553 lines
66 KiB
XML
553 lines
66 KiB
XML
<document id="6A5E24148DB4BD4DCF33B87B88E281BB" ID-DOI="10.11646/zootaxa.3670.4.9" ID-GBIF-Dataset="3e1a3d58-c70c-4aa7-8d97-38ebc0d72b00" ID-ISSN="1175-5326" ID-Zenodo-Dep="220205" ID-ZooBank="EE8234D8-6B6E-406C-9F6D-A82D5021731D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460635130597" checkinUser="plazi" docAuthor="Matsui, Masafumi, Hamidy, Amir & Eto, Koshiro" docDate="2013" docId="03B1011A767EFF87B7BAFD9EFDF5FD2B" docLanguage="en" docName="zt03670p590.pdf" docOrigin="Zootaxa 3670 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microhyla orientalis Matsui, Hamidy & Eto, 2013, sp. nov." docType="treatment" docUuid="58483409-29F9-4BE1-B149-6CD087DAE39E" docUuidSource="ZooBank" docVersion="9" lastPageNumber="588" masterDocId="FF887962767BFF8DB72DFFDBFFD4FF8C" masterDocTitle="Description of a new species of Microhyla from Bali, Indonesia (Amphibia, Anura)" masterLastPageNumber="590" masterPageNumber="579" pageNumber="584" updateTime="1698335142047" updateUser="plazi">
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<mods:mods id="999093F794EE74E1F4E49B96B444F10E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="36A7F7D3E2EB12E0A51231298CF42126">
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<mods:title id="BF7414660F4B764DA414DF34999DD9BB">Description of a new species of Microhyla from Bali, Indonesia (Amphibia, Anura)</mods:title>
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</mods:titleInfo>
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<mods:name id="E964ED25253FBE6227929A247190BF32" type="personal">
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<mods:roleTerm id="D1868D004034518FCA0225AE467CDEAC">Author</mods:roleTerm>
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<mods:namePart id="1BE07A7507F3E9658A9AEC12FBF2A639">Matsui, Masafumi</mods:namePart>
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<mods:roleTerm id="CA3F5B6CA15EA2303C7B9A47C615D996">Author</mods:roleTerm>
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<mods:namePart id="2416EF484436AF2FB3946C9A16610F6D">Hamidy, Amir</mods:namePart>
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</mods:name>
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<mods:roleTerm id="9C984C911554D1FE26BAD088B893B403">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E0462A067EED0D0824AEB4C3D5967AE9">Eto, Koshiro</mods:namePart>
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<mods:typeOfResource id="7812242C2D9629B6CE7B9B17CD2C312B">text</mods:typeOfResource>
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<mods:title id="4F7F6793E60605015148CE8CAB46D4B2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="2546E3109986B15B4331DFDCEE6D72A9">
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<mods:date id="2F3FA96357A18483AF2CB86F24E252EB">2013</mods:date>
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<mods:number id="3E8BC15FBDEE25F844EF2BEEBE38FF86">3670</mods:number>
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<mods:number id="16B2A4537170455E7C347C42500DEB6D">4</mods:number>
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<mods:start id="93C2553E01F414BE92D739D2E7538E1B">579</mods:start>
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<mods:classification id="2E48B0FCB86DE39C5A2C8216C9418264">journal article</mods:classification>
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<mods:identifier id="F9939DF2FE0D0995531607F15FBBAA87" type="DOI">10.11646/zootaxa.3670.4.9</mods:identifier>
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<mods:identifier id="448D8E804EB2580AB435C98993A6EFF9" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B1011A767EFF87B7BAFD9EFDF5FD2B" ID-DOI="http://doi.org/10.5281/zenodo.5624547" ID-GBIF-Taxon="119570763" ID-Zenodo-Dep="5624547" LSID="urn:lsid:zoobank.org:act:58483409-29F9-4BE1-B149-6CD087DAE39E" httpUri="http://treatment.plazi.org/id/03B1011A767EFF87B7BAFD9EFDF5FD2B" lastPageId="10" lastPageNumber="588" pageId="5" pageNumber="584">
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<subSubSection id="C302E387767EFF88B7BAFD9EFD68FD2D" pageId="5" pageNumber="584" type="nomenclature">
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<paragraph id="8BA7B00C767EFF88B7BAFD9EFE25FDD3" blockId="5.[151,497,581,607]" box="[151,497,581,607]" pageId="5" pageNumber="584">
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<heading id="D0EF0760767EFF88B7BAFD9EFE25FDD3" bold="true" box="[151,497,581,607]" fontSize="11" level="1" pageId="5" pageNumber="584" reason="1">
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<emphasis id="B96C6C1E767EFF88B7BAFD9EFE25FDD3" bold="true" box="[151,497,581,607]" pageId="5" pageNumber="584">
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<taxonomicName id="4C18CB8F767EFF88B7BAFD9EFE5AFDD3" ID-CoL="6RGQW" box="[151,398,581,607]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="584" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
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<emphasis id="B96C6C1E767EFF88B7BAFD9EFE5AFDD3" bold="true" box="[151,398,581,607]" italics="true" pageId="5" pageNumber="584">Microhyla orientalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25FD165767EFF88B6B9FD9EFE25FDD3" box="[404,497,581,607]" pageId="5" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA7B00C767EFF88B7BAFD50FD68FD2D" blockId="5.[151,700,651,674]" box="[151,700,651,674]" pageId="5" pageNumber="584">
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<emphasis id="B96C6C1E767EFF88B7BAFD50FEC6FD2E" bold="true" box="[151,274,651,674]" pageId="5" pageNumber="584">Synonymy:</emphasis>
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<taxonomicName id="4C18CB8F767EFF88B63AFD57FDACFD2D" authority="McKay, 2006" authorityName="McKay" authorityYear="2006" box="[279,632,651,674]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="584" phylum="Chordata" rank="species" species="achatina">
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<emphasis id="B96C6C1E767EFF88B63AFD57FE0AFD2E" box="[279,478,652,674]" italics="true" pageId="5" pageNumber="584">Microhyla achatina</emphasis>
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:
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<bibRefCitation id="EF89CDFD767EFF88B6C1FD56FDACFD2D" author="McKay" box="[492,632,651,674]" pageId="5" pageNumber="584" refString="McKay, J. L. (2006) A Field Guide to the Amphibians and Reptiles of Bali. Krieger, Malabar, Florida, 138 pp." type="book" year="2006">McKay, 2006</bibRefCitation>
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</taxonomicName>
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, p. 34.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C302E387767EFF88B7BAFD14FB7AFC48" pageId="5" pageNumber="584" type="materials_examined">
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||
<paragraph id="8BA7B00C767EFF88B7BAFD14FB55FC81" blockId="5.[151,1436,718,2008]" pageId="5" pageNumber="584">
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<emphasis id="B96C6C1E767EFF88B7BAFD14FEDFFD6B" bold="true" box="[151,267,719,743]" pageId="5" pageNumber="584">
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<typeStatus id="54A30EAE767EFF88B7BAFD14FED2FD6B" box="[151,262,719,743]" pageId="5" pageNumber="584" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED0928C9767EFF88B635FD14FE82FD6B" box="[280,342,719,743]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="584">MZB</collectionCode>
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.Amp 20404 (formerly
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<collectionCode id="ED0928C9767EFF88B542FD14FD69FD6A" box="[623,701,719,742]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="584">KUHE</collectionCode>
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55073), an adult male from Wongaya Gede, Bali,
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<collectingCountry id="F30FF09C767EFF88B202FD15FA48FD6A" box="[1327,1436,718,742]" name="Indonesia" pageId="5" pageNumber="584">Indonesia</collectingCountry>
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(
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<geoCoordinate id="EE2CD6CB767EFF88B7B2FD2EFEC9FC81" box="[159,285,754,781]" direction="south" orientation="latitude" pageId="5" pageNumber="584" precision="15" value="-8.4175005">08o25'03''S</geoCoordinate>
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,
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<geoCoordinate id="EE2CD6CB767EFF88B605FD2EFE60FC81" box="[296,436,754,781]" direction="east" orientation="longitude" pageId="5" pageNumber="584" precision="15" value="115.12444">115o07'28''E</geoCoordinate>
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, alt.
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<quantity id="4CE01DE9767EFF88B6C4FD2EFDE4FC81" box="[489,560,757,781]" metricMagnitude="2" metricUnit="m" metricValue="4.35" pageId="5" pageNumber="584" unit="m" value="435.0">435 m</quantity>
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a.s.l.:
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<figureCitation id="1323AC89767EFF88B557FD2EFD6FFC81" box="[634,699,757,781]" captionStart="FIGURE 1" captionStartId="1.[151,250,1921,1944]" captionTargetBox="[151,1436,844,1899]" captionTargetId="figure@1.[151,1436,844,1900]" captionTargetPageId="1" captionText="FIGURE 1. Map of Bali, Indonesia, showing the known distributions of Microhyla orientalis sp. nov. The filled star and filled circle, respectively, indicate the type localities of Wongaya Gede and Batukaru. Microhyla palmipes was sympatric with M. orientalis at both localities. The open circle indicates Ubud, where only M. palmipes was found." httpUri="https://zenodo.org/record/220206/files/figure.png" pageId="5" pageNumber="584">Fig. 1</figureCitation>
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), collected on
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<date id="FFA696CC767EFF88B44FFD2DFBD3FC81" box="[866,1031,757,782]" pageId="5" pageNumber="584" value="2012-08-07">7 August 2012</date>
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by K. Eto.
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</paragraph>
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<paragraph id="8BA7B00C767EFF88B7EAFCC6FD62FC2C" blockId="5.[151,1436,718,2008]" pageId="5" pageNumber="584">
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||
<emphasis id="B96C6C1E767EFF88B7EAFCC6FE92FCB9" bold="true" box="[199,326,797,821]" pageId="5" pageNumber="584">
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||
<typeStatus id="54A30EAE767EFF88B7EAFCC6FE95FCB9" box="[199,321,797,821]" pageId="5" pageNumber="584" type="paratype">Paratypes</typeStatus>
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||
.
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</emphasis>
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<collectionCode id="ED0928C9767EFF88B662FCC7FE5AFCB8" box="[335,398,796,820]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="584">MZB</collectionCode>
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.Amp16259, an adult female from Batukaru, Bali,
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<collectingCountry id="F30FF09C767EFF88B4F9FCC7FB95FCB8" box="[980,1089,796,820]" name="Indonesia" pageId="5" pageNumber="584">Indonesia</collectingCountry>
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(
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<geoCoordinate id="EE2CD6CB767EFF88B37EFCC7FB05FCB8" box="[1107,1233,793,820]" direction="south" orientation="latitude" pageId="5" pageNumber="584" precision="15" value="-8.373334">08o22'24''S</geoCoordinate>
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,
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<geoCoordinate id="EE2CD6CB767EFF88B3F2FCC7FABFFCB8" box="[1247,1387,793,820]" direction="east" orientation="longitude" pageId="5" pageNumber="584" precision="15" value="115.10667">115o06'24''E</geoCoordinate>
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, alt.
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<quantity id="4CE01DE9767EFF88B7BAFC9BFF0AFCD4" box="[151,222,832,856]" metricMagnitude="2" metricUnit="m" metricValue="8.15" pageId="5" pageNumber="584" unit="m" value="815.0">815 m</quantity>
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a.s.l.), collected on
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<date id="FFA696CC767EFF88B6ECFC9BFD86FCD4" box="[449,594,832,856]" pageId="5" pageNumber="584" value="2010-07-12">12 July 2010</date>
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by A. Hamidy, M.D. Kusrini., and U. Arifin;
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<collectionCode id="ED0928C9767EFF88B375FC9AFB72FCD4" box="[1112,1190,833,856]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="584">KUHE</collectionCode>
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55049, 55074, 55075 (three adult males),
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<collectionCode id="ED0928C9767EFF88B651FCBEFE1EFCF0" box="[380,458,869,892]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="584">KUHE</collectionCode>
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55050, 55072 (two adult females), and
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<collectionCode id="ED0928C9767EFF88B4B8FCBEFC37FCF0" box="[917,995,869,892]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="584">KUHE</collectionCode>
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55076 (one subadult female) from the
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<typeStatus id="54A30EAE767EFF88B7BAFC52FF13FC2D" box="[151,199,905,929]" pageId="5" pageNumber="584">type</typeStatus>
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locality, collected from
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<date id="FFA696CC767EFF88B6F5FC52FD66FC2C" box="[472,690,904,929]" pageId="5" pageNumber="584" value="2012-08-07" valueMax="2012-08-08" valueMin="2012-08-07">7 to 8 August 2012</date>
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.
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</paragraph>
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<paragraph id="8BA7B00C767EFF88B7EAFC77FB7AFC48" blockId="5.[151,1436,718,2008]" box="[199,1198,940,965]" pageId="5" pageNumber="584">
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<emphasis id="B96C6C1E767EFF88B7EAFC77FE6DFC49" bold="true" box="[199,441,940,965]" pageId="5" pageNumber="584">Referred specimens.</emphasis>
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<collectionCode id="ED0928C9767EFF88B6EDFC76FDDAFC48" box="[448,526,941,964]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="5" pageNumber="584">KUHE</collectionCode>
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47215 (five tadpoles), collection data same as the
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<typeStatus id="54A30EAE767EFF88B368FC77FB7EFC48" box="[1093,1194,940,964]" pageId="5" pageNumber="584" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C302E387767EFF88B7EAFC0BFBFAFB80" pageId="5" pageNumber="584" type="etymology">
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||
<paragraph id="8BA7B00C767EFF88B7EAFC0BFBFAFB80" blockId="5.[151,1436,718,2008]" pageId="5" pageNumber="584">
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||
<emphasis id="B96C6C1E767EFF88B7EAFC0BFE9AFC64" bold="true" box="[199,334,976,1000]" pageId="5" pageNumber="584">Etymology.</emphasis>
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The specific epithet is from a Latin word
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<emphasis id="B96C6C1E767EFF88B417FC0AFC5BFC64" box="[826,911,977,1000]" italics="true" pageId="5" pageNumber="584">orientis</emphasis>
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denoting eastern, alluding to the fact that the species from Bali represents geographically the easternmost species of
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<taxonomicName id="4C18CB8F767EFF88B499FC2FFBF3FB80" box="[948,1063,1012,1036]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="584" phylum="Chordata" rank="genus">
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<emphasis id="B96C6C1E767EFF88B499FC2FFBF3FB80" box="[948,1063,1012,1036]" italics="true" pageId="5" pageNumber="584">Microhyla</emphasis>
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</taxonomicName>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302E387767EFF88B7EAFBCCFB0DFA30" pageId="5" pageNumber="584" type="diagnosis">
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||
<paragraph id="8BA7B00C767EFF88B7EAFBCCFB0DFA30" blockId="5.[151,1436,718,2008]" pageId="5" pageNumber="584">
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||
<emphasis id="B96C6C1E767EFF88B7EAFBCCFE95FBBC" bold="true" box="[199,321,1047,1072]" pageId="5" pageNumber="584">Diagnosis.</emphasis>
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The new species is assigned to
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<taxonomicName id="4C18CB8F767EFF88B58AFBC3FCEAFBBC" box="[679,830,1048,1072]" class="Amphibia" family="Microhylidae" kingdom="Animalia" order="Anura" pageId="5" pageNumber="584" phylum="Chordata" rank="family">Microhylidae</taxonomicName>
|
||
by the possession of median spiracle in larvae, and to
|
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<taxonomicName id="4C18CB8F767EFF88B7BAFBE7FEDFFBD8" box="[151,267,1084,1108]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="584" phylum="Chordata" rank="genus">
|
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<emphasis id="B96C6C1E767EFF88B7BAFBE7FEDFFBD8" box="[151,267,1084,1108]" italics="true" pageId="5" pageNumber="584">Microhyla</emphasis>
|
||
</taxonomicName>
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||
among members of the family from Southeast Asia by: lack of small spine-like projection of skin at heel and elbow; belly without a brown network; underside of fingers without greatly enlarged tubercles; snout less than twice diameter of eye; inner metatarsal tubercle low, not shovel-like; tympanum hidden by skin (Inger, 1966). A small form of the genus, adult males about
|
||
<quantity id="4CE01DE9767EFF88B556FB73FD20FB4C" box="[635,756,1192,1216]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="5" pageNumber="584" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">16–17 mm</quantity>
|
||
in SVL; snout rounded in profile; first finger short, less than one-fifth of third; tips of three outer fingers weakly dilated, forming weak disks, dorsally with median longitudinal groove; outer palmar tubercle single; tibiotarsal articulation reaching up to center of eye; tips of toes distinctly dilated into disks, dorsally with median longitudinal groove; two or more phalanges on outer side of second toe, more than three phalanges on inner side of third toe, three or more phalanges on outer side of third toe, four or more phalanges on inner and outer sides of fourth toe, and three phalanges on inner side of fifth toe free of web; inner and outer metatarsal tubercles present; eyelid without supraciliary spines; faint vertebral stripe present; a black lateral stripe from behind eye to half length of trunk; tail of larva with a black marking at middle.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302E387767EFF87B7EAFA1CFDF5FD2B" lastPageId="10" lastPageNumber="589" pageId="5" pageNumber="584" type="description">
|
||
<paragraph id="8BA7B00C767EFF88B7EAFA1CFC15F918" blockId="5.[151,1436,718,2008]" pageId="5" pageNumber="584">
|
||
<emphasis id="B96C6C1E767EFF88B7EAFA1CFCD9FA6C" bold="true" box="[199,781,1479,1504]" pageId="5" pageNumber="584">
|
||
Description of
|
||
<typeStatus id="54A30EAE767EFF88B650FA13FE30FA6C" box="[381,484,1480,1504]" pageId="5" pageNumber="584" type="holotype">holotype</typeStatus>
|
||
(measurements in mm).
|
||
</emphasis>
|
||
SVL 16.6; habitus moderate (
|
||
<figureCitation id="1323AC89767EFF88B343FA13FB07FA6C" box="[1134,1235,1480,1504]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="6.[151,250,1315,1338]" captionStartId-1="6.[151,250,1979,2002]" captionTargetBox-0="[293,1274,499,1241]" captionTargetBox-1="[216,1350,1412,1955]" captionTargetId-0="figure@6.[219,1353,399,1310]" captionTargetId-1="figure@6.[216,1350,1412,1955]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionText-0="FIGURE 3. Dorsolateral view of male holotype of Microhyla orientalis sp. nov. (MZB. Amp 20404) from Wongaya Gede, Bali, Indonesia." captionText-1="FIGURE 4. Dorsal (A) and ventral (B) views of male holotype of Microhyla orientalis sp. nov. (MZB. Amp 20404). Scale bar = 10 mm." httpUri-0="https://zenodo.org/record/220208/files/figure.png" httpUri-1="https://zenodo.org/record/220209/files/figure.png" pageId="5" pageNumber="584">Fig. 3–4</figureCitation>
|
||
); head triangular, wider (5.5) than long (5.1); snout rounded dorsally and in profile, projecting beyond lower jaw; eye shorter (2.2) than snout (3.2); canthus rostralis rounded; lore sloping, very weakly concave; nostril lateral, below canthus rostralis, closer to tip of snout (1.0) than to eye (1.2); interorbital distance (1.6) subequal to internarial distance (1.6), the latter larger than upper eyelid (1.2); pineal spot absent; tympanum hidden; upper jaw edentate; tongue oval, without papillae; slit-like openings to a median subgular vocal sac.
|
||
</paragraph>
|
||
<paragraph id="8BA7B00C767EFF88B7EAF97BFAC6F8E0" blockId="5.[151,1436,718,2008]" pageId="5" pageNumber="584">
|
||
Forelimb short (9.9); fingers thin, free of web, but with slight skin fringes on both sides of second and third and inner side of fourth; second finger slightly longer (measured from inner side 1.9, outer 1.3) than fourth (inner 1.7), latter much longer than first (outer 0.6); tips of three outer fingers weakly dilated and forming weak disks slightly wider than basal phalanges, dorsally with median longitudinal groove; diameter of first finger (0.2) one-third that of third finger disk (0.6), latter one and half times width of phalange; a single outer palmar tubercle (0.6) larger than inner (0.4); distinct, rounded subarticular tubercles, formula 1, 1, 2, 2; nuptial pad absent (
|
||
<figureCitation id="1323AC89767EFF88B3EDF88FFAD0F8E0" box="[1216,1284,1876,1900]" captionStart="FIGURE 5" captionStartId="7.[151,250,1104,1127]" captionTargetBox="[370,1216,193,1082]" captionTargetId="figure@7.[370,1217,193,1083]" captionTargetPageId="7" captionText="FIGURE 5. Ventral view of left hand (A) and foot (B) of male holotype of Microhyla orientalis sp. nov. (MZB. Amp 20404). Scale bar = 5 mm." httpUri="https://zenodo.org/record/220210/files/figure.png" pageId="5" pageNumber="584">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA7B00C767EFF8BB7EAF8A3FAFFFE90" blockId="5.[151,1436,718,2008]" lastBlockId="6.[151,1436,151,391]" lastPageId="6" lastPageNumber="585" pageId="5" pageNumber="584">
|
||
Hindlimb moderately long (28.5) about three times length of forelimb; tibia long (9.0), heels overlapping when limbs are held at right angles to body; tibiotarsal articulation of adpressed limb reaching to center of eye; foot (9.4) longer than tibia; tips of toes distinctly dilated into disks, much wider than those of fingers (disk diameter of third toe 0.7), dorsally with median longitudinal groove; third toe longer than fifth; webs between toes poorly developed (
|
||
<figureCitation id="1323AC89767DFF8BB7B2FF67FF36FF58" box="[159,226,188,212]" captionStart="FIGURE 5" captionStartId="7.[151,250,1104,1127]" captionTargetBox="[370,1216,193,1082]" captionTargetId="figure@7.[370,1217,193,1083]" captionTargetPageId="7" captionText="FIGURE 5. Ventral view of left hand (A) and foot (B) of male holotype of Microhyla orientalis sp. nov. (MZB. Amp 20404). Scale bar = 5 mm." httpUri="https://zenodo.org/record/220210/files/figure.png" pageId="6" pageNumber="585">Fig. 5</figureCitation>
|
||
) and absent between first and second toes, formula (the number of phallanges free of web):
|
||
<emphasis id="B96C6C1E767DFF8BB3C6FF67FB22FF58" bold="true" box="[1259,1270,188,212]" pageId="6" pageNumber="585">I</emphasis>
|
||
n.a.–n.a.
|
||
<date id="FFA696CC767DFF8BB24EFF67FF70FF7B" pageId="6" pageNumber="585">
|
||
<emphasis id="B96C6C1E767DFF8BB24EFF67FAADFF58" bold="true" box="[1379,1401,188,212]" pageId="6" pageNumber="585">II</emphasis>
|
||
2– 3
|
||
</date>
|
||
+
|
||
<date id="FFA696CC767DFF8BB791FF3BFEC2FF7B" box="[188,278,223,248]" pageId="6" pageNumber="585">
|
||
<emphasis id="B96C6C1E767DFF8BB791FF3BFF09FF74" bold="true" box="[188,221,224,248]" pageId="6" pageNumber="585">III</emphasis>
|
||
3 –4
|
||
</date>
|
||
+
|
||
<date id="FFA696CC767DFF8BB601FF3BFEB5FF7B" box="[300,353,223,248]" pageId="6" pageNumber="585">
|
||
<emphasis id="B96C6C1E767DFF8BB601FF3BFE9FFF74" bold="true" box="[300,331,224,248]" pageId="6" pageNumber="585">IV</emphasis>
|
||
4
|
||
</date>
|
||
+–
|
||
<date id="FFA696CC767DFF8BB6ADFF04FE72FF74" box="[384,422,223,248]" pageId="6" pageNumber="585">
|
||
3
|
||
<emphasis id="B96C6C1E767DFF8BB6B9FF3BFE72FF74" bold="true" box="[404,422,224,248]" pageId="6" pageNumber="585">V</emphasis>
|
||
</date>
|
||
; subarticular tubercles prominent, rounded, formula 1, 1, 2, 3, 2; inner metatarsal tubercle oval, large, length (0.8) half of first toe (1.6); outer metatarsal tubercle elevated, smaller (0.5) than inner.
|
||
</paragraph>
|
||
<paragraph id="8BA7B00C767DFF8BB7EAFEFCFD14FE0B" blockId="6.[151,1436,151,391]" pageId="6" pageNumber="585">Skin smooth above with a few low tubercles scattered; eyelid without supraciliary spines; no supratympanic fold discernible; side of body sparsely scattered with tubercles or low ridges; hindlimb dorsally scattered with few tubercles; ventral side of body and limbs smooth.</paragraph>
|
||
<caption id="DF67E084767DFF8BB7BAFAF8FEE3FAD4" httpUri="https://zenodo.org/record/220208/files/figure.png" pageId="6" pageNumber="585" targetBox="[293,1274,499,1241]" targetPageId="6">
|
||
<paragraph id="8BA7B00C767DFF8BB7BAFAF8FEE3FAD4" blockId="6.[151,1436,1315,1368]" pageId="6" pageNumber="585">
|
||
<emphasis id="B96C6C1E767DFF8BB7BAFAF8FEC1FAB5" bold="true" box="[151,277,1315,1338]" pageId="6" pageNumber="585">FIGURE 3.</emphasis>
|
||
Dorsolateral view of male holotype of
|
||
<taxonomicName id="4C18CB8F767DFF8BB598FAFFFC53FAB6" box="[693,903,1316,1338]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="585" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="B96C6C1E767DFF8BB598FAFFFC53FAB6" box="[693,903,1316,1338]" italics="true" pageId="6" pageNumber="585">Microhyla orientalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25FD165767DFF8BB4BDFAFFFC37FAB5" box="[912,995,1316,1337]" pageId="6" pageNumber="585" rank="species">
|
||
<emphasis id="B96C6C1E767DFF8BB4BDFAFFFC37FAB5" bold="true" box="[912,995,1316,1337]" pageId="6" pageNumber="585">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(MZB.Amp 20404) from Wongaya Gede, Bali, Indonesia.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF67E084767DFF8BB7BAF860FF2EF87D" httpUri="https://zenodo.org/record/220209/files/figure.png" pageId="6" pageNumber="585" targetBox="[216,1350,1412,1955]" targetPageId="6">
|
||
<paragraph id="8BA7B00C767DFF8BB7BAF860FF2EF87D" blockId="6.[151,1436,1979,2033]" pageId="6" pageNumber="585">
|
||
<emphasis id="B96C6C1E767DFF8BB7BAF860FEC7F85D" bold="true" box="[151,275,1979,2002]" pageId="6" pageNumber="585">FIGURE 4.</emphasis>
|
||
Dorsal (A) and ventral (B) views of male holotype of
|
||
<taxonomicName id="4C18CB8F767DFF8BB411F867FBD9F85E" box="[828,1037,1980,2002]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="585" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="B96C6C1E767DFF8BB411F867FBD9F85E" box="[828,1037,1980,2002]" italics="true" pageId="6" pageNumber="585">Microhyla orientalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25FD165767DFF8BB33EF867FBB0F85D" box="[1043,1124,1980,2001]" pageId="6" pageNumber="585" rank="species">
|
||
<emphasis id="B96C6C1E767DFF8BB33EF867FBB0F85D" bold="true" box="[1043,1124,1980,2001]" pageId="6" pageNumber="585">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(MZB.Amp 20404). Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF67E084767CFF8AB7BAFB8BFE85FB09" httpUri="https://zenodo.org/record/220210/files/figure.png" pageId="7" pageNumber="586" targetBox="[370,1216,193,1082]" targetPageId="7">
|
||
<paragraph id="8BA7B00C767CFF8AB7BAFB8BFE85FB09" blockId="7.[151,1437,1104,1157]" pageId="7" pageNumber="586">
|
||
<emphasis id="B96C6C1E767CFF8AB7BAFB8BFEC7FBEA" bold="true" box="[151,275,1104,1127]" pageId="7" pageNumber="586">FIGURE 5.</emphasis>
|
||
Ventral view of left hand (A) and foot (B) of male holotype of
|
||
<taxonomicName id="4C18CB8F767CFF8AB4B3FB8BFBBBFBEA" box="[926,1135,1104,1126]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="7" pageNumber="586" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="B96C6C1E767CFF8AB4B3FB8BFBBBFBEA" box="[926,1135,1104,1126]" italics="true" pageId="7" pageNumber="586">Microhyla orientalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25FD165767CFF8AB35BFB8AFB13FBEA" box="[1142,1223,1105,1126]" pageId="7" pageNumber="586" rank="species">
|
||
<emphasis id="B96C6C1E767CFF8AB35BFB8AFB13FBEA" bold="true" box="[1142,1223,1105,1126]" pageId="7" pageNumber="586">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(MZB.Amp 20404). Scale bar = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF67E084767CFF8AB7BAF870FC58F86C" httpUri="https://zenodo.org/record/220211/files/figure.png" pageId="7" pageNumber="586" targetBox="[302,1284,1206,1941]" targetPageId="7">
|
||
<paragraph id="8BA7B00C767CFF8AB7BAF870FC58F86C" blockId="7.[151,1436,1963,2017]" pageId="7" pageNumber="586">
|
||
<emphasis id="B96C6C1E767CFF8AB7BAF870FEC7F84C" bold="true" box="[151,275,1963,1986]" pageId="7" pageNumber="586">FIGURE 6.</emphasis>
|
||
Larval
|
||
<taxonomicName id="4C18CB8F767CFF8AB64EF870FDE7F84D" box="[355,563,1963,1985]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="7" pageNumber="586" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="B96C6C1E767CFF8AB64EF870FDE7F84D" box="[355,563,1963,1985]" italics="true" pageId="7" pageNumber="586">Microhyla orientalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25FD165767CFF8AB517F877FD5FF84D" box="[570,651,1964,1985]" pageId="7" pageNumber="586" rank="species">
|
||
<emphasis id="B96C6C1E767CFF8AB517F877FD5FF84D" bold="true" box="[570,651,1964,1985]" pageId="7" pageNumber="586">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from Wongaya Gede, Bali, stage 40 of Shimizu & Ota (2003), total length = 23.2 mm. Dorsal (A), lateral (B), and ventral views (C). Scale bar = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA7B00C7673FF85B7EAFF4CFDD9FE78" blockId="8.[151,1436,151,788]" pageId="8" pageNumber="587">
|
||
<emphasis id="B96C6C1E7673FF85B7EAFF4CFEC4FF3C" bold="true" box="[199,272,151,176]" pageId="8" pageNumber="587">Color.</emphasis>
|
||
Color in life pinkish brown dorsally, with a dark mark medially continued from interorbital bar posteriorly to cloacal opening; interorbital bar, reverse triangle in shape and covering posterior half of upper eyelid; dorsal dark mark forming subtriangle marks on shoulder and behind sacrum, and running laterally to form a widelimbed reverse V-shaped mark in between on middle of back; a narrow, discontinuous vertebral line from tip of snout to above anus; a black lateral stripe extending from above arm to half length of trunk; a paler dark marking connecting interorbital bar and lateral stripe, dorsally bordering the latter; a dark stripe from snout to eye; a cream stripe extending from eye to axilla; except for upper arm, limbs dorsally with narrow dark brown bars; throat and chest darkly pigmented and abdomen cream white; iris dorsally and ventrally golden with black reticulation, darkly pigmented at anterior and posterior corners (
|
||
<figureCitation id="1323AC897673FF85B583FE6CFCF5FE43" box="[686,801,439,463]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="6.[151,250,1315,1338]" captionStartId-1="6.[151,250,1979,2002]" captionStartId-2="7.[151,250,1104,1127]" captionTargetBox-0="[293,1274,499,1241]" captionTargetBox-1="[216,1350,1412,1955]" captionTargetBox-2="[370,1216,193,1082]" captionTargetId-0="figure@6.[219,1353,399,1310]" captionTargetId-1="figure@6.[216,1350,1412,1955]" captionTargetId-2="figure@7.[370,1217,193,1083]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 3. Dorsolateral view of male holotype of Microhyla orientalis sp. nov. (MZB. Amp 20404) from Wongaya Gede, Bali, Indonesia." captionText-1="FIGURE 4. Dorsal (A) and ventral (B) views of male holotype of Microhyla orientalis sp. nov. (MZB. Amp 20404). Scale bar = 10 mm." captionText-2="FIGURE 5. Ventral view of left hand (A) and foot (B) of male holotype of Microhyla orientalis sp. nov. (MZB. Amp 20404). Scale bar = 5 mm." httpUri-0="https://zenodo.org/record/220208/files/figure.png" httpUri-1="https://zenodo.org/record/220209/files/figure.png" httpUri-2="https://zenodo.org/record/220210/files/figure.png" pageId="8" pageNumber="587">Figs. 3–5</figureCitation>
|
||
). In preservative, pattern has not obviously changed, although color has slightly faded.
|
||
</paragraph>
|
||
<paragraph id="8BA7B00C7673FF85B7EAFE24FE20FC98" blockId="8.[151,1436,151,788]" pageId="8" pageNumber="587">
|
||
<emphasis id="B96C6C1E7673FF85B7EAFE24FEEFFD94" bold="true" box="[199,315,511,536]" pageId="8" pageNumber="587">Variation.</emphasis>
|
||
Individuals of the
|
||
<typeStatus id="54A30EAE7673FF85B50AFDDBFD83FD94" box="[551,599,512,536]" pageId="8" pageNumber="587">type</typeStatus>
|
||
series are generally similar in appearance. Variation in size and body proportions is given in
|
||
<tableCitation id="C69A85B77673FF85B6B6FDFFFE24FDB0" box="[411,496,548,572]" captionStart="TABLE 3" captionStartId="8.[151,239,835,859]" captionTargetBox="[159,1148,918,1684]" captionTargetPageId="8" captionText="TABLE 3. Measurements of adult Microhyla orientalis sp. nov. types. SVL (Mean ± 1 SD, in mm) and medians of ratios (R) of other characters to SVL, followed by ranges in parenthesis. See text for character abbreviations." httpUri="http://table.plazi.org/id/DF67E0847673FF85B7BAFC98FB0AFCF7" pageId="8" pageNumber="587" tableUuid="DF67E0847673FF85B7BAFC98FB0AFCF7">Table 3</tableCitation>
|
||
. Female
|
||
<typeStatus id="54A30EAE7673FF85B578FDFEFD15FDB1" box="[597,705,549,573]" pageId="8" pageNumber="587" type="paratype">paratypes</typeStatus>
|
||
have a larger body size (>
|
||
<quantity id="4CE01DE97673FF85B4DCFDFFFB81FDB0" box="[1009,1109,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="8" pageNumber="587" unit="mm" value="18.3">18.3 mm</quantity>
|
||
SVL) than the males (<
|
||
<quantity id="4CE01DE97673FF85B247FDFFFF17FDD3" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" pageId="8" pageNumber="587" unit="mm" value="17.4">17.4 mm</quantity>
|
||
SVL), but tend to have shorter hindlimbs relative to SVL; tibiotarsal articulation of adpressed limbs reaching the posterior border of the eye in females, and the center of the eye in males. Webbing formula was stable for the outer side of the first toe, where web is absent, but slight variations are recognized in other positions, e.g., in a
|
||
<typeStatus id="54A30EAE7673FF85B7BAFD6EFF23FD41" box="[151,247,693,717]" pageId="8" pageNumber="587" type="paratype">paratype</typeStatus>
|
||
(KUHE 55072), webbing is absent from the first toe to the inner side of the third toe, while another (KUHE 55075) had slight webbing on the inner side of the second toe. Throats of females are less darkly pigmented than those of males.
|
||
</paragraph>
|
||
<caption id="DF67E0847673FF85B7BAFC98FB0AFCF7" ID-Table-UUID="DF67E0847673FF85B7BAFC98FB0AFCF7" httpUri="http://table.plazi.org/id/DF67E0847673FF85B7BAFC98FB0AFCF7" pageId="8" pageNumber="587" targetBox="[159,1148,918,1684]" targetIsTable="true" targetPageId="8">
|
||
<paragraph id="8BA7B00C7673FF85B7BAFC98FB0AFCF7" blockId="8.[151,1436,835,891]" pageId="8" pageNumber="587">
|
||
<emphasis id="B96C6C1E7673FF85B7BAFC98FEDDFCD7" bold="true" box="[151,265,835,859]" pageId="8" pageNumber="587">TABLE 3.</emphasis>
|
||
Measurements of adult
|
||
<taxonomicName id="4C18CB8F7673FF85B523FC9FFD3EFCD7" box="[526,746,836,859]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="8" pageNumber="587" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="B96C6C1E7673FF85B523FC9FFD3EFCD7" box="[526,746,836,859]" italics="true" pageId="8" pageNumber="587">Microhyla orientalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25FD1657673FF85B5DCFC98FC93FCD7" box="[753,839,835,859]" pageId="8" pageNumber="587" rank="species">
|
||
<emphasis id="B96C6C1E7673FF85B5DCFC98FC93FCD7" bold="true" box="[753,839,835,859]" pageId="8" pageNumber="587">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
types. SVL (Mean±1SD, in mm) and medians of ratios (R) of other characters to SVL, followed by ranges in parenthesis. See text for character abbreviations.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA7B00C7673FF85B6A8FC4DFBE4F918" pageId="8" pageNumber="587">
|
||
<table id="F91842AC76730072B7B2FC4DFBA8F918" box="[159,1148,918,1684]" gridcols="3" gridrows="19" pageId="8" pageNumber="587">
|
||
<tr id="3528B24E76730072B7B2FC4DFBA8FC21" box="[159,1148,918,941]" gridrow="0" pageId="8" pageNumber="587" rowspan-0="1">
|
||
<th id="76F9DB3276730072B6A8FC4DFDBEFC21" box="[389,618,918,941]" gridcol="1" gridrow="0" pageId="8" pageNumber="587">Males (N = 5)</th>
|
||
<th id="76F9DB3276730072B486FC4DFBA8FC21" box="[939,1148,918,941]" gridcol="2" gridrow="0" pageId="8" pageNumber="587">Females (N = 2)</th>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FC1BFBA8FC5A" box="[159,1148,960,982]" gridrow="1" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FC1BFF29FC5A" box="[159,253,960,982]" gridcol="0" gridrow="1" pageId="8" pageNumber="587">SVL</th>
|
||
<td id="76F9DB3276730072B6A8FC1BFDBEFC5A" box="[389,618,960,982]" gridcol="1" gridrow="1" pageId="8" pageNumber="587">16.7 ± 0.7 (15.8−17.4)</td>
|
||
<td id="76F9DB3276730072B486FC1BFBA8FC5A" box="[939,1148,960,982]" gridcol="2" gridrow="1" pageId="8" pageNumber="587">18.8 (18.3−19.2)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FC32FBA8FB8C" box="[159,1148,1001,1024]" gridrow="2" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FC32FF29FB8C" box="[159,253,1001,1024]" gridcol="0" gridrow="2" pageId="8" pageNumber="587">RHL</th>
|
||
<td id="76F9DB3276730072B6A8FC32FDBEFB8C" box="[389,618,1001,1024]" gridcol="1" gridrow="2" pageId="8" pageNumber="587">31.4 (30.7−35.4)</td>
|
||
<td id="76F9DB3276730072B486FC32FBA8FB8C" box="[939,1148,1001,1024]" gridcol="2" gridrow="2" pageId="8" pageNumber="587">30.7 (29.7−31.7)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FBC9FBA8FBA5" box="[159,1148,1042,1065]" gridrow="3" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FBC9FF29FBA5" box="[159,253,1042,1065]" gridcol="0" gridrow="3" pageId="8" pageNumber="587">RHW</th>
|
||
<td id="76F9DB3276730072B6A8FBC9FDBEFBA5" box="[389,618,1042,1065]" gridcol="1" gridrow="3" pageId="8" pageNumber="587">33.1 (32.2−36.7)</td>
|
||
<td id="76F9DB3276730072B486FBC9FBA8FBA5" box="[939,1148,1042,1065]" gridcol="2" gridrow="3" pageId="8" pageNumber="587">33.6 (32.8−34.4)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FBE0FBA8FBDE" box="[159,1148,1083,1106]" gridrow="4" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FBE0FF29FBDE" box="[159,253,1083,1106]" gridcol="0" gridrow="4" pageId="8" pageNumber="587">RIND</th>
|
||
<td id="76F9DB3276730072B6A8FBE0FDBEFBDE" box="[389,618,1083,1106]" gridcol="1" gridrow="4" pageId="8" pageNumber="587">9.2 (8.8−9.9)</td>
|
||
<td id="76F9DB3276730072B486FBE0FBA8FBDE" box="[939,1148,1083,1106]" gridcol="2" gridrow="4" pageId="8" pageNumber="587">8.1 (7.7−8.5)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FBBEFBA8FBF0" box="[159,1148,1125,1148]" gridrow="5" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FBBEFF29FBF0" box="[159,253,1125,1148]" gridcol="0" gridrow="5" pageId="8" pageNumber="587">RIOD</th>
|
||
<td id="76F9DB3276730072B6A8FBBEFDBEFBF0" box="[389,618,1125,1148]" gridcol="1" gridrow="5" pageId="8" pageNumber="587">9.5 (8.8−11.0)</td>
|
||
<td id="76F9DB3276730072B486FBBEFBA8FBF0" box="[939,1148,1125,1148]" gridcol="2" gridrow="5" pageId="8" pageNumber="587">9.0 (8.9−9.1)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FB55FBA8FB28" box="[159,1148,1166,1188]" gridrow="6" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FB55FF29FB28" box="[159,253,1166,1188]" gridcol="0" gridrow="6" pageId="8" pageNumber="587">RUEW</th>
|
||
<td id="76F9DB3276730072B6A8FB55FDBEFB28" box="[389,618,1166,1188]" gridcol="1" gridrow="6" pageId="8" pageNumber="587">7.8 (7.0−8.6)</td>
|
||
<td id="76F9DB3276730072B486FB55FBA8FB28" box="[939,1148,1166,1188]" gridcol="2" gridrow="6" pageId="8" pageNumber="587">6.7 (6.2−7.3)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FB63FBA8FB42" box="[159,1148,1208,1230]" gridrow="7" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FB63FF29FB42" box="[159,253,1208,1230]" gridcol="0" gridrow="7" pageId="8" pageNumber="587">REL</th>
|
||
<td id="76F9DB3276730072B6A8FB63FDBEFB42" box="[389,618,1208,1230]" gridcol="1" gridrow="7" pageId="8" pageNumber="587">13.9 (13.1−14.9)</td>
|
||
<td id="76F9DB3276730072B486FB63FBA8FB42" box="[939,1148,1208,1230]" gridcol="2" gridrow="7" pageId="8" pageNumber="587">13.5 (13.0−14.0)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FB3AFBA8FB74" box="[159,1148,1249,1272]" gridrow="8" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FB3AFF29FB74" box="[159,253,1249,1272]" gridcol="0" gridrow="8" pageId="8" pageNumber="587">RED</th>
|
||
<td id="76F9DB3276730072B6A8FB3AFDBEFB74" box="[389,618,1249,1272]" gridcol="1" gridrow="8" pageId="8" pageNumber="587">11.4 (10.7−12.9)</td>
|
||
<td id="76F9DB3276730072B486FB3AFBA8FB74" box="[939,1148,1249,1272]" gridcol="2" gridrow="8" pageId="8" pageNumber="587">9.9 (9.5−10.3)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FAD1FBA8FAAD" box="[159,1148,1290,1313]" gridrow="9" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FAD1FF29FAAD" box="[159,253,1290,1313]" gridcol="0" gridrow="9" pageId="8" pageNumber="587">RLAL</th>
|
||
<td id="76F9DB3276730072B6A8FAD1FDBEFAAD" box="[389,618,1290,1313]" gridcol="1" gridrow="9" pageId="8" pageNumber="587">43.0 (39.7−46.0)</td>
|
||
<td id="76F9DB3276730072B486FAD1FBA8FAAD" box="[939,1148,1290,1313]" gridcol="2" gridrow="9" pageId="8" pageNumber="587">41.1 (40.1−42.1)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FAE8FBA8FAC6" box="[159,1148,1331,1354]" gridrow="10" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FAE8FF29FAC6" box="[159,253,1331,1354]" gridcol="0" gridrow="10" pageId="8" pageNumber="587">RFLL</th>
|
||
<td id="76F9DB3276730072B6A8FAE8FDBEFAC6" box="[389,618,1331,1354]" gridcol="1" gridrow="10" pageId="8" pageNumber="587">58.9 (54.1−60.1)</td>
|
||
<td id="76F9DB3276730072B486FAE8FBA8FAC6" box="[939,1148,1331,1354]" gridcol="2" gridrow="10" pageId="8" pageNumber="587">53.3 (53.1−53.6)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FA86FBA8FAFF" box="[159,1148,1373,1395]" gridrow="11" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FA86FF29FAFF" box="[159,253,1373,1395]" gridcol="0" gridrow="11" pageId="8" pageNumber="587">RIPTL</th>
|
||
<td id="76F9DB3276730072B6A8FA86FDBEFAFF" box="[389,618,1373,1395]" gridcol="1" gridrow="11" pageId="8" pageNumber="587">3.1 (2.5−3.8)</td>
|
||
<td id="76F9DB3276730072B486FA86FBA8FAFF" box="[939,1148,1373,1395]" gridcol="2" gridrow="11" pageId="8" pageNumber="587">2.5 (2.3−2.6)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FA5DFBA8FA11" box="[159,1148,1414,1437]" gridrow="12" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FA5DFF29FA11" box="[159,253,1414,1437]" gridcol="0" gridrow="12" pageId="8" pageNumber="587">ROPTL</th>
|
||
<td id="76F9DB3276730072B6A8FA5DFDBEFA11" box="[389,618,1414,1437]" gridcol="1" gridrow="12" pageId="8" pageNumber="587">3.9 (3.1−4.7)</td>
|
||
<td id="76F9DB3276730072B486FA5DFBA8FA11" box="[939,1148,1414,1437]" gridcol="2" gridrow="12" pageId="8" pageNumber="587">3.6 (3.4−3.7)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FA6BFBA8FA4A" box="[159,1148,1456,1478]" gridrow="13" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FA6BFF29FA4A" box="[159,253,1456,1478]" gridcol="0" gridrow="13" pageId="8" pageNumber="587">RTL</th>
|
||
<td id="76F9DB3276730072B6A8FA6BFDBEFA4A" box="[389,618,1456,1478]" gridcol="1" gridrow="13" pageId="8" pageNumber="587">54.0 (50.6−56.3)</td>
|
||
<td id="76F9DB3276730072B486FA6BFBA8FA4A" box="[939,1148,1456,1478]" gridcol="2" gridrow="13" pageId="8" pageNumber="587">51.5 (50.0−53.0)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2FA02FBA8FA63" box="[159,1148,1497,1519]" gridrow="14" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2FA02FF29FA63" box="[159,253,1497,1519]" gridcol="0" gridrow="14" pageId="8" pageNumber="587">RFL</th>
|
||
<td id="76F9DB3276730072B6A8FA02FDBEFA63" box="[389,618,1497,1519]" gridcol="1" gridrow="14" pageId="8" pageNumber="587">56.6 (50.6−57.7)</td>
|
||
<td id="76F9DB3276730072B486FA02FBA8FA63" box="[939,1148,1497,1519]" gridcol="2" gridrow="14" pageId="8" pageNumber="587">52.6 (51.6−53.6)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2F9D9FBA8F994" box="[159,1148,1538,1560]" gridrow="15" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2F9D9FF29F994" box="[159,253,1538,1560]" gridcol="0" gridrow="15" pageId="8" pageNumber="587">RHLL</th>
|
||
<td id="76F9DB3276730072B6A8F9D9FDBEF994" box="[389,618,1538,1560]" gridcol="1" gridrow="15" pageId="8" pageNumber="587">171.5 (159.2−173.0)</td>
|
||
<td id="76F9DB3276730072B486F9D9FBA8F994" box="[939,1148,1538,1560]" gridcol="2" gridrow="15" pageId="8" pageNumber="587">163.0 (160.0−166.1)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2F9F0FBA8F9CE" box="[159,1148,1579,1602]" gridrow="16" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2F9F0FF29F9CE" box="[159,253,1579,1602]" gridcol="0" gridrow="16" pageId="8" pageNumber="587">RIMTL</th>
|
||
<td id="76F9DB3276730072B6A8F9F0FDBEF9CE" box="[389,618,1579,1602]" gridcol="1" gridrow="16" pageId="8" pageNumber="587">5.1 (3.4−5.8)</td>
|
||
<td id="76F9DB3276730072B486F9F0FBA8F9CE" box="[939,1148,1579,1602]" gridcol="2" gridrow="16" pageId="8" pageNumber="587">3.8 (3.7−3.8)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2F98EFBA8F9E0" box="[159,1148,1621,1644]" gridrow="17" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2F98EFF29F9E0" box="[159,253,1621,1644]" gridcol="0" gridrow="17" pageId="8" pageNumber="587">R1TOEL</th>
|
||
<td id="76F9DB3276730072B6A8F98EFDBEF9E0" box="[389,618,1621,1644]" gridcol="1" gridrow="17" pageId="8" pageNumber="587">9.3 (7.6−12.7)</td>
|
||
<td id="76F9DB3276730072B486F98EFBA8F9E0" box="[939,1148,1621,1644]" gridcol="2" gridrow="17" pageId="8" pageNumber="587">8.7 (8.1−9.4)</td>
|
||
</tr>
|
||
<tr id="3528B24E76730072B7B2F9A5FBA8F918" box="[159,1148,1662,1684]" gridrow="18" pageId="8" pageNumber="587">
|
||
<th id="76F9DB3276730072B7B2F9A5FF29F918" box="[159,253,1662,1684]" gridcol="0" gridrow="18" pageId="8" pageNumber="587">ROMTL</th>
|
||
<td id="76F9DB3276730072B6A8F9A5FDBEF918" box="[389,618,1662,1684]" gridcol="1" gridrow="18" pageId="8" pageNumber="587">3.6 (2.4−5.5)</td>
|
||
<td id="76F9DB3276730072B486F9A5FBA8F918" box="[939,1148,1662,1684]" gridcol="2" gridrow="18" pageId="8" pageNumber="587">2.5 (2.4−2.5)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BA7B00C7673FF84B7EAF901FC13FDB0" blockId="8.[151,1436,1753,2029]" lastBlockId="9.[151,1437,151,1040]" lastPageId="9" lastPageNumber="588" pageId="8" pageNumber="587">
|
||
<emphasis id="B96C6C1E7673FF85B7EAF901FEECF97E" bold="true" box="[199,312,1754,1778]" pageId="8" pageNumber="587">Tadpoles.</emphasis>
|
||
A total of five tadpoles from St. 39 (total length=
|
||
<quantity id="4CE01DE97673FF85B45CF902FC03F97D" box="[881,983,1753,1777]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="8" pageNumber="587" unit="mm" value="19.9">19.9 mm</quantity>
|
||
, head-body length=
|
||
<quantity id="4CE01DE97673FF85B394F902FAC4F97D" box="[1209,1296,1753,1777]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="8" pageNumber="587" unit="mm" value="7.1">7.1 mm</quantity>
|
||
) to 40 (
|
||
<quantity id="4CE01DE97673FF85B244F902FF11F899" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="8" pageNumber="587" unit="mm" value="23.2">23.2 mm</quantity>
|
||
,
|
||
<quantity id="4CE01DE97673FF85B7FCF926FEFDF899" box="[209,297,1789,1813]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="8" pageNumber="587" unit="mm" value="7.1">7.1 mm</quantity>
|
||
) from the
|
||
<typeStatus id="54A30EAE7673FF85B68FF925FE06F89A" box="[418,466,1790,1814]" pageId="8" pageNumber="587">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C18CB8F7673FF85B57BF925FD3DF899" box="[598,745,1789,1813]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="8" pageNumber="587" phylum="Chordata" rank="species" species="orientalis">
|
||
<emphasis id="B96C6C1E7673FF85B57BF925FD3DF899" box="[598,745,1789,1813]" italics="true" pageId="8" pageNumber="587">M. orientalis</emphasis>
|
||
</taxonomicName>
|
||
and identified as that species by DNA analyses were closely examined. Head and body flattened above, spheroidal below; maximum head-body width at level of eye 63–66% (median=63%) of head-body length; maximum head-body depth 84–96% (median=92%) of maximum head-body width; snout broadly rounded, almost truncate in profile; eyes lateral, visible from below, eyeball diameter 14–16% (median=15%) of head-body length; interorbital space very wide, 322–371% (median=337%) of eyeball diameter; eye-snout distance 25–31% (median=29%) of head-body length. Oral disk dorso-terminal, small; lower lip moderately expanded with width 21–25% (median=22%) of maximum head-body width, with a prominent lateral papilla on each side; labial teeth and jaw sheaths entirely absent, but lower labium scattered with small papillae on lateral margin. Spiracle opening median, without free flap, opening 70–85% (median=81%) of distance from tip of snout to end of body; vent median, in form of long tube directed nearly vertically downward, small opening at edge of ventral fin; thick loops of gut visible ventrally. Tail long and lanceolate, abruptly tapering in posterior half and drawn out into a short filament; tail length 182–251% (median=221%) of head-body length, maximum depth 25– 28% (median=26%) of length; dorsal fin originating at end of head-body, with a straight margin, sub-parallel with much deeper ventral fin in anterior half of tail; ventral fin deeper than dorsal throughout anterior to tail tip; caudal muscle moderately strong, maximum tail width 36–44% (median=39%) of maximum head-body width; muscle depth maximum at origin, 52–68% (median=57%) of maximum tail depth, but steadily narrowed posteriorly, with depth at middle of tail shallower than fin depths. Color in life (
|
||
<figureCitation id="1323AC897672FF84B472FE07FC77FE78" box="[863,931,476,500]" captionStart="FIGURE 6" captionStartId="7.[151,250,1963,1986]" captionTargetBox="[302,1284,1206,1941]" captionTargetId="figure@7.[302,1284,1206,1941]" captionTargetPageId="7" captionText="FIGURE 6. Larval Microhyla orientalis sp. nov. from Wongaya Gede, Bali, stage 40 of Shimizu & Ota (2003), total length = 23.2 mm. Dorsal (A), lateral (B), and ventral views (C). Scale bar = 5 mm." httpUri="https://zenodo.org/record/220211/files/figure.png" pageId="9" pageNumber="588">Fig. 6</figureCitation>
|
||
) light brown on dorsum and laterally, with a pair of paler interorbital markings and darker mid-dorsal band, and marking at end of flank; venter grey and belly semi-translucent; tail at middle with a black marking dotted with golden.
|
||
</paragraph>
|
||
<paragraph id="8BA7B00C7672FF84B7EAFD93FA89FDD3" blockId="9.[151,1437,151,1040]" box="[199,1373,583,608]" pageId="9" pageNumber="588">
|
||
<emphasis id="B96C6C1E7672FF84B7EAFD93FECEFDEC" bold="true" box="[199,282,584,608]" pageId="9" pageNumber="588">Range.</emphasis>
|
||
The new species is so far known only from Wongaya Gede and Batukaru, Bali Island,
|
||
<collectingCountry id="F30FF09C7672FF84B3C5FD9CFA8DFDD3" box="[1256,1369,583,607]" name="Indonesia" pageId="9" pageNumber="588">Indonesia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA7B00C7672FF84B7EAFDB7FB28FCB4" blockId="9.[151,1437,151,1040]" pageId="9" pageNumber="588">
|
||
<emphasis id="B96C6C1E7672FF84B7EAFDB7FE52FD08" bold="true" box="[199,390,619,644]" pageId="9" pageNumber="588">Natural history.</emphasis>
|
||
<taxonomicName id="4C18CB8F7672FF84B6A3FDB7FDA1FD08" box="[398,629,620,644]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="B96C6C1E7672FF84B6A3FDB7FDA1FD08" box="[398,629,620,644]" italics="true" pageId="9" pageNumber="588">Microhyla orientalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25FD1657672FF84B550FDB7FD03FD08" box="[637,727,620,644]" pageId="9" pageNumber="588" rank="species">
|
||
<emphasis id="B96C6C1E7672FF84B550FDB7FD05FD08" bold="true" box="[637,721,620,644]" pageId="9" pageNumber="588">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is sympatric with
|
||
<taxonomicName id="4C18CB8F7672FF84B486FDB6FBE3FD08" box="[939,1079,620,644]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="palmipes">
|
||
<emphasis id="B96C6C1E7672FF84B486FDB6FBE3FD08" box="[939,1079,620,644]" italics="true" pageId="9" pageNumber="588">M. palmipes</emphasis>
|
||
</taxonomicName>
|
||
on Bali around Wongaya Gede (
|
||
<typeStatus id="54A30EAE7672FF84B7B3FD54FF04FD2B" box="[158,208,655,679]" pageId="9" pageNumber="588">type</typeStatus>
|
||
locality) and Batukaru, while at Ubud, only
|
||
<taxonomicName id="4C18CB8F7672FF84B5FFFD4AFC89FD24" box="[722,861,656,680]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="palmipes">
|
||
<emphasis id="B96C6C1E7672FF84B5FFFD4AFC89FD24" box="[722,861,656,680]" italics="true" pageId="9" pageNumber="588">M. palmipes</emphasis>
|
||
</taxonomicName>
|
||
was found. They are actually syntopic and can be found very close to each other in the localities of sympatry. At Batukaru, calls of
|
||
<taxonomicName id="4C18CB8F7672FF84B361FD6EFB34FD40" box="[1100,1248,692,716]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="orientalis">
|
||
<emphasis id="B96C6C1E7672FF84B361FD6EFB34FD40" box="[1100,1248,692,716]" italics="true" pageId="9" pageNumber="588">M. orientalis</emphasis>
|
||
</taxonomicName>
|
||
were frequently heard in mid July, but at Wongaya Gede, calls were rarely heard in early August, while tadpoles approaching to metamorphosis were observed. In both cases,
|
||
<taxonomicName id="4C18CB8F7672FF84B58BFD26FCE7FC98" box="[678,819,764,788]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="palmipes">
|
||
<emphasis id="B96C6C1E7672FF84B58BFD26FCE7FC98" box="[678,819,764,788]" italics="true" pageId="9" pageNumber="588">M. palmipes</emphasis>
|
||
</taxonomicName>
|
||
was actively calling. Thus the new species may have shorter breeding seasons than
|
||
<taxonomicName id="4C18CB8F7672FF84B6C5FCFAFDA6FCB4" box="[488,626,800,824]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="palmipes">
|
||
<emphasis id="B96C6C1E7672FF84B6C5FCFAFDA6FCB4" box="[488,626,800,824]" italics="true" pageId="9" pageNumber="588">M. palmipes</emphasis>
|
||
</taxonomicName>
|
||
. Tadpoles were found in paddies with larval
|
||
<taxonomicName id="4C18CB8F7672FF84B346FCFAFB21FCB4" box="[1131,1269,800,824]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="palmipes">
|
||
<emphasis id="B96C6C1E7672FF84B346FCFAFB21FCB4" box="[1131,1269,800,824]" italics="true" pageId="9" pageNumber="588">M. palmipes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA7B00C7672FF84B7EAFC98FE8FFB83" blockId="9.[151,1437,151,1040]" pageId="9" pageNumber="588">
|
||
<emphasis id="B96C6C1E7672FF84B7EAFC98FE63FCD7" bold="true" box="[199,439,835,860]" pageId="9" pageNumber="588">Call characteristics.</emphasis>
|
||
Calls were recorded at Batukaru at an air temperature of 26.0°C at 21:00 h on
|
||
<date id="FFA696CC7672FF84B265FC9FFF1BFCF3" pageId="9" pageNumber="588" value="2010-07-12">12 July 2010</date>
|
||
by A. Hamidy. Calls (33 notes from two males were analyzed) consisted of a series of notes each emitted at an interval (between the beginnings of two successive notes) of 0.54±0.13 (0.37−0.97) s (
|
||
<figureCitation id="1323AC897672FF84B3B4FC57FB0BFC28" box="[1177,1247,908,932]" captionStart="FIGURE 7" captionStartId="9.[151,250,1753,1776]" captionTargetBox="[332,1270,1109,1725]" captionTargetId="figure@9.[314,1273,1093,1732]" captionTargetPageId="9" captionText="FIGURE 7. Sonagram (A) and wave form (B) showing four successive notes of an advertisement call of Microhyla orientalis sp. nov. from Batukaru, Bali, recorded at an air temperature of 26.0 ° C." httpUri="https://zenodo.org/record/220212/files/figure.png" pageId="9" pageNumber="588">Fig. 7</figureCitation>
|
||
). Each note was composed of 4.1±0.7 (3−5) short pulses and lasted for 0.07 ± 0.01 (0.01−0.08) s. Frequency bands spread over the 1.0–5.3 kHz range, and the dominant frequency was 3.4±0.1 (3.2−3.6) kHz. Frequency and intensity modulations were not marked.
|
||
</paragraph>
|
||
<caption id="DF67E0847672FF84B7BAF902FCB2F882" httpUri="https://zenodo.org/record/220212/files/figure.png" pageId="9" pageNumber="588" targetBox="[332,1270,1109,1725]" targetPageId="9">
|
||
<paragraph id="8BA7B00C7672FF84B7BAF902FCB2F882" blockId="9.[151,1436,1753,1806]" pageId="9" pageNumber="588">
|
||
<emphasis id="B96C6C1E7672FF84B7BAF902FEC7F963" bold="true" box="[151,275,1753,1776]" pageId="9" pageNumber="588">FIGURE 7.</emphasis>
|
||
Sonagram (A) and wave form (B) showing four successive notes of an advertisement call of
|
||
<taxonomicName id="4C18CB8F7672FF84B3E1F901FA48F97C" box="[1228,1436,1754,1776]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="B96C6C1E7672FF84B3E1F901FA48F97C" box="[1228,1436,1754,1776]" italics="true" pageId="9" pageNumber="588">Microhyla orientalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25FD1657672FF84B7BAF922FF33F882" box="[151,231,1785,1806]" pageId="9" pageNumber="588" rank="species">
|
||
<emphasis id="B96C6C1E7672FF84B7BAF922FF33F882" bold="true" box="[151,231,1785,1806]" pageId="9" pageNumber="588">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from Batukaru, Bali, recorded at an air temperature of 26.0°C.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA7B00C7672FF87B7EAF8E0FDF5FD2B" blockId="9.[151,1436,1851,2020]" lastBlockId="10.[151,1436,151,679]" lastPageId="10" lastPageNumber="589" pageId="9" pageNumber="588">
|
||
<emphasis id="B96C6C1E7672FF84B7EAF8E0FEB9F8D8" bold="true" box="[199,365,1851,1876]" pageId="9" pageNumber="588">Comparisons.</emphasis>
|
||
<taxonomicName id="4C18CB8F7672FF84B658F8E0FD8AF8DF" box="[373,606,1851,1875]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="B96C6C1E7672FF84B658F8E0FD8AF8DF" box="[373,606,1851,1875]" italics="true" pageId="9" pageNumber="588">Microhyla orientalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25FD1657672FF84B54BF8E7FD15F8DF" box="[614,705,1852,1876]" pageId="9" pageNumber="588" rank="species">
|
||
<emphasis id="B96C6C1E7672FF84B54BF8E7FD6EF8D8" bold="true" box="[614,698,1852,1876]" pageId="9" pageNumber="588">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
can be differentiated from genetically close members of the
|
||
<taxonomicName id="4C18CB8F7672FF84B250F8E7FF2DF8FB" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="achatina">
|
||
<emphasis id="B96C6C1E7672FF84B250F8E7FF2DF8FB" italics="true" pageId="9" pageNumber="588">M. achatina</emphasis>
|
||
</taxonomicName>
|
||
group of Matsui
|
||
<emphasis id="B96C6C1E7672FF84B697F8BBFE26F8FB" box="[442,498,1887,1911]" italics="true" pageId="9" pageNumber="588">et al.</emphasis>
|
||
(2011) and sympatric congeners in the following way. The new species differs from
|
||
<taxonomicName id="4C18CB8F7672FF84B7BAF85FFEF5F817" box="[151,289,1923,1947]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="achatina">
|
||
<emphasis id="B96C6C1E7672FF84B7BAF85FFEF5F817" box="[151,289,1923,1947]" italics="true" pageId="9" pageNumber="588">M. achatina</emphasis>
|
||
</taxonomicName>
|
||
by having dorsal dark markings covering the posterior half of the upper eyelid, a lateral dark band beginning at the base of the upper arm and vaguely bordered, and four or five dark bars on limbs (vs. dark marking posterior to upper eyelid, lateral band strong and clearly bordered, and two bars on limbs). From
|
||
<taxonomicName id="4C18CB8F7672FF87B22CF817FF3EFF23" authority="Blyth" authorityName="Blyth" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" lastPageId="10" lastPageNumber="589" order="Anura" pageId="9" pageNumber="588" phylum="Chordata" rank="species" species="berdmorei">
|
||
<emphasis id="B96C6C1E7672FF84B22CF817FA48F86F" box="[1281,1436,1995,2019]" italics="true" pageId="9" pageNumber="588">M. berdmorei</emphasis>
|
||
(Blyth)
|
||
</taxonomicName>
|
||
, the new species differs by poorly developed toe webbing (vs. very well developed webbing). The new species differs from
|
||
<taxonomicName id="4C18CB8F7671FF87B651FF66FDCEFF58" box="[380,538,188,212]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="10" pageNumber="589" phylum="Chordata" rank="species" species="borneensis">
|
||
<emphasis id="B96C6C1E7671FF87B651FF66FDCEFF58" box="[380,538,188,212]" italics="true" pageId="10" pageNumber="589">M. borneensis</emphasis>
|
||
</taxonomicName>
|
||
by larger body size, male SVL
|
||
<quantity id="4CE01DE97671FF87B452FF67FBF1FF58" box="[895,1061,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" metricValueMax="1.7399999999999998" metricValueMin="1.58" pageId="10" pageNumber="589" unit="mm" value="16.6" valueMax="17.4" valueMin="15.8">15.8–17.4 mm</quantity>
|
||
, and less developed toe webbing, with fifth toe with more than three phalanges free (vs. 11.0–
|
||
<quantity id="4CE01DE97671FF87B463FF04FC67FF7B" box="[846,947,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="10" pageNumber="589" unit="mm" value="13.2">13.2 mm</quantity>
|
||
and two phalanges free). The new species differs from
|
||
<taxonomicName id="4C18CB8F7671FF87B608FEDEFDC8FE90" authority="Boulenger" authorityName="Boulenger" box="[293,540,260,284]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="10" pageNumber="589" phylum="Chordata" rank="species" species="fissipes">
|
||
<emphasis id="B96C6C1E7671FF87B608FEDEFE4AFE90" box="[293,414,260,284]" italics="true" pageId="10" pageNumber="589">M. fissipes</emphasis>
|
||
Boulenger
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C18CB8F7671FF87B505FEDEFCE5FE90" authority="Liu and Hu" authorityName="Liu and Hu" box="[552,817,260,284]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="10" pageNumber="589" phylum="Chordata" rank="species" species="mixtura">
|
||
<emphasis id="B96C6C1E7671FF87B505FEDEFD72FE90" box="[552,678,261,284]" italics="true" pageId="10" pageNumber="589">M. mixtura</emphasis>
|
||
Liu and Hu
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C18CB8F7671FF87B413FEDEFB8DFE90" authority="Stejneger" authorityName="Stejneger" box="[830,1113,260,284]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="10" pageNumber="589" phylum="Chordata" rank="species" species="okinavensis">
|
||
<emphasis id="B96C6C1E7671FF87B413FEDEFC33FE90" box="[830,999,260,284]" italics="true" pageId="10" pageNumber="589">M. okinavensis</emphasis>
|
||
Stejneger
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C18CB8F7671FF87B3BBFEDEFA48FE90" authority="Hallowell" authorityName="Hallowell" box="[1174,1436,260,284]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="10" pageNumber="589" phylum="Chordata" rank="species" species="pulchra">
|
||
<emphasis id="B96C6C1E7671FF87B3BBFEDEFAC7FE90" box="[1174,1299,260,284]" italics="true" pageId="10" pageNumber="589">M. pulchra</emphasis>
|
||
(Hallowell)
|
||
</taxonomicName>
|
||
by having finger and toe disks with dorsal median longitudinal grooves (vs. disks and grooves entirely absent or present only on toes). From
|
||
<taxonomicName id="4C18CB8F7671FF87B6CDFE96FD66FEE9" authority="Vogt" authorityName="Vogt" box="[480,690,332,357]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="10" pageNumber="589" phylum="Chordata" rank="species" species="heymonsi">
|
||
<emphasis id="B96C6C1E7671FF87B6CDFE96FDA6FEE8" box="[480,626,332,356]" italics="true" pageId="10" pageNumber="589">M. heymonsi</emphasis>
|
||
Vogt
|
||
</taxonomicName>
|
||
, the new species differs by having two metacarpal tubercles and lacking loreal mask or marking on the vertebral line (vs. having three tubercles, a black band covering side of head, and one or two pairs of black marks on sides of vertebral line). The new species differs from
|
||
<taxonomicName id="4C18CB8F7671FF87B3CEFE4EFA8BFE20" box="[1251,1375,404,428]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="10" pageNumber="589" phylum="Chordata" rank="species" species="malang">
|
||
<emphasis id="B96C6C1E7671FF87B3CEFE4EFB2EFE20" box="[1251,1274,405,428]" italics="true" pageId="10" pageNumber="589">M</emphasis>
|
||
.
|
||
<emphasis id="B96C6C1E7671FF87B226FE4FFA8BFE20" box="[1291,1375,404,428]" italics="true" pageId="10" pageNumber="589">malang</emphasis>
|
||
</taxonomicName>
|
||
by a smaller body size, male SVL
|
||
<quantity id="4CE01DE97671FF87B6CAFE6CFD59FE43" box="[487,653,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" metricValueMax="1.7399999999999998" metricValueMin="1.58" pageId="10" pageNumber="589" unit="mm" value="16.6" valueMax="17.4" valueMin="15.8">15.8–17.4 mm</quantity>
|
||
, and less developed toe webbing, with the fifth toe having more than three phalanges free (vs.
|
||
<quantity id="4CE01DE97671FF87B683FE07FD84FE78" box="[430,592,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="2.0799999999999996" metricValueMax="2.2199999999999998" metricValueMin="1.94" pageId="10" pageNumber="589" unit="mm" value="20.799999999999997" valueMax="22.2" valueMin="19.4">19.4–22.2 mm</quantity>
|
||
and one phalange free), and from
|
||
<taxonomicName id="4C18CB8F7671FF87B4E0FE06FB8DFE78" box="[973,1113,476,500]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="10" pageNumber="589" phylum="Chordata" rank="species" species="mantheyi">
|
||
<emphasis id="B96C6C1E7671FF87B4E0FE06FB8DFE78" box="[973,1113,476,500]" italics="true" pageId="10" pageNumber="589">M. mantheyi</emphasis>
|
||
</taxonomicName>
|
||
by a smaller body size, male SVL
|
||
<quantity id="4CE01DE97671FF87B7FBFE24FEAAFD9B" box="[214,382,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" metricValueMax="1.7399999999999998" metricValueMin="1.58" pageId="10" pageNumber="589" unit="mm" value="16.6" valueMax="17.4" valueMin="15.8">15.8–17.4 mm</quantity>
|
||
, uniformly brown head, and less developed toe webbing, with the fifth toe with more than three phalanges free (vs.
|
||
<quantity id="4CE01DE97671FF87B699FDFFFD8EFDB0" box="[436,602,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.92" metricValueMin="1.88" pageId="10" pageNumber="589" unit="mm" value="24.0" valueMax="29.2" valueMin="18.8">18.8–29.2 mm</quantity>
|
||
, snout cream colored, and one phalange free). Finally, from
|
||
<taxonomicName id="4C18CB8F7671FF87B224FDFEFA41FDB0" box="[1289,1429,548,572]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="10" pageNumber="589" phylum="Chordata" rank="species" species="palmipes">
|
||
<emphasis id="B96C6C1E7671FF87B224FDFEFA41FDB0" box="[1289,1429,548,572]" italics="true" pageId="10" pageNumber="589">M. palmipes</emphasis>
|
||
</taxonomicName>
|
||
, the new species differs by lacking a supraciliary tubercle, having much less developed toe webbing, and the presence of dorsal median grooves on digital disks (vs. having a supraciliary tubercle and fairly developed toe webbing, and lacking the grooves).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |