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<mods:title id="7313BEB9230763D817D6CC4A6C1BE7F5">South American Coccinellidae (Coleoptera), Part XIX: Overview of Cryptognathini and systematic revision of South American Cryptognatha Mulsant</mods:title>
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<mods:namePart id="A3B34C0B4AB87AF713FAE543D631C1C7">F, Guillermo González</mods:namePart>
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<treatment id="4E5F8790C74C95348D90D7D52166FAEC" ID-DOI="http://doi.org/10.5281/zenodo.3680646" ID-GBIF-Taxon="162173278" ID-Zenodo-Dep="3680646" LSID="urn:lsid:plazi:treatment:4E5F8790C74C95348D90D7D52166FAEC" httpUri="http://treatment.plazi.org/id/4E5F8790C74C95348D90D7D52166FAEC" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17">
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17.
<taxonomicName id="01F64D05C74C95358DBDD7D421AAFA78" authority="Mulsant" authorityName="Mulsant" box="[237,707,1496,1521]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="pudibunda">
<emphasis id="F482EA94C74C95358DBDD7D42123FA79" bold="true" box="[237,586,1496,1520]" italics="true" pageId="18" pageNumber="17">Cryptognatha pudibunda</emphasis>
Mulsant
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<taxonomicName id="01F64D05C74C95358D90D40B21DAF994" authority="Mulsant 1850: 499" authorityName="Mulsant" authorityPageNumber="499" authorityYear="1850" box="[192,691,1543,1565]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="pudibunda">
<emphasis id="F482EA94C74C95358D90D40B22B3F995" box="[192,474,1543,1564]" italics="true" pageId="18" pageNumber="17">Cryptognatha pudibunda</emphasis>
<treatmentCitation id="47571097C74C95358CB1D40B21DAF994" author="Mulsant, M. E." box="[481,691,1543,1565]" page="499" pageId="18" pageNumber="17" year="1850">
<bibRefCitation id="A2674B77C74C95358CB1D40B21DAF994" author="Mulsant, M. E." box="[481,691,1543,1565]" pageId="18" pageNumber="17" pagination="1 - 1104" refId="ref19658" refString="Mulsant, M. E. 1850. Species de coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, Lyon 2: 1 - 1104." type="journal article" year="1850">Mulsant 1850: 499</bibRefCitation>
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;
<treatmentCitation id="47571097C74C95358FEED40B2014F994" author="Crotch, G. R." box="[702,893,1543,1565]" page="206" pageId="18" pageNumber="17" year="1874">
<bibRefCitation id="A2674B77C74C95358FEED40B2014F994" author="Crotch, G. R." box="[702,893,1543,1565]" pageId="18" pageNumber="17" refId="ref19186" refString="Crotch, G. R. 1874. A revision of the coleopterous family Coccinellidae. E. W. Janson; London. 311 p." type="book" year="1874">Crotch 1874: 206</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="47571097C74C95358ED7D40B27E8F994" author="Korschefsky, R." box="[903,1153,1543,1565]" page="219" pageId="18" pageNumber="17" year="1931">
<bibRefCitation id="A2674B77C74C95358ED7D40B27E8F994" author="Korschefsky, R." box="[903,1153,1543,1565]" pageId="18" pageNumber="17" refId="ref19600" refString="Korschefsky, R. 1931. Coccinellidae I. Coleopterorum Catalogus. Part 118. W. Junk; Berlin. 224 p." type="book" year="1931">Korschefsky 1931: 219</bibRefCitation>
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;
<treatmentCitation id="47571097C74C953589DCD40B26E0F994" author="Blackwelder, E." box="[1164,1417,1543,1565]" page="449" pageId="18" pageNumber="17" year="1945">
<bibRefCitation id="A2674B77C74C953589DCD40B26E0F994" author="Blackwelder, E." box="[1164,1417,1543,1565]" pageId="18" pageNumber="17" pagination="343 - 550" refId="ref18961" refString="Blackwelder, E. 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 3. Bulletin of the United States National Museum 185: 343 - 550." type="journal article" year="1945">Blackwelder 1945: 449</bibRefCitation>
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.
<taxonomicName id="01F64D05C74C95358D90D428213AF9B3" authority="Crotch 1874: 207" authorityName="Crotch" authorityPageNumber="207" authorityYear="1874" box="[192,595,1572,1594]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="fryii">
<emphasis id="F482EA94C74C95358D90D42822E6F9B0" box="[192,399,1572,1593]" italics="true" pageId="18" pageNumber="17">Cryptognatha fryii</emphasis>
<treatmentCitation id="47571097C74C95358CC6D428213AF9B3" author="Crotch, G. R." box="[406,595,1572,1594]" page="207" pageId="18" pageNumber="17" year="1874">
<bibRefCitation id="A2674B77C74C95358CC6D428213AF9B3" author="Crotch, G. R." box="[406,595,1572,1594]" pageId="18" pageNumber="17" refId="ref19186" refString="Crotch, G. R. 1874. A revision of the coleopterous family Coccinellidae. E. W. Janson; London. 311 p." type="book" year="1874">Crotch 1874: 207</bibRefCitation>
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;
<treatmentCitation id="47571097C74C95358F0ED4282031F9B3" author="Korschefsky, R." box="[606,856,1572,1594]" page="219" pageId="18" pageNumber="17" year="1931">
<bibRefCitation id="A2674B77C74C95358F0ED4282031F9B3" author="Korschefsky, R." box="[606,856,1572,1594]" pageId="18" pageNumber="17" refId="ref19600" refString="Korschefsky, R. 1931. Coccinellidae I. Coleopterorum Catalogus. Part 118. W. Junk; Berlin. 224 p." type="book" year="1931">Korschefsky 1931: 219</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="47571097C74C95358E33D4282709F9B3" author="Blackwelder, E." box="[867,1120,1572,1594]" page="449" pageId="18" pageNumber="17" year="1945">
<bibRefCitation id="A2674B77C74C95358E33D4282709F9B3" author="Blackwelder, E." box="[867,1120,1572,1594]" pageId="18" pageNumber="17" pagination="343 - 550" refId="ref18961" refString="Blackwelder, E. 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 3. Bulletin of the United States National Museum 185: 343 - 550." type="journal article" year="1945">Blackwelder 1945: 449</bibRefCitation>
</treatmentCitation>
.
<emphasis id="F482EA94C74C9535893AD428267EF9B0" bold="true" box="[1130,1303,1572,1593]" pageId="18" pageNumber="17">New synonym</emphasis>
.
<taxonomicName id="01F64D05C74C95358D90D44D21E6F9DF" authority="Brethes 1925 b: 201" authorityName="Brethes" authorityPageNumber="201" authorityYear="1925" box="[192,655,1601,1623]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="F482EA94C74C95358D90D44D22D9F9DF" box="[192,432,1601,1622]" italics="true" pageId="18" pageNumber="17">Cryptognatha bryanti</emphasis>
<treatmentCitation id="47571097C74C95358CE7D44D21E6F9DF" author="Brethes, J." box="[439,655,1601,1623]" page="201" pageId="18" pageNumber="17" year="1925">
<bibRefCitation id="A2674B77C74C95358CE7D44D21E6F9DF" author="Brethes, J." box="[439,655,1601,1623]" pageId="18" pageNumber="17" pagination="195 - 214" refId="ref19084" refString="Brethes, J. 1925 b. Coccinellides du British Museum. Anales Museo Histoire Naturelle 33: 195 - 214." type="journal article" year="1925">Brèthes 1925b: 201</bibRefCitation>
</treatmentCitation>
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;
<treatmentCitation id="47571097C74C95358FCAD44D20FDF9DF" author="Korschefsky, R." box="[666,916,1601,1623]" page="219" pageId="18" pageNumber="17" year="1931">
<bibRefCitation id="A2674B77C74C95358FCAD44D20FDF9DF" author="Korschefsky, R." box="[666,916,1601,1623]" pageId="18" pageNumber="17" refId="ref19600" refString="Korschefsky, R. 1931. Coccinellidae I. Coleopterorum Catalogus. Part 118. W. Junk; Berlin. 224 p." type="book" year="1931">Korschefsky 1931: 219</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="47571097C74C95358ECFD44D27F5F9DF" author="Blackwelder, E." box="[927,1180,1601,1623]" page="449" pageId="18" pageNumber="17" year="1945">
<bibRefCitation id="A2674B77C74C95358ECFD44D27F5F9DF" author="Blackwelder, E." box="[927,1180,1601,1623]" pageId="18" pageNumber="17" pagination="343 - 550" refId="ref18961" refString="Blackwelder, E. 1945. Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Part 3. Bulletin of the United States National Museum 185: 343 - 550." type="journal article" year="1945">Blackwelder 1945: 449</bibRefCitation>
</treatmentCitation>
.
<emphasis id="F482EA94C74C953589F6D44D2633F9DF" bold="true" box="[1190,1370,1601,1622]" pageId="18" pageNumber="17">New synonym.</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74C95348D90D4632736FD5C" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17" type="description">
<paragraph id="C6493686C74C95348D90D4632001FD9C" blockId="18.[192,1441,1647,1929]" lastBlockId="19.[192,1440,221,533]" lastPageId="19" lastPageNumber="18" pageId="18" pageNumber="17">
<emphasis id="F482EA94C74C95358D90D46322D3F901" bold="true" box="[192,442,1647,1672]" pageId="18" pageNumber="17">Description. Male.</emphasis>
Length
<quantity id="010E9B63C74C95358F72D47C21E9F90E" box="[546,640,1648,1672]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="18" pageNumber="17" unit="mm" value="2.3">2.3 mm</quantity>
, width 2.0 mm. Dorsal surface shiny with faint trace of microsculpture on pronotum. Color black; head yellow; pronotum with lateral 1/4 yellow (
<figureCitation id="5ECD2A03C74C953589F3D49C2793F921" box="[1187,1274,1680,1704]" captionStart="Figures 86102" captionStartId="30.[192,287,1820,1842]" captionTargetBox="[209,1423,326,1699]" captionTargetId="figure@30.[192,1438,309,1740]" captionTargetPageId="30" captionText="Figures 86102. Cryptognatha spp. 8690) Cryptognatha yolandi. 86) Habitus. 87) Frons. 88) Phallobase ventral. 89) Phallobase lateral. 90) Penis. 9198) Cryptognatha pudibunda. 91) Habitus. 92) Frons. 93) Maxillary palpus. 94) Antenna. 95) Phallobase ventral. 96) Phallobase lateral. 97) Penis. 98) Spermathecal capsule. 99102) Cryptognatha batesi. 99) Habitus. 100) Frons. 101) Phallobase lateral. 102) Phallobase ventral." figureDoi="http://doi.org/10.5281/zenodo.3674867" httpUri="https://zenodo.org/record/3674867/files/figure.png" pageId="18" pageNumber="17">Fig. 91</figureCitation>
); venter with mouthparts and legs yellow; median portion of basal abdominal ventrite, ventrites 24 brown, ventrites 25 brownish yellow. Head without punctures; pronotal punctures small, separated by a diameter or less; elytral punctures larger than on pronotum, separated by less than a diameter to twice a diameter, becoming gradually larger toward lateral margin; prosternal punctures small, indistinct; mesosternal punctures larger than on prosternum, separated by less than a diameter, metasternal punctures large, separated by two or three times a diameter medially, entirely absent in lateral 1/3; basal abdominal ventrite with coarse punctures separated by about a diameter medially, lateral 1/3 of ventrite lacking punctation, ventrites 24 finely punctured, punctures separated by about a diameter, ventrite 5 with dense, fine punctation. Head with frons parallel-sided, medially depressed, 2.5 times as wide as eye, apex feebly arcuate medially, lateral 1/8 slightly retracted, angle acute (
<figureCitation id="5ECD2A03C74D95348EBBD3112728FEBC" box="[1003,1089,285,309]" captionStart="Figures 86102" captionStartId="30.[192,287,1820,1842]" captionTargetBox="[209,1423,326,1699]" captionTargetId="figure@30.[192,1438,309,1740]" captionTargetPageId="30" captionText="Figures 86102. Cryptognatha spp. 8690) Cryptognatha yolandi. 86) Habitus. 87) Frons. 88) Phallobase ventral. 89) Phallobase lateral. 90) Penis. 9198) Cryptognatha pudibunda. 91) Habitus. 92) Frons. 93) Maxillary palpus. 94) Antenna. 95) Phallobase ventral. 96) Phallobase lateral. 97) Penis. 98) Spermathecal capsule. 99102) Cryptognatha batesi. 99) Habitus. 100) Frons. 101) Phallobase lateral. 102) Phallobase ventral." figureDoi="http://doi.org/10.5281/zenodo.3674867" httpUri="https://zenodo.org/record/3674867/files/figure.png" pageId="19" pageNumber="18">Fig. 92</figureCitation>
); eye canthus long, about 3/4 width of eye. Prosternum with lateral carina on each side extended from apex of intercoxal process more than 2/3 distance to apex of prosternum. Epipleuron moderately descending externally. Postcoxal line on basal abdominal ventrite evenly curved, extended slightly more than 1/2 distance to rear margin of ventrite. Genitalia short, penis guide nearly as long as paramere, slender, evenly narrowed from base to abruptly rounded apex; paramere long, slender, curved to rounded apex (
<figureCitation id="5ECD2A03C74D95348929D3B1266BFE5C" box="[1145,1282,445,469]" captionStart="Figures 86102" captionStartId="30.[192,287,1820,1842]" captionTargetBox="[209,1423,326,1699]" captionTargetId="figure@30.[192,1438,309,1740]" captionTargetPageId="30" captionText="Figures 86102. Cryptognatha spp. 8690) Cryptognatha yolandi. 86) Habitus. 87) Frons. 88) Phallobase ventral. 89) Phallobase lateral. 90) Penis. 9198) Cryptognatha pudibunda. 91) Habitus. 92) Frons. 93) Maxillary palpus. 94) Antenna. 95) Phallobase ventral. 96) Phallobase lateral. 97) Penis. 98) Spermathecal capsule. 99102) Cryptognatha batesi. 99) Habitus. 100) Frons. 101) Phallobase lateral. 102) Phallobase ventral." figureDoi="http://doi.org/10.5281/zenodo.3674867" httpUri="https://zenodo.org/record/3674867/files/figure.png" pageId="19" pageNumber="18">Fig. 95, 96</figureCitation>
); penis long, apex slightly attenuate; penis capsule slender, elongate, with inner arm long, sinuate, apex somewhat truncate, outer arm long, apex slightly convex (
<figureCitation id="5ECD2A03C74D95348E53D3F12032FD9C" box="[771,859,509,533]" captionStart="Figures 86102" captionStartId="30.[192,287,1820,1842]" captionTargetBox="[209,1423,326,1699]" captionTargetId="figure@30.[192,1438,309,1740]" captionTargetPageId="30" captionText="Figures 86102. Cryptognatha spp. 8690) Cryptognatha yolandi. 86) Habitus. 87) Frons. 88) Phallobase ventral. 89) Phallobase lateral. 90) Penis. 9198) Cryptognatha pudibunda. 91) Habitus. 92) Frons. 93) Maxillary palpus. 94) Antenna. 95) Phallobase ventral. 96) Phallobase lateral. 97) Penis. 98) Spermathecal capsule. 99102) Cryptognatha batesi. 99) Habitus. 100) Frons. 101) Phallobase lateral. 102) Phallobase ventral." figureDoi="http://doi.org/10.5281/zenodo.3674867" httpUri="https://zenodo.org/record/3674867/files/figure.png" pageId="19" pageNumber="18">Fig. 97</figureCitation>
).
</paragraph>
<paragraph id="C6493686C74D95348D90D02126C9FD0C" blockId="19.[192,1440,557,645]" pageId="19" pageNumber="18">
<emphasis id="F482EA94C74D95348D90D0212245FDCC" bold="true" box="[192,300,557,581]" pageId="19" pageNumber="18">Female.</emphasis>
Similar to male except head black with dark brown clypeus, often with clypeus entirely black, pronotum entirely black, head feebly impressed medially; spermathecal capsule short, ramus widened, cornu widened from base to abruptly rounded apex, with long, sclerotized portion of sperm duct (
<figureCitation id="5ECD2A03C74D9534886ED06126FAFD0C" box="[1342,1427,621,645]" captionStart="Figures 86102" captionStartId="30.[192,287,1820,1842]" captionTargetBox="[209,1423,326,1699]" captionTargetId="figure@30.[192,1438,309,1740]" captionTargetPageId="30" captionText="Figures 86102. Cryptognatha spp. 8690) Cryptognatha yolandi. 86) Habitus. 87) Frons. 88) Phallobase ventral. 89) Phallobase lateral. 90) Penis. 9198) Cryptognatha pudibunda. 91) Habitus. 92) Frons. 93) Maxillary palpus. 94) Antenna. 95) Phallobase ventral. 96) Phallobase lateral. 97) Penis. 98) Spermathecal capsule. 99102) Cryptognatha batesi. 99) Habitus. 100) Frons. 101) Phallobase lateral. 102) Phallobase ventral." figureDoi="http://doi.org/10.5281/zenodo.3674867" httpUri="https://zenodo.org/record/3674867/files/figure.png" pageId="19" pageNumber="18">Fig. 98</figureCitation>
).
</paragraph>
<paragraph id="C6493686C74D95348D90D0912736FD5C" blockId="19.[192,1440,669,725]" pageId="19" pageNumber="18">
<emphasis id="F482EA94C74D95348D90D0912222FD3C" bold="true" box="[192,331,669,693]" pageId="19" pageNumber="18">Variation.</emphasis>
Length
<quantity id="010E9B63C74D95348CE1D0912133FD3C" box="[433,602,669,693]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="19" pageNumber="18" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.2 to 2.5 mm</quantity>
, width
<quantity id="010E9B63C74D95348FE3D0912034FD3C" box="[691,861,669,693]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="19" pageNumber="18" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.8 to 2.1 mm</quantity>
. Penis guide of male genitalia slightly variable, apex nearly smoothly rounded or with indication of “shoulders” before apex.
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74D95348D90D0E12216FCAC" pageId="19" pageNumber="18" type="distribution">
<paragraph id="C6493686C74D95348D90D0E12216FCAC" blockId="19.[192,1439,749,805]" pageId="19" pageNumber="18">
<emphasis id="F482EA94C74D95348D90D0E12210FC8C" bold="true" box="[192,377,749,773]" pageId="19" pageNumber="18">
<typeStatus id="194D8824C74D95348D90D0E1226DFC8C" box="[192,260,749,773]" pageId="19" pageNumber="18">Type</typeStatus>
locality.
</emphasis>
<location id="C329605DC74D95348C2ED0E122F2FC8C" LSID="urn:lsid:plazi:treatment:4E5F8790C74C95348D90D7D52166FAEC:C329605DC74D95348C2ED0E122F2FC8C" box="[382,411,749,773]" country="Brazil" name="Of" pageId="19" pageNumber="18">Of</location>
<emphasis id="F482EA94C74D95348CF1D0E12139FC8C" box="[417,592,749,773]" italics="true" pageId="19" pageNumber="18">
<taxonomicName id="01F64D05C74D95348CF1D0E12122FC8C" box="[417,587,749,773]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="pudibunda">C. pudibunda</taxonomicName>
,
</emphasis>
<collectingCountry id="BEE17616C74D95348F30D0E121C2FC8C" box="[608,683,749,773]" name="Brazil" pageId="19" pageNumber="18">Brésil</collectingCountry>
”; of
<taxonomicName id="01F64D05C74D95348F8BD0E12047FC8C" box="[731,814,749,773]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="fryii">
<emphasis id="F482EA94C74D95348F8BD0E12047FC8C" box="[731,814,749,773]" italics="true" pageId="19" pageNumber="18">C. fryii</emphasis>
</taxonomicName>
,
<collectingRegion id="0432F864C74D95348E6BD0E1208EFC8C" box="[827,999,749,773]" country="Brazil" name="Rio de Janeiro" pageId="19" pageNumber="18">Rio de Janeiro</collectingRegion>
(
<collectingCountry id="BEE17616C74D95348EA5D0E1272AFC8C" box="[1013,1091,749,773]" name="Brazil" pageId="19" pageNumber="18">Brazil</collectingCountry>
); of
<taxonomicName id="01F64D05C74D95348922D0E12212FCAC" authority=", Santo Amara, Santos" authorityName="Santo Amara, Santos" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="F482EA94C74D95348922D0E12782FC8C" box="[1138,1259,749,773]" italics="true" pageId="19" pageNumber="18">C. bryanti</emphasis>
, Santo Amara, Santos (
<collectingCountry id="BEE17616C74D95348C73D101221BFCAC" box="[291,370,781,805]" name="Brazil" pageId="19" pageNumber="18">Brazil</collectingCountry>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74D95348D90D131210BFC5C" pageId="19" pageNumber="18" type="materials_examined">
<paragraph id="C6493686C74D95348D90D131210BFC5C" blockId="19.[192,1440,829,981]" pageId="19" pageNumber="18">
<emphasis id="F482EA94C74D95348D90D13122E2FCDC" bold="true" box="[192,395,829,853]" pageId="19" pageNumber="18">Type material.</emphasis>
<materialsCitation id="769E3CDBC74D95348CC3D1312640FCDC" ID-GBIF-Occurrence="2569343932" box="[403,1321,829,854]" country="Brazil" location="Of" pageId="19" pageNumber="18" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
Of
<taxonomicName id="01F64D05C74D95348CEAD13121A2FCDC" authority=", Dejean" authorityName="Dejean" box="[442,715,829,853]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="pudibunda">
<emphasis id="F482EA94C74D95348CEAD1312105FCDC" box="[442,620,829,853]" italics="true" pageId="19" pageNumber="18">C. pudibunda,</emphasis>
Dejean
</taxonomicName>
collection, Musée des confluences, Lyon,
<collectingCountry id="BEE17616C74D9534899CD132264DFCDC" box="[1228,1316,830,853]" name="France" pageId="19" pageNumber="18">France</collectingCountry>
;
</materialsCitation>
<materialsCitation id="769E3CDBC74D95348862D131211EFCFC" ID-GBIF-Occurrence="2569343927" country="Brazil" location="Of" pageId="19" pageNumber="18" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="lectotype">
<typeStatus id="194D8824C74D95348862D13126C9FCDC" box="[1330,1440,829,853]" pageId="19" pageNumber="18" type="lectotype">lectotype</typeStatus>
here designated; of
<taxonomicName id="01F64D05C74D95348CE2D1512161FCFC" box="[434,520,861,885]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="fryii">
<emphasis id="F482EA94C74D95348CE2D15122ACFCFD" box="[434,453,861,884]" italics="true" pageId="19" pageNumber="18">C</emphasis>
.
<emphasis id="F482EA94C74D95348C85D1512161FCFC" box="[469,520,861,885]" italics="true" pageId="19" pageNumber="18">fryii</emphasis>
</taxonomicName>
, BMNH,
</materialsCitation>
<materialsCitation id="769E3CDBC74D95348FD0D15127BDFCFC" ID-GBIF-Occurrence="2569343930" box="[640,1236,861,885]" country="Brazil" location="Of" pageId="19" pageNumber="18" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="lectotype">
<typeStatus id="194D8824C74D95348FD0D1512187FCFC" box="[640,750,861,885]" pageId="19" pageNumber="18" type="lectotype">lectotype</typeStatus>
here designated; of
<taxonomicName id="01F64D05C74D95348EB8D151270DFCFC" box="[1000,1124,861,885]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="F482EA94C74D95348EB8D1512092FCFD" box="[1000,1019,861,884]" italics="true" pageId="19" pageNumber="18">C</emphasis>
.
<emphasis id="F482EA94C74D9534895BD151270DFCFC" box="[1035,1124,861,885]" italics="true" pageId="19" pageNumber="18">bryanti</emphasis>
</taxonomicName>
, BMNH.
</materialsCitation>
<materialsCitation id="769E3CDBC74D9534898CD15127C7FC1C" ID-GBIF-Occurrence="2569343926" country="Brazil" location="Of" pageId="19" pageNumber="18" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="lectotype">
The
<typeStatus id="194D8824C74D95348844D15126EBFCFC" box="[1300,1410,861,885]" pageId="19" pageNumber="18" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="01F64D05C74D95348D90D1712203FC1C" box="[192,362,893,917]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="pudibunda">
<emphasis id="F482EA94C74D95348D90D1712203FC1C" box="[192,362,893,917]" italics="true" pageId="19" pageNumber="18">C. pudibunda</emphasis>
</taxonomicName>
is labeled “Type/15139/TYPE/Fry RioJan/Type/Fry Coll. 1905-100.”
</materialsCitation>
<materialsCitation id="769E3CDBC74D953489E8D171210BFC5C" ID-GBIF-Occurrence="2569343940" country="Brazil" location="Of" pageId="19" pageNumber="18" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="lectotype">
The
<typeStatus id="194D8824C74D953489A0D1712637FC1C" box="[1264,1374,893,917]" pageId="19" pageNumber="18" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="01F64D05C74D953488D5D171239AFC3C" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="fryii">
<emphasis id="F482EA94C74D953488D5D171239AFC3C" italics="true" pageId="19" pageNumber="18">C. fryii</emphasis>
</taxonomicName>
the is labeled “Type/Ilha Santo Amaro,nr. Santos,
<collectingCountry id="BEE17616C74D95348E28D19120AFFC3C" box="[888,966,925,949]" name="Brazil" pageId="19" pageNumber="18">Brazil</collectingCountry>
, G. E. Bryant
<date id="B2481046C74D953489D1D19127B0FC3C" box="[1153,1241,925,949]" pageId="19" pageNumber="18" value="1912-04">iv.1912</date>
/G. Bryant Coll. 1919-147/
<taxonomicName id="01F64D05C74D95348C67D1B1213EFC5C" authority="Brethes." authorityName="Brethes." box="[311,599,957,981]" class="Insecta" family="Nymphalidae" genus="Oeneis" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="F482EA94C74D95348C67D1B12280FC5C" box="[311,489,957,981]" italics="true" pageId="19" pageNumber="18">Oeneis bryanti</emphasis>
Brethes.
</taxonomicName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74D95348D90D1E1208FFB8C" box="[192,998,1005,1030]" pageId="19" pageNumber="18" type="distribution">
<paragraph id="C6493686C74D95348D90D1E1208FFB8C" blockId="19.[192,998,1005,1030]" box="[192,998,1005,1030]" pageId="19" pageNumber="18">
<emphasis id="F482EA94C74D95348D90D1E12146FB8C" bold="true" box="[192,559,1005,1029]" pageId="19" pageNumber="18">Geographical distribution.</emphasis>
<collectingCountry id="BEE17616C74D95348F66D1E121EDFB8C" box="[566,644,1005,1029]" name="Brazil" pageId="19" pageNumber="18">Brazil</collectingCountry>
,
<collectingCountry id="BEE17616C74D95348FC0D1E22060FB8F" box="[656,777,1006,1030]" name="Paraguay" pageId="19" pageNumber="18">Paraguay</collectingCountry>
,
<collectingCountry id="BEE17616C74D95348E45D1E2203AFB8C" box="[789,851,1006,1029]" name="Peru" pageId="19" pageNumber="18">Peru</collectingCountry>
,
<collectingCountry id="BEE17616C74D95348E0FD1E1208BFB8C" box="[863,994,1005,1029]" name="Venezuela" pageId="19" pageNumber="18">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74D95348D90D6112230FBDC" pageId="19" pageNumber="18" type="materials_examined">
<paragraph id="C6493686C74D95348D90D6112230FBDC" blockId="19.[192,1440,1053,1109]" pageId="19" pageNumber="18">
<materialsCitation id="769E3CDBC74D95348D90D611223CFBDC" ID-GBIF-Occurrence="2569343943" country="Brazil" location="Of" pageId="19" pageNumber="18" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<emphasis id="F482EA94C74D95348D90D611228EFBBC" bold="true" box="[192,487,1053,1077]" pageId="19" pageNumber="18">Specimens examined.</emphasis>
42. Widely distributed in
<collectingCountry id="BEE17616C74D95348E4CD611200EFBBC" box="[796,871,1053,1077]" name="Brazil" pageId="19" pageNumber="18">Brazil</collectingCountry>
with specimens examined from other countries listed above
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8EEC650DC74D95348D90D6612166FAEC" pageId="19" pageNumber="18" type="discussion">
<paragraph id="C6493686C74D95348D90D66127F2FA8F" blockId="19.[192,1441,1133,1381]" pageId="19" pageNumber="18">
<emphasis id="F482EA94C74D95348D90D661222AFB0C" bold="true" box="[192,323,1133,1157]" pageId="19" pageNumber="18">Remarks.</emphasis>
Of the dorsally black species
<taxonomicName id="01F64D05C74D95348FF4D6612025FB0C" box="[676,844,1133,1157]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="pudibunda">
<emphasis id="F482EA94C74D95348FF4D66121DEFB0D" box="[676,695,1133,1156]" italics="true" pageId="19" pageNumber="18">C</emphasis>
.
<emphasis id="F482EA94C74D95348F95D6612025FB0C" box="[709,844,1133,1157]" italics="true" pageId="19" pageNumber="18">pudibunda</emphasis>
</taxonomicName>
is most widely distributed, particularly in
<collectingCountry id="BEE17616C74D9534881ED66126F4FB0C" box="[1358,1437,1133,1157]" name="Brazil" pageId="19" pageNumber="18">Brazil</collectingCountry>
. Male genitalia must be examined to identify this and all other taxa with entirely black elytra.
<bibRefCitation id="A2674B77C74D95348869D6812265FB4C" author="Mulsant, M. E." pageId="19" pageNumber="18" pagination="1 - 1104" refId="ref19658" refString="Mulsant, M. E. 1850. Species de coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, Lyon 2: 1 - 1104." type="journal article" year="1850">Mulsant (1850)</bibRefCitation>
stated his type material was in the Dejean collection, now housed in Musée des confluences, Lyon. Harold Labrique provided us with images of the type specimen and that specimen labeled “
<collectingCountry id="BEE17616C74D95348860D6C12611FB6C" box="[1328,1400,1229,1253]" name="Brazil" pageId="19" pageNumber="18">Bresil</collectingCountry>
” is here designated as the
<typeStatus id="194D8824C74D95348C8CD6E12124FA8C" box="[476,589,1261,1285]" pageId="19" pageNumber="18" type="lectotype">lectotype</typeStatus>
, because
<bibRefCitation id="A2674B77C74D95348F90D6E12012FA8C" author="Mulsant, M. E." box="[704,891,1261,1285]" pageId="19" pageNumber="18" pagination="1 - 1104" refId="ref19658" refString="Mulsant, M. E. 1850. Species de coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles, Lyon 2: 1 - 1104." type="journal article" year="1850">Mulsant (1850)</bibRefCitation>
specified it as the type.
</paragraph>
<paragraph id="C6493686C74D95348DB8D7012166FAEC" blockId="19.[192,1441,1133,1381]" pageId="19" pageNumber="18">
Specimens identified as
<taxonomicName id="01F64D05C74D95348F43D70121D4FAAC" box="[531,701,1293,1317]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="pudibunda">
<emphasis id="F482EA94C74D95348F43D701214FFAAD" box="[531,550,1293,1316]" italics="true" pageId="19" pageNumber="18">C</emphasis>
.
<emphasis id="F482EA94C74D95348F66D70121D4FAAC" box="[566,701,1293,1317]" italics="true" pageId="19" pageNumber="18">pudibunda</emphasis>
</taxonomicName>
may actually represent more than one species, however, no convincing evidence of this was forthcoming. Both
<taxonomicName id="01F64D05C74D95348E6FD72120FFFACC" box="[831,918,1325,1349]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="fryii">
<emphasis id="F482EA94C74D95348E6FD721203BFACD" box="[831,850,1325,1348]" italics="true" pageId="19" pageNumber="18">C</emphasis>
.
<emphasis id="F482EA94C74D95348E33D72120FFFACC" box="[867,918,1325,1349]" italics="true" pageId="19" pageNumber="18">fryii</emphasis>
</taxonomicName>
(
<typeStatus id="194D8824C74D95348EF9D7212770FACC" box="[937,1049,1325,1349]" pageId="19" pageNumber="18" type="lectotype">lectotype</typeStatus>
) and
<taxonomicName id="01F64D05C74D95348933D7212788FACC" box="[1123,1249,1325,1349]" class="Insecta" family="Nymphalidae" genus="Oeneis" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="F482EA94C74D95348933D721271FFACD" box="[1123,1142,1325,1348]" italics="true" pageId="19" pageNumber="18">C</emphasis>
.
<emphasis id="F482EA94C74D953489D8D7212788FACC" box="[1160,1249,1325,1349]" italics="true" pageId="19" pageNumber="18">bryanti</emphasis>
</taxonomicName>
are considered synonyms of
<taxonomicName id="01F64D05C74D95348C0FD7412161FAEC" box="[351,520,1357,1381]" class="Insecta" family="Coccinellidae" genus="Cryptognatha" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="pudibunda">
<emphasis id="F482EA94C74D95348C0FD741221BFAED" box="[351,370,1357,1380]" italics="true" pageId="19" pageNumber="18">C</emphasis>
.
<emphasis id="F482EA94C74D95348CD1D7412161FAEC" box="[385,520,1357,1381]" italics="true" pageId="19" pageNumber="18">pudibunda</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>