0F6D22024139FFCDFF08FA69B8F23000" blockId="8.[151,1436,1442,1539]" pageId="8" pageNumber="100">The leg IV observed with LM (A) and SEM (B, C). white-filled indent arrowhead and white dashed line indicate the granulation pattern. White-filled arrows indicate the accessory points. Scale bars = μm.</paragraph>
</caption>
<subSubSection id="47C871894139FFCFFF08F99FB953314F" lastPageId="10" lastPageNumber="102" pageId="8" pageNumber="100" type="diagnosis">
<paragraph id="0F6D22024139FFCDFF08F99FBA803048" blockId="8.[151,1437,1585,2007]" box="[151,483,1585,1611]" pageId="8" pageNumber="100">
<emphasis id="3DA6FE104139FFCDFF08F99FBA803048" box="[151,483,1585,1611]" italics="true" pageId="8" pageNumber="100">Genetic differential diagnosis</emphasis>
</paragraph>
<paragraph id="0F6D22024139FFCDFF58F9FBBB893090" blockId="8.[151,1437,1585,2007]" pageId="8" pageNumber="100">Below is a summary of genetic p-distances, whereas detailed matrices are available in Supporting Material SM.02.</paragraph>
<paragraph id="0F6D22024139FFCDFF58F933BE0630DF" blockId="8.[151,1437,1585,2007]" pageId="8" pageNumber="100">
• 
<emphasis id="3DA6FE104139FFCDFF68F933BA1F30B4" bold="true" box="[247,380,1693,1719]" pageId="8" pageNumber="100">18S rDNA</emphasis>
: 0.5%–3.6%, with the most similar species being 
<taxonomicName id="C8D259814139FFCDFC6AF930BFDC30B4" authorityName="Kaczmarek, Mioduchowska, Poprawa &amp; Roszkowska" authorityYear="2020" box="[1013,1215,1693,1719]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="experimentalis">
<emphasis id="3DA6FE104139FFCDFC6AF930BFDC30B4" box="[1013,1215,1693,1719]" italics="true" pageId="8" pageNumber="100">P. experimentalis</emphasis>
</taxonomicName>
from 
<collectingCountry id="77C562924139FFCDFA8AF933BEFF30B4" box="[1301,1436,1693,1719]" name="Madagascar" pageId="8" pageNumber="100">Madagascar</collectingCountry>
(
<accessionNumber id="1081BFE14139FFCDFF00F96FBA4030D8" box="[159,291,1729,1755]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN073467" pageId="8" pageNumber="100">MN073467</accessionNumber>
–8 and 
<accessionNumber id="1081BFE14139FFCDFEEBF96FBA9430DF" box="[372,503,1729,1756]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH664938" pageId="8" pageNumber="100">MH664938</accessionNumber>
) and the least similar being 
<taxonomicName id="C8D259814139FFCDFCADF96CB89330DF" authority="NO." authorityName="NO." box="[818,1008,1729,1756]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="areolatus">
<emphasis id="3DA6FE104139FFCDFCADF96CB82030D8" box="[818,835,1730,1755]" italics="true" pageId="8" pageNumber="100">P</emphasis>
. 
<emphasis id="3DA6FE104139FFCDFCCDF96FB8D830D8" box="[850,955,1729,1755]" italics="true" pageId="8" pageNumber="100">areolatus</emphasis>
NO.
</taxonomicName>
385 from 
<collectingCountry id="77C562924139FFCDFBC0F96FBFA030D8" box="[1119,1219,1729,1755]" name="Svalbard and Jan Mayen" pageId="8" pageNumber="100">Svalbard</collectingCountry>
(
<accessionNumber id="1081BFE14139FFCDFB4DF96FBE3A30DF" box="[1234,1369,1729,1756]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH664931" pageId="8" pageNumber="100">MH664931</accessionNumber>
).
</paragraph>
<paragraph id="0F6D22024139FFCDFF58F94BBE763127" blockId="8.[151,1437,1585,2007]" pageId="8" pageNumber="100">
• 
<emphasis id="3DA6FE104139FFCDFF68F94BBA0C30FC" bold="true" box="[247,367,1765,1791]" pageId="8" pageNumber="100">28S rDNA</emphasis>
: 0.98–7.5%, with the most similar species being 
<taxonomicName id="C8D259814139FFCDFCE4F948BF5B30FC" authorityName="Kaczmarek, Mioduchowska, Poprawa &amp; Roszkowska" authorityYear="2020" box="[891,1080,1765,1791]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="experimentalis">
<emphasis id="3DA6FE104139FFCDFCE4F948BF5B30FC" box="[891,1080,1765,1791]" italics="true" pageId="8" pageNumber="100">P.experimentalis</emphasis>
</taxonomicName>
(
<accessionNumber id="1081BFE14139FFCDFBDCF94BBFA53103" box="[1091,1222,1765,1792]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN073465" pageId="8" pageNumber="100">MN073465</accessionNumber>
–6and 
<accessionNumber id="1081BFE14139FFCDFA8DF94BBEF73103" box="[1298,1428,1765,1792]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH664956" pageId="8" pageNumber="100">MH664956</accessionNumber>
) and the least being 
<taxonomicName id="C8D259814139FFCDFEF0F8A7B8343127" authority="Ramazzotti, 1956" authorityName="Ramazzotti" authorityYear="1956" box="[367,855,1801,1828]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="tonollii">
<emphasis id="3DA6FE104139FFCDFEF0F8A7B9E93120" box="[367,650,1801,1827]" italics="true" pageId="8" pageNumber="100">Paramacrobiotus tonollii</emphasis>
Ramazzotti, 1956
</taxonomicName>
strain 
<collectingCountry id="77C562924139FFCDFC3CF8A7B8AB3127" box="[931,968,1801,1828]" name="United States of America" pageId="8" pageNumber="100">US</collectingCountry>
.035 from 
<collectingCountry id="77C562924139FFCDFBA3F8A7BF113120" box="[1084,1138,1801,1827]" name="United States of America" pageId="8" pageNumber="100">USA</collectingCountry>
(
<accessionNumber id="1081BFE14139FFCDFB1EF8A7BE6A3127" box="[1153,1289,1801,1828]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH664963" pageId="8" pageNumber="100">MH664963</accessionNumber>
).
</paragraph>
<paragraph id="0F6D22024139FFCDFF58F883B8DA316F" blockId="8.[151,1437,1585,2007]" pageId="8" pageNumber="100">
• 
<emphasis id="3DA6FE104139FFCDFF68F883BA5F3144" bold="true" box="[247,316,1837,1863]" pageId="8" pageNumber="100">ITS-2</emphasis>
: 9.9–33.9%, with the most similar species being 
<taxonomicName id="C8D259814139FFCDFCF5F880BF483144" authorityName="Kaczmarek, Mioduchowska, Poprawa &amp; Roszkowska" authorityYear="2020" box="[874,1067,1837,1863]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="experimentalis">
<emphasis id="3DA6FE104139FFCDFCF5F880BF483144" box="[874,1067,1837,1863]" italics="true" pageId="8" pageNumber="100">P. experimentalis</emphasis>
</taxonomicName>
(MH073463–4 and 
<accessionNumber id="1081BFE14139FFCDFA8DF883BEF7314B" box="[1298,1428,1837,1864]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH666087" pageId="8" pageNumber="100">MH666087</accessionNumber>
) and the least similar species being 
<taxonomicName id="C8D259814139FFCDFD83F8FCB9EF3168" authorityName="Ramazzotti" authorityYear="1956" box="[540,652,1873,1899]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="tonollii">
<emphasis id="3DA6FE104139FFCDFD83F8FCB94E3168" box="[540,557,1874,1899]" italics="true" pageId="8" pageNumber="100">P</emphasis>
. 
<emphasis id="3DA6FE104139FFCDFDA4F8FFB9EF3168" box="[571,652,1873,1899]" italics="true" pageId="8" pageNumber="100">tonollii</emphasis>
</taxonomicName>
(
<accessionNumber id="1081BFE14139FFCDFD04F8FFB87F3168" box="[667,796,1873,1899]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GQ403679" pageId="8" pageNumber="100">GQ403679</accessionNumber>
, 
<accessionNumber id="1081BFE14139FFCDFCB8F8FFB8CE316F" box="[807,941,1873,1900]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH666096" pageId="8" pageNumber="100">MH666096</accessionNumber>
).
</paragraph>
<paragraph id="0F6D22024139FFCDFF58F8DBB91F31D4" blockId="8.[151,1437,1585,2007]" pageId="8" pageNumber="100">
• 
<emphasis id="3DA6FE104139FFCDFF68F8DBBA4F318C" bold="true" box="[247,300,1909,1935]" pageId="8" pageNumber="100">COI</emphasis>
: 21.2–32.6%, with the most similar species being 
<taxonomicName id="C8D259814139FFCDFCF8F8DBBF31318C" box="[871,1106,1909,1935]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="undetermined">
<emphasis id="3DA6FE104139FFCDFCF8F8DBBF49318C" box="[871,1066,1909,1935]" italics="true" pageId="8" pageNumber="100">Paramacrobiotus</emphasis>
sp.
</taxonomicName>
strain TZ.018 from 
<collectingCountry id="77C562924139FFCDFAA7F8DBBEFE318C" box="[1336,1437,1909,1935]" name="Tanzania" pageId="8" pageNumber="100">Tanzania</collectingCountry>
(
<accessionNumber id="1081BFE14139FFCDFF00F837BA4031B7" box="[159,291,1945,1972]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH676017" pageId="8" pageNumber="100">MH676017</accessionNumber>
) and with the least similar species being 
<taxonomicName id="C8D259814139FFCDFD7CF837BA0131D4" authority="Guidetti, Cesari, Bertolani, Altiero &amp; Rebecchi, 2019" authorityName="Guidetti, Cesari, Bertolani, Altiero &amp; Rebecchi" authorityYear="2019" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="100" phylum="Tardigrada" rank="species" species="depressus">
<emphasis id="3DA6FE104139FFCDFD7CF837BF7B31B0" box="[739,1048,1945,1971]" italics="true" pageId="8" pageNumber="100">Paramacrobiotus depressus</emphasis>
<bibRefCitation id="6B435FF34139FFCDFB81F837BA0131D4" author="Guidetti, R. &amp; Cesari, M. &amp; Bertolani, R. &amp; Altiero, T. &amp; Rebecchi, L." pageId="8" pageNumber="100" pagination="1 - 28" refId="ref8619" refString="Guidetti, R., Cesari, M., Bertolani, R., Altiero, T. &amp; Rebecchi, L. (2019) High diversity in species, reproductive modes and distribution within the Paramacrobiotus richtersi complex (Eutardigrada, Macrobiotidae). Zoological letters, 5 (1), 1 - 28. https: // doi. org / 10.1186 / s 40851 - 018 - 0113 - z" type="journal article" year="2019">Guidetti, Cesari, Bertolani, Altiero &amp; Rebecchi, 2019</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="77C562924139FFCDFE38F813BAB931D4" box="[423,474,1981,2007]" name="Italy" pageId="8" pageNumber="100">Italy</collectingCountry>
(
<accessionNumber id="1081BFE14139FFCDFE76F813B91331D4" box="[489,624,1981,2007]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK041013" pageId="8" pageNumber="100">MK041013</accessionNumber>
).
</paragraph>
<paragraph id="0F6D22024138FFCCFF08FF39BA6E36B2" blockId="9.[151,1436,151,322]" box="[151,269,151,177]" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFF08FF39BA6E36B2" box="[151,269,151,177]" italics="true" pageId="9" pageNumber="101">Phylogeny</emphasis>
</paragraph>
<paragraph id="0F6D22024138FFCCFF58FF15B9963741" blockId="9.[151,1436,151,322]" pageId="9" pageNumber="101">
The supported bootstrap values of ML trees were usually low (&lt;50%), thus the phylogenetic analyses were based on the BI trees (
<figureCitation id="97E93E874138FFCCFE0FFF71BAB636FA" box="[400,469,223,249]" captionStart="FIGURE 7" captionStartId="12.[151,250,1665,1689]" captionTargetBox="[184,1394,222,1613]" captionTargetId="figure-18@12.[151,1436,198,1641]" captionTargetPageId="12" captionText="FIGURE 7A. Phylogenetic position of the new species based on the concatenated 18S+28S+ITS-2+COI dataset. Values at nodes indicate Bayesian posterior probabilities. Scale bars = substitutions per position." figureDoi="http://doi.org/10.5281/zenodo.6531356" httpUri="https://zenodo.org/record/6531356/files/figure.png" pageId="9" pageNumber="101">Fig. 7</figureCitation>
). Three phylogenetic trees indicated that the new species is sister to a clade containing tropical species from the Afrotropic (
<taxonomicName id="C8D259814138FFCCFDAAFEADB9D7371E" authorityName="Schill, Forster, Dandekar &amp; Wolf" authorityYear="2010" box="[565,692,259,285]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="101" phylum="Tardigrada" rank="species" species="kenianus">
<emphasis id="3DA6FE104138FFCCFDAAFEADB9D7371E" box="[565,692,259,285]" italics="true" pageId="9" pageNumber="101">P. kenianus</emphasis>
</taxonomicName>
, 
<taxonomicName id="C8D259814138FFCCFD5EFEADB8E2371E" authorityName="Kaczmarek, Mioduchowska, Poprawa &amp; Roszkowska" authorityYear="2020" box="[705,897,259,285]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="101" phylum="Tardigrada" rank="species" species="experimentalis">
<emphasis id="3DA6FE104138FFCCFD5EFEADB8E2371E" box="[705,897,259,285]" italics="true" pageId="9" pageNumber="101">P. experimentalis</emphasis>
</taxonomicName>
, and an undescribed species from 
<collectingCountry id="77C562924138FFCCFB60FEADBE05371D" box="[1279,1382,259,286]" name="Tanzania" pageId="9" pageNumber="101">Tanzania</collectingCountry>
) and the Neotropics (two unidentified species from 
<collectingCountry id="77C562924138FFCCFD3FFE89B98A3741" box="[672,745,295,322]" name="Brazil" pageId="9" pageNumber="101">Brazil</collectingCountry>
).
</paragraph>
<paragraph id="0F6D22024138FFCCFF08FEC1BAAA378A" blockId="9.[151,1436,367,465]" box="[151,457,367,393]" pageId="9" pageNumber="101">Genetic species delineation</paragraph>
<paragraph id="0F6D22024138FFCCFF58FE3DBF1A37D2" blockId="9.[151,1436,367,465]" pageId="9" pageNumber="101">
Both, the ITS-2 and the COI BI trees indicated that the cluster of 
<taxonomicName id="C8D259814138FFCCFC08FE3DBF3937AE" authority="Sugiura &amp; Matsumoto &amp; Kunieda, 2022" authorityName="Sugiura &amp; Matsumoto &amp; Kunieda" authorityYear="2022" box="[919,1114,403,429]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="101" phylum="Tardigrada" rank="species" species="metropolitanus" status="sp. nov.">
<emphasis id="3DA6FE104138FFCCFC08FE3DBF3937AE" box="[919,1114,403,429]" italics="true" pageId="9" pageNumber="101">P. metropolitanus</emphasis>
</taxonomicName>
<emphasis id="3DA6FE104138FFCCFBFFFE3DBFDA37AE" bold="true" box="[1120,1209,403,429]" pageId="9" pageNumber="101">
<taxonomicNameLabel id="2695436B4138FFCCFBFFFE3DBFDA37AE" box="[1120,1209,403,429]" pageId="9" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a separate species with&gt;93% probabilities by both maximum likelihood and Bayesian supported solutions.
</paragraph>
<caption id="5BAD728A4138FFCCFF08FDAEBF70341B" ID-Table-UUID="5BAD728A4138FFCCFF08FDAEBF70341B" box="[151,1043,512,537]" httpUri="http://table.plazi.org/id/5BAD728A4138FFCCFF08FDAEBF70341B" pageId="9" pageNumber="101" startId="9.[151,239,512,536]" targetBox="[163,1420,553,1946]" targetIsTable="true" targetPageId="9">
<paragraph id="0F6D22024138FFCCFF08FDAEBF70341B" blockId="9.[151,1043,512,537]" box="[151,1043,512,537]" pageId="9" pageNumber="101">
TABLE 2. Morphometrics of 
<taxonomicName id="C8D259814138FFCCFE79FDAEB832341B" authority="Sugiura &amp; Matsumoto &amp; Kunieda, 2022" authorityName="Sugiura &amp; Matsumoto &amp; Kunieda" authorityYear="2022" box="[486,849,512,536]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="101" phylum="Tardigrada" rank="species" species="metropolitanus" status="sp. nov.">
<emphasis id="3DA6FE104138FFCCFE79FDAEB832341B" bold="true" box="[486,849,512,536]" italics="true" pageId="9" pageNumber="101">Paramacrobiotus metropolitanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2695436B4138FFCCFCC8FDAFB8CE341A" box="[855,941,513,537]" pageId="9" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
animals.
</paragraph>
</caption>
<paragraph id="0F6D22024138FFCCFF3CFD90BEEB3199" pageId="9" pageNumber="101">
<table id="7DD2D0A24138003AFF3CFD87BEEF3199" box="[163,1420,553,1946]" gridcols="10" gridrows="34" pageId="9" pageNumber="101">
<tr id="B1E220404138003AFF3CFD87BEEF3469" box="[163,1420,553,618]" gridrow="0" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFD87BABF3469" box="[163,476,553,618]" gridcol="0" gridrow="0" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFF3CFD90BA2C3455" bold="true" box="[163,335,574,598]" pageId="9" pageNumber="101">CHARACTER</emphasis>
</th>
<th id="F233493C4138003AFD99FD87B97C3469" box="[518,543,553,618]" gridcol="1" gridrow="0" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD96FD90B97F3455" bold="true" box="[521,540,574,598]" pageId="9" pageNumber="101">N</emphasis>
</th>
<th id="F233493C4138003AFDD5FD87B8213469" box="[586,834,553,618]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD05FD87B87A346A" bold="true" pageId="9" pageNumber="101">
RANGE μm 
<emphasis id="3DA6FE104138FFCCFC9AFDFCB87A346A" bold="true" box="[773,793,594,617]" italics="true" pageId="9" pageNumber="101">pt</emphasis>
</emphasis>
</th>
<th id="F233493C4138003AFCEDFD87B89F3469" box="[882,1020,553,618]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC11FD87B88E346A" bold="true" pageId="9" pageNumber="101">
MEAN μm 
<emphasis id="3DA6FE104138FFCCFC46FDFCB88E346A" bold="true" box="[985,1005,594,617]" italics="true" pageId="9" pageNumber="101">pt</emphasis>
</emphasis>
</th>
<th id="F233493C4138003AFBAEFD87BFD43469" box="[1073,1207,553,618]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFBFCFD87BFD2346A" bold="true" pageId="9" pageNumber="101">
SD μm 
<emphasis id="3DA6FE104138FFCCFB02FDFCBFD2346A" bold="true" box="[1181,1201,594,617]" italics="true" pageId="9" pageNumber="101">pt</emphasis>
</emphasis>
</th>
<th id="F233493C4138003AFB66FD87BEEF3469" box="[1273,1420,553,618]" colspan="2" colspanRight="1" gridcol="8" gridrow="0" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFA97FD84BE1E346A" bold="true" pageId="9" pageNumber="101">
Holotype μm 
<emphasis id="3DA6FE104138FFCCFAF6FDFCBE1E346A" bold="true" box="[1385,1405,594,617]" italics="true" pageId="9" pageNumber="101">pt</emphasis>
</emphasis>
</th>
</tr>
<tr id="B1E220404138003AFF3CFDD4BEEF3491" box="[163,1420,634,658]" gridrow="1" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFDD4BABF3491" box="[163,476,634,658]" gridcol="0" gridrow="1" pageId="9" pageNumber="101">Body length</th>
<td id="F233493C4138003AFD99FDD4B97C3491" box="[518,543,634,658]" gridcol="1" gridrow="1" pageId="9" pageNumber="101">40</td>
<td id="F233493C4138003AFDD5FDD4B9D33491" box="[586,688,634,658]" gridcol="2" gridrow="1" pageId="9" pageNumber="101">393–711</td>
<td id="F233493C4138003AFD43FDD4B8213491" box="[732,834,634,658]" gridcol="3" gridrow="1" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FDD4B8223491" box="[732,833,634,658]" italics="true" pageId="9" pageNumber="101">962–1306</emphasis>
</td>
<td id="F233493C4138003AFCEDFDD4B8FD3491" box="[882,926,634,658]" gridcol="4" gridrow="1" pageId="9" pageNumber="101">540</td>
<td id="F233493C4138003AFC54FDD4B89F3491" box="[971,1020,634,658]" gridcol="5" gridrow="1" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC54FDD4B89F3491" box="[971,1020,634,658]" italics="true" pageId="9" pageNumber="101">1135</emphasis>
</td>
<td id="F233493C4138003AFBAEFDD4BF303491" box="[1073,1107,634,658]" gridcol="6" gridrow="1" pageId="9" pageNumber="101">75</td>
<td id="F233493C4138003AFB08FDD4BFD43491" box="[1175,1207,634,658]" gridcol="7" gridrow="1" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB05FDD4BFD03491" box="[1178,1203,634,658]" italics="true" pageId="9" pageNumber="101">80</emphasis>
</td>
<td id="F233493C4138003AFB66FDD4BE463491" box="[1273,1317,634,658]" gridcol="8" gridrow="1" pageId="9" pageNumber="101">485</td>
<td id="F233493C4138003AFAC6FDD4BEEF3491" box="[1369,1420,634,658]" gridcol="9" gridrow="1" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC6FDD4BEEF3491" box="[1369,1420,634,658]" italics="true" pageId="9" pageNumber="101">1058</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFD0CBEEF34B9" box="[163,1420,674,698]" gridrow="2" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFD0CBEEF34B9" box="[163,1420,674,698]" colspan="10" colspanRight="9" gridcol="0" gridrow="2" pageId="9" pageNumber="101">Buccal apparatus</th>
</tr>
<tr id="B1E220404138003AFF3CFD64BEEF34E1" box="[163,1420,714,738]" gridrow="3" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFD64BABF34E1" box="[163,476,714,738]" gridcol="0" gridrow="3" pageId="9" pageNumber="101">Buccal tube length</th>
<td id="F233493C4138003AFD99FD64B97C34E1" box="[518,543,714,738]" gridcol="1" gridrow="3" pageId="9" pageNumber="101">37</td>
<td id="F233493C4138003AFDD5FD64B9D334E1" box="[586,688,714,738]" gridcol="2" gridrow="3" pageId="9" pageNumber="101">40.0–58.4</td>
<td id="F233493C4138003AFD43FD64B82134E1" box="[732,834,714,738]" gridcol="3" gridrow="3" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC96FD65B87534E1" box="[777,790,715,738]" italics="true" pageId="9" pageNumber="101">–</emphasis>
</td>
<td id="F233493C4138003AFCEDFD64B8FD34E1" box="[882,926,714,738]" gridcol="4" gridrow="3" pageId="9" pageNumber="101">48.1</td>
<td id="F233493C4138003AFC54FD64B89F34E1" box="[971,1020,714,738]" gridcol="5" gridrow="3" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC42FD65B88934E1" box="[989,1002,715,738]" italics="true" pageId="9" pageNumber="101">–</emphasis>
</td>
<td id="F233493C4138003AFBAEFD64BF3034E1" box="[1073,1107,714,738]" gridcol="6" gridrow="3" pageId="9" pageNumber="101">4.3</td>
<td id="F233493C4138003AFB08FD64BFD434E1" box="[1175,1207,714,738]" gridcol="7" gridrow="3" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB3FFD65BFCE34E1" box="[1184,1197,715,738]" italics="true" pageId="9" pageNumber="101">–</emphasis>
</td>
<td id="F233493C4138003AFB66FD64BE4634E1" box="[1273,1317,714,738]" gridcol="8" gridrow="3" pageId="9" pageNumber="101">45.9</td>
<td id="F233493C4138003AFAC6FD64BEEF34E1" box="[1369,1420,714,738]" gridcol="9" gridrow="3" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAF3FD65BE1A34E1" box="[1388,1401,715,738]" italics="true" pageId="9" pageNumber="101">–</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFD5CBEEF3508" box="[163,1420,754,779]" gridrow="4" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFD5CBABF3508" box="[163,476,754,779]" gridcol="0" gridrow="4" pageId="9" pageNumber="101">Stylet support insertion point</th>
<td id="F233493C4138003AFD99FD5CB97C3508" box="[518,543,754,779]" gridcol="1" gridrow="4" pageId="9" pageNumber="101">37</td>
<td id="F233493C4138003AFDD5FD5CB9D33508" box="[586,688,754,779]" gridcol="2" gridrow="4" pageId="9" pageNumber="101">33.4–45.3</td>
<td id="F233493C4138003AFD43FD5CB8213508" box="[732,834,754,779]" gridcol="3" gridrow="4" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FD5CB8223509" box="[732,833,754,778]" italics="true" pageId="9" pageNumber="101">75.2–83.6</emphasis>
</td>
<td id="F233493C4138003AFCEDFD5CB8FD3508" box="[882,926,754,779]" gridcol="4" gridrow="4" pageId="9" pageNumber="101">38.8</td>
<td id="F233493C4138003AFC54FD5CB89F3508" box="[971,1020,754,779]" gridcol="5" gridrow="4" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FD5DB89A3508" box="[973,1017,755,779]" italics="true" pageId="9" pageNumber="101">80.8</emphasis>
</td>
<td id="F233493C4138003AFBAEFD5CBF303508" box="[1073,1107,754,779]" gridcol="6" gridrow="4" pageId="9" pageNumber="101">3.5</td>
<td id="F233493C4138003AFB08FD5CBFD43508" box="[1175,1207,754,779]" gridcol="7" gridrow="4" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FD5DBFD43508" box="[1175,1207,755,779]" italics="true" pageId="9" pageNumber="101">1.9</emphasis>
</td>
<td id="F233493C4138003AFB66FD5CBE463508" box="[1273,1317,754,779]" gridcol="8" gridrow="4" pageId="9" pageNumber="101">38.0</td>
<td id="F233493C4138003AFAC6FD5CBEEF3508" box="[1369,1420,754,779]" gridcol="9" gridrow="4" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FD5DBEEB3508" box="[1372,1416,755,779]" italics="true" pageId="9" pageNumber="101">82.8</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFCB5BEEF3530" box="[163,1420,795,819]" gridrow="5" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFCB5BABF3530" box="[163,476,795,819]" gridcol="0" gridrow="5" pageId="9" pageNumber="101">Buccal tube external width</th>
<td id="F233493C4138003AFD99FCB5B97C3530" box="[518,543,795,819]" gridcol="1" gridrow="5" pageId="9" pageNumber="101">37</td>
<td id="F233493C4138003AFDD5FCB5B9D33530" box="[586,688,795,819]" gridcol="2" gridrow="5" pageId="9" pageNumber="101">7.6–12.0</td>
<td id="F233493C4138003AFD43FCB5B8213530" box="[732,834,795,819]" gridcol="3" gridrow="5" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FCB5B8213530" box="[732,834,795,819]" italics="true" pageId="9" pageNumber="101">16.8–25.2</emphasis>
</td>
<td id="F233493C4138003AFCEDFCB5B8FD3530" box="[882,926,795,819]" gridcol="4" gridrow="5" pageId="9" pageNumber="101">10.0</td>
<td id="F233493C4138003AFC54FCB5B89F3530" box="[971,1020,795,819]" gridcol="5" gridrow="5" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FCB5B89A3530" box="[973,1017,795,819]" italics="true" pageId="9" pageNumber="101">20.8</emphasis>
</td>
<td id="F233493C4138003AFBAEFCB5BF303530" box="[1073,1107,795,819]" gridcol="6" gridrow="5" pageId="9" pageNumber="101">1.3</td>
<td id="F233493C4138003AFB08FCB5BFD43530" box="[1175,1207,795,819]" gridcol="7" gridrow="5" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FCB5BFD43530" box="[1175,1207,795,819]" italics="true" pageId="9" pageNumber="101">1.9</emphasis>
</td>
<td id="F233493C4138003AFB66FCB5BE463530" box="[1273,1317,795,819]" gridcol="8" gridrow="5" pageId="9" pageNumber="101">8.5</td>
<td id="F233493C4138003AFAC6FCB5BEEF3530" box="[1369,1420,795,819]" gridcol="9" gridrow="5" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FCB5BEEB3530" box="[1372,1416,795,819]" italics="true" pageId="9" pageNumber="101">18.5</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFCEDBEEF3558" box="[163,1420,835,859]" gridrow="6" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFCEDBABF3558" box="[163,476,835,859]" gridcol="0" gridrow="6" pageId="9" pageNumber="101">Buccal tube internal width</th>
<td id="F233493C4138003AFD99FCEDB97C3558" box="[518,543,835,859]" gridcol="1" gridrow="6" pageId="9" pageNumber="101">37</td>
<td id="F233493C4138003AFDD5FCEDB9D33558" box="[586,688,835,859]" gridcol="2" gridrow="6" pageId="9" pageNumber="101">4.8–8.9</td>
<td id="F233493C4138003AFD43FCEDB8213558" box="[732,834,835,859]" gridcol="3" gridrow="6" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FCEDB8213558" box="[732,834,835,859]" italics="true" pageId="9" pageNumber="101">10.8–17.4</emphasis>
</td>
<td id="F233493C4138003AFCEDFCEDB8FD3558" box="[882,926,835,859]" gridcol="4" gridrow="6" pageId="9" pageNumber="101">6.9</td>
<td id="F233493C4138003AFC54FCEDB89F3558" box="[971,1020,835,859]" gridcol="5" gridrow="6" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FCEDB89A3558" box="[973,1017,835,859]" italics="true" pageId="9" pageNumber="101">14.2</emphasis>
</td>
<td id="F233493C4138003AFBAEFCEDBF303558" box="[1073,1107,835,859]" gridcol="6" gridrow="6" pageId="9" pageNumber="101">1.1</td>
<td id="F233493C4138003AFB08FCEDBFD43558" box="[1175,1207,835,859]" gridcol="7" gridrow="6" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FCEDBFD43558" box="[1175,1207,835,859]" italics="true" pageId="9" pageNumber="101">1.7</emphasis>
</td>
<td id="F233493C4138003AFB66FCEDBE463558" box="[1273,1317,835,859]" gridcol="8" gridrow="6" pageId="9" pageNumber="101">6.2</td>
<td id="F233493C4138003AFAC6FCEDBEEF3558" box="[1369,1420,835,859]" gridcol="9" gridrow="6" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FCEDBEEB3558" box="[1372,1416,835,859]" italics="true" pageId="9" pageNumber="101">13.5</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFCC5BEEF3580" box="[163,1420,875,899]" gridrow="7" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFCC5BABF3580" box="[163,476,875,899]" gridcol="0" gridrow="7" pageId="9" pageNumber="101">Ventral lamina length</th>
<td id="F233493C4138003AFD99FCC5B97C3580" box="[518,543,875,899]" gridcol="1" gridrow="7" pageId="9" pageNumber="101">37</td>
<td id="F233493C4138003AFDD5FCC5B9D33580" box="[586,688,875,899]" gridcol="2" gridrow="7" pageId="9" pageNumber="101">23.1–43.4</td>
<td id="F233493C4138003AFD43FCC5B8213580" box="[732,834,875,899]" gridcol="3" gridrow="7" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FCC5B8223580" box="[732,833,875,899]" italics="true" pageId="9" pageNumber="101">53.9–74.5</emphasis>
</td>
<td id="F233493C4138003AFCEDFCC5B8FD3580" box="[882,926,875,899]" gridcol="4" gridrow="7" pageId="9" pageNumber="101">31.5</td>
<td id="F233493C4138003AFC54FCC5B89F3580" box="[971,1020,875,899]" gridcol="5" gridrow="7" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FCC5B89A3580" box="[973,1017,875,899]" italics="true" pageId="9" pageNumber="101">65.5</emphasis>
</td>
<td id="F233493C4138003AFBAEFCC5BF303580" box="[1073,1107,875,899]" gridcol="6" gridrow="7" pageId="9" pageNumber="101">3.7</td>
<td id="F233493C4138003AFB08FCC5BFD43580" box="[1175,1207,875,899]" gridcol="7" gridrow="7" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FCC5BFD43580" box="[1175,1207,875,899]" italics="true" pageId="9" pageNumber="101">5.0</emphasis>
</td>
<td id="F233493C4138003AFB66FCC5BE463580" box="[1273,1317,875,899]" gridcol="8" gridrow="7" pageId="9" pageNumber="101">32.6</td>
<td id="F233493C4138003AFAC6FCC5BEEF3580" box="[1369,1420,875,899]" gridcol="9" gridrow="7" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FCC5BEEB3580" box="[1372,1416,875,899]" italics="true" pageId="9" pageNumber="101">71.0</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFC3DBEEF35AF" box="[163,1420,915,940]" gridrow="8" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFC3DBABF35AF" box="[163,476,915,940]" gridcol="0" gridrow="8" pageId="9" pageNumber="101">Macroplacoid 1</th>
<td id="F233493C4138003AFD99FC3DB97C35AF" box="[518,543,915,940]" gridcol="1" gridrow="8" pageId="9" pageNumber="101">37</td>
<td id="F233493C4138003AFDD5FC3DB9D335AF" box="[586,688,915,940]" gridcol="2" gridrow="8" pageId="9" pageNumber="101">4.0–9.0</td>
<td id="F233493C4138003AFD43FC3DB82135AF" box="[732,834,915,940]" gridcol="3" gridrow="8" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD7CFC3DB85835A8" box="[739,827,915,939]" italics="true" pageId="9" pageNumber="101">8.6–18.4</emphasis>
</td>
<td id="F233493C4138003AFCEDFC3DB8FD35AF" box="[882,926,915,940]" gridcol="4" gridrow="8" pageId="9" pageNumber="101">6.1</td>
<td id="F233493C4138003AFC54FC3DB89F35AF" box="[971,1020,915,940]" gridcol="5" gridrow="8" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FC3AB89A35AF" box="[973,1017,916,940]" italics="true" pageId="9" pageNumber="101">12.7</emphasis>
</td>
<td id="F233493C4138003AFBAEFC3DBF3035AF" box="[1073,1107,915,940]" gridcol="6" gridrow="8" pageId="9" pageNumber="101">1.2</td>
<td id="F233493C4138003AFB08FC3DBFD435AF" box="[1175,1207,915,940]" gridcol="7" gridrow="8" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FC3ABFD435AF" box="[1175,1207,916,940]" italics="true" pageId="9" pageNumber="101">2.2</emphasis>
</td>
<td id="F233493C4138003AFB66FC3DBE4635AF" box="[1273,1317,915,940]" gridcol="8" gridrow="8" pageId="9" pageNumber="101">6.2</td>
<td id="F233493C4138003AFAC6FC3DBEEF35AF" box="[1369,1420,915,940]" gridcol="9" gridrow="8" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FC3ABEEB35AF" box="[1372,1416,916,940]" italics="true" pageId="9" pageNumber="101">13.5</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFC12BEEF35D7" box="[163,1420,956,980]" gridrow="9" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFC12BABF35D7" box="[163,476,956,980]" gridcol="0" gridrow="9" pageId="9" pageNumber="101">Macroplacoid 2</th>
<td id="F233493C4138003AFD99FC12B97C35D7" box="[518,543,956,980]" gridcol="1" gridrow="9" pageId="9" pageNumber="101">37</td>
<td id="F233493C4138003AFDD5FC12B9D335D7" box="[586,688,956,980]" gridcol="2" gridrow="9" pageId="9" pageNumber="101">2.6–7.0</td>
<td id="F233493C4138003AFD43FC12B82135D7" box="[732,834,956,980]" gridcol="3" gridrow="9" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD7CFC12B85F35D7" box="[739,828,956,980]" italics="true" pageId="9" pageNumber="101">5.7–13.2</emphasis>
</td>
<td id="F233493C4138003AFCEDFC12B8FD35D7" box="[882,926,956,980]" gridcol="4" gridrow="9" pageId="9" pageNumber="101">4.5</td>
<td id="F233493C4138003AFC54FC12B89F35D7" box="[971,1020,956,980]" gridcol="5" gridrow="9" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC4CFC12B89035D7" box="[979,1011,956,980]" italics="true" pageId="9" pageNumber="101">9.3</emphasis>
</td>
<td id="F233493C4138003AFBAEFC12BF3035D7" box="[1073,1107,956,980]" gridcol="6" gridrow="9" pageId="9" pageNumber="101">1.0</td>
<td id="F233493C4138003AFB08FC12BFD435D7" box="[1175,1207,956,980]" gridcol="7" gridrow="9" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FC12BFD435D7" box="[1175,1207,956,980]" italics="true" pageId="9" pageNumber="101">1.7</emphasis>
</td>
<td id="F233493C4138003AFB66FC12BE4635D7" box="[1273,1317,956,980]" gridcol="8" gridrow="9" pageId="9" pageNumber="101">4.0</td>
<td id="F233493C4138003AFAC6FC12BEEF35D7" box="[1369,1420,956,980]" gridcol="9" gridrow="9" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAFCFC12BEE035D7" box="[1379,1411,956,980]" italics="true" pageId="9" pageNumber="101">8.7</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFC4ABEEF35FF" box="[163,1420,996,1020]" gridrow="10" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFC4ABABF35FF" box="[163,476,996,1020]" gridcol="0" gridrow="10" pageId="9" pageNumber="101">Macroplacoid 3</th>
<td id="F233493C4138003AFD99FC4AB97C35FF" box="[518,543,996,1020]" gridcol="1" gridrow="10" pageId="9" pageNumber="101">36</td>
<td id="F233493C4138003AFDD5FC4AB9D335FF" box="[586,688,996,1020]" gridcol="2" gridrow="10" pageId="9" pageNumber="101">4.0–11.4</td>
<td id="F233493C4138003AFD43FC4AB82135FF" box="[732,834,996,1020]" gridcol="3" gridrow="10" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD7CFC4AB85835FF" box="[739,827,996,1020]" italics="true" pageId="9" pageNumber="101">9.1–20.9</emphasis>
</td>
<td id="F233493C4138003AFCEDFC4AB8FD35FF" box="[882,926,996,1020]" gridcol="4" gridrow="10" pageId="9" pageNumber="101">7.3</td>
<td id="F233493C4138003AFC54FC4AB89F35FF" box="[971,1020,996,1020]" gridcol="5" gridrow="10" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FC4AB89A35FF" box="[973,1017,996,1020]" italics="true" pageId="9" pageNumber="101">15.1</emphasis>
</td>
<td id="F233493C4138003AFBAEFC4ABF3035FF" box="[1073,1107,996,1020]" gridcol="6" gridrow="10" pageId="9" pageNumber="101">1.6</td>
<td id="F233493C4138003AFB08FC4ABFD435FF" box="[1175,1207,996,1020]" gridcol="7" gridrow="10" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FC4ABFD435FF" box="[1175,1207,996,1020]" italics="true" pageId="9" pageNumber="101">2.4</emphasis>
</td>
<td id="F233493C4138003AFB66FC4ABE4635FF" box="[1273,1317,996,1020]" gridcol="8" gridrow="10" pageId="9" pageNumber="101">7.1</td>
<td id="F233493C4138003AFAC6FC4ABEEF35FF" box="[1369,1420,996,1020]" gridcol="9" gridrow="10" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FC4ABEEB35FF" box="[1372,1416,996,1020]" italics="true" pageId="9" pageNumber="101">15.5</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFBA2BEEF3227" box="[163,1420,1036,1060]" gridrow="11" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFBA2BABF3227" box="[163,476,1036,1060]" gridcol="0" gridrow="11" pageId="9" pageNumber="101">Microplacoid</th>
<td id="F233493C4138003AFD99FBA2B97C3227" box="[518,543,1036,1060]" gridcol="1" gridrow="11" pageId="9" pageNumber="101">37</td>
<td id="F233493C4138003AFDD5FBA2B9D33227" box="[586,688,1036,1060]" gridcol="2" gridrow="11" pageId="9" pageNumber="101">1.1–4.2</td>
<td id="F233493C4138003AFD43FBA2B8213227" box="[732,834,1036,1060]" gridcol="3" gridrow="11" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD76FBA2B8563227" box="[745,821,1036,1060]" italics="true" pageId="9" pageNumber="101">2.5–8.7</emphasis>
</td>
<td id="F233493C4138003AFCEDFBA2B8FD3227" box="[882,926,1036,1060]" gridcol="4" gridrow="11" pageId="9" pageNumber="101">2.6</td>
<td id="F233493C4138003AFC54FBA2B89F3227" box="[971,1020,1036,1060]" gridcol="5" gridrow="11" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC4CFBA2B8903227" box="[979,1011,1036,1060]" italics="true" pageId="9" pageNumber="101">5.3</emphasis>
</td>
<td id="F233493C4138003AFBAEFBA2BF303227" box="[1073,1107,1036,1060]" gridcol="6" gridrow="11" pageId="9" pageNumber="101">0.8</td>
<td id="F233493C4138003AFB08FBA2BFD43227" box="[1175,1207,1036,1060]" gridcol="7" gridrow="11" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FBA2BFD43227" box="[1175,1207,1036,1060]" italics="true" pageId="9" pageNumber="101">1.4</emphasis>
</td>
<td id="F233493C4138003AFB66FBA2BE463227" box="[1273,1317,1036,1060]" gridcol="8" gridrow="11" pageId="9" pageNumber="101">2.7</td>
<td id="F233493C4138003AFAC6FBA2BEEF3227" box="[1369,1420,1036,1060]" gridcol="9" gridrow="11" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAFCFBA2BEE03227" box="[1379,1411,1036,1060]" italics="true" pageId="9" pageNumber="101">5.9</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFB9ABEEF324E" box="[163,1420,1076,1101]" gridrow="12" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFB9ABABF324E" box="[163,476,1076,1101]" gridcol="0" gridrow="12" pageId="9" pageNumber="101">Macroplacoid row</th>
<td id="F233493C4138003AFD99FB9AB97C324E" box="[518,543,1076,1101]" gridcol="1" gridrow="12" pageId="9" pageNumber="101">37</td>
<td id="F233493C4138003AFDD5FB9AB9D3324E" box="[586,688,1076,1101]" gridcol="2" gridrow="12" pageId="9" pageNumber="101">18.5–34.6</td>
<td id="F233493C4138003AFD43FB9AB821324E" box="[732,834,1076,1101]" gridcol="3" gridrow="12" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FB9BB822324E" box="[732,833,1077,1101]" italics="true" pageId="9" pageNumber="101">40.3–61.8</emphasis>
</td>
<td id="F233493C4138003AFCEDFB9AB8FD324E" box="[882,926,1076,1101]" gridcol="4" gridrow="12" pageId="9" pageNumber="101">24.4</td>
<td id="F233493C4138003AFC54FB9AB89F324E" box="[971,1020,1076,1101]" gridcol="5" gridrow="12" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FB9BB89A324E" box="[973,1017,1077,1101]" italics="true" pageId="9" pageNumber="101">50.8</emphasis>
</td>
<td id="F233493C4138003AFBAEFB9ABF30324E" box="[1073,1107,1076,1101]" gridcol="6" gridrow="12" pageId="9" pageNumber="101">3.5</td>
<td id="F233493C4138003AFB08FB9ABFD4324E" box="[1175,1207,1076,1101]" gridcol="7" gridrow="12" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FB9BBFD4324E" box="[1175,1207,1077,1101]" italics="true" pageId="9" pageNumber="101">5.3</emphasis>
</td>
<td id="F233493C4138003AFB66FB9ABE46324E" box="[1273,1317,1076,1101]" gridcol="8" gridrow="12" pageId="9" pageNumber="101">23.9</td>
<td id="F233493C4138003AFAC6FB9ABEEF324E" box="[1369,1420,1076,1101]" gridcol="9" gridrow="12" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FB9BBEEB324E" box="[1372,1416,1077,1101]" italics="true" pageId="9" pageNumber="101">52.0</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFBF3BEEF3276" box="[163,1420,1117,1141]" gridrow="13" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFBF3BABF3276" box="[163,476,1117,1141]" gridcol="0" gridrow="13" pageId="9" pageNumber="101">Placoid row</th>
<td id="F233493C4138003AFD99FBF3B97C3276" box="[518,543,1117,1141]" gridcol="1" gridrow="13" pageId="9" pageNumber="101">37</td>
<td id="F233493C4138003AFDD5FBF3B9D33276" box="[586,688,1117,1141]" gridcol="2" gridrow="13" pageId="9" pageNumber="101">26.8–44.5</td>
<td id="F233493C4138003AFD43FBF3B8213276" box="[732,834,1117,1141]" gridcol="3" gridrow="13" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FBF3B8223276" box="[732,833,1117,1141]" italics="true" pageId="9" pageNumber="101">58.3–79.7</emphasis>
</td>
<td id="F233493C4138003AFCEDFBF3B8FD3276" box="[882,926,1117,1141]" gridcol="4" gridrow="13" pageId="9" pageNumber="101">32.9</td>
<td id="F233493C4138003AFC54FBF3B89F3276" box="[971,1020,1117,1141]" gridcol="5" gridrow="13" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FBF3B89A3276" box="[973,1017,1117,1141]" italics="true" pageId="9" pageNumber="101">68.4</emphasis>
</td>
<td id="F233493C4138003AFBAEFBF3BF303276" box="[1073,1107,1117,1141]" gridcol="6" gridrow="13" pageId="9" pageNumber="101">4.2</td>
<td id="F233493C4138003AFB08FBF3BFD43276" box="[1175,1207,1117,1141]" gridcol="7" gridrow="13" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FBF3BFD43276" box="[1175,1207,1117,1141]" italics="true" pageId="9" pageNumber="101">5.8</emphasis>
</td>
<td id="F233493C4138003AFB66FBF3BE463276" box="[1273,1317,1117,1141]" gridcol="8" gridrow="13" pageId="9" pageNumber="101">31.6</td>
<td id="F233493C4138003AFAC6FBF3BEEF3276" box="[1369,1420,1117,1141]" gridcol="9" gridrow="13" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FBF3BEEB3276" box="[1372,1416,1117,1141]" italics="true" pageId="9" pageNumber="101">68.8</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFB2BBEEF329E" box="[163,1420,1157,1181]" gridrow="14" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFB2BBEEF329E" box="[163,1420,1157,1181]" colspan="10" colspanRight="9" gridcol="0" gridrow="14" pageId="9" pageNumber="101">Claw I heights</th>
</tr>
<tr id="B1E220404138003AFF3CFB03BEEF32C5" box="[163,1420,1197,1222]" gridrow="15" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFB03BABF32C5" box="[163,476,1197,1222]" gridcol="0" gridrow="15" pageId="9" pageNumber="101">External primary branch</th>
<td id="F233493C4138003AFD99FB03B97C32C5" box="[518,543,1197,1222]" gridcol="1" gridrow="15" pageId="9" pageNumber="101">40</td>
<td id="F233493C4138003AFDD5FB03B9D332C5" box="[586,688,1197,1222]" gridcol="2" gridrow="15" pageId="9" pageNumber="101">8.1–16.1</td>
<td id="F233493C4138003AFD43FB03B82132C5" box="[732,834,1197,1222]" gridcol="3" gridrow="15" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FB00B82232C5" box="[732,833,1198,1222]" italics="true" pageId="9" pageNumber="101">17.1–30.5</emphasis>
</td>
<td id="F233493C4138003AFCEDFB03B8FD32C5" box="[882,926,1197,1222]" gridcol="4" gridrow="15" pageId="9" pageNumber="101">11.4</td>
<td id="F233493C4138003AFC54FB03B89F32C5" box="[971,1020,1197,1222]" gridcol="5" gridrow="15" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FB00B89A32C5" box="[973,1017,1198,1222]" italics="true" pageId="9" pageNumber="101">23.8</emphasis>
</td>
<td id="F233493C4138003AFBAEFB03BF3032C5" box="[1073,1107,1197,1222]" gridcol="6" gridrow="15" pageId="9" pageNumber="101">1.8</td>
<td id="F233493C4138003AFB08FB03BFD432C5" box="[1175,1207,1197,1222]" gridcol="7" gridrow="15" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FB00BFD432C5" box="[1175,1207,1198,1222]" italics="true" pageId="9" pageNumber="101">3.0</emphasis>
</td>
<td id="F233493C4138003AFB66FB03BE4632C5" box="[1273,1317,1197,1222]" gridcol="8" gridrow="15" pageId="9" pageNumber="101">10.2</td>
<td id="F233493C4138003AFAC6FB03BEEF32C5" box="[1369,1420,1197,1222]" gridcol="9" gridrow="15" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FB00BEEB32C5" box="[1372,1416,1198,1222]" italics="true" pageId="9" pageNumber="101">22.3</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFB7BBEEF32ED" box="[163,1420,1237,1262]" gridrow="16" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFB7BBABF32ED" box="[163,476,1237,1262]" gridcol="0" gridrow="16" pageId="9" pageNumber="101">External secondary branch</th>
<td id="F233493C4138003AFD99FB7BB97C32ED" box="[518,543,1237,1262]" gridcol="1" gridrow="16" pageId="9" pageNumber="101">40</td>
<td id="F233493C4138003AFDD5FB7BB9D332ED" box="[586,688,1237,1262]" gridcol="2" gridrow="16" pageId="9" pageNumber="101">4.4–12.0</td>
<td id="F233493C4138003AFD43FB7BB82132ED" box="[732,834,1237,1262]" gridcol="3" gridrow="16" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD7CFB78B85F32ED" box="[739,828,1238,1262]" italics="true" pageId="9" pageNumber="101">9.3–22.4</emphasis>
</td>
<td id="F233493C4138003AFCEDFB7BB8FD32ED" box="[882,926,1237,1262]" gridcol="4" gridrow="16" pageId="9" pageNumber="101">7.5</td>
<td id="F233493C4138003AFC54FB7BB89F32ED" box="[971,1020,1237,1262]" gridcol="5" gridrow="16" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FB78B89A32ED" box="[973,1017,1238,1262]" italics="true" pageId="9" pageNumber="101">15.6</emphasis>
</td>
<td id="F233493C4138003AFBAEFB7BBF3032ED" box="[1073,1107,1237,1262]" gridcol="6" gridrow="16" pageId="9" pageNumber="101">1.5</td>
<td id="F233493C4138003AFB08FB7BBFD432ED" box="[1175,1207,1237,1262]" gridcol="7" gridrow="16" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FB78BFD432ED" box="[1175,1207,1238,1262]" italics="true" pageId="9" pageNumber="101">3.1</emphasis>
</td>
<td id="F233493C4138003AFB66FB7BBE4632ED" box="[1273,1317,1237,1262]" gridcol="8" gridrow="16" pageId="9" pageNumber="101">6.5</td>
<td id="F233493C4138003AFAC6FB7BBEEF32ED" box="[1369,1420,1237,1262]" gridcol="9" gridrow="16" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FB78BEEB32ED" box="[1372,1416,1238,1262]" italics="true" pageId="9" pageNumber="101">14.1</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFB50BEEF3315" box="[163,1420,1278,1302]" gridrow="17" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFB50BABF3315" box="[163,476,1278,1302]" gridcol="0" gridrow="17" pageId="9" pageNumber="101">Internal primary branch</th>
<td id="F233493C4138003AFD99FB50B97C3315" box="[518,543,1278,1302]" gridcol="1" gridrow="17" pageId="9" pageNumber="101">39</td>
<td id="F233493C4138003AFDD5FB50B9D33315" box="[586,688,1278,1302]" gridcol="2" gridrow="17" pageId="9" pageNumber="101">5.8–16.0</td>
<td id="F233493C4138003AFD43FB50B8213315" box="[732,834,1278,1302]" gridcol="3" gridrow="17" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FB50B8213315" box="[732,834,1278,1302]" italics="true" pageId="9" pageNumber="101">12.7–32.8</emphasis>
</td>
<td id="F233493C4138003AFCEDFB50B8FD3315" box="[882,926,1278,1302]" gridcol="4" gridrow="17" pageId="9" pageNumber="101">11.0</td>
<td id="F233493C4138003AFC54FB50B89F3315" box="[971,1020,1278,1302]" gridcol="5" gridrow="17" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FB50B89A3315" box="[973,1017,1278,1302]" italics="true" pageId="9" pageNumber="101">23.0</emphasis>
</td>
<td id="F233493C4138003AFBAEFB50BF303315" box="[1073,1107,1278,1302]" gridcol="6" gridrow="17" pageId="9" pageNumber="101">2.0</td>
<td id="F233493C4138003AFB08FB50BFD43315" box="[1175,1207,1278,1302]" gridcol="7" gridrow="17" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FB50BFD43315" box="[1175,1207,1278,1302]" italics="true" pageId="9" pageNumber="101">4.0</emphasis>
</td>
<td id="F233493C4138003AFB66FB50BE463315" box="[1273,1317,1278,1302]" gridcol="8" gridrow="17" pageId="9" pageNumber="101">5.8</td>
<td id="F233493C4138003AFAC6FB50BEEF3315" box="[1369,1420,1278,1302]" gridcol="9" gridrow="17" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FB50BEEB3315" box="[1372,1416,1278,1302]" italics="true" pageId="9" pageNumber="101">12.7</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFA88BEEF333D" box="[163,1420,1318,1342]" gridrow="18" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFA88BABF333D" box="[163,476,1318,1342]" gridcol="0" gridrow="18" pageId="9" pageNumber="101">Internal secondary branch</th>
<td id="F233493C4138003AFD99FA88B97C333D" box="[518,543,1318,1342]" gridcol="1" gridrow="18" pageId="9" pageNumber="101">39</td>
<td id="F233493C4138003AFDD5FA88B9D3333D" box="[586,688,1318,1342]" gridcol="2" gridrow="18" pageId="9" pageNumber="101">5.2–12.4</td>
<td id="F233493C4138003AFD43FA88B821333D" box="[732,834,1318,1342]" gridcol="3" gridrow="18" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD42FA88B823333D" box="[733,832,1318,1342]" italics="true" pageId="9" pageNumber="101">11.6–25.8</emphasis>
</td>
<td id="F233493C4138003AFCEDFA88B8FD333D" box="[882,926,1318,1342]" gridcol="4" gridrow="18" pageId="9" pageNumber="101">7.9</td>
<td id="F233493C4138003AFC54FA88B89F333D" box="[971,1020,1318,1342]" gridcol="5" gridrow="18" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FA88B89A333D" box="[973,1017,1318,1342]" italics="true" pageId="9" pageNumber="101">16.6</emphasis>
</td>
<td id="F233493C4138003AFBAEFA88BF30333D" box="[1073,1107,1318,1342]" gridcol="6" gridrow="18" pageId="9" pageNumber="101">1.7</td>
<td id="F233493C4138003AFB08FA88BFD4333D" box="[1175,1207,1318,1342]" gridcol="7" gridrow="18" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FA88BFD4333D" box="[1175,1207,1318,1342]" italics="true" pageId="9" pageNumber="101">3.4</emphasis>
</td>
<td id="F233493C4138003AFB66FA88BE46333D" box="[1273,1317,1318,1342]" gridcol="8" gridrow="18" pageId="9" pageNumber="101">5.3</td>
<td id="F233493C4138003AFAC6FA88BEEF333D" box="[1369,1420,1318,1342]" gridcol="9" gridrow="18" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC2FA88BEE4333D" box="[1373,1415,1318,1342]" italics="true" pageId="9" pageNumber="101">11.6</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFAE0BEEF3365" box="[163,1420,1358,1382]" gridrow="19" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFAE0BEEF3365" box="[163,1420,1358,1382]" colspan="10" colspanRight="9" gridcol="0" gridrow="19" pageId="9" pageNumber="101">Claw II heights</th>
</tr>
<tr id="B1E220404138003AFF3CFAD9BEEF338C" box="[163,1420,1399,1423]" gridrow="20" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFAD9BABF338C" box="[163,476,1399,1423]" gridcol="0" gridrow="20" pageId="9" pageNumber="101">External primary branch</th>
<td id="F233493C4138003AFD99FAD9B97C338C" box="[518,543,1399,1423]" gridcol="1" gridrow="20" pageId="9" pageNumber="101">38</td>
<td id="F233493C4138003AFDD5FAD9B9D3338C" box="[586,688,1399,1423]" gridcol="2" gridrow="20" pageId="9" pageNumber="101">7.6–16.3</td>
<td id="F233493C4138003AFD43FAD9B821338C" box="[732,834,1399,1423]" gridcol="3" gridrow="20" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FAD9B822338C" box="[732,833,1399,1423]" italics="true" pageId="9" pageNumber="101">16.4–36.2</emphasis>
</td>
<td id="F233493C4138003AFCEDFAD9B8FD338C" box="[882,926,1399,1423]" gridcol="4" gridrow="20" pageId="9" pageNumber="101">11.9</td>
<td id="F233493C4138003AFC54FAD9B89F338C" box="[971,1020,1399,1423]" gridcol="5" gridrow="20" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FAD9B89A338C" box="[973,1017,1399,1423]" italics="true" pageId="9" pageNumber="101">24.8</emphasis>
</td>
<td id="F233493C4138003AFBAEFAD9BF30338C" box="[1073,1107,1399,1423]" gridcol="6" gridrow="20" pageId="9" pageNumber="101">2.0</td>
<td id="F233493C4138003AFB08FAD9BFD4338C" box="[1175,1207,1399,1423]" gridcol="7" gridrow="20" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FAD9BFD4338C" box="[1175,1207,1399,1423]" italics="true" pageId="9" pageNumber="101">4.3</emphasis>
</td>
<td id="F233493C4138003AFB66FAD9BE46338C" box="[1273,1317,1399,1423]" gridcol="8" gridrow="20" pageId="9" pageNumber="101">9.9</td>
<td id="F233493C4138003AFAC6FAD9BEEF338C" box="[1369,1420,1399,1423]" gridcol="9" gridrow="20" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FAD9BEEB338C" box="[1372,1416,1399,1423]" italics="true" pageId="9" pageNumber="101">21.6</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFA31BEEF33B4" box="[163,1420,1439,1463]" gridrow="21" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFA31BABF33B4" box="[163,476,1439,1463]" gridcol="0" gridrow="21" pageId="9" pageNumber="101">External secondary branch</th>
<td id="F233493C4138003AFD99FA31B97C33B4" box="[518,543,1439,1463]" gridcol="1" gridrow="21" pageId="9" pageNumber="101">38</td>
<td id="F233493C4138003AFDD5FA31B9D333B4" box="[586,688,1439,1463]" gridcol="2" gridrow="21" pageId="9" pageNumber="101">5.4–11.7</td>
<td id="F233493C4138003AFD43FA31B82133B4" box="[732,834,1439,1463]" gridcol="3" gridrow="21" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD42FA31B82233B4" box="[733,833,1439,1463]" italics="true" pageId="9" pageNumber="101">11.8–23.9</emphasis>
</td>
<td id="F233493C4138003AFCEDFA31B8FD33B4" box="[882,926,1439,1463]" gridcol="4" gridrow="21" pageId="9" pageNumber="101">8.4</td>
<td id="F233493C4138003AFC54FA31B89F33B4" box="[971,1020,1439,1463]" gridcol="5" gridrow="21" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FA31B89A33B4" box="[973,1017,1439,1463]" italics="true" pageId="9" pageNumber="101">17.5</emphasis>
</td>
<td id="F233493C4138003AFBAEFA31BF3033B4" box="[1073,1107,1439,1463]" gridcol="6" gridrow="21" pageId="9" pageNumber="101">1.4</td>
<td id="F233493C4138003AFB08FA31BFD433B4" box="[1175,1207,1439,1463]" gridcol="7" gridrow="21" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FA31BFD433B4" box="[1175,1207,1439,1463]" italics="true" pageId="9" pageNumber="101">2.9</emphasis>
</td>
<td id="F233493C4138003AFB66FA31BE4633B4" box="[1273,1317,1439,1463]" gridcol="8" gridrow="21" pageId="9" pageNumber="101">8.0</td>
<td id="F233493C4138003AFAC6FA31BEEF33B4" box="[1369,1420,1439,1463]" gridcol="9" gridrow="21" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FA31BEEB33B4" box="[1372,1416,1439,1463]" italics="true" pageId="9" pageNumber="101">17.4</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFA69BEEF33DC" box="[163,1420,1479,1503]" gridrow="22" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFA69BABF33DC" box="[163,476,1479,1503]" gridcol="0" gridrow="22" pageId="9" pageNumber="101">Internal primary branch</th>
<td id="F233493C4138003AFD99FA69B97C33DC" box="[518,543,1479,1503]" gridcol="1" gridrow="22" pageId="9" pageNumber="101">38</td>
<td id="F233493C4138003AFDD5FA69B9D333DC" box="[586,688,1479,1503]" gridcol="2" gridrow="22" pageId="9" pageNumber="101">7.5–15.3</td>
<td id="F233493C4138003AFD43FA69B82133DC" box="[732,834,1479,1503]" gridcol="3" gridrow="22" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FA69B82233DC" box="[732,833,1479,1503]" italics="true" pageId="9" pageNumber="101">16.3–30.6</emphasis>
</td>
<td id="F233493C4138003AFCEDFA69B8FD33DC" box="[882,926,1479,1503]" gridcol="4" gridrow="22" pageId="9" pageNumber="101">11.0</td>
<td id="F233493C4138003AFC54FA69B89F33DC" box="[971,1020,1479,1503]" gridcol="5" gridrow="22" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FA69B89A33DC" box="[973,1017,1479,1503]" italics="true" pageId="9" pageNumber="101">22.9</emphasis>
</td>
<td id="F233493C4138003AFBAEFA69BF3033DC" box="[1073,1107,1479,1503]" gridcol="6" gridrow="22" pageId="9" pageNumber="101">2.0</td>
<td id="F233493C4138003AFB08FA69BFD433DC" box="[1175,1207,1479,1503]" gridcol="7" gridrow="22" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FA69BFD433DC" box="[1175,1207,1479,1503]" italics="true" pageId="9" pageNumber="101">3.6</emphasis>
</td>
<td id="F233493C4138003AFB66FA69BE4633DC" box="[1273,1317,1479,1503]" gridcol="8" gridrow="22" pageId="9" pageNumber="101">9.3</td>
<td id="F233493C4138003AFAC6FA69BEEF33DC" box="[1369,1420,1479,1503]" gridcol="9" gridrow="22" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FA69BEEB33DC" box="[1372,1416,1479,1503]" italics="true" pageId="9" pageNumber="101">20.2</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CFA41BEEF300B" box="[163,1420,1519,1544]" gridrow="23" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CFA41BABF300B" box="[163,476,1519,1544]" gridcol="0" gridrow="23" pageId="9" pageNumber="101">Internal secondary branch</th>
<td id="F233493C4138003AFD99FA41B97C300B" box="[518,543,1519,1544]" gridcol="1" gridrow="23" pageId="9" pageNumber="101">38</td>
<td id="F233493C4138003AFDD5FA41B9D3300B" box="[586,688,1519,1544]" gridcol="2" gridrow="23" pageId="9" pageNumber="101">4.4–11.6</td>
<td id="F233493C4138003AFD43FA41B821300B" box="[732,834,1519,1544]" gridcol="3" gridrow="23" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43FA5EB821300B" box="[732,834,1520,1544]" italics="true" pageId="9" pageNumber="101">10.4–22.0</emphasis>
</td>
<td id="F233493C4138003AFCEDFA41B8FD300B" box="[882,926,1519,1544]" gridcol="4" gridrow="23" pageId="9" pageNumber="101">7.6</td>
<td id="F233493C4138003AFC54FA41B89F300B" box="[971,1020,1519,1544]" gridcol="5" gridrow="23" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52FA5EB89A300B" box="[973,1017,1520,1544]" italics="true" pageId="9" pageNumber="101">15.9</emphasis>
</td>
<td id="F233493C4138003AFBAEFA41BF30300B" box="[1073,1107,1519,1544]" gridcol="6" gridrow="23" pageId="9" pageNumber="101">1.7</td>
<td id="F233493C4138003AFB08FA41BFD4300B" box="[1175,1207,1519,1544]" gridcol="7" gridrow="23" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08FA5EBFD4300B" box="[1175,1207,1520,1544]" italics="true" pageId="9" pageNumber="101">3.2</emphasis>
</td>
<td id="F233493C4138003AFB66FA41BE46300B" box="[1273,1317,1519,1544]" gridcol="8" gridrow="23" pageId="9" pageNumber="101">5.9</td>
<td id="F233493C4138003AFAC6FA41BEEF300B" box="[1369,1420,1519,1544]" gridcol="9" gridrow="23" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3FA5EBEEB300B" box="[1372,1416,1520,1544]" italics="true" pageId="9" pageNumber="101">12.9</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CF9B6BEEF3033" box="[163,1420,1560,1584]" gridrow="24" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CF9B6BEEF3033" box="[163,1420,1560,1584]" colspan="10" colspanRight="9" gridcol="0" gridrow="24" pageId="9" pageNumber="101">Claw III heights</th>
</tr>
<tr id="B1E220404138003AFF3CF9EEBEEF305B" box="[163,1420,1600,1624]" gridrow="25" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CF9EEBABF305B" box="[163,476,1600,1624]" gridcol="0" gridrow="25" pageId="9" pageNumber="101">External primary branch</th>
<td id="F233493C4138003AFD99F9EEB97C305B" box="[518,543,1600,1624]" gridcol="1" gridrow="25" pageId="9" pageNumber="101">40</td>
<td id="F233493C4138003AFDD5F9EEB9D3305B" box="[586,688,1600,1624]" gridcol="2" gridrow="25" pageId="9" pageNumber="101">7.4–15.0</td>
<td id="F233493C4138003AFD43F9EEB821305B" box="[732,834,1600,1624]" gridcol="3" gridrow="25" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43F9EEB821305B" box="[732,834,1600,1624]" italics="true" pageId="9" pageNumber="101">16.3–31.5</emphasis>
</td>
<td id="F233493C4138003AFCEDF9EEB8FD305B" box="[882,926,1600,1624]" gridcol="4" gridrow="25" pageId="9" pageNumber="101">11.7</td>
<td id="F233493C4138003AFC54F9EEB89F305B" box="[971,1020,1600,1624]" gridcol="5" gridrow="25" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52F9EEB89A305B" box="[973,1017,1600,1624]" italics="true" pageId="9" pageNumber="101">24.6</emphasis>
</td>
<td id="F233493C4138003AFBAEF9EEBF30305B" box="[1073,1107,1600,1624]" gridcol="6" gridrow="25" pageId="9" pageNumber="101">1.9</td>
<td id="F233493C4138003AFB08F9EEBFD4305B" box="[1175,1207,1600,1624]" gridcol="7" gridrow="25" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08F9EEBFD4305B" box="[1175,1207,1600,1624]" italics="true" pageId="9" pageNumber="101">3.3</emphasis>
</td>
<td id="F233493C4138003AFB66F9EEBE46305B" box="[1273,1317,1600,1624]" gridcol="8" gridrow="25" pageId="9" pageNumber="101">11.4</td>
<td id="F233493C4138003AFAC6F9EEBEEF305B" box="[1369,1420,1600,1624]" gridcol="9" gridrow="25" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3F9EEBEEB305B" box="[1372,1416,1600,1624]" italics="true" pageId="9" pageNumber="101">24.8</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CF9C6BEEF3083" box="[163,1420,1640,1664]" gridrow="26" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CF9C6BABF3083" box="[163,476,1640,1664]" gridcol="0" gridrow="26" pageId="9" pageNumber="101">External secondary branch</th>
<td id="F233493C4138003AFD99F9C6B97C3083" box="[518,543,1640,1664]" gridcol="1" gridrow="26" pageId="9" pageNumber="101">40</td>
<td id="F233493C4138003AFDD5F9C6B9D33083" box="[586,688,1640,1664]" gridcol="2" gridrow="26" pageId="9" pageNumber="101">5.2–11.5</td>
<td id="F233493C4138003AFD43F9C6B8213083" box="[732,834,1640,1664]" gridcol="3" gridrow="26" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43F9C6B8223083" box="[732,833,1640,1664]" italics="true" pageId="9" pageNumber="101">10.6–21.8</emphasis>
</td>
<td id="F233493C4138003AFCEDF9C6B8FD3083" box="[882,926,1640,1664]" gridcol="4" gridrow="26" pageId="9" pageNumber="101">8.2</td>
<td id="F233493C4138003AFC54F9C6B89F3083" box="[971,1020,1640,1664]" gridcol="5" gridrow="26" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52F9C6B89A3083" box="[973,1017,1640,1664]" italics="true" pageId="9" pageNumber="101">17.1</emphasis>
</td>
<td id="F233493C4138003AFBAEF9C6BF303083" box="[1073,1107,1640,1664]" gridcol="6" gridrow="26" pageId="9" pageNumber="101">1.6</td>
<td id="F233493C4138003AFB08F9C6BFD43083" box="[1175,1207,1640,1664]" gridcol="7" gridrow="26" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08F9C6BFD43083" box="[1175,1207,1640,1664]" italics="true" pageId="9" pageNumber="101">3.1</emphasis>
</td>
<td id="F233493C4138003AFB66F9C6BE463083" box="[1273,1317,1640,1664]" gridcol="8" gridrow="26" pageId="9" pageNumber="101">9.5</td>
<td id="F233493C4138003AFAC6F9C6BEEF3083" box="[1369,1420,1640,1664]" gridcol="9" gridrow="26" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3F9C6BEEB3083" box="[1372,1416,1640,1664]" italics="true" pageId="9" pageNumber="101">20.7</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CF93EBEEF30AA" box="[163,1420,1680,1705]" gridrow="27" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CF93EBABF30AA" box="[163,476,1680,1705]" gridcol="0" gridrow="27" pageId="9" pageNumber="101">Internal primary branch</th>
<td id="F233493C4138003AFD99F93EB97C30AA" box="[518,543,1680,1705]" gridcol="1" gridrow="27" pageId="9" pageNumber="101">40</td>
<td id="F233493C4138003AFDD5F93EB9D330AA" box="[586,688,1680,1705]" gridcol="2" gridrow="27" pageId="9" pageNumber="101">8.0–14.9</td>
<td id="F233493C4138003AFD43F93EB82130AA" box="[732,834,1680,1705]" gridcol="3" gridrow="27" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43F93FB82230AA" box="[732,833,1681,1705]" italics="true" pageId="9" pageNumber="101">15.9–31.1</emphasis>
</td>
<td id="F233493C4138003AFCEDF93EB8FD30AA" box="[882,926,1680,1705]" gridcol="4" gridrow="27" pageId="9" pageNumber="101">10.9</td>
<td id="F233493C4138003AFC54F93EB89F30AA" box="[971,1020,1680,1705]" gridcol="5" gridrow="27" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52F93FB89A30AA" box="[973,1017,1681,1705]" italics="true" pageId="9" pageNumber="101">22.9</emphasis>
</td>
<td id="F233493C4138003AFBAEF93EBF3030AA" box="[1073,1107,1680,1705]" gridcol="6" gridrow="27" pageId="9" pageNumber="101">1.7</td>
<td id="F233493C4138003AFB08F93EBFD430AA" box="[1175,1207,1680,1705]" gridcol="7" gridrow="27" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08F93FBFD430AA" box="[1175,1207,1681,1705]" italics="true" pageId="9" pageNumber="101">3.4</emphasis>
</td>
<td id="F233493C4138003AFB66F93EBE4630AA" box="[1273,1317,1680,1705]" gridcol="8" gridrow="27" pageId="9" pageNumber="101">11.8</td>
<td id="F233493C4138003AFAC6F93EBEEF30AA" box="[1369,1420,1680,1705]" gridcol="9" gridrow="27" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3F93FBEEB30AA" box="[1372,1416,1681,1705]" italics="true" pageId="9" pageNumber="101">25.7</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CF917BEEF30D2" box="[163,1420,1721,1745]" gridrow="28" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CF917BABF30D2" box="[163,476,1721,1745]" gridcol="0" gridrow="28" pageId="9" pageNumber="101">Internal secondary branch</th>
<td id="F233493C4138003AFD99F917B97C30D2" box="[518,543,1721,1745]" gridcol="1" gridrow="28" pageId="9" pageNumber="101">40</td>
<td id="F233493C4138003AFDD5F917B9D330D2" box="[586,688,1721,1745]" gridcol="2" gridrow="28" pageId="9" pageNumber="101">4.5–10.1</td>
<td id="F233493C4138003AFD43F917B82130D2" box="[732,834,1721,1745]" gridcol="3" gridrow="28" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD7CF917B85830D2" box="[739,827,1721,1745]" italics="true" pageId="9" pageNumber="101">8.5–22.4</emphasis>
</td>
<td id="F233493C4138003AFCEDF917B8FD30D2" box="[882,926,1721,1745]" gridcol="4" gridrow="28" pageId="9" pageNumber="101">8.0</td>
<td id="F233493C4138003AFC54F917B89F30D2" box="[971,1020,1721,1745]" gridcol="5" gridrow="28" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52F917B89A30D2" box="[973,1017,1721,1745]" italics="true" pageId="9" pageNumber="101">16.7</emphasis>
</td>
<td id="F233493C4138003AFBAEF917BF3030D2" box="[1073,1107,1721,1745]" gridcol="6" gridrow="28" pageId="9" pageNumber="101">1.3</td>
<td id="F233493C4138003AFB08F917BFD430D2" box="[1175,1207,1721,1745]" gridcol="7" gridrow="28" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08F917BFD430D2" box="[1175,1207,1721,1745]" italics="true" pageId="9" pageNumber="101">2.8</emphasis>
</td>
<td id="F233493C4138003AFB66F917BE4630D2" box="[1273,1317,1721,1745]" gridcol="8" gridrow="28" pageId="9" pageNumber="101">8.8</td>
<td id="F233493C4138003AFAC6F917BEEF30D2" box="[1369,1420,1721,1745]" gridcol="9" gridrow="28" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3F917BEEB30D2" box="[1372,1416,1721,1745]" italics="true" pageId="9" pageNumber="101">19.2</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CF94FBEEF30FA" box="[163,1420,1761,1785]" gridrow="29" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CF94FBEEF30FA" box="[163,1420,1761,1785]" colspan="10" colspanRight="9" gridcol="0" gridrow="29" pageId="9" pageNumber="101">Claw IV heights</th>
</tr>
<tr id="B1E220404138003AFF3CF8A7BEEF3122" box="[163,1420,1801,1825]" gridrow="30" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CF8A7BABF3122" box="[163,476,1801,1825]" gridcol="0" gridrow="30" pageId="9" pageNumber="101">Anterior primary branch</th>
<td id="F233493C4138003AFD99F8A7B97C3122" box="[518,543,1801,1825]" gridcol="1" gridrow="30" pageId="9" pageNumber="101">40</td>
<td id="F233493C4138003AFDD5F8A7B9D33122" box="[586,688,1801,1825]" gridcol="2" gridrow="30" pageId="9" pageNumber="101">9.1–16.8</td>
<td id="F233493C4138003AFD43F8A7B8213122" box="[732,834,1801,1825]" gridcol="3" gridrow="30" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43F8A7B8223122" box="[732,833,1801,1825]" italics="true" pageId="9" pageNumber="101">16.4–32.6</emphasis>
</td>
<td id="F233493C4138003AFCEDF8A7B8FD3122" box="[882,926,1801,1825]" gridcol="4" gridrow="30" pageId="9" pageNumber="101">12.7</td>
<td id="F233493C4138003AFC54F8A7B89F3122" box="[971,1020,1801,1825]" gridcol="5" gridrow="30" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52F8A7B89A3122" box="[973,1017,1801,1825]" italics="true" pageId="9" pageNumber="101">26.3</emphasis>
</td>
<td id="F233493C4138003AFBAEF8A7BF303122" box="[1073,1107,1801,1825]" gridcol="6" gridrow="30" pageId="9" pageNumber="101">1.8</td>
<td id="F233493C4138003AFB08F8A7BFD43122" box="[1175,1207,1801,1825]" gridcol="7" gridrow="30" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08F8A7BFD43122" box="[1175,1207,1801,1825]" italics="true" pageId="9" pageNumber="101">3.4</emphasis>
</td>
<td id="F233493C4138003AFB66F8A7BE463122" box="[1273,1317,1801,1825]" gridcol="8" gridrow="30" pageId="9" pageNumber="101">12.7</td>
<td id="F233493C4138003AFAC6F8A7BEEF3122" box="[1369,1420,1801,1825]" gridcol="9" gridrow="30" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3F8A7BEEB3122" box="[1372,1416,1801,1825]" italics="true" pageId="9" pageNumber="101">27.7</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CF89FBEEF3149" box="[163,1420,1841,1866]" gridrow="31" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CF89FBABF3149" box="[163,476,1841,1866]" gridcol="0" gridrow="31" pageId="9" pageNumber="101">Anterior secondary branch</th>
<td id="F233493C4138003AFD99F89FB97C3149" box="[518,543,1841,1866]" gridcol="1" gridrow="31" pageId="9" pageNumber="101">40</td>
<td id="F233493C4138003AFDD5F89FB9D33149" box="[586,688,1841,1866]" gridcol="2" gridrow="31" pageId="9" pageNumber="101">6.0–11.9</td>
<td id="F233493C4138003AFD43F89FB8213149" box="[732,834,1841,1866]" gridcol="3" gridrow="31" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43F89FB821314A" box="[732,834,1841,1865]" italics="true" pageId="9" pageNumber="101">12.5–26.9</emphasis>
</td>
<td id="F233493C4138003AFCEDF89FB8FD3149" box="[882,926,1841,1866]" gridcol="4" gridrow="31" pageId="9" pageNumber="101">8.8</td>
<td id="F233493C4138003AFC54F89FB89F3149" box="[971,1020,1841,1866]" gridcol="5" gridrow="31" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52F89CB89A3149" box="[973,1017,1842,1866]" italics="true" pageId="9" pageNumber="101">18.4</emphasis>
</td>
<td id="F233493C4138003AFBAEF89FBF303149" box="[1073,1107,1841,1866]" gridcol="6" gridrow="31" pageId="9" pageNumber="101">1.4</td>
<td id="F233493C4138003AFB08F89FBFD43149" box="[1175,1207,1841,1866]" gridcol="7" gridrow="31" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08F89CBFD43149" box="[1175,1207,1842,1866]" italics="true" pageId="9" pageNumber="101">3.2</emphasis>
</td>
<td id="F233493C4138003AFB66F89FBE463149" box="[1273,1317,1841,1866]" gridcol="8" gridrow="31" pageId="9" pageNumber="101">9.8</td>
<td id="F233493C4138003AFAC6F89FBEEF3149" box="[1369,1420,1841,1866]" gridcol="9" gridrow="31" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3F89CBEEB3149" box="[1372,1416,1842,1866]" italics="true" pageId="9" pageNumber="101">21.4</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CF8F4BEEF3171" box="[163,1420,1882,1906]" gridrow="32" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CF8F4BABF3171" box="[163,476,1882,1906]" gridcol="0" gridrow="32" pageId="9" pageNumber="101">Posterior primary branch</th>
<td id="F233493C4138003AFD99F8F4B97C3171" box="[518,543,1882,1906]" gridcol="1" gridrow="32" pageId="9" pageNumber="101">40</td>
<td id="F233493C4138003AFDD5F8F4B9D33171" box="[586,688,1882,1906]" gridcol="2" gridrow="32" pageId="9" pageNumber="101">8.9–17.1</td>
<td id="F233493C4138003AFD43F8F4B8213171" box="[732,834,1882,1906]" gridcol="3" gridrow="32" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43F8F4B8223171" box="[732,833,1882,1906]" italics="true" pageId="9" pageNumber="101">17.4–37.3</emphasis>
</td>
<td id="F233493C4138003AFCEDF8F4B8FD3171" box="[882,926,1882,1906]" gridcol="4" gridrow="32" pageId="9" pageNumber="101">13.3</td>
<td id="F233493C4138003AFC54F8F4B89F3171" box="[971,1020,1882,1906]" gridcol="5" gridrow="32" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52F8F4B89A3171" box="[973,1017,1882,1906]" italics="true" pageId="9" pageNumber="101">27.9</emphasis>
</td>
<td id="F233493C4138003AFBAEF8F4BF303171" box="[1073,1107,1882,1906]" gridcol="6" gridrow="32" pageId="9" pageNumber="101">2.1</td>
<td id="F233493C4138003AFB08F8F4BFD43171" box="[1175,1207,1882,1906]" gridcol="7" gridrow="32" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08F8F4BFD43171" box="[1175,1207,1882,1906]" italics="true" pageId="9" pageNumber="101">4.3</emphasis>
</td>
<td id="F233493C4138003AFB66F8F4BE463171" box="[1273,1317,1882,1906]" gridcol="8" gridrow="32" pageId="9" pageNumber="101">13.2</td>
<td id="F233493C4138003AFAC6F8F4BEEF3171" box="[1369,1420,1882,1906]" gridcol="9" gridrow="32" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFAC3F8F4BEEB3171" box="[1372,1416,1882,1906]" italics="true" pageId="9" pageNumber="101">28.8</emphasis>
</td>
</tr>
<tr id="B1E220404138003AFF3CF82CBEEF3199" box="[163,1420,1922,1946]" gridrow="33" pageId="9" pageNumber="101">
<th id="F233493C4138003AFF3CF82CBABF3199" box="[163,476,1922,1946]" gridcol="0" gridrow="33" pageId="9" pageNumber="101">Posterior secondary branch</th>
<td id="F233493C4138003AFD99F82CB97C3199" box="[518,543,1922,1946]" gridcol="1" gridrow="33" pageId="9" pageNumber="101">40</td>
<td id="F233493C4138003AFDD5F82CB9D33199" box="[586,688,1922,1946]" gridcol="2" gridrow="33" pageId="9" pageNumber="101">6.3–12.6</td>
<td id="F233493C4138003AFD43F82CB8213199" box="[732,834,1922,1946]" gridcol="3" gridrow="33" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFD43F82CB8223199" box="[732,833,1922,1946]" italics="true" pageId="9" pageNumber="101">14.0–25.3</emphasis>
</td>
<td id="F233493C4138003AFCEDF82CB8FD3199" box="[882,926,1922,1946]" gridcol="4" gridrow="33" pageId="9" pageNumber="101">9.5</td>
<td id="F233493C4138003AFC54F82CB89F3199" box="[971,1020,1922,1946]" gridcol="5" gridrow="33" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFC52F82CB89A3199" box="[973,1017,1922,1946]" italics="true" pageId="9" pageNumber="101">19.8</emphasis>
</td>
<td id="F233493C4138003AFBAEF82CBF303199" box="[1073,1107,1922,1946]" gridcol="6" gridrow="33" pageId="9" pageNumber="101">1.5</td>
<td id="F233493C4138003AFB08F82CBFD43199" box="[1175,1207,1922,1946]" gridcol="7" gridrow="33" pageId="9" pageNumber="101">
<emphasis id="3DA6FE104138FFCCFB08F82CBFD43199" box="[1175,1207,1922,1946]" italics="true" pageId="9" pageNumber="101">2.9</emphasis>
</td>
<td id="F233493C4138003AFB66F82CBE463199" box="[1273,1317,1922,1946]" gridcol="8" gridrow="33" pageId="9" pageNumber="101">8.7</td>
<td id="F233493C4138003AFAC6F82CBEEF3199" box="[1369,1420,1922,1946]" gridcol="9" gridrow="33" pageId="9" pageNumber="101">19.0</td>
</tr>
</table>
</paragraph>
<caption id="5BAD728A413BFFCFFF08F969B8A530DC" ID-DOI="http://doi.org/10.5281/zenodo.6531352" ID-Zenodo-Dep="6531352" box="[151,966,1735,1760]" httpUri="https://zenodo.org/record/6531352/files/figure.png" pageId="10" pageNumber="102" startId="10.[151,250,1735,1759]" targetBox="[151,1435,181,1711]" targetPageId="10">
<paragraph id="0F6D2202413BFFCFFF08F969B8A530DC" blockId="10.[151,1436,1735,1868]" box="[151,966,1735,1760]" pageId="10" pageNumber="102">FIGURE 5. Dorsal cuticle apodemes (muscle attachments/cribriform areas)</paragraph>
</caption>
<paragraph id="0F6D2202413BFFCFFF08F942B953314F" blockId="10.[151,1436,1735,1868]" pageId="10" pageNumber="102">
(A) SEM image of the postero-dorsal surface of 
<taxonomicName id="C8D25981413BFFCFFD0FF942B8203100" authority="Sugiura &amp; Matsumoto &amp; Kunieda, 2022" authorityName="Sugiura &amp; Matsumoto &amp; Kunieda" authorityYear="2022" box="[656,835,1771,1795]" class="Eutardigrada" family="Macrobiotidae" genus="Paramacrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="102" phylum="Tardigrada" rank="species" species="metropolitanus" status="sp. nov.">
<emphasis id="3DA6FE10413BFFCFFD0FF942B8203100" box="[656,835,1771,1795]" italics="true" pageId="10" pageNumber="102">P. metropolitanus</emphasis>
</taxonomicName>
<emphasis id="3DA6FE10413BFFCFFCD3F942B8FD3107" bold="true" box="[844,926,1772,1796]" pageId="10" pageNumber="102">
<taxonomicNameLabel id="2695436B413BFFCFFCD3F942B8FD3107" box="[844,926,1772,1796]" pageId="10" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(B) Magnified image of dorsal cuticle apodemes corresponding to the white dashed box in A. (C) Magnified image of the left apodemes in B. White-filled indented arrowheads indicate the apodemes. Scale bars = μm.
</paragraph>
</subSubSection>
</treatment>
</document>