treatments-xml/data/3A/6C/87/3A6C87E6FF8A4C61FF0DD92D039700B1.xml
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<document id="C58C727AB78EBBA36DE8EFE9ECD8C9FB" ID-DOI="10.21248/contrib.entomol.71.1.029-085" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742999" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638270695310" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2021" docId="3A6C87E6FF8A4C61FF0DD92D039700B1" docLanguage="en" docName="ContribEntomol.71.1.29-85.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 71 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Lithocharodes fuscula SHARP 1885" docType="treatment" docVersion="3" lastPageNumber="47" masterDocId="C655FF9EFF984C73FFEEDB7E00260230" masterDocTitle="The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)" masterLastPageNumber="85" masterPageNumber="29" pageNumber="47" updateTime="1699437350205" updateUser="plazi">
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<mods:title id="55311723C743E26808FF2936C2102CD3">The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)</mods:title>
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<mods:namePart id="F568C866A3ED90C5041FDC33E4307FB8">Irmler, Ulrich</mods:namePart>
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<mods:title id="0F9093D14EAFCBEADBB4FD91C76375C4">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="2BA42AA245C89D2ADBC5DBF95DC1EE1E">2021</mods:date>
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<subSubSection id="FADF657BFF8A4C61FF0DD92D026C005E" box="[227,586,595,622]" pageId="18" pageNumber="47" type="nomenclature">
<paragraph id="B27A36F0FF8A4C61FF0DD92D026C005E" blockId="18.[227,586,595,655]" box="[227,586,595,622]" pageId="18" pageNumber="47">
<heading id="E932819CFF8A4C61FF0DD92D026C005E" box="[227,586,595,622]" centered="true" fontSize="12" level="2" pageId="18" pageNumber="47" reason="2">
<taxonomicName id="75C54D73FF8A4C61FF0DD92D026C005E" ID-CoL="3VKFS" authority="SHARP, 1885" authorityName="SHARP" authorityYear="1885" box="[227,586,595,622]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="fuscula">
<emphasis id="80B1EAE2FF8A4C61FF0DD92D01E5005E" box="[227,451,595,622]" italics="true" pageId="18" pageNumber="47">Lithocharodes fuscula</emphasis>
<bibRefCitation id="D6544B01FF8A4C61FE24D92D026C005E" author="SHARP, D." box="[458,586,595,622]" pageId="18" pageNumber="47" pagination="393 - 536" refId="ref41459" refString="SHARP, D. 1885: Fam. Staphylinidae. - In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2): pp. 393 - 536. - London, Taylor &amp; Francis." type="journal article" year="1885">SHARP, 1885</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="FADF657BFF8A4C61FF0DD90501B500BF" box="[227,403,635,655]" pageId="18" pageNumber="47" type="description">
<paragraph id="B27A36F0FF8A4C61FF0DD90501B500BF" blockId="18.[227,586,595,655]" box="[227,403,635,655]" pageId="18" pageNumber="47">
<figureCitation id="2AFE2A75FF8A4C61FF0DD905013F00BF" box="[227,281,635,655]" captionStart="Fig" captionStartId="45.[151,184,2040,2062]" captionTargetBox="[167,1375,247,2020]" captionTargetId="figure-105@45.[156,1388,232,2032]" captionTargetPageId="45" captionText="Fig. 3742: L.-??-group: Lithocharodes compacta (37), L. elegans (38), L. elongata (39), L. fuscula (40), L. rufula (41); aedeagus (a)," figureDoi="http://doi.org/10.5281/zenodo.5743019" httpUri="https://zenodo.org/record/5743019/files/figure.png" pageId="18" pageNumber="47">Figs 40</figureCitation>
a-d, 43 M, 47 D
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF8A4C61FF53D9D5021700F0" box="[189,561,683,705]" pageId="18" pageNumber="47" type="reference_group">
<paragraph id="B27A36F0FF8A4C61FF53D9D5021700F0" blockId="18.[189,561,683,705]" box="[189,561,683,705]" pageId="18" pageNumber="47">
<treatmentCitationGroup id="92D511DEFF8A4C61FF53D9D5021700F0" box="[189,561,683,705]" pageId="18" pageNumber="47">
<treatmentCitation id="336410E1FF8A4C61FF53D9D5021700F0" author="SHARP, D." box="[189,561,683,705]" page="494" pageId="18" pageNumber="47" year="1885">
<taxonomicName id="75C54D73FF8A4C61FF53D9D5021700F0" ID-CoL="3VKFS" authority="SHARP, 1885: 494" authorityName="SHARP" authorityPageNumber="494" authorityYear="1885" box="[189,561,683,705]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="fuscula">
<emphasis id="80B1EAE2FF8A4C61FF53D9D501AE00F0" box="[189,392,683,704]" italics="true" pageId="18" pageNumber="47">Lithocharodes fuscula</emphasis>
<bibRefCitation id="D6544B01FF8A4C61FE63D9D5021700F0" author="SHARP, D." box="[397,561,683,705]" pageId="18" pageNumber="47" pagination="393 - 536" refId="ref41459" refString="SHARP, D. 1885: Fam. Staphylinidae. - In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2): pp. 393 - 536. - London, Taylor &amp; Francis." type="journal article" year="1885">SHARP, 1885: 494</bibRefCitation>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF8A4C61FF53D99701D90151" pageId="18" pageNumber="47" type="materials_examined">
<paragraph id="B27A36F0FF8A4C61FF53D99701D90151" blockId="18.[189,794,745,865]" pageId="18" pageNumber="47">
<emphasis id="80B1EAE2FF8A4C61FF53D99701E00131" bold="true" box="[189,454,745,769]" pageId="18" pageNumber="47">Type material examined</emphasis>
:
<materialsCitation id="02AD3CADFF8A4C61FE3DD99701DA0151" collectionCode="BMNH" collectorName="San Geronimo &amp; Champion" country="Guatemala" location="Guatemala City" pageId="18" pageNumber="47" specimenCount="2" stateProvince="Guatemala" typeStatus="lectotype">
male,
<typeStatus id="6D7E8852FF8A4C61FDFAD99702530131" box="[532,629,745,769]" pageId="18" pageNumber="47" type="lectotype">lectotype</typeStatus>
(present designation):
<collectingCountry id="CAD27660FF8A4C61FECAD87701BC0111" box="[292,410,777,801]" name="Guatemala" pageId="18" pageNumber="47">Guatemala</collectingCountry>
,
<collectorName id="1F305326FF8A4C61FE5FD877027C0111" box="[433,602,777,801]" pageId="18" pageNumber="47">San Geronimo</collectorName>
, leg.
<collectorName id="1F305326FF8A4C61FD46D877033F0111" box="[680,793,777,801]" pageId="18" pageNumber="47">Champion</collectorName>
(
<collectionCode id="D4D4AE35FF8A4C61FF2AD857013F0171" box="[196,281,809,833]" pageId="18" pageNumber="47">BMNH</collectionCode>
); female,
<typeStatus id="6D7E8852FF8A4C61FE68D85401FD0172" box="[390,475,810,834]" pageId="18" pageNumber="47" type="syntype">syntype</typeStatus>
:
<collectingCountry id="CAD27660FF8A4C61FE07D85702790171" box="[489,607,809,833]" name="Guatemala" pageId="18" pageNumber="47">Guatemala</collectingCountry>
, near
<collectingRegion id="7001F812FF8A4C61FD48D85700CC0151" country="Guatemala" name="Guatemala" pageId="18" pageNumber="47">Guatemala City</collectingRegion>
, leg.
<collectorName id="1F305326FF8A4C61FEF1D83701B60151" box="[287,400,841,865]" pageId="18" pageNumber="47">Champion</collectorName>
(
<collectionCode id="D4D4AE35FF8A4C61FE70D83701D20151" box="[414,500,841,865]" pageId="18" pageNumber="47">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF8A4C61FF53D8F7023706B1" pageId="18" pageNumber="47" type="diagnosis">
<paragraph id="B27A36F0FF8A4C61FF53D8F7023706B1" blockId="18.[189,794,905,1153]" pageId="18" pageNumber="47">
<emphasis id="80B1EAE2FF8A4C61FF53D8F7010C0191" bold="true" box="[189,298,905,929]" pageId="18" pageNumber="47">Diagnosis</emphasis>
: Among the
<specimenCount id="A4C3FD79FF8A4C61FE2DD8F402740192" box="[451,594,906,930]" pageId="18" pageNumber="47" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
from the BMNH studied,
<specimenCount id="A4C3FD79FF8A4C61FEF8D8D4015301F1" box="[278,373,937,961]" pageId="18" pageNumber="47" type="male">one male</specimenCount>
was found. This specimen was dissected and labelled as
<typeStatus id="6D7E8852FF8A4C61FEB0D8B701E401D1" box="[350,450,969,993]" pageId="18" pageNumber="47" type="lectotype">lectotype</typeStatus>
.
<taxonomicName id="75C54D73FF8A4C61FE24D8B7028D01D1" authorityName="SHARP" authorityYear="1885" box="[458,683,969,993]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="fuscula">
<emphasis id="80B1EAE2FF8A4C61FE24D8B7028D01D1" box="[458,683,969,993]" italics="true" pageId="18" pageNumber="47">Lithocharodes fuscula</emphasis>
</taxonomicName>
resembles a group of dark unicoloured species, smaller than
<quantity id="753D9B15FF8A4C61FD35D897033C0631" box="[731,794,1001,1025]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="47" unit="mm" value="4.0">4 mm</quantity>
in size:
<taxonomicName id="75C54D73FF8A4C61FEFDDF7401B30611" authorityName="Irmler" authorityYear="2021" box="[275,405,1033,1057]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="verhaaghi">
<emphasis id="80B1EAE2FF8A4C61FEFDDF7401B30611" box="[275,405,1033,1057]" italics="true" pageId="18" pageNumber="47">L. verhaaghi</emphasis>
</taxonomicName>
and
<taxonomicName id="75C54D73FF8A4C61FE21DF74021A0611" authorityName="Irmler" authorityYear="2021" box="[463,572,1033,1057]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="silvicola">
<emphasis id="80B1EAE2FF8A4C61FE21DF74021A0611" box="[463,572,1033,1057]" italics="true" pageId="18" pageNumber="47">L. silvicola</emphasis>
</taxonomicName>
. The species of this group are extremely similar and can hardly be separated without studying the aedeagus. Male endophallus of
<taxonomicName id="75C54D73FF8A4C61FF53DF14010506B1" authorityName="SHARP" authorityYear="1885" box="[189,291,1129,1153]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="fuscula">
<emphasis id="80B1EAE2FF8A4C61FF53DF14010506B1" box="[189,291,1129,1153]" italics="true" pageId="18" pageNumber="47">L. fuscula</emphasis>
</taxonomicName>
has no spiral torsions.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF8A4C61FF53DFD7039700B1" pageId="18" pageNumber="47" type="description">
<paragraph id="B27A36F0FF8A4C61FF53DFD701810731" blockId="18.[189,795,1193,2081]" pageId="18" pageNumber="47">
<emphasis id="80B1EAE2FF8A4C61FF53DFD7011906F1" bold="true" box="[189,319,1193,1217]" pageId="18" pageNumber="47">Description</emphasis>
: Length:
<quantity id="753D9B15FF8A4C61FE2ADFD7023106F1" box="[452,535,1193,1217]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="18" pageNumber="47" unit="mm" value="3.7">3.7 mm</quantity>
. Colouration: Totally brown; elytra slightly darker brown; legs and antennae slightly lighter brown.
</paragraph>
<paragraph id="B27A36F0FF8A4C61FF53DE77039700B1" blockId="18.[189,795,1193,2081]" lastBlockId="18.[831,1437,233,641]" pageId="18" pageNumber="47">
Head:
<quantity id="753D9B15FF8A4C61FEEDDE7701470711" box="[259,353,1289,1313]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="47" unit="mm" value="0.7">0.70 mm</quantity>
long,
<quantity id="753D9B15FF8A4C61FE4DDE7702240711" box="[419,514,1289,1313]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="18" pageNumber="47" unit="mm" value="0.58">0.58 mm</quantity>
wide; eyes slightly prominent; postocular sides twice as long as eyes; posterior angles widely rounded; posterior margin slightly emarginate; interantennal furrows distinct; reaching posterior third of eyes; setiferous punctation moderately dense and deep; on average, interstices between punctures as wide as diameter of punctures; interstices on anterior vertex slightly shorter, on posterior vertex slightly wider; wide midline impunctate; surface with extremely weak microsculpture; shiny. Antennae with first antennomere slightly shorter than half-length of head; second and third antennomere conical; second slightly longer than wide; third as long as apical width; subsequent antennomeres wider than long and increasing in width; fourth antennomere twice as wide as long; tenth antennomere 2.5 times as wide as long; all antennomeres pubescent. Pronotum:
<quantity id="753D9B15FF8A4C61FF53DC77013A0511" box="[189,284,1801,1825]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="18" pageNumber="47" unit="mm" value="0.75">0.75 mm</quantity>
long,
<quantity id="753D9B15FF8A4C61FE8BDC7701E50511" box="[357,451,1801,1825]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="18" pageNumber="47" unit="mm" value="0.53">0.53 mm</quantity>
wide; widest in anterior third; anteriad, conically narrowed toward neck; posteriad, distinctly narrowed in middle third; in posterior third nearly parallel; posterior angles sub-rectangular; posterior margin nearly straight; setiferous punctation deeper than on head but partly sparser; wide midline impunctate; surface without microsculpture; glossy. Elytra:
<quantity id="753D9B15FF8A4C61FF53DC97013A0A31" box="[189,284,2025,2049]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="47" unit="mm" value="0.7">0.70 mm</quantity>
long,
<quantity id="753D9B15FF8A4C61FE9DDC9701F70A31" box="[371,465,2025,2049]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="18" pageNumber="47" unit="mm" value="0.62">0.62 mm</quantity>
wide; humeral angles subrectangular; sides slightly divergent posteriad; posterior angles sub-rectangular; posterior margin nearly straight; setiferous punctation deep and dense; on average, interstices between punctures half as wide as diameter of punctures; surface with extremely weak microsculpture; shiny. Abdomen with finer setiferous punctation than on fore-body; punctation at base of segments denser than at apex; surface with weak irregular microsculpture; shiny; sternite VII of male evenly narrowed toward obtuse apex; posterior margin of tergite VII of male straight. Aedeagus oval with anterior angles sub-rectangular; endophallus in anterior half weakly sclerotised; posterior half distinctly sclerotised; partly covered by minute teeth and partly by long teeth.
</paragraph>
</subSubSection>
</treatment>
</document>