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<document id="AE2995EE83A5A3C8DCE8CC1E7AD90835" ID-CLB-Dataset="46584" ID-DOI="10.11646/zootaxa.3755.4.5" ID-GBIF-Dataset="e8a9a9bf-63a1-48e5-9fc1-bdbbd0c8be8f" ID-ISSN="1175-5326" ID-Zenodo-Dep="231257" ID-ZooBank="13446D49-76A1-4C12-975E-F59106AF4BD3" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460374538766" checkinUser="plazi" docAuthor="Shockley, Floyd W. &amp; Alekseev, Vitaly I." docDate="2014" docId="03CB3C15FFBA3F7EFF435495164039A7" docLanguage="en" docName="zt03755p400.pdf" docOrigin="Zootaxa 3755 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glesirhanis bercioi Shockley &amp; Alekseev, 2014, sp. nov." docType="treatment" docVersion="8" lastPageNumber="396" masterDocId="FFF2446DFFB83F7BFFD4527F12503B22" masterDocTitle="Glesirhanis bercioi, a new genus and species from Baltic amber (Coleoptera: Endomychidae: Leiestinae) with a checklist and nomenclatural notes regarding fossil Endomychidae" masterLastPageNumber="400" masterPageNumber="391" pageNumber="393" updateTime="1698296157067" updateUser="ExternalLinkService">
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<mods:title id="65DAD9B5C4633609C155B7A652658EE4">Glesirhanis bercioi, a new genus and species from Baltic amber (Coleoptera: Endomychidae: Leiestinae) with a checklist and nomenclatural notes regarding fossil Endomychidae</mods:title>
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<mods:namePart id="8B907C9EF25F287C729799BCF04529D9">Shockley, Floyd W.</mods:namePart>
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<mods:namePart id="96ACF9D3CD53FD266E3E29D59284DDDD">Alekseev, Vitaly I.</mods:namePart>
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<mods:date id="5ACBF520244409F3A55EF5047FB36206">2014</mods:date>
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<mods:number id="DC7107A6A949B9304CC55AE7C18EEF70">3755</mods:number>
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<mods:identifier id="19DAD3C376B4E53AE767A51F0CB409E4" type="CLB-Dataset">46584</mods:identifier>
<mods:identifier id="4659A68C2F297EECD471A4176A3169DA" type="DOI">10.11646/zootaxa.3755.4.5</mods:identifier>
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<treatment id="03CB3C15FFBA3F7EFF435495164039A7" ID-DOI="http://doi.org/10.5281/zenodo.6136834" ID-GBIF-Taxon="119412847" ID-Zenodo-Dep="6136834" LSID="urn:lsid:plazi:treatment:03CB3C15FFBA3F7EFF435495164039A7" httpUri="http://treatment.plazi.org/id/03CB3C15FFBA3F7EFF435495164039A7" lastPageId="5" lastPageNumber="396" pageId="2" pageNumber="393">
<subSubSection id="C378DE88FFBA3F79FF43549513423C04" pageId="2" pageNumber="393" type="nomenclature">
<paragraph id="8BDD8D03FFBA3F79FF43549513B23C27" blockId="2.[151,482,1770,1830]" box="[151,482,1770,1797]" pageId="2" pageNumber="393">
<heading id="D0953A6FFFBA3F79FF43549513B23C27" bold="true" box="[151,482,1770,1797]" fontSize="11" level="1" pageId="2" pageNumber="393" reason="1">
<emphasis id="B9165111FFBA3F79FF43549513B23C27" bold="true" box="[151,482,1770,1797]" pageId="2" pageNumber="393">
<taxonomicName id="4C62F680FFBA3F79FF435495132D3C26" box="[151,381,1770,1796]" class="Insecta" family="Endomychidae" genus="Glesirhanis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="bercioi" status="sp. nov.">
<emphasis id="B9165111FFBA3F79FF435495132D3C26" bold="true" box="[151,381,1770,1796]" italics="true" pageId="2" pageNumber="393">Glesirhanis bercioi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A225EC6AFFBA3F79FE50549413B23C27" box="[388,482,1771,1797]" pageId="2" pageNumber="393" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDD8D03FFBA3F79FF43557213423C04" blockId="2.[151,482,1770,1830]" box="[151,274,1805,1830]" pageId="2" pageNumber="393">
(
<figureCitation id="13599186FFBA3F79FF4B557213583C04" box="[159,264,1805,1830]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1775,1797]" captionStartId-1="4.[151,250,1739,1761]" captionTargetBox-0="[331,1255,195,1751]" captionTargetBox-1="[183,1416,429,1707]" captionTargetId-0="figure@3.[331,1255,195,1751]" captionTargetId-1="figure@4.[170,1426,421,1713]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Glesirhanis bercioi Shockley &amp; Alekseev, sp. nov. A. Dorsal habitus. B. Lateral habitus." captionText-1="FIGURE 2. Glesirhanis bercioi Shockley &amp; Alekseev, sp. nov. (punctures and setae not illustrated). A. Left antenna. B. Left metathoracic leg. C. Head, prothorax and scutellum." httpUri-0="https://zenodo.org/record/231258/files/figure.png" httpUri-1="https://zenodo.org/record/231259/files/figure.png" pageId="2" pageNumber="393">Figs. 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C378DE88FFBA3F7EFF43552A17593A47" lastPageId="5" lastPageNumber="396" pageId="2" pageNumber="393" type="description">
<paragraph id="8BDD8D03FFBA3F79FF43552A13DB3C94" blockId="2.[151,1436,1877,1974]" pageId="2" pageNumber="393">
<emphasis id="B9165111FFBA3F79FF43552A13793C4C" bold="true" box="[151,297,1877,1902]" pageId="2" pageNumber="393">Description.</emphasis>
Body length =
<quantity id="4C9A20E6FFBA3F79FE0F552A10653C4C" box="[475,565,1877,1902]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="393" unit="mm" value="2.5">2.5 mm</quantity>
; width (at widest point) =
<quantity id="4C9A20E6FFBA3F79FCB7552911F83C4C" box="[867,936,1878,1902]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="393" unit="mm" value="1.0">1 mm</quantity>
. Highly elongate habitus; dorsal and ventral surfaces, head, and appendages are dark brown, almost black (
<figureCitation id="13599186FFBA3F79FCB7550611F63CB0" box="[867,934,1913,1938]" captionStart="FIGURE 1" captionStartId="3.[151,250,1775,1797]" captionTargetBox="[331,1255,195,1751]" captionTargetId="figure@3.[331,1255,195,1751]" captionTargetPageId="3" captionText="FIGURE 1. Glesirhanis bercioi Shockley &amp; Alekseev, sp. nov. A. Dorsal habitus. B. Lateral habitus." httpUri="https://zenodo.org/record/231258/files/figure.png" pageId="2" pageNumber="393">Fig. 1</figureCitation>
). Head, pronotum and elytra conspicuously but variably punctate.
</paragraph>
<caption id="DF1DDD8BFFBB3F78FF43549016CB3C27" box="[151,1179,1775,1797]" httpUri="https://zenodo.org/record/231258/files/figure.png" pageId="3" pageNumber="394" targetBox="[331,1255,195,1751]" targetPageId="3">
<paragraph id="8BDD8D03FFBB3F78FF43549016CB3C27" blockId="3.[151,1179,1775,1797]" box="[151,1179,1775,1797]" pageId="3" pageNumber="394">
<emphasis id="B9165111FFBB3F78FF43549013433C26" bold="true" box="[151,275,1775,1797]" pageId="3" pageNumber="394">FIGURE 1.</emphasis>
<taxonomicName id="4C62F680FFBB3F78FECD549010903C27" authority="Shockley &amp; Alekseev" authorityName="Shockley &amp; Alekseev" box="[281,704,1775,1797]" class="Insecta" family="Endomychidae" genus="Glesirhanis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="bercioi" status="sp. nov.">
<emphasis id="B9165111FFBB3F78FECD5490138C3C27" box="[281,476,1775,1797]" italics="true" pageId="3" pageNumber="394">Glesirhanis bercioi</emphasis>
Shockley &amp; Alekseev
</taxonomicName>
,
<emphasis id="B9165111FFBB3F78FD1E548F116A3C26" bold="true" box="[714,826,1775,1797]" pageId="3" pageNumber="394">
<taxonomicNameLabel id="A225EC6AFFBB3F78FD1E548F114A3C27" box="[714,794,1776,1797]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Dorsal habitus.
<emphasis id="B9165111FFBB3F78FC0B548F11A73C27" bold="true" box="[991,1015,1776,1797]" pageId="3" pageNumber="394">B.</emphasis>
Lateral habitus.
</paragraph>
</caption>
<paragraph id="8BDD8D03FFBB3F7FFF13553417503AAA" blockId="3.[151,1436,1867,2037]" lastBlockId="4.[151,1436,151,392]" lastPageId="4" lastPageNumber="395" pageId="3" pageNumber="394">
Head with coarse, sparse punctation; eyes large, prominent. Frontoclypeal ridge weakly arcuate. Antennae relatively long, extending to base of elytra; 11-segmented with loose, conspicuous 3-segmented club (
<figureCitation id="13599186FFBB3F78FAE6551017283CAA" box="[1330,1400,1903,1928]" captionStart="FIGURE 2" captionStartId="4.[151,250,1739,1761]" captionTargetBox="[183,1416,429,1707]" captionTargetId="figure@4.[170,1426,421,1713]" captionTargetPageId="4" captionText="FIGURE 2. Glesirhanis bercioi Shockley &amp; Alekseev, sp. nov. (punctures and setae not illustrated). A. Left antenna. B. Left metathoracic leg. C. Head, prothorax and scutellum." httpUri="https://zenodo.org/record/231259/files/figure.png" pageId="3" pageNumber="394">Fig. 2</figureCitation>
A). Antennomere length proportions according to the formula: 2.0: 1.3: 1.3: 1.0: 1.0: 1.0: 1.0: 1.0: 1.7: 1.7: 2.0. Antennomeres 3–11 distinctly pubescent. Antennomeres 9–11 twice as wide as antennomere 8, as wide as long. Terminal antennomere subconical and narrowing apically. Maxillary palpomeres 3 and 4 only visible; palpomere 3 approximately twice as wide as 4; palpomere 4 conical, weakly rounded apically, 2.3 times as long as palpomere 3. Pronotum weakly convex, transverse, 0.71X as long as wide; disc coarsely and moderately densely punctate; interspaces 1.5–2.0X puncture diameter. Lateral sulci clearly visible, triangular in form; basal sulcus distinct and deep medially, becoming indistinct laterally (
<figureCitation id="13599186FFBC3F7FFD5A537B10803A3E" box="[654,720,260,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1739,1761]" captionTargetBox="[183,1416,429,1707]" captionTargetId="figure@4.[170,1426,421,1713]" captionTargetPageId="4" captionText="FIGURE 2. Glesirhanis bercioi Shockley &amp; Alekseev, sp. nov. (punctures and setae not illustrated). A. Left antenna. B. Left metathoracic leg. C. Head, prothorax and scutellum." httpUri="https://zenodo.org/record/231259/files/figure.png" pageId="4" pageNumber="395">Fig. 2</figureCitation>
C). Lateral margins of pronotum arcuate from anterior margin to midlength, weakly convergent just beyond midlength, and subparallel at posterior angles. Anterior pronotal angles appearing obtuse when viewed dorsally, posterior angles nearly right-angled (
<figureCitation id="13599186FFBC3F7FFC2A533316123A46" box="[1022,1090,332,357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1739,1761]" captionTargetBox="[183,1416,429,1707]" captionTargetId="figure@4.[170,1426,421,1713]" captionTargetPageId="4" captionText="FIGURE 2. Glesirhanis bercioi Shockley &amp; Alekseev, sp. nov. (punctures and setae not illustrated). A. Left antenna. B. Left metathoracic leg. C. Head, prothorax and scutellum." httpUri="https://zenodo.org/record/231259/files/figure.png" pageId="4" pageNumber="395">Fig. 2</figureCitation>
C). Prosternal process partially obscured ventrally, but apparently narrow, short. Procoxal cavities externally open, internally closed.
</paragraph>
<caption id="DF1DDD8BFFBC3F7FFF4354B410FA3DDD" httpUri="https://zenodo.org/record/231259/files/figure.png" pageId="4" pageNumber="395" targetBox="[183,1416,429,1707]" targetPageId="4">
<paragraph id="8BDD8D03FFBC3F7FFF4354B410FA3DDD" blockId="4.[151,1436,1739,1791]" pageId="4" pageNumber="395">
<emphasis id="B9165111FFBC3F7FFF4354B413433DC2" bold="true" box="[151,275,1739,1761]" pageId="4" pageNumber="395">FIGURE 2.</emphasis>
<taxonomicName id="4C62F680FFBC3F7FFECF54B410963DC3" authority="Shockley &amp; Alekseev" authorityName="Shockley &amp; Alekseev" box="[283,710,1739,1761]" class="Insecta" family="Endomychidae" genus="Glesirhanis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="bercioi" status="sp. nov.">
<emphasis id="B9165111FFBC3F7FFECF54B4138F3DC3" box="[283,479,1739,1761]" italics="true" pageId="4" pageNumber="395">Glesirhanis bercioi</emphasis>
Shockley &amp; Alekseev
</taxonomicName>
,
<taxonomicNameLabel id="A225EC6AFFBC3F7FFD0554B411723DC2" box="[721,802,1739,1760]" pageId="4" pageNumber="395" rank="species">
<emphasis id="B9165111FFBC3F7FFD0554B411723DC2" bold="true" box="[721,802,1739,1760]" pageId="4" pageNumber="395">sp. nov.</emphasis>
</taxonomicNameLabel>
(punctures and setae not illustrated).
<emphasis id="B9165111FFBC3F7FFB7154B416EE3DC2" bold="true" box="[1189,1214,1739,1760]" pageId="4" pageNumber="395">A.</emphasis>
Left antenna.
<emphasis id="B9165111FFBC3F7FFA8754B4173A3DC2" bold="true" box="[1363,1386,1739,1760]" pageId="4" pageNumber="395">B.</emphasis>
Left metathoracic leg.
<emphasis id="B9165111FFBC3F7FFE98549613343DDD" bold="true" box="[332,356,1769,1791]" pageId="4" pageNumber="395">C.</emphasis>
Head, prothorax and scutellum.
</paragraph>
</caption>
<paragraph id="8BDD8D03FFBC3F7FFF13553A177E3CCC" blockId="4.[151,1436,1861,2030]" pageId="4" pageNumber="395">
Elytra elongate, moderately convex, shining; disc coarsely, densely punctate, punctures lacking visible pubescence; punctures forming irregular rows, interspaces 1.0–2.0X, denser medially, sparser laterally. Sutural striae present, fine, entire (running from base to apex). Scutellum visible, highly transverse (about 2 times wider than long), widely rounded apically (
<figureCitation id="13599186FFBC3F7FFD8F55CE10FC3CE8" box="[603,684,1969,1994]" captionStart="FIGURE 1" captionStartId="3.[151,250,1775,1797]" captionTargetBox="[331,1255,195,1751]" captionTargetId="figure@3.[331,1255,195,1751]" captionTargetPageId="3" captionText="FIGURE 1. Glesirhanis bercioi Shockley &amp; Alekseev, sp. nov. A. Dorsal habitus. B. Lateral habitus." httpUri="https://zenodo.org/record/231258/files/figure.png" pageId="4" pageNumber="395">Figs. 1</figureCitation>
A, 2C). Mesoventrite bicarinate, mesocoxa circular in outline, trochantin concealed. Mesoventral process boat-shaped, subequal in width to mesocoxal cavity diameter.
</paragraph>
<paragraph id="8BDD8D03FFBD3F7EFF1352E711233BDA" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="396">
Legs (
<figureCitation id="13599186FFBD3F7EFED952E813023B92" box="[269,338,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1739,1761]" captionTargetBox="[183,1416,429,1707]" captionTargetId="figure@4.[170,1426,421,1713]" captionTargetPageId="4" captionText="FIGURE 2. Glesirhanis bercioi Shockley &amp; Alekseev, sp. nov. (punctures and setae not illustrated). A. Left antenna. B. Left metathoracic leg. C. Head, prothorax and scutellum." httpUri="https://zenodo.org/record/231259/files/figure.png" pageId="5" pageNumber="396">Fig. 2</figureCitation>
B) with tarsal formula 4-4-4. Femur subclavate, sparsely punctate. All tibiae simple (without any angulations or projections), but conspicuously curved basally with short spines apically. Tarsomere 1 small but distinct; tarsomere 2 weakly lobed ventrally. Tarsal claws simple.
</paragraph>
<paragraph id="8BDD8D03FFBD3F7EFF13537B17593A47" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="396">
Abdomen with five ventrites (
<figureCitation id="13599186FFBD3F7EFDCA537B100F3A3E" box="[542,607,260,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1775,1797]" captionTargetBox="[331,1255,195,1751]" captionTargetId="figure@3.[331,1255,195,1751]" captionTargetPageId="3" captionText="FIGURE 1. Glesirhanis bercioi Shockley &amp; Alekseev, sp. nov. A. Dorsal habitus. B. Lateral habitus." httpUri="https://zenodo.org/record/231258/files/figure.png" pageId="5" pageNumber="396">Fig. 1</figureCitation>
B). Ventrite 1 longer than
<date id="FFDCABC3FFBD3F7EFC5A537B165C3A3E" box="[910,1036,260,285]" pageId="5" pageNumber="396">V2 and V3</date>
combined;
<date id="FFDCABC3FFBD3F7EFB40537B16E73A3E" box="[1172,1207,260,284]" pageId="5" pageNumber="396">V2</date>
slightly longer than
<date id="FFDCABC3FFBD3F7EFF43535712EB3A62" box="[151,187,296,320]" pageId="5" pageNumber="396">V3</date>
;
<date id="FFDCABC3FFBD3F7EFF19535713033A62" box="[205,339,295,320]" pageId="5" pageNumber="396">V3 and V4</date>
subequal in length;
<date id="FFDCABC3FFBD3F7EFD9F5358103D3A62" box="[587,621,295,320]" pageId="5" pageNumber="396">V5</date>
1.5 times longer than
<date id="FFDCABC3FFBD3F7EFC56535711FB3A62" box="[898,939,296,320]" pageId="5" pageNumber="396">V4.</date>
Apex of terminal tergite widely rounded. Unfortunately, the reproductive system is obscured, making definite sexual determination impossible.
</paragraph>
</subSubSection>
<subSubSection id="C378DE88FFBD3F7EFF1353101018391F" pageId="5" pageNumber="396" type="materials_examined">
<paragraph id="8BDD8D03FFBD3F7EFF135310166C3A8F" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="396">
<emphasis id="B9165111FFBD3F7EFF13531013E03AAA" bold="true" box="[199,432,367,392]" pageId="5" pageNumber="396">Material examined.</emphasis>
<typeStatus id="54D933A1FFBD3F7EFE63531010743AAA" box="[439,548,367,392]" pageId="5" pageNumber="396" type="holotype">Holotype</typeStatus>
: Nr. AWI-035. Sex unknown. Locality data are as follows:
<collectingCountry id="F375CD93FFBD3F7EFB695310175C3AAA" box="[1213,1292,367,392]" name="Russia" pageId="5" pageNumber="396">Russia</collectingCountry>
: Kaliningrad region: the Sambian [Samland] peninsula: Yantarny village [formerly Palmnicken].
</paragraph>
<paragraph id="8BDD8D03FFBD3F7EFF1353C817263AF2" blockId="5.[151,1437,151,645]" box="[199,1398,439,464]" pageId="5" pageNumber="396">
<emphasis id="B9165111FFBD3F7EFF1353C813053AF2" bold="true" box="[199,341,439,464]" pageId="5" pageNumber="396">
<typeStatus id="54D933A1FFBD3F7EFF1353C813523AF2" box="[199,258,439,464]" pageId="5" pageNumber="396">Type</typeStatus>
strata.
</emphasis>
Baltic Amber. Upper Eocene, Prussian Formation (Priabonian). Estimated age: 37.2–33.9 my.
</paragraph>
<paragraph id="8BDD8D03FFBD3F7EFF1353A31018391F" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="396">
<emphasis id="B9165111FFBD3F7EFF1353A313C73AD7" bold="true" box="[199,407,476,501]" pageId="5" pageNumber="396">
<typeStatus id="54D933A1FFBD3F7EFF1353A313523AD7" box="[199,258,476,501]" pageId="5" pageNumber="396">Type</typeStatus>
Depository.
</emphasis>
The
<typeStatus id="54D933A1FFBD3F7EFE0853A2105C3AD7" box="[476,524,477,501]" pageId="5" pageNumber="396">type</typeStatus>
is currently housed in the private collection of the second author, Vitaly I. Alekseev (Kaliningrad,
<collectingCountry id="F375CD93FFBD3F7EFE76538013A1393A" box="[418,497,511,536]" name="Russia" pageId="5" pageNumber="396">Russia</collectingCountry>
), but will be deposited in the Paleontological Institute, Russian Academy of Science (Moscow) for permanent preservation.
</paragraph>
</subSubSection>
<subSubSection id="C378DE88FFBD3F7EFF135038164039A7" pageId="5" pageNumber="396" type="etymology">
<paragraph id="8BDD8D03FFBD3F7EFF135038164039A7" blockId="5.[151,1437,151,645]" pageId="5" pageNumber="396">
<emphasis id="B9165111FFBD3F7EFF135038131E3942" bold="true" box="[199,334,583,608]" pageId="5" pageNumber="396">Etymology.</emphasis>
The specific epithet is a patronym for Dr. Hans Bercio, an Eastern Prussian coleopterologist and one of the authors of the last catalogue of beetles of the former Eastern Prussia.
</paragraph>
</subSubSection>
</treatment>
</document>
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