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<document id="2AF9986ADEB7DB5442F7F98BFC0FC8E6" ID-CLB-Dataset="80832" ID-DOI="10.5281/zenodo.6707142" ID-GBIF-Dataset="ab66b2b7-9544-4411-bf61-5bc3651d7bca" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6707142" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1656002532852" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="03F06D13FF1820D10D8A17370CE8F577" docLanguage="en" docName="hbmw_7_Cricetidae_0204.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Kunsia tomentosus" docType="treatment" docVersion="12" lastPageNumber="461" masterDocId="FFC9156BFFAE20670D37145C0837FFDB" masterDocTitle="Cricetidae" masterLastPageNumber="535" masterPageNumber="204" pageNumber="461" updateTime="1718799336328" updateUser="carolina">
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<mods:namePart id="F5037837150A1F7EB3B2B597A3BC4B5E">Don E. Wilson</mods:namePart>
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<mods:namePart id="DC69DEB146CB967738E47D5FF4EA973A">Thomas E. Lacher, Jr</mods:namePart>
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<treatment id="03F06D13FF1820D10D8A17370CE8F577" ID-DOI="http://doi.org/10.5281/zenodo.6727374" ID-GBIF-Taxon="196221528" ID-Zenodo-Dep="6727374" LSID="urn:lsid:plazi:treatment:03F06D13FF1820D10D8A17370CE8F577" httpUri="http://treatment.plazi.org/id/03F06D13FF1820D10D8A17370CE8F577" lastPageNumber="461" pageId="182" pageNumber="461">
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<heading id="D0AE6B69FF1820D10D8A17370938FC42" box="[189,271,875,921]" pageId="182" pageNumber="461">
<figureCitation id="1362C080FF1820D10D8A17370938FC42" box="[189,271,875,921]" captionStart="Plate 24: Cricetidae" captionStartId="173.[132,162,3175,3200]" captionTargetBox="[11,2744,13,3647]" captionTargetPageId="172" captionText="487. Common Marsh Rice Rat (Oryzomys palustris), 488. Texas Marsh Rice Rat (Oryzomys texensus), 489. Couess Marsh Rice Rat (Oryzomys couesi), 490. White-bellied Marsh Rice Rat (Oryzomys albiventer), 491. Nicaraguan Marsh Rice Rat (Oryzomys dimidiatus), 492. Gorgass Marsh Rice Rat (Oryzomys gorgasi), 493. Santiago Galapagos Mouse (Nesoryzomys swarthi), 494. Small Fernandina Galapagos Mouse (Nesoryzomys fernandinae), 495. Large Fernandina Galapagos Mouse (Nesoryzomys narboroughi), 496. Galapagos Rice Rat (Aegialomys galapagoensis), 497. Yellowish Rice Rat (Aegialomys xanthaeolus), 498. Baron's Rice Rat (Aegialomys baroni), 499. Ica Rice Rat (Aegialomys ica), 500. Alfaros Water Rat (Sigmodontomys alfari), 501. Harris's Rice Water Rat (Tanyuromys aphrastus), 502. Black-and-Yellow Rice Rat (Melanomys chrysomelas), 503. Cinnamon-rufous Rice Rat (Melanomys idoneus), 504. Colombian Rice Rat (Melanomys columbianus), 505. Dusky Rice Rat (Melanomys caliginosus), 506. Robust Dark Rice Rat (Melanomys robustulus), 507. Zunigas Dark Rice Rat (Melanomys zunigae), 508. Intermediate Lesser Grass Mouse (Microakodontomys transitorius), 509. Lagoa Santa Pink-lipped Mouse (Bibimys labiosus), 510. Chacoan Pink-lipped Mouse (Bibimys chacoensis), 511. Torress Pink-lipped Mouse (Bibimys torresi), 512. Brazilian Swamp Rat (Scapteromys meridionalis), 513. Argentinean Swamp Rat (Scapteromys aquaticus), 514. Uruguay Swamp Rat (Scapteromys tumidus), 515. Cerrado Giant Rat (Gyldenstolpia planaltensis), 516. Fossorial Giant Rat (Gyldenstolpia fronto), 517. Woolly Giant Rat (Kunsia tomentosus), 518. Andean Rat (Lenoxus apicalis), 519. Atlantic Forest Burrowing Mouse (Blarinomys breviceps), 520. Gray-bellied Brucie (Brucepattersonius griserufescens), 521. Short-tailed Brucie (Brucepattersonius soricinus), 522. Iherings Brucie (Brucepattersonius theringi)" figureDoi="http://doi.org/10.5281/zenodo.6708535" httpUri="https://zenodo.org/record/6708535/files/figure.png" pageId="182" pageNumber="461">517.</figureCitation>
</heading>
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<subSubSection id="C3438F8EFF1820D10C2917370AB6FC42" box="[286,641,875,921]" pageId="182" pageNumber="461" type="vernacular_names">
<paragraph id="8BE6DC05FF1820D10C2917370AB6FC42" blockId="182.[187,1048,875,999]" box="[286,641,875,921]" pageId="182" pageNumber="461">
<heading id="D0AE6B69FF1820D10C2917370AB6FC42" box="[286,641,875,921]" pageId="182" pageNumber="461">
<vernacularName id="055AAC2BFF1820D10C2917370AB6FC42" ID-CoL="3RFPB" baseAuthorityName="Lichtenstein" baseAuthorityYear="1830" box="[286,641,875,921]" class="Mammalia" family="Cricetidae" genus="Kunsia" kingdom="Animalia" language="eng" order="Rodentia" pageId="182" pageNumber="461" phylum="Chordata" rank="species" species="tomentosus">Woolly Giant Rat</vernacularName>
</heading>
</paragraph>
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<subSubSection id="C3438F8EFF1820D10F8417370C2FFC42" box="[691,1048,875,921]" pageId="182" pageNumber="461" type="nomenclature">
<paragraph id="8BE6DC05FF1820D10F8417370C2FFC42" blockId="182.[187,1048,875,999]" box="[691,1048,875,921]" pageId="182" pageNumber="461">
<heading id="D0AE6B69FF1820D10F8417370C2FFC42" box="[691,1048,875,921]" pageId="182" pageNumber="461">
<taxonomicName id="4C59A786FF1820D10F8417370C2FFC42" ID-CoL="3RFPB" baseAuthorityName="Lichtenstein" baseAuthorityYear="1830" box="[691,1048,875,921]" class="Mammalia" family="Cricetidae" genus="Kunsia" kingdom="Animalia" order="Rodentia" pageId="182" pageNumber="461" phylum="Chordata" rank="species" species="tomentosus">
<emphasis id="B92D0017FF1820D10F8417370C2FFC42" box="[691,1048,875,921]" italics="true" pageId="182" pageNumber="461">Kunsia tomentosus</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3438F8EFF1820D10D8B17F70A72FC3C" pageId="182" pageNumber="461" type="vernacular_names">
<paragraph id="8BE6DC05FF1820D10D8B17F70C36FC1B" blockId="182.[187,1048,875,999]" box="[188,1025,939,960]" pageId="182" pageNumber="461">
<heading id="D0AE6B69FF1820D10D8B17F70C36FC1B" box="[188,1025,939,960]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10D8B17F70930FC1B" bold="true" box="[188,263,939,960]" pageId="182" pageNumber="461">French:</emphasis>
<vernacularName id="055AAC2BFF1820D10C3817F7099FFC1B" ID-CoL="3RFPB" baseAuthorityName="Lichtenstein" baseAuthorityYear="1830" box="[271,424,939,960]" class="Mammalia" family="Cricetidae" genus="Kunsia" kingdom="Animalia" language="fra" order="Rodentia" pageId="182" pageNumber="461" phylum="Chordata" rank="species" species="tomentosus">Akodon laineux</vernacularName>
/
<emphasis id="B92D0017FF1820D10C8A17F70A20FC1B" bold="true" box="[445,535,939,960]" pageId="182" pageNumber="461">German:</emphasis>
<vernacularName id="055AAC2BFF1820D10F2917F70AF4FC1B" ID-CoL="3RFPB" baseAuthorityName="Lichtenstein" baseAuthorityYear="1830" box="[542,707,939,960]" class="Mammalia" family="Cricetidae" genus="Kunsia" kingdom="Animalia" language="deu" order="Rodentia" pageId="182" pageNumber="461" phylum="Chordata" rank="species" species="tomentosus">Woll-Riesenratte</vernacularName>
/
<emphasis id="B92D0017FF1820D10FEF17F70B04FC1B" bold="true" box="[728,819,939,960]" pageId="182" pageNumber="461">Spanish:</emphasis>
<vernacularName id="055AAC2BFF1820D10E0A17F70C36FC1B" ID-CoL="3RFPB" baseAuthorityName="Lichtenstein" baseAuthorityYear="1830" box="[829,1025,939,960]" class="Mammalia" family="Cricetidae" genus="Kunsia" kingdom="Animalia" language="esp" order="Rodentia" pageId="182" pageNumber="461" phylum="Chordata" rank="species" species="tomentosus">Rata gigante lanuda</vernacularName>
</heading>
</paragraph>
<paragraph id="8BE6DC05FF1820D10D8C178E0A72FC3C" blockId="182.[187,1048,875,999]" box="[187,581,978,999]" pageId="182" pageNumber="461">
<heading id="D0AE6B69FF1820D10D8C178E0A72FC3C" box="[187,581,978,999]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10D8C178E0985FC3C" bold="true" box="[187,434,978,999]" pageId="182" pageNumber="461">Other common names:</emphasis>
<vernacularName id="055AAC2BFF1820D10C8E178E0A72FC3C" ID-CoL="3RFPB" baseAuthorityName="Lichtenstein" baseAuthorityYear="1830" box="[441,581,978,999]" class="Mammalia" family="Cricetidae" genus="Kunsia" kingdom="Animalia" language="eng" order="Rodentia" pageId="182" pageNumber="461" phylum="Chordata" rank="species" species="tomentosus">Woolly Kunsia</vernacularName>
</heading>
</paragraph>
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<subSubSection id="C3438F8EFF1820D10E2810400C90FB21" pageId="182" pageNumber="461" type="reference_group">
<paragraph id="8BE6DC05FF1820D10E2810400C90FB21" blockId="182.[798,1392,1052,1471]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10E2810400B8CFBEE" bold="true" box="[799,955,1052,1077]" pageId="182" pageNumber="461">Taxonomy.</emphasis>
<taxonomicName id="4C59A786FF1820D10EE610400B53FB86" authority="Lichtenstein, 1830" authorityName="Lichtenstein" authorityYear="1830" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="182" pageNumber="461" phylum="Chordata" rank="species" species="tomentosus">Mus tomentosus Lichtenstein, 1830</taxonomicName>
,
<materialsCitation id="3B31D658FF1820D10E4A101C0C90FB21" ID-GBIF-Occurrence="3818814769" pageId="182" pageNumber="461">
“waldigen Gegenden am
<collectingCountry id="F34E9C95FF1820D109C0101C0D47FB86" box="[1271,1392,1088,1117]" name="Uruguay" pageId="182" pageNumber="461">Uruguay</collectingCountry>
[= wooded areas of
<collectingCountry id="F34E9C95FF1820D1090C103F0C8CFB5F" box="[1083,1211,1123,1156]" name="Uruguay" pageId="182" pageNumber="461">Uruguay</collectingCountry>
].” Restricted by P. Hershkovitz in 1966 to the Rio
<collectingCountry id="F34E9C95FF1820D1080710D60B57FB08" name="Uruguay" pageId="182" pageNumber="461">Uruguay</collectingCountry>
in south-eastern
<collectingCountry id="F34E9C95FF1820D1094910EA0CFBFB08" box="[1150,1228,1206,1235]" name="Brazil" pageId="182" pageNumber="461">Brazil</collectingCountry>
(probably state of
<collectingRegion id="499D12E7FF1820D10EB910810CAEFB21" box="[910,1177,1245,1274]" country="Brazil" name="Rio Grande do Sul" pageId="182" pageNumber="461">Rio Grande do Sul</collectingRegion>
).
</materialsCitation>
</paragraph>
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<subSubSection id="C3438F8EFF1820D10E2911590C5CFA43" pageId="182" pageNumber="461" type="discussion">
<paragraph id="8BE6DC05FF1820D10E2911590C5CFA43" blockId="182.[798,1392,1052,1471]" pageId="182" pageNumber="461">Validity of subspecies principalis named by P. W. Lund in 1840 for speciments near [Lagoa Santa, Minas Gerais State, Brazil, is questioned. Monotypic.</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF1820D10E1711FA0ACCFA3C" pageId="182" pageNumber="461" type="distribution">
<caption id="DF268C8DFF1820D10E1711FA0ACCFA3C" ID-DOI="http://doi.org/10.5281/zenodo.6707224" ID-Zenodo-Dep="6707224" httpUri="https://zenodo.org/record/6707224/files/figure.png" inLine="true" pageId="182" pageNumber="461" targetBox="[184,776,1051,1466]" targetPageId="182">
<paragraph id="8BE6DC05FF1820D10E1711FA0ACCFA3C" blockId="182.[798,1392,1052,1471]" lastBlockId="182.[183,1392,1478,2732]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10E1711FA0BE7FA64" bold="true" box="[800,976,1446,1471]" pageId="182" pageNumber="461">Distribution.</emphasis>
N &amp; E Bolivia and C Brazil; it probably also occurs in extreme SE Peru.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C3438F8EFF1820D10D8B11B1094DF843" pageId="182" pageNumber="461" type="description">
<paragraph id="8BE6DC05FF1820D10D8B11B1094DF843" blockId="182.[183,1392,1478,2732]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10D8B11B10980F9D5" bold="true" box="[188,439,1517,1550]" pageId="182" pageNumber="461">Descriptive notes.</emphasis>
Head—body 185-287 mm, tail 147-196 mm, ear 28-36 mm, hindfoot 43-54 mm; weight 241-630 g. The Woolly Giant Rat is the largest extant Sigmodontinae. It has dense and coarse fur, proportionally short limbs with large and powerful feet and claws, and moderately short tail (¢.65% of head-body length). Rhinarium is large and naked and highlighted by ring of whitish hair. Ears are small and round (c.13% of head-body length), and pinnae are covered with short hairs that match color of dorsal pelage. Tail is unicolored, scarcely covered with short rigid hairs, and has large subrectangular scales. General pelage is dark gray or dark brown on dorsal surface and grayish on gular and ventral surfaces. Eight mammae are arranged in pectoral, postaxial, abdominal, and inguinal pairs. Chromosomal complement is 2n = 44, FN = 42.
</paragraph>
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<subSubSection id="C3438F8EFF1820D10D8B13FB0B22F7D5" pageId="182" pageNumber="461" type="biology_ecology">
<paragraph id="8BE6DC05FF1820D10D8B13FB0B22F7D5" blockId="182.[183,1392,1478,2732]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10D8B13FB091DF81B" bold="true" box="[188,298,1959,1984]" pageId="182" pageNumber="461">Habitat.</emphasis>
Tropical savanna and grasslands from Cerrado and Beni Savanna biomes; two records are also known for Amazon savannas. The Woolly Giant Rat reportedly occurs almost exclusively in unflooded grasslands.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF1820D10D8B1C480B9DF777" pageId="182" pageNumber="461" type="food_feeding">
<paragraph id="8BE6DC05FF1820D10D8B1C480B9DF777" blockId="182.[183,1392,1478,2732]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10D8B1C4809F1F7EE" bold="true" box="[188,454,2068,2101]" pageId="182" pageNumber="461">Food and Feeding.</emphasis>
Direct field observations indicate that The Woolly Giant Rat eats roots of grasses. One captive individual ate insects (
<taxonomicName id="4C59A786FF1820D10EB61C1C0C15F786" authorityName="Latreille" authorityYear="1810" box="[897,1058,2112,2141]" class="Insecta" kingdom="Animalia" order="Orthoptera" pageId="182" pageNumber="461" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
and Isoptera) that were moving in leaflitter inside the vivarium where it was kept; insects were located by smell, captured with forefeet, and transferred to the mouth.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF1820D10D8B1CEE0A17F692" pageId="182" pageNumber="461" type="breeding">
<paragraph id="8BE6DC05FF1820D10D8B1CEE0A17F692" blockId="182.[183,1392,1478,2732]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10D8B1CEE0975F708" bold="true" box="[188,322,2226,2259]" pageId="182" pageNumber="461">Breeding.</emphasis>
Breeding of the Woolly Giant Rat might occur twice a year. A pregnant female was recorded in July (dry season) with three embryos; another pregnant female was captured in October (early wet season) with one embryo. Juveniles were caught in June-July and November.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF1820D10D8D1D130D13F643" pageId="182" pageNumber="461" type="activity">
<paragraph id="8BE6DC05FF1820D10D8D1D130D13F643" blockId="182.[183,1392,1478,2732]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10D8D1D130991F6AB" bold="true" box="[186,422,2383,2416]" pageId="182" pageNumber="461">Activity patterns.</emphasis>
The Woolly Giant Ratis active
<taxonomicName id="4C59A786FF1820D10E501D130BA0F6AB" box="[871,919,2383,2416]" class="Mammalia" family="Cricetidae" genus="Ondatra" kingdom="Animalia" order="Rodentia" pageId="182" pageNumber="461" phylum="Chordata" rank="species" species="day">day</taxonomicName>
and night, semi-fossorial, and reportedly builds small galleries with grasses pulled up from the surrounding area.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF1820D10D8B1DC20C50F664" box="[188,1127,2462,2495]" pageId="182" pageNumber="461" type="biology_ecology">
<paragraph id="8BE6DC05FF1820D10D8B1DC20C50F664" blockId="182.[183,1392,1478,2732]" box="[188,1127,2462,2495]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10D8B1DC20B4FF664" bold="true" box="[188,888,2462,2495]" pageId="182" pageNumber="461">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF1820D10D8A1D950D5DF5D5" pageId="182" pageNumber="461" type="conservation">
<paragraph id="8BE6DC05FF1820D10D8A1D950D5DF5D5" blockId="182.[183,1392,1478,2732]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10D8A1D950A2DF63D" bold="true" box="[189,538,2505,2534]" pageId="182" pageNumber="461">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. Nevertheless, the Woolly Giant Rat suffered recent regional extinctions in south-eastern Brazil.
</paragraph>
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<subSubSection id="C3438F8EFF1820D10D8A1E7D0CE8F577" pageId="182" pageNumber="461" type="bibRefCitation_list">
<paragraph id="8BE6DC05FF1820D10D8A1E7D0CE8F577" blockId="182.[183,1392,1478,2732]" pageId="182" pageNumber="461">
<emphasis id="B92D0017FF1820D10D8A1E7D0962F5ED" bold="true" box="[189,341,2593,2614]" pageId="182" pageNumber="461">Bibliography.</emphasis>
Anderson (1997), Andrades-Miranda et al. (1999), Bezerra (2005, 2015), Bezerra &amp; Pardinas (2016), Bezerra et al. (2007), Emmons (1998), Hadler et al. (2016), Hershkovitz (1966a), Ibanez et al. (1994), Lichtenstein (1830), Lund (1840b), Marinho-Filho &amp; Vieira (2016), Massoia &amp; Fornes (1965d), Miranda-Ribeiro (1914), Pardinas, D'Elia &amp; Teta (2009), Santos-Filho et al. (2001), Teran et al. (2008), Winge (1887).
</paragraph>
</subSubSection>
</treatment>
</document>