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<document ID-DOI="10.11646/zootaxa.4766.1.5" ID-GBIF-Dataset="d16178ec-2278-4c7a-a479-53423c68e52c" ID-ISSN="1175-5334" ID-Zenodo-Dep="3763991" ID-ZooBank="urn:lsid:zoobank.org:pub:0E43BB66-03FD-443E-9D6E-1BEE52B0C459" checkinTime="1587670939645" checkinUser="carolina" docAuthor="Guinot, Danièle &amp; Bakel, Barry Van" docDate="2020" docId="663987C6FFA7A600B6F8F98EFDE3F95B" docLanguage="en" docName="zootaxa.4766.1.5.pdf.imf" docOrigin="Zootaxa 4766 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@1ef1a562" docStyleName="zootaxa.v2_0.2013.journal_article" docTitle="Paulitinae Guinot &amp; Van Bakel" docType="treatment" docVersion="5" lastPageNumber="116" masterDocId="9A00FFBEFFA9A60FB66FFFF5FFC5FFB8" masterDocTitle="Extraordinary majoid crabs: the genus Esopus A. Milne-Edwards, 1875 in the new subfamily Esopinae subfam. nov., and erection of Paulitinae subfam. nov. (Crustacea, Decapoda, Brachyura, Majoidea, Inachoididae Dana, 1851)" masterLastPageNumber="127" masterPageNumber="101" pageNumber="115" updateTime="1643448219730" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Extraordinary majoid crabs: the genus Esopus A. Milne-Edwards, 1875 in the new subfamily Esopinae subfam. nov., and erection of Paulitinae subfam. nov. (Crustacea, Decapoda, Brachyura, Majoidea, Inachoididae Dana, 1851)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Guinot, Danièle</mods:namePart>
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<mods:namePart>Bakel, Barry Van</mods:namePart>
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<mods:date>2020</mods:date>
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<mods:number>2020-04-17</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3803779" ID-GBIF-Taxon="163737259" ID-Zenodo-Dep="3803779" LSID="urn:lsid:plazi:treatment:663987C6FFA7A600B6F8F98EFDE3F95B" httpUri="http://treatment.plazi.org/id/663987C6FFA7A600B6F8F98EFDE3F95B" lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115">
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<heading box="[151,850,1659,1685]" pageId="14" pageNumber="115">
<emphasis bold="true" box="[151,850,1659,1685]" pageId="14" pageNumber="115">
Subfamily
<taxonomicName authority="Guinot &amp; Van Bakel" authorityName="Guinot &amp; Van Bakel" box="[287,678,1659,1685]" pageId="14" pageNumber="115" rank="subFamily" subFamily="Paulitinae">Paulitinae Guinot &amp; Van Bakel</taxonomicName>
, subfam. nov.
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</paragraph>
</subSubSection>
<subSubSection box="[151,1405,1731,1758]" pageId="14" pageNumber="115" type="type_taxon">
<paragraph blockId="14.[151,1437,1731,2046]" box="[151,1405,1731,1758]" pageId="14" pageNumber="115">
<emphasis bold="true" box="[151,285,1731,1757]" pageId="14" pageNumber="115">
<typeStatus box="[151,210,1731,1757]" pageId="14" pageNumber="115">Type</typeStatus>
genus
</emphasis>
.
<taxonomicName authority="Guinot, 2012" authorityName="Guinot" authorityYear="2012" box="[299,536,1731,1757]" class="Magnoliopsida" family="Apiaceae" genus="Paulita" kingdom="Plantae" order="Apiales" pageId="14" pageNumber="115" phylum="Tracheophyta" rank="genus">
<emphasis box="[299,381,1731,1757]" italics="true" pageId="14" pageNumber="115">Paulita</emphasis>
<bibRefCitation author="Guinot" box="[389,536,1731,1757]" firstAuthor="Guinot" pageId="14" pageNumber="115" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot, 2012</bibRefCitation>
</taxonomicName>
(
<typeStatus box="[551,601,1731,1757]" pageId="14" pageNumber="115">type</typeStatus>
species by monotypy:
<taxonomicName authority="Lemos de Castro, 1949" authorityName="Lemos de Castro" authorityYear="1949" box="[859,1393,1731,1758]" class="Malacostraca" family="Inachoididae" genus="Dasygyius" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[859,1122,1731,1757]" italics="true" pageId="14" pageNumber="115">Dasygyius tuberculatus</emphasis>
<bibRefCitation author="Lemos de Castro" box="[1129,1393,1731,1758]" firstAuthor="Lemos de Castro" pageId="14" pageNumber="115" pagination="349 - 352" refId="ref23072" refString="Lemos de Castro, A. (1949) &quot; Dasygyius tuberculatus &quot;, uma nova especie de Crustaceo do Brasil (Decapoda, Majidae). Revista Brasileira de Biologia, 9 (3), 349 - 352." type="journal article" year="1949">Lemos de Castro, 1949</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115" type="description">
<paragraph blockId="14.[151,1437,1731,2046]" lastBlockId="15.[151,1437,153,1763]" lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115">
<emphasis bold="true" box="[199,337,1766,1793]" pageId="14" pageNumber="115">Description</emphasis>
. Body moderately flat. Carapace urn-shaped, with moderate anterior narrowing, resting on setting gutter (
<bibRefCitation author="Lemos de Castro" box="[234,494,1803,1830]" firstAuthor="Lemos de Castro" pageId="14" pageNumber="115" pagination="349 - 352" refId="ref23072" refString="Lemos de Castro, A. (1949) &quot; Dasygyius tuberculatus &quot;, uma nova especie de Crustaceo do Brasil (Decapoda, Majidae). Revista Brasileira de Biologia, 9 (3), 349 - 352." type="journal article" year="1949">Lemos de Castro 1949</bibRefCitation>
: figs. 8, 9, as
<taxonomicName authorityName="Lemos de Castro" authorityYear="1949" box="[653,917,1803,1829]" class="Malacostraca" family="Inachoididae" genus="Dasygyius" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[653,917,1803,1829]" italics="true" pageId="14" pageNumber="115">Dasygyius tuberculatus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Drach &amp; Guinot" box="[933,1186,1803,1829]" firstAuthor="Drach" pageId="14" pageNumber="115" pagination="715 - 720" refId="ref20996" refString="Drach, P. &amp; Guinot, D. (1982) Connexions morphologiques et fonctionnelles d'un type nouveau dans le squelette des Brachyoures du genre Paradasygius [sic] Garth (carapace, pleurites, sternites, pleon). Comptes rendus hebdomadaires des Seances de l'Academie des Sciences, Serie 3, 295, 715 - 720." type="journal article" year="1982">Drach &amp; Guinot 1982</bibRefCitation>
: pl. 1, fig. 1, as
<taxonomicName baseAuthorityName="Lemos de Castro" baseAuthorityYear="1949" class="Malacostraca" family="Inachoididae" genus="Paradasygyius" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis italics="true" pageId="14" pageNumber="115">Paradasygyius tuberculatus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Santana, W. R. A." box="[422,578,1839,1865]" pageId="14" pageNumber="115" refId="ref24839" refString="Santana, W. R. A. (2008) Revisao taxonomica e relacoes filogeneticas em Inachoididae Dana, 1851 (Crustacea, Brachyura, Majoidea). Unpublished doctoral thesis, Universidade de Sao Paulo, Sao Paulo, 244 pp." type="book" year="2008">Santana 2008</bibRefCitation>
: figs. 50C, 51B, C, as
<taxonomicName baseAuthorityName="Lemos de Castro" baseAuthorityYear="1949" box="[828,1140,1839,1865]" class="Malacostraca" family="Inachoididae" genus="Paradasygyius" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[828,1140,1839,1865]" italics="true" pageId="14" pageNumber="115">Paradasygyius tuberculatus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Guinot" box="[1155,1300,1839,1865]" firstAuthor="Guinot" pageId="14" pageNumber="115" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: figs. 1, 2A;
<bibRefCitation author="Guinot" box="[151,361,1875,1901]" etAl="et al." firstAuthor="Guinot" pageId="14" pageNumber="115" pagination="285 - 322" refId="ref22322" refString="Guinot, D., Carbot-Chanona, G. &amp; Vega, F. J. (2019) Archaeochiapasidae n. fam., a new early Cenomanian brachyuran family from Chiapas, Mexico, new insights on Lecythocaridae Schweitzer &amp; Feldmann, 2009, and phylogenetic implications (Crustacea, Decapoda, Brachyura, Eubrachyura). Geodiversitas, 41 (7), 285 - 322. https: // doi. org / 10.5252 / geodiversitas 2019 v 41 a 7" type="journal article" year="2019">
Guinot
<emphasis box="[237,294,1875,1901]" italics="true" pageId="14" pageNumber="115">et al.</emphasis>
2019
</bibRefCitation>
: fig. 16A). Dorsal surface (
<bibRefCitation author="Guinot" box="[670,817,1875,1901]" firstAuthor="Guinot" pageId="14" pageNumber="115" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: figs. 1, 2A, B) divided into convex regions delineated by several parallel, deep, broad grooves: cervical groove well marked, bearing median gastric pits; branchial groove delimiting main part of branchial region; branchiocardiac groove marked. Dorsal surface uniformly covered with prominent tubercles. Lateral margins rounded, unarmed. Pleurites 58 (
<bibRefCitation author="Guinot" box="[955,1102,1983,2009]" firstAuthor="Guinot" pageId="14" pageNumber="115" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: fig. 2A, B) exposed on same level as carapace, calcified, ornamented as carapace; first pleonal somite in prolongation of cephalothorax, dorsally visible, calcified, ornamented; insertion of carapace into setting gutter (
<bibRefCitation author="Drach &amp; Guinot" box="[946,1195,153,179]" firstAuthor="Drach" pageId="15" pageNumber="116" pagination="715 - 720" refId="ref20996" refString="Drach, P. &amp; Guinot, D. (1982) Connexions morphologiques et fonctionnelles d'un type nouveau dans le squelette des Brachyoures du genre Paradasygius [sic] Garth (carapace, pleurites, sternites, pleon). Comptes rendus hebdomadaires des Seances de l'Academie des Sciences, Serie 3, 295, 715 - 720." type="journal article" year="1982">Drach &amp; Guinot 1982</bibRefCitation>
: pl. 1, fig. 3, as
<taxonomicName baseAuthorityName="Lemos de Castro" baseAuthorityYear="1949" class="Malacostraca" family="Inachoididae" genus="Paradasygyius" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis italics="true" pageId="15" pageNumber="116">Paradasygyius tuberculatus</emphasis>
</taxonomicName>
); branchiostegite mostly absent. Rostrum simple, broadly triangular, pentagonal (
<bibRefCitation author="Lemos de Castro" firstAuthor="Lemos de Castro" pageId="15" pageNumber="116" pagination="349 - 352" refId="ref23072" refString="Lemos de Castro, A. (1949) &quot; Dasygyius tuberculatus &quot;, uma nova especie de Crustaceo do Brasil (Decapoda, Majidae). Revista Brasileira de Biologia, 9 (3), 349 - 352." type="journal article" year="1949">Lemos de Castro 1949</bibRefCitation>
: fig. 4, as
<taxonomicName authorityName="Lemos de Castro" authorityYear="1949" box="[417,682,225,251]" class="Malacostraca" family="Inachoididae" genus="Dasygyius" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[417,682,225,251]" italics="true" pageId="15" pageNumber="116">Dasygyius tuberculatus</emphasis>
</taxonomicName>
), its ventral portion (proepistome) joining the epistomial process. Antennules vertically folded beneath rostrum. Eyestalk narrow. Antenna: urinary article adjacent to basal article (2+3) (thus not sunken in epistome); basal article large, fused to front, unarmed; articles 4, 5 free; flagellum short (
<bibRefCitation author="Lemos de Castro" box="[161,417,333,359]" firstAuthor="Lemos de Castro" pageId="15" pageNumber="116" pagination="349 - 352" refId="ref23072" refString="Lemos de Castro, A. (1949) &quot; Dasygyius tuberculatus &quot;, uma nova especie de Crustaceo do Brasil (Decapoda, Majidae). Revista Brasileira de Biologia, 9 (3), 349 - 352." type="journal article" year="1949">Lemos de Castro 1949</bibRefCitation>
: fig. 5, as
<taxonomicName authorityName="Lemos de Castro" authorityYear="1949" box="[532,795,333,359]" class="Malacostraca" family="Inachoididae" genus="Dasygyius" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[532,795,333,359]" italics="true" pageId="15" pageNumber="116">Dasygyius tuberculatus</emphasis>
</taxonomicName>
). Supraorbital margin with median tooth. Exorbital tooth blunt. Pterygostomial lobe well marked (
<bibRefCitation author="Guinot" box="[607,754,369,395]" firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: fig. 3B). Mxp3: ischium with inner margin coarsely dentate, produced into lobe at inner distal angle; merus cordiform, with outer angle slightly rounded laterally and inner angle produced; palp stocky, with carpus enlarged, three-fifths as thick as long (
<bibRefCitation author="Lemos de Castro" box="[1064,1325,441,467]" firstAuthor="Lemos de Castro" pageId="15" pageNumber="116" pagination="349 - 352" refId="ref23072" refString="Lemos de Castro, A. (1949) &quot; Dasygyius tuberculatus &quot;, uma nova especie de Crustaceo do Brasil (Decapoda, Majidae). Revista Brasileira de Biologia, 9 (3), 349 - 352." type="journal article" year="1949">Lemos de Castro 1949</bibRefCitation>
: fig. 3, as
<taxonomicName authorityName="Lemos de Castro" authorityYear="1949" box="[151,413,477,503]" class="Malacostraca" family="Inachoididae" genus="Dasygyius" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[151,413,477,503]" italics="true" pageId="15" pageNumber="116">Dasygyius tuberculatus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Guinot" box="[428,572,477,503]" firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: fig. 3B). Thoracic sternum markedly wide, inflated; anterior shield (sternites 13) inserted between mxp3, as a pentagonal shield; sternite 3 developed, much inflated; suture 3/4 laterally visible, continuing into depression (
<bibRefCitation author="Santana, W. R. A." box="[467,625,549,575]" pageId="15" pageNumber="116" refId="ref24839" refString="Santana, W. R. A. (2008) Revisao taxonomica e relacoes filogeneticas em Inachoididae Dana, 1851 (Crustacea, Brachyura, Majoidea). Unpublished doctoral thesis, Universidade de Sao Paulo, Sao Paulo, 244 pp." type="book" year="2008">Santana 2008</bibRefCitation>
: figs. 51D, 52B, as
<taxonomicName baseAuthorityName="Lemos de Castro" baseAuthorityYear="1949" box="[852,1165,549,575]" class="Malacostraca" family="Inachoididae" genus="Paradasygyius" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[852,1165,549,575]" italics="true" pageId="15" pageNumber="116">Paradasygyius tuberculatus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Guinot" box="[1181,1327,549,575]" firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: figs. 2C, 3A;
<bibRefCitation author="Guinot" box="[201,413,585,611]" etAl="et al." firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="285 - 322" refId="ref22322" refString="Guinot, D., Carbot-Chanona, G. &amp; Vega, F. J. (2019) Archaeochiapasidae n. fam., a new early Cenomanian brachyuran family from Chiapas, Mexico, new insights on Lecythocaridae Schweitzer &amp; Feldmann, 2009, and phylogenetic implications (Crustacea, Decapoda, Brachyura, Eubrachyura). Geodiversitas, 41 (7), 285 - 322. https: // doi. org / 10.5252 / geodiversitas 2019 v 41 a 7" type="journal article" year="2019">
Guinot
<emphasis box="[288,345,585,611]" italics="true" pageId="15" pageNumber="116">et al.</emphasis>
2019
</bibRefCitation>
: fig. 16C). Sutures 4/57/8 all interrupted, with distant interruption points. Sternite 8 with basal median line, continuing by depression. Sternum/pterygostome junction complete thanks to curved extension of sternite 4. Milne-Edwards openings separated from chelipeds, large, entirely filled by mxp3 coxa (
<bibRefCitation author="Guinot" box="[1286,1432,657,683]" firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: fig. 3B). Wide sternal extensions joining exposed pleurites (sternum/pleurites connections) between P1/P2, P2/P3, P3/P4, P4/P5 (
<bibRefCitation author="Guinot" box="[317,466,729,755]" firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: figs. 2C, 3A). Sternal device for pleonal-locking system represented by 2 or several granules disposed between sutures 4/5, 5/6; pleonal edge of somites 46 markedly thickened, ending in deep socket. Male, female pleons with first somite developed, entirely dorsally visible, ornamented like carapace, not produced into spine. Male pleon (
<bibRefCitation author="Lemos de Castro" box="[420,679,837,863]" firstAuthor="Lemos de Castro" pageId="15" pageNumber="116" pagination="349 - 352" refId="ref23072" refString="Lemos de Castro, A. (1949) &quot; Dasygyius tuberculatus &quot;, uma nova especie de Crustaceo do Brasil (Decapoda, Majidae). Revista Brasileira de Biologia, 9 (3), 349 - 352." type="journal article" year="1949">Lemos de Castro 1949</bibRefCitation>
: fig. 6, as
<taxonomicName authorityName="Lemos de Castro" authorityYear="1949" box="[796,1059,837,863]" class="Malacostraca" family="Inachoididae" genus="Dasygyius" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[796,1059,837,863]" italics="true" pageId="15" pageNumber="116">Dasygyius tuberculatus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Guinot" box="[1074,1220,837,863]" firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: figs. 2C, 3A) with all somites free, except for somite 6 fused to telson (pleotelson); somite 1 narrow but high; somites 2, 3 widened; somites 4, 5, pleotelson narrow; pleotelson base laterally inflated at location of sockets. Female pleon (
<bibRefCitation author="Lemos de Castro" firstAuthor="Lemos de Castro" pageId="15" pageNumber="116" pagination="349 - 352" refId="ref23072" refString="Lemos de Castro, A. (1949) &quot; Dasygyius tuberculatus &quot;, uma nova especie de Crustaceo do Brasil (Decapoda, Majidae). Revista Brasileira de Biologia, 9 (3), 349 - 352." type="journal article" year="1949">Lemos de Castro 1949</bibRefCitation>
: fig. 7, as
<taxonomicName authorityName="Lemos de Castro" authorityYear="1949" box="[358,620,945,971]" class="Malacostraca" family="Inachoididae" genus="Dasygyius" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[358,620,945,971]" italics="true" pageId="15" pageNumber="116">Dasygyius tuberculatus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Guinot" box="[633,777,945,971]" firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: fig. 2D) with somites 14 free, not high; somites 5, 6 fused to pleotelson, forming large, flat or convex disc, inserted in sterno-pleonal cavity; brood cavity closed like a box, limited by elevated sternal arch formed by raised sternites all around. Male chelipeds (
<bibRefCitation author="Lemos de Castro" box="[1107,1361,1017,1043]" firstAuthor="Lemos de Castro" pageId="15" pageNumber="116" pagination="349 - 352" refId="ref23072" refString="Lemos de Castro, A. (1949) &quot; Dasygyius tuberculatus &quot;, uma nova especie de Crustaceo do Brasil (Decapoda, Majidae). Revista Brasileira de Biologia, 9 (3), 349 - 352." type="journal article" year="1949">Lemos de Castro 1949</bibRefCitation>
: fig. 2, as
<taxonomicName authorityName="Lemos de Castro" authorityYear="1949" box="[181,443,1053,1079]" class="Malacostraca" family="Inachoididae" genus="Dasygyius" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[181,443,1053,1079]" italics="true" pageId="15" pageNumber="116">Dasygyius tuberculatus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Guinot" box="[457,601,1053,1079]" firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: fig. 1) equal, unarmed, with minute granules; propodus narrow, elongated, slightly inflated in larger males; fingers long, tapering, gaping in proximal half, distally joined; prehensile border finely denticulated in proximal half, distinctly toothed distally. Female chelipeds (
<bibRefCitation author="Lemos de Castro" box="[1071,1329,1125,1151]" firstAuthor="Lemos de Castro" pageId="15" pageNumber="116" pagination="349 - 352" refId="ref23072" refString="Lemos de Castro, A. (1949) &quot; Dasygyius tuberculatus &quot;, uma nova especie de Crustaceo do Brasil (Decapoda, Majidae). Revista Brasileira de Biologia, 9 (3), 349 - 352." type="journal article" year="1949">Lemos de Castro 1949</bibRefCitation>
: fig. 1, as
<taxonomicName authorityName="Lemos de Castro" authorityYear="1949" box="[151,414,1161,1187]" class="Malacostraca" family="Inachoididae" genus="Dasygyius" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[151,414,1161,1187]" italics="true" pageId="15" pageNumber="116">Dasygyius tuberculatus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Guinot" box="[429,574,1161,1187]" firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: fig. 2D) narrow, fingers joined. Pereopods long, thin, cylindrical throughout length, unarmed; P2 densely fringed with hairs in males; P2-P5 with additional setae in females (
<bibRefCitation author="Lemos de Castro" firstAuthor="Lemos de Castro" pageId="15" pageNumber="116" pagination="349 - 352" refId="ref23072" refString="Lemos de Castro, A. (1949) &quot; Dasygyius tuberculatus &quot;, uma nova especie de Crustaceo do Brasil (Decapoda, Majidae). Revista Brasileira de Biologia, 9 (3), 349 - 352." type="journal article" year="1949">Lemos de Castro 1949</bibRefCitation>
: figs. 811 as
<taxonomicName authorityName="Lemos de Castro" authorityYear="1949" box="[373,636,1233,1259]" class="Malacostraca" family="Inachoididae" genus="Dasygyius" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[373,636,1233,1259]" italics="true" pageId="15" pageNumber="116">Dasygyius tuberculatus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Guinot" box="[652,797,1233,1259]" firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: fig. 1). Coxal male gonopore large, located far from suture 7/8, thus in posteriormost location; opening on anterior border of coxo-sternal condyle, thus partially condylar, partially coxal (
<bibRefCitation author="Guinot" box="[329,600,1305,1331]" etAl="et al." firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="1 - 414" refId="ref22407" refString="Guinot, D., Tavares, M. &amp; Castro, P. (2013) Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa. Zootaxa, 3665 (1), 1 - 414. https: // doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
Guinot
<emphasis box="[417,467,1305,1331]" italics="true" pageId="15" pageNumber="116">et al</emphasis>
. 2013: 127
</bibRefCitation>
); penis short, emerging on lateral side of coxo-sternal condyle. G1 gently curved, with elongated, twisted lobe (
<bibRefCitation author="Guinot-Dumortier" box="[575,849,1341,1367]" firstAuthor="Guinot-Dumortier" pageId="15" pageNumber="116" pagination="177 - 187" refId="ref22477" refString="Guinot-Dumortier, D. (1960) Sur une collection de Crustaces (Decapodes Reptantia) de Guyane francaise. II. Brachyura Oxyrhyncha et Macrura. Bulletin du Museum national d'Histoire naturelle, Paris, Series 2, 32 (2), 177 - 187." type="journal article" year="1960">Guinot-Dumortier 1960</bibRefCitation>
: fig. 22ac, as
<taxonomicName baseAuthorityName="Lemos de Castro" baseAuthorityYear="1949" box="[1019,1332,1341,1367]" class="Malacostraca" family="Inachoididae" genus="Paradasygyius" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[1019,1332,1341,1367]" italics="true" pageId="15" pageNumber="116">Paradasygyius tuberculatus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Santana, W. R. A." pageId="15" pageNumber="116" refId="ref24839" refString="Santana, W. R. A. (2008) Revisao taxonomica e relacoes filogeneticas em Inachoididae Dana, 1851 (Crustacea, Brachyura, Majoidea). Unpublished doctoral thesis, Universidade de Sao Paulo, Sao Paulo, 244 pp." type="book" year="2008">Santana 2008</bibRefCitation>
: figs. 50C, 51D, as
<taxonomicName baseAuthorityName="Lemos de Castro" baseAuthorityYear="1949" box="[430,742,1377,1403]" class="Malacostraca" family="Inachoididae" genus="Paradasygyius" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[430,742,1377,1403]" italics="true" pageId="15" pageNumber="116">Paradasygyius tuberculatus</emphasis>
</taxonomicName>
). G2 conspicuously short. Vulvae much protruded, located on sternite 6 displaced anteriorly to suture 4/5; sternal vulvar cover developed. Axial skeleton with lateral compartment together with dorsoventral partition (developed junction plate), pleurites being almost horizontal (
<bibRefCitation author="Guinot" box="[1242,1388,1449,1475]" firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="22 - 40" refId="ref21874" refString="Guinot, D. (2012) Remarks on Inachoididae Dana, 1851, with the description of a new genus and the resurrection of Stenorhynchinae Dana, 1851, and recognition of the inachid subfamily Podochelinae Neumann, 1878 (Crustacea, Decapoda, Brachyura, Majoidea). Zootaxa, 3416 (1), 22 - 40. https: // doi. org / 10.11646 / zootaxa. 3416.1.2" type="journal article" year="2012">Guinot 2012</bibRefCitation>
: fig. 2B); in anterior region, dorsal edges of pleurites connected to internal surface of carapace by vertical pillars (
<bibRefCitation author="Drach &amp; Guinot" firstAuthor="Drach" pageId="15" pageNumber="116" pagination="715 - 720" refId="ref20996" refString="Drach, P. &amp; Guinot, D. (1982) Connexions morphologiques et fonctionnelles d'un type nouveau dans le squelette des Brachyoures du genre Paradasygius [sic] Garth (carapace, pleurites, sternites, pleon). Comptes rendus hebdomadaires des Seances de l'Academie des Sciences, Serie 3, 295, 715 - 720." type="journal article" year="1982">Drach &amp; Guinot 1982</bibRefCitation>
: pl. 1, fig. 6, as
<taxonomicName baseAuthorityName="Lemos de Castro" baseAuthorityYear="1949" box="[513,826,1521,1547]" class="Malacostraca" family="Inachoididae" genus="Paradasygyius" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis box="[513,826,1521,1547]" italics="true" pageId="15" pageNumber="116">Paradasygyius tuberculatus</emphasis>
</taxonomicName>
). A bridge with median line (with posteriorly and anteriorly a depression) in middle of sternite 7 (corresponding to raised median plate). Presence of thick sella turcica (
<bibRefCitation author="Guinot" box="[159,368,1593,1619]" etAl="et al." firstAuthor="Guinot" pageId="15" pageNumber="116" pagination="1 - 414" refId="ref22407" refString="Guinot, D., Tavares, M. &amp; Castro, P. (2013) Significance of the sexual openings and supplementary structures on the phylogeny of brachyuran crabs (Crustacea, Decapoda, Brachyura), with new nomina for higher-ranked podotreme taxa. Zootaxa, 3665 (1), 1 - 414. https: // doi. org / 10.11646 / zootaxa. 3665.1.1" type="journal article" year="2013">
Guinot
<emphasis box="[245,295,1593,1619]" italics="true" pageId="15" pageNumber="116">et al</emphasis>
. 2013
</bibRefCitation>
: fig. 47G, H).
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="116" type="discussion">
<paragraph blockId="15.[151,1437,153,1763]" pageId="15" pageNumber="116">
<emphasis bold="true" box="[199,315,1629,1655]" pageId="15" pageNumber="116">Remarks.</emphasis>
<taxonomicName box="[326,438,1629,1655]" pageId="15" pageNumber="115" rank="subFamily" subFamily="Paulitinae">Paulitinae</taxonomicName>
subfam. nov. may be differentiated from the
<taxonomicName authorityName="Holmes" authorityYear="1900" box="[978,1118,1629,1655]" pageId="15" pageNumber="114" rank="subFamily" subFamily="Dasygyiinae">Dasygyiinae</taxonomicName>
by a number of characters mainly related to the carapace, thoracic sternum, links between sternum and pterygostome, Milne-Edwards openings, and first gonopod.
</paragraph>
</subSubSection>
<subSubSection box="[199,550,1736,1763]" pageId="15" pageNumber="116">
<paragraph blockId="15.[151,1437,153,1763]" box="[199,550,1736,1763]" pageId="15" pageNumber="116">
<emphasis bold="true" box="[199,468,1736,1763]" pageId="15" pageNumber="116">Other included genera</emphasis>
. None.
</paragraph>
</subSubSection>
</treatment>
</document>