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<document id="0D37B059F9FA086EF970E8F366264671" ID-DOI="10.5281/zenodo.202972" ID-GBIF-Dataset="0a8fcf00-25e6-4353-9c76-e3ba4a9d9f40" ID-ISSN="1175-5326" ID-Zenodo-Dep="202972" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460219602799" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2011" docId="667AC75A174CFF9A66E0B17447AFC88A" docLanguage="en" docName="zt02734p022.pdf" docOrigin="Zootaxa 2734" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Axianassa heardi Anker, 2011, n. sp." docType="treatment" docVersion="10" lastPageNumber="19" masterDocId="9A43BF221741FF8E6677B6554566C90D" masterDocTitle="Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia" masterLastPageNumber="22" masterPageNumber="1" pageNumber="14" updateTime="1698237460885" updateUser="plazi">
<mods:mods id="8B6AD647FF723F90A54AB810FC7306E6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="48DB9DCB9987126A4B6A4146D661571C">
<mods:title id="03949917528E91EBAEE1841DD0265215">Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia</mods:title>
</mods:titleInfo>
<mods:name id="617261A71266465DD9746F9091E415B2" type="personal">
<mods:role id="EC7E47C93CD26FC2AAC6FEE1866CA889">
<mods:roleTerm id="0D54E3FBF070E5EBADCA2E01FBEAD592">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D13D70A4BE9CDE26019B4331DF2C7FCB">Anker, Arthur</mods:namePart>
</mods:name>
<mods:typeOfResource id="A2D153014F8CF6B6D0532FDD119E6DF3">text</mods:typeOfResource>
<mods:relatedItem id="5D4A8254A95E0A66E38556D1914694FD" type="host">
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<mods:title id="B734D0AF66FBC2AEB296B0AEE87C3A16">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9E9B24C2EA0291FBA6E122574BFBD726">
<mods:date id="17118AB6931A8B871240009AE07FAA43">2011</mods:date>
<mods:detail id="997BC12F8A268D5BAF7ED8865F88A830" type="volume">
<mods:number id="35DC7ADFCB6C533808EBA18F72407668">2734</mods:number>
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<mods:classification id="CA1D4245ABD97687B36E796350230EE5">journal article</mods:classification>
<mods:identifier id="AF286A123BC6363763935AA9F8421F8D" type="DOI">10.5281/zenodo.202972</mods:identifier>
<mods:identifier id="FF4A3DFA91478461D1DD0981DAD1E042" type="GBIF-Dataset">0a8fcf00-25e6-4353-9c76-e3ba4a9d9f40</mods:identifier>
<mods:identifier id="350405251D092BED77E07D01045FAC16" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8F3DD12FEEC13D8D6E233B45465829A7" type="Zenodo-Dep">202972</mods:identifier>
</mods:mods>
<treatment id="667AC75A174CFF9A66E0B17447AFC88A" ID-DOI="http://doi.org/10.5281/zenodo.6186746" ID-GBIF-Taxon="119385027" ID-Zenodo-Dep="6186746" LSID="urn:lsid:plazi:treatment:667AC75A174CFF9A66E0B17447AFC88A" httpUri="http://treatment.plazi.org/id/667AC75A174CFF9A66E0B17447AFC88A" lastPageId="20" lastPageNumber="19" pageId="13" pageNumber="14">
<subSubSection id="A6C925C7174CFF8366E0B174447ECE51" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="EE6C764C174CFF8366E0B17444D5CE36" blockId="13.[151,435,1825,1884]" box="[151,435,1825,1851]" pageId="13" pageNumber="14">
<heading id="B524C120174CFF8366E0B17444D5CE36" bold="true" box="[151,435,1825,1851]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="DCA7AA5E174CFF8366E0B17444D5CE36" bold="true" box="[151,435,1825,1851]" pageId="13" pageNumber="14">
<taxonomicName id="29D30DCF174CFF8366E0B174440CCE36" ID-CoL="a5b9e104-656f-4a9e-87f7-a9c650984dd3" box="[151,362,1825,1851]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E174CFF8366E0B174440CCE36" bold="true" box="[151,362,1825,1851]" italics="true" pageId="13" pageNumber="14">Axianassa heardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7941725174CFF836706B17444D5CE36" box="[369,435,1825,1851]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="EE6C764C174CFF8366E0B111447ECE51" blockId="13.[151,435,1825,1884]" box="[151,280,1860,1884]" pageId="13" pageNumber="14">
(
<figureCitation id="76E86AC9174CFF8366E8B1114477CE51" box="[159,273,1860,1884]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStart-4="FIGURE 13" captionStartId-0="14.[151,255,1883,1907]" captionStartId-1="15.[151,255,1810,1834]" captionStartId-2="16.[151,255,1492,1516]" captionStartId-3="17.[151,255,1797,1821]" captionStartId-4="19.[151,255,1970,1994]" captionTargetBox-0="[154,1430,590,1850]" captionTargetBox-1="[182,1432,200,1756]" captionTargetBox-2="[157,1430,193,1472]" captionTargetBox-3="[167,1423,540,1783]" captionTargetBox-4="[155,1433,179,1936]" captionTargetId-0="figure@14.[151,1436,590,1862]" captionTargetId-1="figure@15.[151,1436,194,1788]" captionTargetId-2="figure@16.[151,1436,186,1479]" captionTargetId-3="figure@17.[167,1424,540,1783]" captionTargetId-4="figure@19.[155,1434,179,1936]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionTargetPageId-3="17" captionTargetPageId-4="19" captionText-0="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." captionText-1="FIGURE 10. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, mandible, lateral; B, same, mesial; C, maxillule, lateral; D, maxilla, lateral; E, same, endopod, mesial view; F, first maxilliped, lateral; G, same, mesial; H, second maxilliped, lateral." captionText-2="FIGURE 11. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, third maxilliped, lateral; B, same, detail of crista dentata, mesial; C, major cheliped, mesial; D, same, lateral; E, same, fingers of chela (setae omitted); F, minor cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral; I, fifth pereiopod, lateral." captionText-3="FIGURE 12. Habitus and colour in life of Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, dorsal; B, lateral." captionText-4="FIGURE 13. Habitus and colour in life of Axianassa heardi n. sp., non-type female from Moorea, French Polynesia (FLMNH UF 23957): A, dorsal, B, lateral." httpUri-0="https://zenodo.org/record/202981/files/figure.png" httpUri-1="https://zenodo.org/record/202982/files/figure.png" httpUri-2="https://zenodo.org/record/202983/files/figure.png" httpUri-3="https://zenodo.org/record/202984/files/figure.png" httpUri-4="https://zenodo.org/record/202985/files/figure.png" pageId="13" pageNumber="14">Figs 9–13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A6C925C7174CFF8066E0B1D841B2C9FA" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="EE6C764C174CFF8366E0B1D840E2CEE1" blockId="13.[151,1436,1932,2029]" pageId="13" pageNumber="14">
<emphasis id="DCA7AA5E174CFF8366E0B1D84421CEA8" bold="true" box="[151,327,1932,1957]" pageId="13" pageNumber="14">
<typeStatus id="3168C8EE174CFF8366E0B1D845B4CEA8" box="[151,210,1933,1957]" pageId="13" pageNumber="14">Type</typeStatus>
material.
</emphasis>
<typeStatus id="3168C8EE174CFF836738B1D944DDCEA9" box="[335,443,1932,1956]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
: male (CL
<quantity id="292BDBA9174CFF836449B1D947CECEA9" box="[574,680,1932,1956]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" pageId="13" pageNumber="14" unit="mm" value="4.65">4.65 mm</quantity>
), QM W29049,
<collectingCountry id="96C436DC174CFF836514B1D946ABCEA9" box="[867,973,1932,1956]" name="Australia" pageId="13" pageNumber="14">Australia</collectingCountry>
, Queensland, Great Barrier Reef, Lizard Island,
<taxonomicName id="29D30DCF174CFF83669EB1E544C0CEC5" authority="Beach" authorityName="Beach" box="[233,422,1968,1992]" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="genus">Casuarina Beach</taxonomicName>
near Lizard Island marine station,
<geoCoordinate id="8BE7108B174CFF83655AB1E546AFCEC5" box="[813,969,1968,1992]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="-14.675389">14°40’31.4”S</geoCoordinate>
,
<geoCoordinate id="8BE7108B174CFF8365A3B1E54119CEC5" box="[980,1151,1968,1992]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="145.4443">145°26’39.5”E</geoCoordinate>
, sand flat with corals and coral rubble, depth
<quantity id="292BDBA9174CFF836707B18144FBCEE1" box="[368,413,2004,2028]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="1.0">1 m</quantity>
, suction (yabby) pump, from burrow, coll. A. Anker,
<date id="9A6D508C174CFF836583B18141D8CEE1" box="[1012,1214,2004,2028]" pageId="13" pageNumber="14" value="2009-02-24">24 February 2009</date>
, fcn AUST-1797.
</paragraph>
<paragraph id="EE6C764C174FFF8066B0B6C241B2C9FA" blockId="14.[151,1436,151,535]" pageId="14" pageNumber="15">
<emphasis id="DCA7AA5E174FFF8066B0B6C244D9C9BD" bold="true" box="[199,447,151,176]" pageId="14" pageNumber="15">Additional material.</emphasis>
<specimenCount id="F8D5BDC5174FFF8067BDB6C24756C9A2" box="[458,560,151,175]" pageId="14" pageNumber="15" type="female">1 female</specimenCount>
(CL
<quantity id="292BDBA9174FFF806403B6C247B5C9A2" box="[628,723,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="14" pageNumber="15" unit="mm" value="5.3">5.3 mm</quantity>
),
<collectionCode id="88C2EE89174FFF80649CB6CD462BC9A2" box="[747,845,152,175]" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="14" pageNumber="15">FLMNH</collectionCode>
UF Arthropoda 23957,
<collectingCountry id="96C436DC174FFF806219B6C2405EC9A2" box="[1134,1336,151,175]" name="French Polynesia" pageId="14" pageNumber="15">French Polynesia</collectingCountry>
, Society Islands, Moorea, channel between NW Motu islands,
<geoCoordinate id="8BE7108B174FFF806498B6E946EDC9D9" box="[751,907,188,212]" direction="south" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="-17.489305">17°29’21.5”S</geoCoordinate>
,
<geoCoordinate id="8BE7108B174FFF8065E1B6E9412FC9D9" box="[918,1097,188,212]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="-149.9132">149°54’47.5”W</geoCoordinate>
, sand flat with some large and small rocks and coral “bommies”,
<quantity id="292BDBA9174FFF80646DB68A4705C9FA" box="[538,611,223,247]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="14" pageNumber="15" unit="m" value="1.0" valueMax="2.0" valueMin="0.0">0–2 m</quantity>
, hand, A. Anker,
<date id="9A6D508C174FFF80655FB68A4165C9FA" box="[808,1027,223,247]" pageId="14" pageNumber="15" value="2009-11-10">10 November 2009</date>
, fcn
<collectionCode id="88C2EE89174FFF80624EB68A41EBC9FA" box="[1081,1165,223,247]" pageId="14" pageNumber="15">BMOO</collectionCode>
8770.
</paragraph>
</subSubSection>
<subSubSection id="A6C925C7174FFF9C66B0B75646A9CA72" lastPageId="18" lastPageNumber="19" pageId="14" pageNumber="15" type="description">
<paragraph id="EE6C764C174FFF8066B0B756405BC8A1" blockId="14.[151,1436,151,535]" pageId="14" pageNumber="15">
<emphasis id="DCA7AA5E174FFF8066B0B756443FC811" bold="true" box="[199,345,259,284]" pageId="14" pageNumber="15">Description.</emphasis>
[based on male
<typeStatus id="3168C8EE174FFF806461B751471DC811" box="[534,635,260,284]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
] Body and appendages sparsely covered with conspicuous, long, erect setae (
<figureCitation id="76E86AC9174FFF806696B7724440C832" box="[225,294,295,319]" captionStart="FIGURE 9" captionStartId="14.[151,255,1883,1907]" captionTargetBox="[154,1430,590,1850]" captionTargetId="figure@14.[151,1436,590,1862]" captionTargetPageId="14" captionText="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
A, 12). Carapace with straight linea thalassinica and well-defined, crescent-shaped cervical groove; pterygostomial region broadly rounded, fringed with setae (
<figureCitation id="76E86AC9174FFF806548B71946E5C869" box="[831,899,332,356]" captionStart="FIGURE 9" captionStartId="14.[151,255,1883,1907]" captionTargetBox="[154,1430,590,1850]" captionTargetId="figure@14.[151,1436,590,1862]" captionTargetPageId="14" captionText="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
A, C). Rostrum about as long as broad at base, reaching far beyond anterior margin of eyestalks, with rounded apical tooth and three teeth on each lateral margin, most posterior lateral tooth being much less than conspicuous than most anterior lateral tooth (
<figureCitation id="76E86AC9174FFF8062CEB7C1419AC8A1" box="[1209,1276,404,428]" captionStart="FIGURE 9" captionStartId="14.[151,255,1883,1907]" captionTargetBox="[154,1430,590,1850]" captionTargetId="figure@14.[151,1436,590,1862]" captionTargetPageId="14" captionText="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
B, E).
</paragraph>
<paragraph id="EE6C764C174FFF8066BEB7E246A8CB1A" blockId="14.[151,1436,151,535]" pageId="14" pageNumber="15">
Abdomen thinly sclerotised, smooth; first abdominal somite with pleuron ventrally produced into fairly strong blunt process (
<figureCitation id="76E86AC9174FFF806748B78944E3C8F9" box="[319,389,476,500]" captionStart="FIGURE 9" captionStartId="14.[151,255,1883,1907]" captionTargetBox="[154,1430,590,1850]" captionTargetId="figure@14.[151,1436,590,1862]" captionTargetPageId="14" captionText="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
A); posterior somites without dense lateral fringes of plumose setae. Telson broad, tapering posteriorly, without armature, posterior margin broadly rounded (
<figureCitation id="76E86AC9174FFF806505B7AA46D1CB1A" box="[882,951,511,535]" captionStart="FIGURE 9" captionStartId="14.[151,255,1883,1907]" captionTargetBox="[154,1430,590,1850]" captionTargetId="figure@14.[151,1436,590,1862]" captionTargetPageId="14" captionText="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="14" pageNumber="15">
Fig.
<date id="9A6D508C174FFF8065D0B7AA46D1CB1A" box="[935,951,511,535]" pageId="14" pageNumber="15">9</date>
</figureCitation>
I).
</paragraph>
<caption id="BAAC26C4174FFF8066E0B10E4723CEDE" httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="14" pageNumber="15" targetBox="[154,1430,590,1850]" targetPageId="14">
<paragraph id="EE6C764C174FFF8066E0B10E4723CEDE" blockId="14.[151,1436,1883,2003]" pageId="14" pageNumber="15">
<emphasis id="DCA7AA5E174FFF8066E0B10E447ACE7E" bold="true" box="[151,284,1883,1907]" pageId="14" pageNumber="15">FIGURE 9.</emphasis>
<taxonomicName id="29D30DCF174FFF806752B1094487CE7E" box="[293,481,1884,1907]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E174FFF806752B1094487CE7E" box="[293,481,1884,1907]" italics="true" pageId="14" pageNumber="15">Axianassa heardi</emphasis>
</taxonomicName>
<emphasis id="DCA7AA5E174FFF80679DB10E474FCE7E" bold="true" box="[490,553,1883,1907]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="C7941725174FFF80679DB10E474FCE7E" box="[490,553,1883,1907]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male from Lizard Island, Australia (QM W29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal.
</paragraph>
</caption>
<caption id="BAAC26C4174EFF8166E0B147470FCE64" httpUri="https://zenodo.org/record/202982/files/figure.png" pageId="15" pageNumber="16" targetBox="[182,1432,200,1756]" targetPageId="15">
<paragraph id="EE6C764C174EFF8166E0B147470FCE64" blockId="15.[151,1436,1810,1897]" pageId="15" pageNumber="16">
<emphasis id="DCA7AA5E174EFF8166E0B1474440CE27" bold="true" box="[151,294,1810,1834]" pageId="15" pageNumber="16">FIGURE 10.</emphasis>
<taxonomicName id="29D30DCF174EFF81675BB1474482CE24" box="[300,484,1810,1833]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E174EFF81675BB1474482CE24" box="[300,484,1810,1833]" italics="true" pageId="15" pageNumber="16">Axianassa heardi</emphasis>
</taxonomicName>
<emphasis id="DCA7AA5E174EFF81679CB1474741CE27" bold="true" box="[491,551,1810,1834]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="C7941725174EFF81679CB1474741CE27" box="[491,551,1810,1834]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male from Lizard Island, Australia (QM W29049): A, mandible, lateral; B, same, mesial; C, maxillule, lateral; D, maxilla, lateral; E, same, endopod, mesial view; F, first maxilliped, lateral; G, same, mesial; H, second maxilliped, lateral.
</paragraph>
</caption>
<caption id="BAAC26C41751FF9E66E0B3814587CF41" httpUri="https://zenodo.org/record/202983/files/figure.png" pageId="16" pageNumber="17" targetBox="[157,1430,193,1472]" targetPageId="16">
<paragraph id="EE6C764C1751FF9E66E0B3814587CF41" blockId="16.[151,1436,1492,1612]" pageId="16" pageNumber="17">
<emphasis id="DCA7AA5E1751FF9E66E0B3814443CCE1" bold="true" box="[151,293,1492,1516]" pageId="16" pageNumber="17">FIGURE 11.</emphasis>
<taxonomicName id="29D30DCF1751FF9E675BB3804480CCE1" box="[300,486,1493,1516]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E1751FF9E675BB3804480CCE1" box="[300,486,1493,1516]" italics="true" pageId="16" pageNumber="17">Axianassa heardi</emphasis>
</taxonomicName>
<emphasis id="DCA7AA5E1751FF9E679BB381474FCCE1" bold="true" box="[492,553,1492,1516]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="C79417251751FF9E679BB381474FCCE1" box="[492,553,1492,1516]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male from Lizard Island, Australia (QM W29049): A, third maxilliped, lateral; B, same, detail of crista dentata, mesial; C, major cheliped, mesial; D, same, lateral; E, same, fingers of chela (setae omitted); F, minor cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral; I, fifth pereiopod, lateral.
</paragraph>
</caption>
<paragraph id="EE6C764C1751FF9E66BEB03B462DCFA7" blockId="16.[151,1436,1646,2031]" pageId="16" pageNumber="17">
Eyestalks rounded, with large cornea occupying entire terminal portion (
<figureCitation id="76E86AC91751FF9E6255B03B410DCF8B" box="[1058,1131,1646,1670]" captionStart="FIGURE 9" captionStartId="14.[151,255,1883,1907]" captionTargetBox="[154,1430,590,1850]" captionTargetId="figure@14.[151,1436,590,1862]" captionTargetPageId="14" captionText="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
D), pigmented blackish in alcohol-preserved specimen, white-yellowish in life (
<figureCitation id="76E86AC91751FF9E649CB0C74658CFA7" box="[747,830,1682,1706]" captionStart="FIGURE 12" captionStartId="17.[151,255,1797,1821]" captionTargetBox="[167,1423,540,1783]" captionTargetId="figure@17.[167,1424,540,1783]" captionTargetPageId="17" captionText="FIGURE 12. Habitus and colour in life of Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, dorsal; B, lateral." httpUri="https://zenodo.org/record/202984/files/figure.png" pageId="16" pageNumber="17">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="EE6C764C1751FF9E66E0B0E341E8CE37" blockId="16.[151,1436,1646,2031]" pageId="16" pageNumber="17">
Antennular peduncle with third article subcylindrical, slender, not reaching half-length of fourth article of antennal peduncle (
<figureCitation id="76E86AC91751FF9E677CB08F442BCFFF" box="[267,333,1754,1778]" captionStart="FIGURE 9" captionStartId="14.[151,255,1883,1907]" captionTargetBox="[154,1430,590,1850]" captionTargetId="figure@14.[151,1436,590,1862]" captionTargetPageId="14" captionText="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
A, B). Antennal acicle dagger-like, tapering distally, reaching to about 0.3 length of third article of antennular peduncle; fourth article subcylindrical, very long, slender, furnished with elongate setae on lateral and ventrolateral surface (
<figureCitation id="76E86AC91751FF9E67FBB17744A9CE37" box="[396,463,1826,1850]" captionStart="FIGURE 9" captionStartId="14.[151,255,1883,1907]" captionTargetBox="[154,1430,590,1850]" captionTargetId="figure@14.[151,1436,590,1862]" captionTargetPageId="14" captionText="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
A); flagellum well developed, reaching well beyond chelipeds.
</paragraph>
<paragraph id="EE6C764C1751FF9F66BEB1134764C8F9" blockId="16.[151,1436,1646,2031]" lastBlockId="17.[151,1436,151,500]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Mouthparts more or less typical for genus (cf.
<bibRefCitation id="8A420BBD1751FF9E64A2B11346B8CE53" author="Kensley" box="[725,990,1862,1886]" pageId="16" pageNumber="22" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">
Kensley &amp;
<collectingCountry id="96C436DC1751FF9E6521B11346FCCE53" box="[854,922,1862,1886]" name="Heard Island and McDonald Islands" pageId="16" pageNumber="17">Heard</collectingCountry>
1990
</bibRefCitation>
: fig. 2D–I;
<bibRefCitation id="8A420BBD1751FF9E6217B11340FECE53" author="Rodrigues" box="[1120,1432,1862,1886]" pageId="16" pageNumber="22" refString="Rodrigues, S. A. &amp; Shimizu, R. M. (1992) Description of a new Axianassa (Crustacea: Decapoda: Thalassinidea) from Brazil, and its first larval stage. Proceedings of the Biological Society of Washington, 105, 317 - 323." type="journal article" year="1992">Rodrigues &amp; Shimizu 1992</bibRefCitation>
: figs. 4–8; Anker 2010: fig. 2). Mandible with triarticulated palp (first two articles possibly partly fused); distal article densely furnished with setae, most distal setae brown-blackish in colour; molar and incisor processes partly fused; molar edge without teeth (except for one blunt distal bump); incisor edge with 11 small subacute teeth (
<figureCitation id="76E86AC91751FF9E6319B1E745D5CEE3" captionStart="FIGURE 10" captionStartId="15.[151,255,1810,1834]" captionTargetBox="[182,1432,200,1756]" captionTargetId="figure@15.[151,1436,194,1788]" captionTargetPageId="15" captionText="FIGURE 10. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, mandible, lateral; B, same, mesial; C, maxillule, lateral; D, maxilla, lateral; E, same, endopod, mesial view; F, first maxilliped, lateral; G, same, mesial; H, second maxilliped, lateral." httpUri="https://zenodo.org/record/202982/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
A, B). Maxillule with biarticulated palp, coxal endite very setose; basal endite with stout spine-like setae in addition to more slender setae (
<figureCitation id="76E86AC91750FF9F6474B6C2473EC9A2" box="[515,600,151,175]" captionStart="FIGURE 10" captionStartId="15.[151,255,1810,1834]" captionTargetBox="[182,1432,200,1756]" captionTargetId="figure@15.[151,1436,194,1788]" captionTargetPageId="15" captionText="FIGURE 10. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, mandible, lateral; B, same, mesial; C, maxillule, lateral; D, maxilla, lateral; E, same, endopod, mesial view; F, first maxilliped, lateral; G, same, mesial; H, second maxilliped, lateral." httpUri="https://zenodo.org/record/202982/files/figure.png" pageId="17" pageNumber="18">Fig. 10</figureCitation>
C). Maxilla with broad scaphognathite, bearing at least six long, flexible setae on ventral (= posterior) margin; endopod biarticulated, distal article slightly narrower than proximal, tapering (
<figureCitation id="76E86AC91750FF9F66E8B68A4593C9FA" box="[159,245,223,247]" captionStart="FIGURE 10" captionStartId="15.[151,255,1810,1834]" captionTargetBox="[182,1432,200,1756]" captionTargetId="figure@15.[151,1436,194,1788]" captionTargetPageId="15" captionText="FIGURE 10. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, mandible, lateral; B, same, mesial; C, maxillule, lateral; D, maxilla, lateral; E, same, endopod, mesial view; F, first maxilliped, lateral; G, same, mesial; H, second maxilliped, lateral." httpUri="https://zenodo.org/record/202982/files/figure.png" pageId="17" pageNumber="18">Fig. 10</figureCitation>
D, E). First maxilliped with broad, paddle-shaped endopod; exopod triarticulated, basal article broad, paddle-shaped, distal articles much narrower; epipod present as large oval-shaped lobe (
<figureCitation id="76E86AC91750FF9F62D3B751419EC811" box="[1188,1272,260,284]" captionStart="FIGURE 10" captionStartId="15.[151,255,1810,1834]" captionTargetBox="[182,1432,200,1756]" captionTargetId="figure@15.[151,1436,194,1788]" captionTargetPageId="15" captionText="FIGURE 10. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, mandible, lateral; B, same, mesial; C, maxillule, lateral; D, maxilla, lateral; E, same, endopod, mesial view; F, first maxilliped, lateral; G, same, mesial; H, second maxilliped, lateral." httpUri="https://zenodo.org/record/202982/files/figure.png" pageId="17" pageNumber="18">Fig. 10</figureCitation>
F, G). Second maxilliped typical for
<taxonomicName id="29D30DCF1750FF9F67E4B77D4760C84D" box="[403,518,296,320]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1750FF9F67E4B77D4760C84D" box="[403,518,296,320]" italics="true" pageId="17" pageNumber="18">Axianassa</emphasis>
</taxonomicName>
, with elongate epipod bearing well-developed podobranch; endopod robust, with apical “head” formed by propodus and dactylus; exopod triarticulated (
<figureCitation id="76E86AC91750FF9F65ACB7194157C869" box="[987,1073,332,356]" captionStart="FIGURE 10" captionStartId="15.[151,255,1810,1834]" captionTargetBox="[182,1432,200,1756]" captionTargetId="figure@15.[151,1436,194,1788]" captionTargetPageId="15" captionText="FIGURE 10. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, mandible, lateral; B, same, mesial; C, maxillule, lateral; D, maxilla, lateral; E, same, endopod, mesial view; F, first maxilliped, lateral; G, same, mesial; H, second maxilliped, lateral." httpUri="https://zenodo.org/record/202982/files/figure.png" pageId="17" pageNumber="18">Fig. 10</figureCitation>
H). Third maxilliped pediform; epipod bifid, with well-developed podobranch; ischium with crista dentata bearing at least 14 teeth; merus with small acute tooth distolaterally; bands of fusiform setae grouped in well-delimited areas present on mesial side of merus, carpus, propodus and dactylus, ventral margin of these articles with long simple setae, latter especially dense on dactylus (
<figureCitation id="76E86AC91750FF9F671AB78944DAC8F9" box="[365,444,476,500]" captionStart="FIGURE 11" captionStartId="16.[151,255,1492,1516]" captionTargetBox="[157,1430,193,1472]" captionTargetId="figure@16.[151,1436,186,1479]" captionTargetPageId="16" captionText="FIGURE 11. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, third maxilliped, lateral; B, same, detail of crista dentata, mesial; C, major cheliped, mesial; D, same, lateral; E, same, fingers of chela (setae omitted); F, minor cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202983/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
A, B).
</paragraph>
<caption id="BAAC26C41750FF9F66E0B1504484CE30" httpUri="https://zenodo.org/record/202984/files/figure.png" pageId="17" pageNumber="18" targetBox="[167,1423,540,1783]" targetPageId="17">
<paragraph id="EE6C764C1750FF9F66E0B1504484CE30" blockId="17.[151,1436,1797,1853]" pageId="17" pageNumber="18">
<emphasis id="DCA7AA5E1750FF9F66E0B150444ECE10" bold="true" box="[151,296,1797,1821]" pageId="17" pageNumber="18">FIGURE 12.</emphasis>
Habitus and colour in life of
<taxonomicName id="29D30DCF1750FF9F641AB153464ECE10" box="[621,808,1798,1821]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E1750FF9F641AB153464ECE10" box="[621,808,1798,1821]" italics="true" pageId="17" pageNumber="18">Axianassa heardi</emphasis>
</taxonomicName>
<emphasis id="DCA7AA5E1750FF9F6547B150460BCE10" bold="true" box="[816,877,1797,1821]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="C79417251750FF9F6547B150460BCE10" box="[816,877,1797,1821]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male from Lizard Island, Australia (QM W29049): A, dorsal; B, lateral.
</paragraph>
</caption>
<paragraph id="EE6C764C1750FF9C66BEB13F4115C811" blockId="17.[151,1437,1898,2030]" lastBlockId="18.[151,1437,151,1977]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
First pereiopods (chelipeds) stout, unequal in size by about 20%, slightly asymmetrical in shape. Major cheliped very robust; ischium with about eight small teeth on ventral margin, most distal tooth largest and most conspicuous; merus with ventral and dorsal margins inflated, ventral margin with at least eight small teeth proximally and one larger projecting tooth at about mid-length; carpus cup-shaped, distally widening, unarmed; chela ovate, compressed laterally, smooth, with fingers about 0.7 palm length, stout, with crossing tips; cutting edge of pollex with two large subtriangular teeth and a series of smaller teeth proximally; cutting edge of dactylus with at least one larger tooth (
<figureCitation id="76E86AC91753FF9C6476B68A4732C9FA" box="[513,596,223,247]" captionStart="FIGURE 11" captionStartId="16.[151,255,1492,1516]" captionTargetBox="[157,1430,193,1472]" captionTargetId="figure@16.[151,1436,186,1479]" captionTargetPageId="16" captionText="FIGURE 11. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, third maxilliped, lateral; B, same, detail of crista dentata, mesial; C, major cheliped, mesial; D, same, lateral; E, same, fingers of chela (setae omitted); F, minor cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202983/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
C–E). Minor cheliped generally similar to major cheliped, with slightly smaller, more slender chela; teeth on cutting edges of pollex less pronounced (
<figureCitation id="76E86AC91753FF9C6273B7514135C811" box="[1028,1107,260,284]" captionStart="FIGURE 11" captionStartId="16.[151,255,1492,1516]" captionTargetBox="[157,1430,193,1472]" captionTargetId="figure@16.[151,1436,186,1479]" captionTargetPageId="16" captionText="FIGURE 11. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, third maxilliped, lateral; B, same, detail of crista dentata, mesial; C, major cheliped, mesial; D, same, lateral; E, same, fingers of chela (setae omitted); F, minor cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202983/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
F).
</paragraph>
<paragraph id="EE6C764C1753FF9C66BEB7724057CB31" blockId="18.[151,1437,151,1977]" pageId="18" pageNumber="19">
Second pereiopod pediform, robust; merus, carpus and propodus with long flexible setae along ventral margin; dactylus about half as long as propodus, simple (
<figureCitation id="76E86AC91753FF9C64CBB719466BC869" box="[700,781,332,356]" captionStart="FIGURE 11" captionStartId="16.[151,255,1492,1516]" captionTargetBox="[157,1430,193,1472]" captionTargetId="figure@16.[151,1436,186,1479]" captionTargetPageId="16" captionText="FIGURE 11. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, third maxilliped, lateral; B, same, detail of crista dentata, mesial; C, major cheliped, mesial; D, same, lateral; E, same, fingers of chela (setae omitted); F, minor cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202983/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
G). Third pereiopod moderately stout; ischium, merus and carpus smooth, moderately setose; propodus with distoventral brush(es) of stiff setae; dactylus slightly shorter than propodus, somewhat sickle-shaped; dorsal margin bearing at least six stout corneous spines; distoventral margin slightly expanded, carrying row of short spiniform setae (
<figureCitation id="76E86AC91753FF9C655FB7E2461FC8C2" box="[808,889,439,463]" captionStart="FIGURE 11" captionStartId="16.[151,255,1492,1516]" captionTargetBox="[157,1430,193,1472]" captionTargetId="figure@16.[151,1436,186,1479]" captionTargetPageId="16" captionText="FIGURE 11. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, third maxilliped, lateral; B, same, detail of crista dentata, mesial; C, major cheliped, mesial; D, same, lateral; E, same, fingers of chela (setae omitted); F, minor cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202983/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
H). Fourth pereiopod similar to third pereiopod, slightly shorter and more slender. Fifth pereiopod much more slender than third or fourth pereiopods; propodus subchelate, ending in short subacute tooth (obscured by setae); distal half of propodus with brush of stiff setae (
<figureCitation id="76E86AC91753FF9C6319B7AA45D5CB31" captionStart="FIGURE 11" captionStartId="16.[151,255,1492,1516]" captionTargetBox="[157,1430,193,1472]" captionTargetId="figure@16.[151,1436,186,1479]" captionTargetPageId="16" captionText="FIGURE 11. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, third maxilliped, lateral; B, same, detail of crista dentata, mesial; C, major cheliped, mesial; D, same, lateral; E, same, fingers of chela (setae omitted); F, minor cheliped, lateral; G, second pereiopod, lateral; H, third pereiopod, lateral; I, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/202983/files/figure.png" pageId="18" pageNumber="19">
Fig.
<date id="9A6D508C1753FF9C66E0B47145D5CB31" box="[151,179,548,572]" pageId="18" pageNumber="19" value="1911-01">11</date>
</figureCitation>
I); dactylus subspatulate, with deep ventroproximal excavation opposed to subacute tooth of propodus.
</paragraph>
<paragraph id="EE6C764C1753FF9C66BEB4124641CBE2" blockId="18.[151,1437,151,1977]" pageId="18" pageNumber="19">
First pleopod absent. Second to fifth pleopods similar, biramous, typical for genus. Uropods with broadly ovoid exopod and endopod; exopod with blunt distolateral tooth adjacent to movable spiniform seta, sometimes with a second much smaller tooth on lateral margin above distolateral tooth, and with distinct transverse suture furnished with about 12 small denticles; endopod with one strong subacute tooth on lateral margin and with several subacute teeth on dorsal surface as illustrated (
<figureCitation id="76E86AC91753FF9C64D6B4824784CBE2" box="[673,738,727,751]" captionStart="FIGURE 9" captionStartId="14.[151,255,1883,1907]" captionTargetBox="[154,1430,590,1850]" captionTargetId="figure@14.[151,1436,590,1862]" captionTargetPageId="14" captionText="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
G, H).
</paragraph>
<paragraph id="EE6C764C1753FF9C66BEB4A9447ECA51" blockId="18.[151,1437,151,1977]" pageId="18" pageNumber="19">
Gill/exopod formula possibly* as described for genus (cf.
<bibRefCitation id="8A420BBD1753FF9C652CB4A94102CA19" author="Kensley" box="[859,1124,764,788]" pageId="18" pageNumber="22" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">
Kensley &amp;
<collectingCountry id="96C436DC1753FF9C65ABB4A94146CA19" box="[988,1056,764,788]" name="Heard Island and McDonald Islands" pageId="18" pageNumber="19">Heard</collectingCountry>
1990
</bibRefCitation>
), except for the presence of well-developed (vs. rudimentary) epipod on the first maxilliped (*note: some arthrobranchs were detached during dissection).
</paragraph>
<paragraph id="EE6C764C1753FF9C66BEB53246A9CA72" blockId="18.[151,1437,151,1977]" box="[201,975,871,897]" pageId="18" pageNumber="19">
<emphasis id="DCA7AA5E1753FF9C66BEB53244E2CA8D" bold="true" box="[201,388,871,896]" pageId="18" pageNumber="19">Colour pattern.</emphasis>
Uniform hyaline white; ovary orange (
<figureCitation id="76E86AC91753FF9C654DB53246F1CA72" box="[826,919,871,895]" captionStart="FIGURE 12" captionStartId="17.[151,255,1797,1821]" captionTargetBox="[167,1423,540,1783]" captionTargetId="figure@17.[167,1424,540,1783]" captionTargetPageId="17" captionText="FIGURE 12. Habitus and colour in life of Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, dorsal; B, lateral." httpUri="https://zenodo.org/record/202984/files/figure.png" pageId="18" pageNumber="19">Figs. 12</figureCitation>
,
<figureCitation id="76E86AC91753FF9C65D4B53246A4CA72" box="[931,962,871,895]" captionStart="FIGURE 13" captionStartId="19.[151,255,1970,1994]" captionTargetBox="[155,1433,179,1936]" captionTargetId="figure@19.[155,1434,179,1936]" captionTargetPageId="19" captionText="FIGURE 13. Habitus and colour in life of Axianassa heardi n. sp., non-type female from Moorea, French Polynesia (FLMNH UF 23957): A, dorsal, B, lateral." httpUri="https://zenodo.org/record/202985/files/figure.png" pageId="18" pageNumber="19">13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A6C925C71753FF9C66BEB5D9466ACD55" pageId="18" pageNumber="19" type="etymology">
<paragraph id="EE6C764C1753FF9C66BEB5D9468DCAE1" blockId="18.[151,1437,151,1977]" pageId="18" pageNumber="19">
<emphasis id="DCA7AA5E1753FF9C66BEB5D94429CAA9" bold="true" box="[201,335,908,932]" pageId="18" pageNumber="19">Etymology.</emphasis>
Named after Dr. Richard
<collectingCountry id="96C436DC1753FF9C64EEB5D947BBCAA9" box="[665,733,908,932]" name="Heard Island and McDonald Islands" pageId="18" pageNumber="19">Heard</collectingCountry>
(Gulf Coast Research Laboratory, University of Southern Mississippi, Ocean Springs, MS,
<collectingCountry id="96C436DC1753FF9C6478B5FA472DCACA" box="[527,587,943,967]" name="United States of America" pageId="18" pageNumber="19">USA</collectingCountry>
), for his contributions to decapod taxonomy, including the co-authorship in the revision of the American species of
<taxonomicName id="29D30DCF1753FF9C6423B58147A1CAE1" box="[596,711,980,1004]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1753FF9C6423B58147A1CAE1" box="[596,711,980,1004]" italics="true" pageId="18" pageNumber="19">Axianassa</emphasis>
</taxonomicName>
(
<bibRefCitation id="8A420BBD1753FF9C64A1B58146B9CAE1" author="Kensley" box="[726,991,980,1004]" pageId="18" pageNumber="22" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">
Kensley &amp;
<collectingCountry id="96C436DC1753FF9C6520B58146FDCAE1" box="[855,923,980,1004]" name="Heard Island and McDonald Islands" pageId="18" pageNumber="19">Heard</collectingCountry>
1990
</bibRefCitation>
).
</paragraph>
<paragraph id="EE6C764C1753FF9C66BEB5A2466ACD55" blockId="18.[151,1437,151,1977]" pageId="18" pageNumber="19">
<emphasis id="DCA7AA5E1753FF9C66BEB5A2444ACD1D" bold="true" box="[201,300,1015,1040]" pageId="18" pageNumber="19">Habitat.</emphasis>
Lizard Island: sand flat with large coral heads and coral rubble, at depth of about
<quantity id="292BDBA91753FF9C6293B5A24075CD1D" box="[1252,1299,1015,1040]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="m" value="1.0">1 m</quantity>
, in burrows (same collection site as for
<taxonomicName id="29D30DCF1753FF9C67A4B24947C5CD39" box="[467,675,1052,1076]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="saurus" status="sp. nov.">
<emphasis id="DCA7AA5E1753FF9C67A4B24947C5CD39" box="[467,675,1052,1076]" italics="true" pageId="18" pageNumber="19">Athanopsis saurus</emphasis>
</taxonomicName>
<emphasis id="DCA7AA5E1753FF9C64DBB249478ACD39" bold="true" box="[684,748,1052,1076]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="C79417251753FF9C64DBB249478ACD39" box="[684,748,1052,1076]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
); Moorea: shallow channel between two small islands, with large areas of sand between rocks and coral “bommies”.
</paragraph>
</subSubSection>
<subSubSection id="A6C925C71753FF9C66BEB2314169CD71" box="[201,1039,1123,1148]" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="EE6C764C1753FF9C66BEB2314169CD71" blockId="18.[151,1437,151,1977]" box="[201,1039,1123,1148]" pageId="18" pageNumber="19">
<emphasis id="DCA7AA5E1753FF9C66BEB2314403CD71" bold="true" box="[201,357,1123,1148]" pageId="18" pageNumber="19">
<typeStatus id="3168C8EE1753FF9C66BEB2314464CD71" box="[201,258,1124,1148]" pageId="18" pageNumber="19">Type</typeStatus>
locality.
</emphasis>
Lizard Island, Great Barrier Reef off Queensland,
<collectingCountry id="96C436DC1753FF9C65D6B231416ACD71" box="[929,1036,1124,1148]" name="Australia" pageId="18" pageNumber="19">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A6C925C71753FF9C66BEB2D244DBCDC9" pageId="18" pageNumber="19" type="distribution">
<paragraph id="EE6C764C1753FF9C66BEB2D244DBCDC9" blockId="18.[151,1437,151,1977]" pageId="18" pageNumber="19">
<emphasis id="DCA7AA5E1753FF9C66BEB2D24404CDAD" bold="true" box="[201,354,1159,1184]" pageId="18" pageNumber="19">Distribution.</emphasis>
Lizard Island, northern section of the Great Barrier Reef (
<typeStatus id="3168C8EE1753FF9C6271B2DD415ECDAD" box="[1030,1080,1160,1184]" pageId="18" pageNumber="19">type</typeStatus>
locality), and Moorea, Society Islands,
<collectingCountry id="96C436DC1753FF9C6683B2F944DFCDC9" box="[244,441,1196,1220]" name="French Polynesia" pageId="18" pageNumber="19">French Polynesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A6C925C71753FF9C66BEB29A404ECC3D" pageId="18" pageNumber="19" type="description">
<paragraph id="EE6C764C1753FF9C66BEB29A404ECC3D" blockId="18.[151,1437,151,1977]" pageId="18" pageNumber="19">
<emphasis id="DCA7AA5E1753FF9C66BEB29A4458CDE5" bold="true" box="[201,318,1231,1256]" pageId="18" pageNumber="19">Variation.</emphasis>
The female specimen from Moorea generally agrees with the male
<typeStatus id="3168C8EE1753FF9C623AB28541D4CDE5" box="[1101,1202,1232,1256]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
, except for having a stouter major cheliped, with a more pronounced tooth on the cutting edge of the dactylus (
<figureCitation id="76E86AC91753FF9C62EEB2A1418ACC01" box="[1177,1260,1268,1292]" captionStart="FIGURE 13" captionStartId="19.[151,255,1970,1994]" captionTargetBox="[155,1433,179,1936]" captionTargetId="figure@19.[155,1434,179,1936]" captionTargetPageId="19" captionText="FIGURE 13. Habitus and colour in life of Axianassa heardi n. sp., non-type female from Moorea, French Polynesia (FLMNH UF 23957): A, dorsal, B, lateral." httpUri="https://zenodo.org/record/202985/files/figure.png" pageId="18" pageNumber="19">Fig. 13</figureCitation>
), and bearing a pair of slender uniramous pleopods on the first abdominal somite, as is typical for females of
<taxonomicName id="29D30DCF1753FF9C62D9B34D4047CC3D" box="[1198,1313,1304,1328]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1753FF9C62D9B34D4047CC3D" box="[1198,1313,1304,1328]" italics="true" pageId="18" pageNumber="19">Axianassa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A6C925C71753FF9A66BEB36947AFC88A" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="discussion">
<paragraph id="EE6C764C1753FF9C66BEB36941D9CFB1" blockId="18.[151,1437,151,1977]" pageId="18" pageNumber="19">
<emphasis id="DCA7AA5E1753FF9C66BEB369445BCC59" bold="true" box="[201,317,1340,1364]" pageId="18" pageNumber="19">Remarks.</emphasis>
<taxonomicName id="29D30DCF1753FF9C6730B369476BCC59" box="[327,525,1340,1364]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E1753FF9C6730B369476BCC59" box="[327,525,1340,1364]" italics="true" pageId="18" pageNumber="19">Axianassa heardi</emphasis>
</taxonomicName>
<emphasis id="DCA7AA5E1753FF9C646FB369473CCC59" bold="true" box="[536,602,1340,1364]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="C79417251753FF9C646FB369473CCC59" box="[536,602,1340,1364]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is presently the only species in the genus
<taxonomicName id="29D30DCF1753FF9C6226B36941A2CC59" box="[1105,1220,1340,1364]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1753FF9C6226B36941A2CC59" box="[1105,1220,1340,1364]" italics="true" pageId="18" pageNumber="19">Axianassa</emphasis>
</taxonomicName>
with the uropodal exopod bearing a conspicuous transverse suture furnished with row of small subacute denticles (
<figureCitation id="76E86AC91753FF9C62A8B3354044CC75" box="[1247,1314,1376,1400]" captionStart="FIGURE 9" captionStartId="14.[151,255,1883,1907]" captionTargetBox="[154,1430,590,1850]" captionTargetId="figure@14.[151,1436,590,1862]" captionTargetPageId="14" captionText="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
G, H). The presence of a dentate transverse suture immediately separates the new species from all the other species of the genus
<taxonomicName id="29D30DCF1753FF9C6691B3FD443FCCCD" box="[230,345,1448,1472]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1753FF9C6691B3FD443FCCCD" box="[230,345,1448,1472]" italics="true" pageId="18" pageNumber="19">Axianassa</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="8A420BBD1753FF9C67ECB3FD47D0CCCD" author="Kensley" box="[411,694,1448,1472]" pageId="18" pageNumber="22" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">
Kensley &amp;
<collectingCountry id="96C436DC1753FF9C645FB3FD470BCCCD" box="[552,621,1448,1472]" name="Heard Island and McDonald Islands" pageId="18" pageNumber="19">Heard</collectingCountry>
1990
</bibRefCitation>
;
<bibRefCitation id="8A420BBD1753FF9C64BFB3FD4175CCCD" author="Rodrigues" box="[712,1043,1448,1472]" pageId="18" pageNumber="22" refString="Rodrigues, S. A. &amp; Shimizu, R. M. (1992) Description of a new Axianassa (Crustacea: Decapoda: Thalassinidea) from Brazil, and its first larval stage. Proceedings of the Biological Society of Washington, 105, 317 - 323." type="journal article" year="1992">Rodrigues &amp; Shimizu 1992</bibRefCitation>
; Anker 2010;
<bibRefCitation id="8A420BBD1753FF9C62B0B3FD40F7CCCD" author="Liu" box="[1223,1425,1448,1472]" pageId="18" pageNumber="22" refString="Liu, W. &amp; Liu, R. (2010) First record of the gebiidean genus Axianassa Schmitt, 1924 (Crustacea: Decapoda: Gebiidea: Axianassidae) in the western Pacific, with description of a new species from the South China Sea. The Raffles Bulletin of Zoology, 58, 193 - 197." type="journal article" year="2010">Liu &amp; Liu 2010</bibRefCitation>
).
<taxonomicName id="29D30DCF1753FF9C66E0B399443FCCE9" box="[151,345,1484,1508]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E1753FF9C66E0B399443FCCE9" box="[151,345,1484,1508]" italics="true" pageId="18" pageNumber="19">Axianassa heardi</emphasis>
</taxonomicName>
<emphasis id="DCA7AA5E1753FF9C6716B39944F9CCE9" bold="true" box="[353,415,1484,1508]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="C79417251753FF9C6716B39944F9CCE9" box="[353,415,1484,1508]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is also the third member of
<taxonomicName id="29D30DCF1753FF9C64A9B3994637CCE9" box="[734,849,1484,1508]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1753FF9C64A9B3994637CCE9" box="[734,849,1484,1508]" italics="true" pageId="18" pageNumber="19">Axianassa</emphasis>
</taxonomicName>
reported from the Indo-West Pacific, differing from the recently described
<taxonomicName id="29D30DCF1753FF9C67E5B3A547C8CF05" authority="Anker, 2010" authorityName="Anker" authorityYear="2010" box="[402,686,1520,1544]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ngochoae">
<emphasis id="DCA7AA5E1753FF9C67E5B3A5477BCF05" box="[402,541,1520,1544]" italics="true" pageId="18" pageNumber="19">A. ngochoae</emphasis>
Anker, 2010
</taxonomicName>
and
<taxonomicName id="29D30DCF1753FF9C6494B3A5469CCF05" authority="Liu &amp; Liu 2010" authorityName="Liu &amp; Liu" authorityYear="2010" box="[739,1018,1520,1544]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="DCA7AA5E1753FF9C6494B3A54623CF05" box="[739,837,1520,1544]" italics="true" pageId="18" pageNumber="19">A. sinica</emphasis>
<bibRefCitation id="8A420BBD1753FF9C653CB3A5469CCF05" author="Liu" box="[843,1018,1520,1544]" pageId="18" pageNumber="22" refString="Liu, W. &amp; Liu, R. (2010) First record of the gebiidean genus Axianassa Schmitt, 1924 (Crustacea: Decapoda: Gebiidea: Axianassidae) in the western Pacific, with description of a new species from the South China Sea. The Raffles Bulletin of Zoology, 58, 193 - 197." type="journal article" year="2010">
Liu &amp; Liu
<quantity id="292BDBA91753FF9C65B5B3A5469CCF05" box="[962,1018,1520,1544]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="18" pageNumber="19" unit="in" value="2010.0">2010</quantity>
</bibRefCitation>
</taxonomicName>
in a number of other characters, such as the marginally dentate rostrum and a much shorter antennal acicle (cf. Anker 2010: fig. 1d;
<bibRefCitation id="8A420BBD1753FF9C62C0B041400DCF21" author="Liu" box="[1207,1387,1556,1580]" pageId="18" pageNumber="22" refString="Liu, W. &amp; Liu, R. (2010) First record of the gebiidean genus Axianassa Schmitt, 1924 (Crustacea: Decapoda: Gebiidea: Axianassidae) in the western Pacific, with description of a new species from the South China Sea. The Raffles Bulletin of Zoology, 58, 193 - 197." type="journal article" year="2010">Liu &amp; Liu 2010</bibRefCitation>
: fig. 2A). All American species of
<taxonomicName id="29D30DCF1753FF9C6799B06D4707CF5D" box="[494,609,1592,1616]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1753FF9C6799B06D4707CF5D" box="[494,609,1592,1616]" italics="true" pageId="18" pageNumber="19">Axianassa</emphasis>
</taxonomicName>
differ from the new species by at least two morphological characters, in addition to the toothed diaeresis on the uropodal exopod (cf.
<bibRefCitation id="8A420BBD1753FF9C6533B009412ACF79" author="Kensley" box="[836,1100,1628,1652]" pageId="18" pageNumber="22" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">
Kensley &amp;
<collectingCountry id="96C436DC1753FF9C65B2B009416CCF79" box="[965,1034,1628,1652]" name="Heard Island and McDonald Islands" pageId="18" pageNumber="19">Heard</collectingCountry>
1990
</bibRefCitation>
;
<bibRefCitation id="8A420BBD1753FF9C622FB00940F6CF79" author="Rodrigues" box="[1112,1424,1628,1652]" pageId="18" pageNumber="22" refString="Rodrigues, S. A. &amp; Shimizu, R. M. (1992) Description of a new Axianassa (Crustacea: Decapoda: Thalassinidea) from Brazil, and its first larval stage. Proceedings of the Biological Society of Washington, 105, 317 - 323." type="journal article" year="1992">Rodrigues &amp; Shimizu 1992</bibRefCitation>
). Noteworthy,
<taxonomicName id="29D30DCF1753FF9C6759B0D544FDCF95" box="[302,411,1664,1688]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E1753FF9C6759B0D544FDCF95" box="[302,411,1664,1688]" italics="true" pageId="18" pageNumber="19">A. heardi</emphasis>
</taxonomicName>
<emphasis id="DCA7AA5E1753FF9C67D0B0D5448ACF95" bold="true" box="[423,492,1664,1688]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="C79417251753FF9C67D0B0D5448ACF95" box="[423,492,1664,1688]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
appears to be confined to sandy reef patches, whereas all the other species of
<taxonomicName id="29D30DCF1753FF9C66E0B0F1446CCFB1" box="[151,266,1700,1724]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1753FF9C66E0B0F1446CCFB1" box="[151,266,1700,1724]" italics="true" pageId="18" pageNumber="19">Axianassa</emphasis>
</taxonomicName>
inhabit muddier bottoms (cf.
<bibRefCitation id="8A420BBD1753FF9C642FB0F14638CFB1" author="Kensley" box="[600,862,1700,1724]" pageId="18" pageNumber="22" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">
Kensley &amp;
<collectingCountry id="96C436DC1753FF9C64AFB0F1467BCFB1" box="[728,797,1700,1724]" name="Heard Island and McDonald Islands" pageId="18" pageNumber="19">Heard</collectingCountry>
1990
</bibRefCitation>
; Anker 2010;
<bibRefCitation id="8A420BBD1753FF9C6589B0F141D5CFB1" author="Liu" box="[1022,1203,1700,1724]" pageId="18" pageNumber="22" refString="Liu, W. &amp; Liu, R. (2010) First record of the gebiidean genus Axianassa Schmitt, 1924 (Crustacea: Decapoda: Gebiidea: Axianassidae) in the western Pacific, with description of a new species from the South China Sea. The Raffles Bulletin of Zoology, 58, 193 - 197." type="journal article" year="2010">Liu &amp; Liu 2010</bibRefCitation>
).
</paragraph>
<paragraph id="EE6C764C1753FF9A66BEB09D47AFC88A" blockId="18.[151,1437,151,1977]" lastBlockId="20.[151,1436,151,391]" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
Several morphological features of
<taxonomicName id="29D30DCF1753FF9C6427B09D47DECFED" box="[592,696,1736,1760]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E1753FF9C6427B09D47DECFED" box="[592,696,1736,1760]" italics="true" pageId="18" pageNumber="19">A. heardi</emphasis>
</taxonomicName>
<emphasis id="DCA7AA5E1753FF9C64B7B09D4666CFED" bold="true" box="[704,768,1736,1760]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="C79417251753FF9C64B7B09D4666CFED" box="[704,768,1736,1760]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
disagree with the generic diagnosis of
<taxonomicName id="29D30DCF1753FF9C62CAB09D4056CFED" box="[1213,1328,1736,1760]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1753FF9C62CAB09D4056CFED" box="[1213,1328,1736,1760]" italics="true" pageId="18" pageNumber="19">Axianassa</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="8A420BBD1753FF9C66CAB0B94486CE09" author="Kensley" box="[189,480,1772,1796]" pageId="18" pageNumber="22" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">
Kensley &amp;
<collectingCountry id="96C436DC1753FF9C6734B0B944E1CE09" box="[323,391,1772,1796]" name="Heard Island and McDonald Islands" pageId="18" pageNumber="19">Heard</collectingCountry>
(1990)
</bibRefCitation>
. For example, the eyestalks were described as “short, cornea poorly defined, eyes poorly pigmented ...”. In
<taxonomicName id="29D30DCF1753FF9C67C0B1454779CE25" box="[439,543,1808,1832]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E1753FF9C67C0B1454779CE25" box="[439,543,1808,1832]" italics="true" pageId="18" pageNumber="19">A. heardi</emphasis>
</taxonomicName>
<emphasis id="DCA7AA5E1753FF9C6450B1454701CE25" bold="true" box="[551,615,1808,1832]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="C79417251753FF9C6450B1454701CE25" box="[551,615,1808,1832]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the eyes have a relatively large, globular cornea occupying most of the eyestalk’s apical area; the cornea seems to be relatively well-pigmented, although in life its colour is yellowish white (
<figureCitation id="76E86AC91753FF9C6690B10D445ACE7D" box="[231,316,1880,1904]" captionStart="FIGURE 12" captionStartId="17.[151,255,1797,1821]" captionTargetBox="[167,1423,540,1783]" captionTargetId="figure@17.[167,1424,540,1783]" captionTargetPageId="17" captionText="FIGURE 12. Habitus and colour in life of Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, dorsal; B, lateral." httpUri="https://zenodo.org/record/202984/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
), becoming blackish in alcohol (
<figureCitation id="76E86AC91753FF9C64C4B10D479FCE7D" box="[691,761,1880,1904]" captionStart="FIGURE 9" captionStartId="14.[151,255,1883,1907]" captionTargetBox="[154,1430,590,1850]" captionTargetId="figure@14.[151,1436,590,1862]" captionTargetPageId="14" captionText="FIGURE 9. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, general lateral view of carapace and abdomen; B, frontal region, dorsal (left antennule regenerating); C, same, lateral; D, rostrum and eyestalks, lateral; E, same, dorsal; F, antennal acicle, dorsal; G, uropod, dorsal; H, same, detail of posterior portion of exopod (setae omitted); I, telson, dorsal." httpUri="https://zenodo.org/record/202981/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
D). More importantly, in all other species of
<taxonomicName id="29D30DCF1753FF9C628AB10D4016CE7D" box="[1277,1392,1880,1904]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1753FF9C628AB10D4016CE7D" box="[1277,1392,1880,1904]" italics="true" pageId="18" pageNumber="19">Axianassa</emphasis>
</taxonomicName>
the uropodal exopod and endopod are “lacking sutures”, with the exopod bearing a “variable weak spination” (
<bibRefCitation id="8A420BBD1753FF9C6349B129445DCEB5" author="Kensley" pageId="18" pageNumber="22" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">
Kensley &amp;
<collectingCountry id="96C436DC1753FF9C66C3B1F5459FCEB5" box="[180,249,1952,1976]" name="Heard Island and McDonald Islands" pageId="18" pageNumber="19">Heard</collectingCountry>
1990
</bibRefCitation>
). As emphasised above, in
<taxonomicName id="29D30DCF1753FF9C64F6B1F5478ECEB5" box="[641,744,1952,1976]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E1753FF9C64F6B1F5478ECEB5" box="[641,744,1952,1976]" italics="true" pageId="18" pageNumber="19">A. heardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C79417251753FF9C6481B1F54652CEB5" box="[758,820,1952,1976]" pageId="18" pageNumber="19" rank="species">
<emphasis id="DCA7AA5E1753FF9C6481B1F54652CEB5" bold="true" box="[758,820,1952,1976]" pageId="18" pageNumber="19">n. sp.</emphasis>
</taxonomicNameLabel>
, the distal area of the exopod has a distinct suture furnished with small acute denticles. Such a suture is absent from the endopod, which only bears a few acute or subacute denticles dorsally and one marginal tooth laterally. In addition, the first maxilliped of
<taxonomicName id="29D30DCF1755FF9A6291B6E94036C9D9" box="[1254,1360,188,212]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E1755FF9A6291B6E94036C9D9" box="[1254,1360,188,212]" italics="true" pageId="20" pageNumber="21">A. heardi</emphasis>
</taxonomicName>
<emphasis id="DCA7AA5E1755FF9A632DB6E940FAC9D9" bold="true" box="[1370,1436,188,212]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="C79417251755FF9A632DB6E940FAC9D9" box="[1370,1436,188,212]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
bears a fairly large epipod (
<figureCitation id="76E86AC91755FF9A67ACB68A4748C9FA" box="[475,558,223,247]" captionStart="FIGURE 10" captionStartId="15.[151,255,1810,1834]" captionTargetBox="[182,1432,200,1756]" captionTargetId="figure@15.[151,1436,194,1788]" captionTargetPageId="15" captionText="FIGURE 10. Axianassa heardi n. sp., holotype, male from Lizard Island, Australia (QM W 29049): A, mandible, lateral; B, same, mesial; C, maxillule, lateral; D, maxilla, lateral; E, same, endopod, mesial view; F, first maxilliped, lateral; G, same, mesial; H, second maxilliped, lateral." httpUri="https://zenodo.org/record/202982/files/figure.png" pageId="20" pageNumber="21">Fig. 10</figureCitation>
E, F), which seems to be rudimentary in all other species of
<taxonomicName id="29D30DCF1755FF9A628FB6B5400DC9F5" box="[1272,1387,224,248]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1755FF9A628FB6B5400DC9F5" box="[1272,1387,224,248]" italics="true" pageId="20" pageNumber="21">Axianassa</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="8A420BBD1755FF9A66E0B75144F9C811" author="Kensley" box="[151,415,260,284]" pageId="20" pageNumber="22" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">
Kensley &amp;
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1990
</bibRefCitation>
: fig. 2H; Anker 2010: fig. 2e). Other possible particularities in the mouthparts of
<taxonomicName id="29D30DCF1755FF9A6343B75140FAC811" box="[1332,1436,260,284]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="heardi" status="sp. nov.">
<emphasis id="DCA7AA5E1755FF9A6343B75140FAC811" box="[1332,1436,260,284]" italics="true" pageId="20" pageNumber="21">A. heardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C79417251755FF9A66E0B77D45BFC832" box="[151,217,296,320]" pageId="20" pageNumber="21" rank="species">
<emphasis id="DCA7AA5E1755FF9A66E0B77D45B4C84D" bold="true" box="[151,210,296,320]" pageId="20" pageNumber="21">n. sp</emphasis>
.
</taxonomicNameLabel>
include the absence of teeth on the molar process and a biarticulated endopod of the maxilla. With more Indo-West Pacific species discovered in the future,
<taxonomicName id="29D30DCF1755FF9A6574B7194610C869" box="[771,886,332,356]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="DCA7AA5E1755FF9A6574B7194610C869" box="[771,886,332,356]" italics="true" pageId="20" pageNumber="21">Axianassa</emphasis>
</taxonomicName>
may eventually require a redefinition, which, however, is beyond the scope of the present study.
</paragraph>
</subSubSection>
</treatment>
</document>
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