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<document id="CC8D94401ACEE539487874AD19DBBFC9" ID-CLB-Dataset="26390" ID-DOI="http://dx.doi.org/10.3897/evolsyst.3.35737" ID-GBIF-Dataset="5b2f91e0-bcd0-4faa-80cd-e2ed2561e8b2" ID-Pensoft-Pub="2535-0730-1-85" ID-ZooBank="AF37CE1C27174410803DFCBD6C41258A" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="2535-0730-1-85" ModsDocOrigin="Evolutionary Systematics 3 (1)" ModsDocTitle="The genus Syrrhoe (Crustacea, Amphipoda, Synopiidae) from the North Atlantic" checkinTime="1561619019251" checkinUser="pensoft" docAuthor="Fuchs, Luisa, Coleman, Charles Oliver &amp; Loerz, Anne-Nina" docDate="2019" docId="3A483667B3B475931CB4B2D114059CA8" docLanguage="en" docName="EvolutSyst 3(1): 85-108" docOrigin="Evolutionary Systematics 3 (1)" docSource="http://dx.doi.org/10.3897/evolsyst.3.35737" docTitle="Syrrhoe anneheleneae Fuchs, Coleman &amp; Loerz, 2019, sp. nov." docType="treatment" docUuid="6B43618F-571D-4A45-9EE8-4F8903A2DD61" docUuidSource="ZooBank" docVersion="6" lastPageNumber="85" masterDocId="FF9A474BFFAEFF87FFACFFF3FFC2FFB3" masterDocTitle="The genus Syrrhoe (Crustacea, Amphipoda, Synopiidae) from the North Atlantic" masterLastPageNumber="108" masterPageNumber="85" pageNumber="85" updateTime="1701379700084" updateUser="ExternalLinkService">
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<mods:title id="EBE6B074A0C6560ECF16C30B99720A0B">The genus Syrrhoe (Crustacea, Amphipoda, Synopiidae) from the North Atlantic</mods:title>
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<mods:namePart id="D53691A7FEE6E6976B21740D349B61DE">Fuchs, Luisa</mods:namePart>
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<mods:namePart id="9545817B234395CD810C1F36954F2F0B">Coleman, Charles Oliver</mods:namePart>
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<mods:namePart id="76BED410AFD0D39E51F2479A8874A039">Loerz, Anne-Nina</mods:namePart>
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<mods:title id="9D4D1626D4AE1DECB03F3C99248833C5">Evolutionary Systematics</mods:title>
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<mods:date id="4D1A6EBCE4DA5C2ED6131AEDD359E356">2019</mods:date>
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<treatment id="3A483667B3B475931CB4B2D114059CA8" ID-GBIF-Taxon="158407475" LSID="urn:lsid:zoobank.org:act:6B43618F-571D-4A45-9EE8-4F8903A2DD61" httpUri="http://treatment.plazi.org/id/3A483667B3B475931CB4B2D114059CA8" lastPageNumber="85" pageId="0" pageNumber="85">
<subSubSection id="0A683B659D7454CCFC05A6EA7BBD500E" pageId="0" pageNumber="85" type="nomenclature">
<paragraph id="D90B8E45B77C0D3483611C44780DE268" pageId="0" pageNumber="85">
<taxonomicName id="33695F7A58954E938418B7EB74EBC543" ID-CoL="7ZL6J" LSID="http://zoobank.org/6B43618F-571D-4A45-9EE8-4F8903A2DD61" class="Malacostraca" family="Synopiidae" genus="Syrrhoe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrrhoe anneheleneae" order="Amphipoda" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="anneheleneae">Syrrhoe anneheleneae</taxonomicName>
<taxonomicNameLabel id="DD5461AFE1050D034569972897B1108A" pageId="0" pageNumber="85">sp. nov.</taxonomicNameLabel>
Figs 6, 7, 8, 9, 10, 11
</paragraph>
</subSubSection>
<subSubSection id="1E6F47DA08B49A5498C5C1DB1869D947" pageId="0" pageNumber="85" type="materials_examined">
<paragraph id="E7F0A40EFD0DD957AF1E01CBC4134586" pageId="0" pageNumber="85">Material examined.</paragraph>
<paragraph id="A0091A36121003A30EACD80354D028B1" pageId="0" pageNumber="85">
Type material: 1 specimen, presumably subadult male, holotype, 9 mm, UNIS 2009 St. 12, AB 321,
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79°1
<normalizedToken id="83321A18FD01F59687B13733372588AC" originalValue="2”">'2&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="91807258A9C42611E09704456409A3E6" direction="east" orientation="longitude" precision="15" value="11.766111">
11°45
<normalizedToken id="856BFA24B05940FAE38EA5EB627D1067" originalValue="58”">'58&quot;</normalizedToken>
E
</geoCoordinate>
, 232 m, 1.9.2009, RP sledge, ZMBN 112487; 5 paratypes, 2 females, 7.5 and 8 mm; 3 without sex determination, 7-8.7 mm, same as type locality, ZMBN 127920.
</paragraph>
<paragraph id="1B9B35755151BB19EACA0FFBBC40244E" pageId="0" pageNumber="85">
Additional material: 1 female, 2 specimens, without sex determination, Deutsche Expedition in das
<normalizedToken id="2D75EAEF7BED1D5F1B69618E56C0A77C" originalValue="Nördliche">Noerdliche</normalizedToken>
Eismeer, 1898, St. 34,
<normalizedToken id="BFFEC80AC7836EC0FA935D1EBBAC912D" originalValue="König">Koenig</normalizedToken>
Karls Land (Svalbard), leg.
<normalizedToken id="BE7FB5349C6CC432A924EBBF01D1FA7A" originalValue="Römer">Roemer</normalizedToken>
&amp; Schaudinn, ZMB 19834; 1 male, Deutsche Expedition in das
<normalizedToken id="203E90E0070D6ADF284671AC3CF57901" originalValue="Nördliche">Noerdliche</normalizedToken>
Eismeer, 1898, St. 47, Thymen
<normalizedToken id="6F361F8A07FCE9FC9C0B696ACF47C241" originalValue="Straße">Strasse</normalizedToken>
(Freeman Strait),
<geoCoordinate id="1F2EA65ECDC12CC481260658AC2EC687" direction="north" orientation="latitude" precision="925" value="78.21667">78°13'N</geoCoordinate>
<geoCoordinate id="EEB3F34715864CA189F4F94B3767FD48" direction="east" orientation="longitude" precision="925" value="21.316668">21°19'E</geoCoordinate>
, leg.
<normalizedToken id="6081C351EF6C55983E114D52793CBFA3" originalValue="Römer">Roemer</normalizedToken>
&amp; Schaudinn, ZMB 19835; 3 specimens, without sex determination, UNIS 2015 St. 825, Svalbard,
<geoCoordinate id="78D17C232B73C35E950C30EA69409F7F" direction="north" orientation="latitude" precision="15" value="78.148605">
78°8
<normalizedToken id="B577AE4989DDB9EA1A343125D43E88F5" originalValue="55”">'55&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="30B1DD7974CEBCD1F162D23F412BF079" direction="east" orientation="longitude" precision="15" value="13.125556">
13°7
<normalizedToken id="D47EE200C9B5B5546803789D3556FFDA" originalValue="32”">'32&quot;</normalizedToken>
E
</geoCoordinate>
, 243 m, 13.5.2015, RP sledge, ZMBN 112488; 3 specimens, without sex determination, UNIS 2009 St. 69, Svalbard,
<geoCoordinate id="E123A97F5F0E9CDB37AA7542778033FC" direction="north" orientation="latitude" precision="15" value="81.02945">
81°1
<normalizedToken id="1CCD7FA4B62C61233DD2065090E03C4C" originalValue="46”">'46&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="0B56AB7EC2DCDEBA0D3FCE2A24C720A8" direction="east" orientation="longitude" precision="15" value="19.155277">
19°9
<normalizedToken id="1CC8609111A046F28F4537E73F5E0AE6" originalValue="19”">'19&quot;</normalizedToken>
E
</geoCoordinate>
, 174 m, 1.9.2009, RP sledge, ZMBN 112490; 1 specimen, without sex determination, UNIS 2009 St. 12 AB 321, Svalbard,
<geoCoordinate id="B01F035355E1AABF9DFC8F600C49F13E" direction="north" orientation="latitude" precision="15" value="79.01723">
79°1
<normalizedToken id="6549757CF1A9503E6DB75C3172CC3087" originalValue="2”">'2&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="2CBC6A1864508363962DF51FE2D995CE" direction="east" orientation="longitude" precision="15" value="11.766111">
11°45
<normalizedToken id="F9627FBAB7E70E6771A523E5E1302EE4" originalValue="58”">'58&quot;</normalizedToken>
E
</geoCoordinate>
, 232 m, 1.9.2009, RP sledge, ZMBN 112486; 1 adult male, 10 mm, UNIS 2009, Adventfjord,
<geoCoordinate id="141B30F24B41AACAA576ABAC2AEDC5C2" direction="north" orientation="latitude" precision="15" value="78.23">
78°13
<normalizedToken id="83B4F5B624993C129621FE017D778AFC" originalValue="48”">'48&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="2DAECAFE6CBC219FD3C3013D7EF8F3D4" direction="east" orientation="longitude" precision="15" value="15.56">
15°33
<normalizedToken id="BA3923837BFB98EA0F3F90D4578ED66D" originalValue="36”">'36&quot;</normalizedToken>
E
</geoCoordinate>
, 6.9.2009, RP sledge, ZMBN 112426; 1 specimen, without sex determination, UNIS 2015 St. 825,
<geoCoordinate id="02B4F367969E605B313D947C630364E5" direction="north" orientation="latitude" precision="15" value="78.148605">
78°8
<normalizedToken id="999DD892F76180CAE494C0A6FCC5FCB1" originalValue="55”">'55&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="A318C09C15312BF1AA3FF0B2F68EEAB4" direction="east" orientation="longitude" precision="15" value="13.125556">
13°7
<normalizedToken id="6115338FFB1BDF4F58ADF504F172953C" originalValue="32”">'32&quot;</normalizedToken>
E
</geoCoordinate>
, 243 m, 13.5.2015, RP sledge, ZMBN 112447; 3 males, 1 female, 2 specimens without sex determination, UNIS 2009 St. 50(2), Svalbard,
<geoCoordinate id="8AC335AFEA9326D50854C8961FB80384" direction="north" orientation="latitude" precision="15" value="80.10861">
80° 6
<normalizedToken id="77C15D63CA1D60C2668E8DB71CDB3B7F" originalValue="31”">'31&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="D07D22D5E3D8FF726936C0BC08A45F9A" direction="east" orientation="longitude" precision="15" value="22.141388">
22°8
<normalizedToken id="A48B4457B0640171DE3BB85640612DD1" originalValue="29”">'29&quot;</normalizedToken>
E
</geoCoordinate>
, 216 m, 1.9.2009, RP sledge, ZMBN 112489; 12 males, 10 females, 211 without sex determination, UNIS 2015 St. 781, Kapp Mitra,
<geoCoordinate id="074F660584F95CFBB4D0FAB008270DA3" direction="north" orientation="latitude" precision="15" value="79.11806">
79°7
<normalizedToken id="418AA859DD2D5C68E9371780D2F0B436" originalValue="5”">'5&quot;</normalizedToken>
N
</geoCoordinate>
,
<geoCoordinate id="7B64DCC7047FEBAADB49D1AC4EB7CF06" direction="east" orientation="longitude" precision="925" value="11.083333">11°5'E</geoCoordinate>
, 110 m, 8.5.2015, RP sledge, ZMBN 112451; 1 specimen ZMH-K 56185 (GenBank MK972331), 1 specimen ZMH-K 56186 (GenBank MK972330), 1 specimen ZMH-K 56187 (GenBank MK972327), East Iceland, Norwegian Sea, IceAGE st. 1219,
<geoCoordinate id="B37B981BCFF82956E40B79F7F0A3550A" direction="north" orientation="latitude" precision="9" value="66.289">66°17.34'N</geoCoordinate>
,
<geoCoordinate id="5FCD238B522CD37B16D773E7268967F0" direction="west" orientation="longitude" precision="92" value="-12.346666">12°20.8'W</geoCoordinate>
, 579 m, 22.09.2011.
</paragraph>
<paragraph id="527AE797D26121E068CFE00FE9687C40" pageId="0" pageNumber="85">Length ranges: females 7-9 mm, males 6-8 mm, without sex determination 4-9 mm.</paragraph>
</subSubSection>
<subSubSection id="7D94C87688C6A956269A6A92CE2E07B7" pageId="0" pageNumber="85" type="description">
<paragraph id="1AF3DFD9575BE36A9DE36FC2B0F18DD4" pageId="0" pageNumber="85">Description</paragraph>
<paragraph id="C64FDF36AFD908A60C9D19728C152E2B" pageId="0" pageNumber="85">based on: male holotype, 9 mm, ZMBN 112487.</paragraph>
<paragraph id="153B20CEA46B6B9C8237C42EAA825454" pageId="0" pageNumber="85">
Head. Rostrum reaching half the length of article 1 of antenna 1; lateral cephalic lobe present, pointed; eyes present, large, oval, on dorsal part of head, unpigmented (Fig. 6a, b). Antenna 1 (Fig. 6e) article 1 with 2 teeth of different length, 1 large distally curved tooth and 1 shorter and straight tooth; article 2 without distomedial tooth, slightly shorter than article 1; article 3 somewhat shorter than article 2; accessory flagellum 3-articulate, surpassing 3rd flagellar article, 3rd article of accessory flagellum less than a quarter length of 1st accessory flagellar article; flagellum shorter than pereon, 14 articles. Antenna 2 (Fig. 7a) longer than antenna 1, not surpassing pereon; article 5 longest; flagellum 13-articulate, shorter than peduncle. Labrum (Fig. 6c) apically rounded, tapering distally. Mandible (Fig. 7b, c, d) incisor multidentate; left lacinia mobilis wide and multidentate; accessory setal row with 5 weakly denticulate setae; molar small, columnar, weakly triturative; palp article 1 smooth, shorter than half the length of article 2, article 2 with slender setae; article 3 half the length of article 1, with long 6 slender setae apically. Lower lip (Fig. 6d) with wide gap; inner lobes present, outer lobes apically setose; hypopharyngeal lobes wide. Maxilla 1 (Fig. 7e) inner plate with 8 plumose setae; outer plate with 11 apical robust setae; bifurcate setae present; palp article 2 outer margin smooth, with apical and medial setation. Maxilla 2 (Fig. 7f) inner plate width about 2
<normalizedToken id="4518C467AE7D232E0FF179C12B9F9807" originalValue="×">x</normalizedToken>
outer plate, with row of facial setae. Maxilliped (Fig. 8
<normalizedToken id="42C5258751EC52C17920D05BAF280919" originalValue="ae">a-e</normalizedToken>
) inner plate reaching 1st palp article, with 10 strong setae; outer plate distally and medially with simple robust setae; palp article 2 is 1.5
<normalizedToken id="8E8F63B0A165CDF37198A14B37C753FE" originalValue="×">x</normalizedToken>
the length of article 3; palp article 3 slightly inflated; article 4 weakly curved, with long unguis.
</paragraph>
<caption id="B2A2AC2D97278F13BAB1F1282EEBE0A5" pageId="0" pageNumber="85">
<paragraph id="8BC091CA0F58619737BFCFC735543ED9" pageId="0" pageNumber="85">
Figure 6.
<taxonomicName id="77A6AECAAD89BF7020ED8FE3A7DEAB98" class="Malacostraca" family="Synopiidae" genus="Syrrhoe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrrhoe anneheleneae" order="Amphipoda" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="anneheleneae">Syrrhoe anneheleneae</taxonomicName>
sp. nov., male holotype, 9 mm, ZMBN 112487. a) habitus; b) head; c) upper lip (labrum); d) lower lip; e) antenna 1. Scale bars: 1 mm (a); 100
<normalizedToken id="F5AD88CC4BE831E9619452F8B685CA26" originalValue="µm">µm</normalizedToken>
(
<normalizedToken id="F3E8A0CB8238123F5BAF7A96C7541833" originalValue="ce">c-e</normalizedToken>
).
</paragraph>
</caption>
<caption id="EE2F0AC3D291E5C0538DD24E95951E4F" pageId="0" pageNumber="85">
<paragraph id="F80AC60513789CB08F2834937B2C8D5F" pageId="0" pageNumber="85">
Figure 7.
<taxonomicName id="9043243DECD7DC26B221262E5DE449C8" class="Malacostraca" family="Synopiidae" genus="Syrrhoe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrrhoe anneheleneae" order="Amphipoda" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="anneheleneae">Syrrhoe anneheleneae</taxonomicName>
sp. nov., male holotype, 9 mm, ZMBN 112487. a) antenna 2; b) mandible; c) mandibular palp; d) mandible; e) maxilla 1; f) maxilla 2, setulation shown on one exemplary seta. Scale bars: 100
<normalizedToken id="2F3541A06C3ACE75BF9BC5185DB15971" originalValue="µm">µm</normalizedToken>
(
<normalizedToken id="43CD4158F319D3B939D92EB41A884039" originalValue="af">a-f</normalizedToken>
).
</paragraph>
</caption>
<caption id="9444423DE68D06EECB782AC7C8746B93" pageId="0" pageNumber="85">
<paragraph id="F1315D8087DF4150156DB9B2BF071316" pageId="0" pageNumber="85">
Figure 8.
<taxonomicName id="8837980929C24ED910CD9A8DE04652F0" class="Malacostraca" family="Synopiidae" genus="Syrrhoe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrrhoe anneheleneae" order="Amphipoda" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="anneheleneae">Syrrhoe anneheleneae</taxonomicName>
sp. nov., male holotype, 9 mm, ZMBN 112487. a) outlines of maxilliped; b) inner plate of maxilliped; c) outer plate of maxilliped; d) maxillipedal palp; e) dactylus of maxillipedal palp; f) chela of gnathopod 1. Scale bars: 100
<normalizedToken id="09C86B5E99292C1A697F7A1E245F5430" originalValue="µm">µm</normalizedToken>
(
<normalizedToken id="BD4545DE58967C1BB4F718589ABF8773" originalValue="af">a-f</normalizedToken>
).
</paragraph>
</caption>
<paragraph id="70537804AEBB38E73F6F660EEDD6AB0A" pageId="0" pageNumber="85">
Pereon. Pereonite 1-6 dorsally, dorsolaterally and laterally smooth (Fig. 6a). Pereonite 7 posterior margin mid-dorsally serrate, with short point; posterolateral corner angular and not produced. Gnathopod 1 (Fig. 9a) coxa not tapering distally, apex strongly directed anteriorly; basis longer than carpus; carpus subrectangular, with smooth setae along the posterior margin; propodus palm transverse, defined by 1 serrate robust seta. Gnathopod 2 (Fig. 9b) coxa distally tapering; basis twice the length of carpus; carpus serrate setae along the posterior margin at ventral part; propodus palm nearly oblique, defined by 1 serrate robust setae; dactylus of gnathopod 2 well developed, unguis similar length to dactylus. Pereopod 3 (Fig. 10a) coxa anterodistal lobe present, obliquely produced, posterodistal lobe present, truncate, half the depth of the coxa, posterodistal margin with minute setae; basis width equal to breadth of ischium; ischium subquadrate, 1/4 of merus length; dactylus weakly curved. Pereopod 3-4 coxa pelagont. Pereopod 4 (Fig. 10b) coxa much shorter than coxa 3, lobate in shape, slightly curved posteriorly; ischium subquadrate, 1/4 of merus length; dactylus weakly curved. Pereopod 5-7 (Fig. 11
<normalizedToken id="C38B4A182739CA2CAA105B335E14E104" originalValue="ac">a-c</normalizedToken>
) basis anterodistal corner weakly produced into a recurved hook, posterior margin strongly serrate. Pereopod 5-6 basis subovoid, expanded, posterodistal lobe weakly developed, not extending past ischium. Pereopod 7 basis ovate, posterodistal lobe well developed, rounded, extending below ischium.
</paragraph>
<caption id="302A411812DA2889847FC56BF9F683A5" pageId="0" pageNumber="85">
<paragraph id="E6522374E1BE7D8449C05B2E5C917EAB" pageId="0" pageNumber="85">
Figure 9.
<taxonomicName id="4CB71E080A44FCBDC7CD7980DB1971F2" class="Malacostraca" family="Synopiidae" genus="Syrrhoe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrrhoe anneheleneae" order="Amphipoda" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="anneheleneae">Syrrhoe anneheleneae</taxonomicName>
sp. nov., male holotype, 9 mm, ZMBN 112487. a) gnathopod 1; b) merus to dactylus of gnathopod 1; c) ischium to dactylus of gnathopod 2; d) gnathopod 2. Scale bars: 100
<normalizedToken id="B0BDEABA1C445E2126DBF38AAA4F21F9" originalValue="µm">µm</normalizedToken>
(a, b, d).
</paragraph>
</caption>
<caption id="5AAAF79D606B34A9E632951BE3A00912" pageId="0" pageNumber="85">
<paragraph id="E6A4C00F5DEDCA593BB519FF147F2B64" pageId="0" pageNumber="85">
Figure 10.
<taxonomicName id="6AD232B377B797EB848C9D3FED45B4EC" class="Malacostraca" family="Synopiidae" genus="Syrrhoe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrrhoe anneheleneae" order="Amphipoda" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="anneheleneae">Syrrhoe anneheleneae</taxonomicName>
sp. nov., male holotype, 9 mm, ZMBN 112487. a) pereopod 3; b) pereopod 4; c) uropod 1; d) uropod 2; e) uropod 3, f) telson. Scale bars: 100
<normalizedToken id="F89BF59CD5DA7935AE43B5A460D65F5C" originalValue="µm">µm</normalizedToken>
(
<normalizedToken id="C52D12BCC4DCAB50BB9E4DAB1D6A49A2" originalValue="af">a-f</normalizedToken>
).
</paragraph>
</caption>
<caption id="744DBC7E494EA189D1F59C74E46C8C55" pageId="0" pageNumber="85">
<paragraph id="CBF8A8DAEBC0E1C5EBD5D0E9803F2718" pageId="0" pageNumber="85">
Figure 11.
<taxonomicName id="DD06841C8BDCDD817C8D165FB8BEF4DE" class="Malacostraca" family="Synopiidae" genus="Syrrhoe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrrhoe anneheleneae" order="Amphipoda" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="anneheleneae">Syrrhoe anneheleneae</taxonomicName>
sp. nov., male holotype, 9 mm, ZMBN 112487. a) pereopod 5; b) pereopod 6; c) pereopod 7; d) pleopod 1. Scale bars: 100
<normalizedToken id="AAA511D8D14A4FBF124926C834AA38D7" originalValue="µm">µm</normalizedToken>
(
<normalizedToken id="D8E307113DC117912275780E0E110A01" originalValue="ad">a-d</normalizedToken>
).
</paragraph>
</caption>
<paragraph id="9367BCBC53EE1B663EAB0F963A1F90E7" pageId="0" pageNumber="85">Pleon. Pleonite 1-3 (Fig. 6a) dorsally round, without carination, posterior margin serrate and mid-dorsally with short point. Epimeron 1 posterior margin smooth, posterodistal corner weakly pointed. Epimeron 2 posterior margin smooth, posterodistal corner pointed. Epimeron 3 posterior margin serrate, posteroventral corner subquadrate.</paragraph>
<paragraph id="A5FCFAABB06F77FDBF6829D2041DD41D" pageId="0" pageNumber="85">Pleopod (Fig. 11d) rami similar length, 13 and 15 articles, all articles with setulated setae.</paragraph>
<paragraph id="2BEF5D1B6A7A290B80AECAEBA551B320" pageId="0" pageNumber="85">Urosomite 1 (Fig. 6a) mid-dorsal posterior and lateral margin serrate. Urosomite 2 (Fig. 6a) mid-dorsal posterior margin smooth. Urosomite 3 (Fig. 6a) mid-dorsal posterior margin smooth.</paragraph>
<paragraph id="47996CD60DD2B52173B9C16D4F0F8803" pageId="0" pageNumber="85">
Uropod 1 (Fig. 10c) peduncle with long distal process, the process reaching half the length of outer ramus; inner ramus same length as peduncle; apical robust setae on tip of both rami present. Uropod 2 (Fig. 10d) apical robust setae on tip of outer ramus present, outer ramus half the length and half the with of inner ramus. Uropod 3 (Fig. 10d) length not exceeding uropods 1-2, outer ramus 2.5
<normalizedToken id="8EB6985941C881A1CF35E2370535F932" originalValue="×">x</normalizedToken>
length of peduncle. Telson (Fig. 10f) almost as long as uropod 3 peduncle, 3
<normalizedToken id="5360B609C9CE5BE3201538645224D2BC" originalValue="×">x</normalizedToken>
as long as wide; with no setae along the lateral margin, with apical slender setae; lobes abutting, deeply cleft, more than 75%.
</paragraph>
</subSubSection>
<subSubSection id="D11DD381695C83490CF425B0B02B46D1" pageId="0" pageNumber="85" type="distribution">
<paragraph id="4E777723542C693EFDA734D5B3CD105A" pageId="0" pageNumber="85">Distribution.</paragraph>
<paragraph id="4102D0D91C628310EFA52766DE8E56E9" pageId="0" pageNumber="85">North Atlantic, 110-579 m.</paragraph>
</subSubSection>
<subSubSection id="31918FA798C07527CC00349B62CED5DB" pageId="0" pageNumber="85" type="etymology">
<paragraph id="2400D873160A79AD55C03349455ADD19" pageId="0" pageNumber="85">Etymology.</paragraph>
<paragraph id="5B4EB41E992BDA5D21A1896C2B4EB175" pageId="0" pageNumber="85">Named for Dr. Anne Helene Tandberg, great amphipodologist and a friend of the authors, who organized the wonderful workshop in Bergen, where the new species was discovered.</paragraph>
</subSubSection>
<subSubSection id="973D7CC72F17B7B233CD5B59AA3F02F2" pageId="0" pageNumber="85" type="remarks">
<paragraph id="34715B2D43793E79CCA72B9833A7B9E7" pageId="0" pageNumber="85">Remarks.</paragraph>
<paragraph id="D81625383722ACD22D8E5C57F4939C0B" pageId="0" pageNumber="85">
<taxonomicName id="900C2EA4AF37C7865EA74391429C8A67" class="Malacostraca" family="Synopiidae" genus="Syrrhoe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrrhoe anneheleneae" order="Amphipoda" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="anneheleneae">Syrrhoe anneheleneae</taxonomicName>
sp. nov. is rather similar to
<taxonomicName id="44CDAA149F35853BA485B8F75AED3440" class="Malacostraca" family="Synopiidae" genus="Syrrhoe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrrhoe crenulata" order="Amphipoda" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="crenulata">Syrrhoe crenulata</taxonomicName>
. Both species have a narrow straight to excavate space between the dorsal serration and the serration of the epimeron 3.
<taxonomicName id="A1EF98829F7013AA5A133B87A0DD868D" class="Malacostraca" family="Synopiidae" genus="Syrrhoe" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Syrrhoe anneheleneae" order="Amphipoda" pageId="0" pageNumber="85" phylum="Arthropoda" rank="species" species="anneheleneae">Syrrhoe anneheleneae</taxonomicName>
sp. nov. has a serrate margin of urosomite 1 (vs. smooth in
<taxonomicName id="CF26BEEF5ABF2DEE571841BBA609A400" lsidName="S. crenulata" pageId="0" pageNumber="85" rank="species" species="crenulata">S. crenulata</taxonomicName>
) and in ethanol white eyes (vs. dark pigmented in
<taxonomicName id="38F3337AC2C42035E74305C871FAE1D6" lsidName="S. crenulata" pageId="0" pageNumber="85" rank="species" species="crenulata">S. crenulata</taxonomicName>
). A serration on urosomite 1 also occurs in
<taxonomicName id="AF110C6A4B64CE490861F8E053C3DFF3" lsidName="S. papyracea" pageId="0" pageNumber="85" rank="species" species="papyracea">S. papyracea</taxonomicName>
Stebbing, 1888, but this species, only known from the type locality Culebra Island, Porto Rico (
<bibRefCitation id="5AC283BF185F72E618EE377946412E4F" author="Shoemaker, CR" journalOrPublisher="Scientific Survey of Porto Rico and the Virgin Islands (New York Academy of Sciences)" pageId="0" pageNumber="85" pagination="229 - 253" title="The Amphipoda of Porto Rico and the Virgin Islands." volume="15" year="1935">Shoemaker 1935</bibRefCitation>
), differs in several respect from
<taxonomicName id="2F1C196F83FEAAC9C1D600A6CC138DA9" lsidName="S. anneheleneae" pageId="0" pageNumber="85" rank="species" species="anneheleneae">S. anneheleneae</taxonomicName>
sp. nov.: head anteriorly produced; no eyes present; epimeral plates without posteromarginal serration and the palm of both gnathopods has an additional stout spine aside the serrate spine.
</paragraph>
</subSubSection>
</treatment>
</document>