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<document id="9CB2800734CBE9F63D3AA8447A3D76B5" ID-DOI="10.5281/zenodo.194025" ID-GBIF-Dataset="49a7308c-eda0-4400-be68-481fe331e358" ID-ISSN="1175-5326" ID-Zenodo-Dep="194025" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460184633881" checkinUser="plazi" docAuthor="Chen, Chao-Chun &amp; Chang, Hseuh-Wen" docDate="2010" docId="E7573659FFBD4D6FC2C0FA472BF3F8D7" docLanguage="en" docName="zt02399p030.pdf" docOrigin="Zootaxa 2399" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anoplodesmus aspinosus Chen &amp; Chang, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="1B6E4E21FFB94D67C257FFBA2A0DFFD7" masterDocTitle="The millipede genus Anoplodesmus Pocock, 1895, recorded in Taiwan for the first time, with descriptions of two new species (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini)" masterLastPageNumber="30" masterPageNumber="20" pageNumber="24" updateTime="1698236258056" updateUser="plazi">
<mods:mods id="0BB2363BC88292920CE106A9F875E01F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6597E786F7997730E96D2CA9AC75390D">
<mods:title id="6DFDD9E27512E30DE8ABED1C8B5E9F24">The millipede genus Anoplodesmus Pocock, 1895, recorded in Taiwan for the first time, with descriptions of two new species (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini)</mods:title>
</mods:titleInfo>
<mods:name id="5DDF114423A0A9E173985EF8E63CDABA" type="personal">
<mods:role id="DFD96218CA88608F92FE3FC19C10AF16">
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</mods:role>
<mods:namePart id="F8290378271FE1683BEC227F01FAB628">Chen, Chao-Chun</mods:namePart>
</mods:name>
<mods:name id="1B9866F380B13DF0153F65293F3E5235" type="personal">
<mods:role id="B4B148EC55EACD167A9CB2DE97BAA454">
<mods:roleTerm id="AA8496D7D57FD074F69D028A302138BA">Author</mods:roleTerm>
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<mods:namePart id="23B821BA93925D027A67604D101E9F7D">Chang, Hseuh-Wen</mods:namePart>
</mods:name>
<mods:typeOfResource id="565643395DBF0C86DF364B9B4322DC90">text</mods:typeOfResource>
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<mods:title id="9FAC3366CE02ADDD2024ABDB854CF2D7">Zootaxa</mods:title>
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<mods:part id="BAA30D0CFFECBAB1E2FDA5E6BC767222">
<mods:date id="C826A0C111DB06C80F1E2205666085C9">2010</mods:date>
<mods:detail id="F96226F62C6E3BF6B6719D6C6F931A42" type="volume">
<mods:number id="5652CD63AFCFBDB8151ACE7F5F144384">2399</mods:number>
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<mods:classification id="FE9FDA0BFA08B4B7BA2C7805DFA1B526">journal article</mods:classification>
<mods:identifier id="350455B41700BFB878D6C5BC4AD1B1F0" type="DOI">10.5281/zenodo.194025</mods:identifier>
<mods:identifier id="048E7AD6A508AB956919DE2B8C8A7522" type="GBIF-Dataset">49a7308c-eda0-4400-be68-481fe331e358</mods:identifier>
<mods:identifier id="23A34960193EBB5BD16432607C6910D9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="62846985B4599EFFFB23821DB13E197B" type="Zenodo-Dep">194025</mods:identifier>
</mods:mods>
<treatment id="E7573659FFBD4D6FC2C0FA472BF3F8D7" ID-DOI="http://doi.org/10.5281/zenodo.6204225" ID-GBIF-Taxon="124956657" ID-Zenodo-Dep="6204225" LSID="urn:lsid:plazi:treatment:E7573659FFBD4D6FC2C0FA472BF3F8D7" httpUri="http://treatment.plazi.org/id/E7573659FFBD4D6FC2C0FA472BF3F8D7" lastPageId="8" lastPageNumber="28" pageId="4" pageNumber="24">
<subSubSection id="27E4D4C4FFBD4D63C2C0FA472B24F9EA" pageId="4" pageNumber="24" type="nomenclature">
<paragraph id="6F41874FFFBD4D63C2C0FA472826F9C0" blockId="4.[151,555,1533,1597]" box="[151,555,1533,1559]" pageId="4" pageNumber="24">
<heading id="34093023FFBD4D63C2C0FA472826F9C0" bold="true" box="[151,555,1533,1559]" fontSize="11" level="1" pageId="4" pageNumber="24" reason="1">
<emphasis id="5D8A5B5DFFBD4D63C2C0FA472826F9C0" bold="true" box="[151,555,1533,1559]" pageId="4" pageNumber="24">
<taxonomicName id="A8FEFCCCFFBD4D63C2C0FA472BCAF9C0" ID-CoL="6798D" box="[151,455,1533,1559]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="24" phylum="Arthropoda" rank="species" species="aspinosus" status="sp. nov.">
<emphasis id="5D8A5B5DFFBD4D63C2C0FA472BCAF9C0" bold="true" box="[151,455,1533,1559]" italics="true" pageId="4" pageNumber="24">Anoplodesmus aspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46B9E626FFBD4D63C399FA472826F9C0" box="[462,555,1533,1559]" pageId="4" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6F41874FFFBD4D63C2C0F9992B24F9EA" blockId="4.[151,555,1533,1597]" box="[151,297,1571,1597]" pageId="4" pageNumber="24">
<figureCitation id="F7C59BCAFFBD4D63C2C0F9992B28F9EA" box="[151,293,1571,1597]" captionStart-0="FIGURES 17 22" captionStart-1="FIGURES 23 29" captionStart-2="FIGURES 30 33" captionStartId-0="5.[151,272,1435,1459]" captionStartId-1="6.[151,272,1853,1877]" captionStartId-2="7.[151,274,1573,1597]" captionTargetBox-0="[157,1431,302,1405]" captionTargetBox-1="[163,1434,190,1833]" captionTargetBox-2="[160,1436,186,1559]" captionTargetId-0="figure@5.[151,1436,279,1426]" captionTargetId-1="figure@6.[155,1440,183,1840]" captionTargetId-2="figure@7.[155,1440,181,1566]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 17 22. Anoplodesmus aspinosus sp. nov., male paratype from GuanU. 17, segment 10 (lateral view); 18, sternal lobe between coxae 4 (subventral view); 19 22, left gonopod (medial, ventral, lateral and dorsal views, respectively). Scale bar = 1.0 mm for figures 17 &amp; 18; 0.5 mm for figures 19 22." captionText-1="FIGURES 23 29. Anoplodesmus aspinosus sp. nov., paratypes from GuanU. 23, habitus of male and female (dorsal view); 24, anterior part of male body (dorsolateral view); 25 &amp; 26, middle part of male body (dorsal and lateral views, respectively); 27, caudal part of male body (lateral view); 28, telson (dorsal view); 29, sternal region of male midbody segments. Scale bar = 1.0 mm." captionText-2="FIGURES 30 33. Anoplodesmus aspinosus sp. nov., right gonopod of male paratype from FuShan (dorsomedial, medial, ventral and lateral views, respectively). Scale bar = 0.5 mm." httpUri-0="https://zenodo.org/record/194029/files/figure.png" httpUri-1="https://zenodo.org/record/194030/files/figure.png" httpUri-2="https://zenodo.org/record/194031/files/figure.png" pageId="4" pageNumber="24">Figs. 1733</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="27E4D4C4FFBD4D63C2C0F9CB2ADCF824" pageId="4" pageNumber="24" type="materials_examined">
<paragraph id="6F41874FFFBD4D63C2C0F9CB2ADCF824" blockId="4.[151,1437,1649,2035]" pageId="4" pageNumber="24">
<emphasis id="5D8A5B5DFFBD4D63C2C0F9CB2B91F95C" bold="true" box="[151,412,1649,1675]" pageId="4" pageNumber="24">Material examined:</emphasis>
<emphasis id="5D8A5B5DFFBD4D63C3F2F9CB281AF95D" box="[421,535,1649,1674]" italics="true" pageId="4" pageNumber="24">
<typeStatus id="B04539EDFFBD4D63C3F2F9CB281AF95D" box="[421,535,1649,1674]" pageId="4" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
male (NMNS-6190-001),
<collectingCountry id="17E9C7DFFFBD4D63C134F9CB29CCF95C" box="[867,961,1649,1675]" name="Taiwan" pageId="4" pageNumber="24">Taiwan</collectingCountry>
, Hsinchu County (), UFong Township (!), GuanU (&quot;#),
<quantity id="A8062AAAFFBD4D63C008F92028BAF963" box="[607,695,1690,1716]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="24" unit="m" value="2000.0">2000 m</quantity>
a.s.l.,
<date id="1B40A18FFFBD4D63C155F92029B3F963" box="[770,958,1690,1716]" pageId="4" pageNumber="24" value="2002-08-12">12 August 2002</date>
, leg. C. C. Chen, Y. H. Lin, J. N. Huang.
<emphasis id="5D8A5B5DFFBD4D63C2C0F97A2B19F90E" box="[151,276,1728,1753]" italics="true" pageId="4" pageNumber="24">
<typeStatus id="B04539EDFFBD4D63C2C0F97A2B19F90E" box="[151,276,1728,1753]" pageId="4" pageNumber="24" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="79F84CC6FFBD4D63C371F97A2B84F90D" box="[294,393,1728,1754]" pageId="4" pageNumber="24" type="male">2 males</specimenCount>
,
<specimenCount id="79F84CC6FFBD4D63C3C0F97A2802F90D" box="[407,527,1728,1754]" pageId="4" pageNumber="24" type="female">6 females</specimenCount>
(NMNS-6190-002), same locality and collectors as
<typeStatus id="B04539EDFFBD4D63C6C8F97A2F1DF90D" box="[1183,1296,1728,1754]" pageId="4" pageNumber="24" type="holotype">holotype</typeStatus>
.
<specimenCount id="79F84CC6FFBD4D63C74BF97A2F72F90D" box="[1308,1407,1728,1754]" pageId="4" pageNumber="24" type="male">5 males</specimenCount>
,
<specimenCount id="79F84CC6FFBD4D63C7DAF97A2AFFF8D4" pageId="4" pageNumber="24" type="female">4 females</specimenCount>
(
<collectionCode id="09EF1F8AFFBD4D63C356F9532B4BF8D4" box="[257,326,1769,1795]" httpUri="http://grbio.org/cool/fitg-90ys" name="Taiwan Fisheries Research Institute" pageId="4" pageNumber="24">TFRI</collectionCode>
), Taipei County ($%), ULai Township (&amp;), FuShan ('(),
<quantity id="A8062AAAFFBD4D63C6D5F9532EC6F8D4" box="[1154,1227,1769,1795]" metricMagnitude="2" metricUnit="m" metricValue="7.26" pageId="4" pageNumber="24" unit="m" value="726.0">726 m</quantity>
a.s.l.,
<date id="1B40A18FFFBD4D63C741F9532AD9F8FB" pageId="4" pageNumber="24" value="2001-06-19" valueMax="2001-06-26" valueMin="2001-06-19">1926 June 2001</date>
, leg. W. B. Huang.
<specimenCount id="79F84CC6FFBD4D63C3E1F8A8280AF8FB" box="[438,519,1810,1836]" pageId="4" pageNumber="24" type="male">1 male</specimenCount>
,
<specimenCount id="79F84CC6FFBD4D63C044F8A82876F8FB" box="[531,635,1810,1836]" pageId="4" pageNumber="24" type="female">1 female</specimenCount>
,
<specimenCount id="79F84CC6FFBD4D63C0D1F8A82908F8FB" box="[646,773,1810,1836]" pageId="4" pageNumber="24" type="juvenile">4 juveniles</specimenCount>
(NSYSUB), Taipei County ($%), ULai Township (&amp;), NeiDong (),
<quantity id="A8062AAAFFBD4D63C3BBF881283AF882" box="[492,567,1851,1877]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="24" unit="m" value="500.0">500 m</quantity>
a.s.l.,
<date id="1B40A18FFFBD4D63C0D4F881294DF882" box="[643,832,1851,1877]" pageId="4" pageNumber="24" value="2002-08-16">16 August 2002</date>
, leg. C. C. Chen &amp; Y. H. Lin.
<specimenCount id="79F84CC6FFBD4D63C6F7F8812F0BF882" box="[1184,1286,1851,1877]" pageId="4" pageNumber="24" type="female">1 female</specimenCount>
(NSYSUB), Nantou County ()), LuGu Township (*+), FengHuangGu (,-+),
<quantity id="A8062AAAFFBD4D63C66EF8DF2E8FF8A8" box="[1081,1154,1893,1919]" metricMagnitude="2" metricUnit="m" metricValue="7.34" pageId="4" pageNumber="24" unit="m" value="734.0">734 m</quantity>
a.s.l.,
<date id="1B40A18FFFBD4D63C69AF8DF2F68F8A8" box="[1229,1381,1893,1919]" pageId="4" pageNumber="24" value="1996-07-24">24 July 1996</date>
, leg. W. H. Chou.
<specimenCount id="79F84CC6FFBD4D63C378F8342BADF87F" box="[303,416,1934,1960]" pageId="4" pageNumber="24" type="female">2 females</specimenCount>
(NSYSUB), Ilan County (./), JiaoXi Township (0 1), Experimental Forest Plantation of National Ilan University,
<quantity id="A8062AAAFFBD4D63C030F80928BEF81A" box="[615,691,1971,1997]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="24" unit="m" value="200.0">200 m</quantity>
a.s.l.,
<geoCoordinate id="0ACAE188FFBD4D63C155F809299DF81A" box="[770,912,1971,1997]" direction="north" orientation="latitude" pageId="4" pageNumber="24" precision="15" value="24.7875">24°47'15&quot;N</geoCoordinate>
,
<geoCoordinate id="0ACAE188FFBD4D63C1CAF8092E3AF81A" box="[925,1079,1971,1997]" direction="east" orientation="longitude" pageId="4" pageNumber="24" precision="15" value="121.67">121°40'12&quot;E</geoCoordinate>
,
<date id="1B40A18FFFBD4D63C613F8092F12F81A" box="[1092,1311,1971,1997]" pageId="4" pageNumber="24" value="2009-09-07">7 September 2009</date>
, leg. W. J. Wen.
</paragraph>
</subSubSection>
<subSubSection id="27E4D4C4FFBC4D62C293FF2D288AFF2B" pageId="5" pageNumber="25" type="diagnosis">
<paragraph id="6F41874FFFBC4D62C293FF2D288AFF2B" blockId="5.[151,1437,151,252]" pageId="5" pageNumber="25">
<emphasis id="5D8A5B5DFFBC4D62C293FF2D2B4BFF66" bold="true" box="[196,326,151,177]" pageId="5" pageNumber="25">Diagnosis:</emphasis>
Differs from congeners in the absence of a distofemoral process of the gonopod, coupled with an especially complex solenophore, as well as with evidently oblong traces of paraterga and a subtruncate sternal lamina between male fourth coxae.
</paragraph>
</subSubSection>
<caption id="3B81D7C7FFBC4D62C2C0FA2129F4FA25" httpUri="https://zenodo.org/record/194029/files/figure.png" pageId="5" pageNumber="25" targetBox="[157,1431,302,1405]" targetPageId="5">
<paragraph id="6F41874FFFBC4D62C2C0FA2129F4FA25" blockId="5.[151,1436,1435,1522]" pageId="5" pageNumber="25">
<emphasis id="5D8A5B5DFFBC4D62C2C0FA212B6FFA64" bold="true" box="[151,354,1435,1459]" pageId="5" pageNumber="25">FIGURES 1722.</emphasis>
<taxonomicName id="A8FEFCCCFFBC4D62C33DFA212870FA65" box="[362,637,1435,1458]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="aspinosus" status="sp. nov.">
<emphasis id="5D8A5B5DFFBC4D62C33DFA212870FA65" box="[362,637,1435,1458]" italics="true" pageId="5" pageNumber="25">Anoplodesmus aspinosus</emphasis>
</taxonomicName>
<emphasis id="5D8A5B5DFFBC4D62C0D3FA2128EEFA64" bold="true" box="[644,739,1435,1459]" pageId="5" pageNumber="25">
<taxonomicNameLabel id="46B9E626FFBC4D62C0D3FA2128D3FA64" box="[644,734,1435,1459]" pageId="5" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male paratype from GuanU. 17, segment 10 (lateral view); 18, sternal lobe between coxae 4 (subventral view); 1922, left gonopod (medial, ventral, lateral and dorsal views, respectively). Scale bar = 1.0 mm for figures 17 &amp; 18; 0.5 mm for figures 1922.
</paragraph>
</caption>
<subSubSection id="27E4D4C4FFBC4D6FC293F99A2E24FF65" lastPageId="8" lastPageNumber="28" pageId="5" pageNumber="25" type="description">
<paragraph id="6F41874FFFBC4D62C293F99A2963F9B7" blockId="5.[151,1437,1568,2005]" pageId="5" pageNumber="25">
<emphasis id="5D8A5B5DFFBC4D62C293F99A2B6EF9ED" bold="true" box="[196,355,1568,1594]" pageId="5" pageNumber="25">Description:</emphasis>
Adult body
<quantity id="A8062AAAFFBC4D62C3AAF99A2873F9ED" box="[509,638,1568,1594]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="5" pageNumber="25" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">2021 mm</quantity>
long and 2.0 mm wide (male),
<quantity id="A8062AAAFFBC4D62C657F99A2E8CF9ED" box="[1024,1153,1568,1594]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="5" pageNumber="25" unit="mm" value="20.0" valueMax="22.0" valueMin="18.0">1822 mm</quantity>
long and
<quantity id="A8062AAAFFBC4D62C6ABF99A2F91F9ED" box="[1276,1436,1568,1594]" metricMagnitude="-3" metricUnit="m" metricValue="2.275" metricValueMax="2.5" metricValueMin="2.05" pageId="5" pageNumber="25" unit="mm" value="2.275" valueMax="2.5" valueMin="2.05">2.052.5 mm</quantity>
wide (female).
<typeStatus id="B04539EDFFBC4D62C31EF9FC2BBBF9B7" box="[329,438,1606,1632]" pageId="5" pageNumber="25" type="holotype">Holotype</typeStatus>
about
<quantity id="A8062AAAFFBC4D62C051F9FC2855F9B7" box="[518,600,1606,1632]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="25" unit="mm" value="20.0">20 mm</quantity>
long and 2.0 mm wide.
</paragraph>
<paragraph id="6F41874FFFBC4D62C29BF9D12925F97D" blockId="5.[151,1437,1568,2005]" pageId="5" pageNumber="25">
General coloration of adults usually dark blackish-brown to light brown in both sexes, sometimes faded to nearly whitish; pattern same as in
<taxonomicName id="A8FEFCCCFFBC4D62C012F92B28CBF97D" box="[581,710,1681,1706]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="spiniger" status="sp. nov.">
<emphasis id="5D8A5B5DFFBC4D62C012F92B28CBF97D" box="[581,710,1681,1706]" italics="true" pageId="5" pageNumber="25">A. spiniger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46B9E626FFBC4D62C09AF92A2925F97D" box="[717,808,1680,1706]" pageId="5" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="6F41874FFFBC4D62C293F90C28E1F802" blockId="5.[151,1437,1568,2005]" pageId="5" pageNumber="25">
Body subcylindrical, non-moniliform, slightly higher than wide. Postcollar constriction very faint, particularly in female; width of head = collum &lt;segments 2=4 &lt;5=16, thereafter body gradually tapering towards telson (
<figureCitation id="F7C59BCAFFBC4D62C307F8BA2BA5F8CD" box="[336,424,1792,1818]" captionStart="FIGURES 23 29" captionStartId="6.[151,272,1853,1877]" captionTargetBox="[163,1434,190,1833]" captionTargetId="figure@6.[155,1440,183,1840]" captionTargetPageId="6" captionText="FIGURES 23 29. Anoplodesmus aspinosus sp. nov., paratypes from GuanU. 23, habitus of male and female (dorsal view); 24, anterior part of male body (dorsolateral view); 25 &amp; 26, middle part of male body (dorsal and lateral views, respectively); 27, caudal part of male body (lateral view); 28, telson (dorsal view); 29, sternal region of male midbody segments. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/194030/files/figure.png" pageId="5" pageNumber="25">Fig. 23</figureCitation>
). All other peripheral characters as in
<taxonomicName id="A8FEFCCCFFBC4D62C132F8BB29E8F8CD" box="[869,997,1793,1818]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="25" phylum="Arthropoda" rank="species" species="spiniger" status="sp. nov.">
<emphasis id="5D8A5B5DFFBC4D62C132F8BB29E8F8CD" box="[869,997,1793,1818]" italics="true" pageId="5" pageNumber="25">A. spiniger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46B9E626FFBC4D62C1BBF8BA2E44F8CD" box="[1004,1097,1792,1818]" pageId="5" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, except as follows. Paraterga nearly wanting, expressed as very faint, mainly oblong bulges only on pore-bearing segments (
<figureCitation id="F7C59BCAFFBC4D62C752F89C2F6FF897" box="[1285,1378,1830,1856]" captionStart="FIGURES 17 22" captionStartId="5.[151,272,1435,1459]" captionTargetBox="[157,1431,302,1405]" captionTargetId="figure@5.[151,1436,279,1426]" captionTargetPageId="5" captionText="FIGURES 17 22. Anoplodesmus aspinosus sp. nov., male paratype from GuanU. 17, segment 10 (lateral view); 18, sternal lobe between coxae 4 (subventral view); 19 22, left gonopod (medial, ventral, lateral and dorsal views, respectively). Scale bar = 1.0 mm for figures 17 &amp; 18; 0.5 mm for figures 19 22." httpUri="https://zenodo.org/record/194029/files/figure.png" pageId="5" pageNumber="25">Figs 17</figureCitation>
,
<figureCitation id="F7C59BCAFFBC4D62C727F89C2AB4F8B2" captionStart="FIGURES 23 29" captionStartId="6.[151,272,1853,1877]" captionTargetBox="[163,1434,190,1833]" captionTargetId="figure@6.[155,1440,183,1840]" captionTargetPageId="6" captionText="FIGURES 23 29. Anoplodesmus aspinosus sp. nov., paratypes from GuanU. 23, habitus of male and female (dorsal view); 24, anterior part of male body (dorsolateral view); 25 &amp; 26, middle part of male body (dorsal and lateral views, respectively); 27, caudal part of male body (lateral view); 28, telson (dorsal view); 29, sternal region of male midbody segments. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/194030/files/figure.png" pageId="5" pageNumber="25">
24
<emphasis id="5D8A5B5DFFBC4D62C7DAF89F2F91F8E8" bold="true" box="[1421,1436,1829,1855]" pageId="5" pageNumber="25"></emphasis>
27
</figureCitation>
). Pleurosternal carinae very well developed, like very evident, arcuated ridges on segments 216 (male) (
<figureCitation id="F7C59BCAFFBC4D62C2C8F8CA2AF5F85D" box="[159,248,1904,1930]" captionStart="FIGURES 17 22" captionStartId="5.[151,272,1435,1459]" captionTargetBox="[157,1431,302,1405]" captionTargetId="figure@5.[151,1436,279,1426]" captionTargetPageId="5" captionText="FIGURES 17 22. Anoplodesmus aspinosus sp. nov., male paratype from GuanU. 17, segment 10 (lateral view); 18, sternal lobe between coxae 4 (subventral view); 19 22, left gonopod (medial, ventral, lateral and dorsal views, respectively). Scale bar = 1.0 mm for figures 17 &amp; 18; 0.5 mm for figures 19 22." httpUri="https://zenodo.org/record/194029/files/figure.png" pageId="5" pageNumber="25">Figs 17</figureCitation>
&amp;
<figureCitation id="F7C59BCAFFBC4D62C348F8CA2B33F85D" box="[287,318,1904,1930]" captionStart="FIGURES 23 29" captionStartId="6.[151,272,1853,1877]" captionTargetBox="[163,1434,190,1833]" captionTargetId="figure@6.[155,1440,183,1840]" captionTargetPageId="6" captionText="FIGURES 23 29. Anoplodesmus aspinosus sp. nov., paratypes from GuanU. 23, habitus of male and female (dorsal view); 24, anterior part of male body (dorsolateral view); 25 &amp; 26, middle part of male body (dorsal and lateral views, respectively); 27, caudal part of male body (lateral view); 28, telson (dorsal view); 29, sternal region of male midbody segments. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/194030/files/figure.png" pageId="5" pageNumber="25">26</figureCitation>
) or 214/15 (female), onward missing; caudal corner of most of carinae in male like a distinct rounded tooth. Epiproct long, conical, tip subtruncate or slightly emarginate in dorsal view; pre-apical papillae rather faint, placed close to apex (
<figureCitation id="F7C59BCAFFBC4D62C0D0F80128D2F802" box="[647,735,1979,2005]" captionStart="FIGURES 23 29" captionStartId="6.[151,272,1853,1877]" captionTargetBox="[163,1434,190,1833]" captionTargetId="figure@6.[155,1440,183,1840]" captionTargetPageId="6" captionText="FIGURES 23 29. Anoplodesmus aspinosus sp. nov., paratypes from GuanU. 23, habitus of male and female (dorsal view); 24, anterior part of male body (dorsolateral view); 25 &amp; 26, middle part of male body (dorsal and lateral views, respectively); 27, caudal part of male body (lateral view); 28, telson (dorsal view); 29, sternal region of male midbody segments. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/194030/files/figure.png" pageId="5" pageNumber="25">Fig. 28</figureCitation>
).
</paragraph>
<caption id="3B81D7C7FFBF4D61C2C0F8872BD2F862" httpUri="https://zenodo.org/record/194030/files/figure.png" pageId="6" pageNumber="26" targetBox="[163,1434,190,1833]" targetPageId="6">
<paragraph id="6F41874FFFBF4D61C2C0F8872BD2F862" blockId="6.[151,1437,1853,1973]" pageId="6" pageNumber="26">
<emphasis id="5D8A5B5DFFBF4D61C2C0F8872B6FF882" bold="true" box="[151,354,1853,1877]" pageId="6" pageNumber="26">FIGURES 2329.</emphasis>
<taxonomicName id="A8FEFCCCFFBF4D61C33DF8842871F882" box="[362,636,1854,1877]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="26" phylum="Arthropoda" rank="species" species="aspinosus" status="sp. nov.">
<emphasis id="5D8A5B5DFFBF4D61C33DF8842871F882" box="[362,636,1854,1877]" italics="true" pageId="6" pageNumber="26">Anoplodesmus aspinosus</emphasis>
</taxonomicName>
<emphasis id="5D8A5B5DFFBF4D61C0D4F88728EFF882" bold="true" box="[643,738,1853,1877]" pageId="6" pageNumber="26">
<taxonomicNameLabel id="46B9E626FFBF4D61C0D4F88728D0F882" box="[643,733,1853,1877]" pageId="6" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
paratypes from GuanU. 23, habitus of male and female (dorsal view); 24, anterior part of male body (dorsolateral view); 25 &amp; 26, middle part of male body (dorsal and lateral views, respectively); 27, caudal part of male body (lateral view); 28, telson (dorsal view); 29, sternal region of male midbody segments. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="3B81D7C7FFBE4D60C2C0F99F2963F9B7" httpUri="https://zenodo.org/record/194031/files/figure.png" pageId="7" pageNumber="27" targetBox="[160,1436,186,1559]" targetPageId="7">
<paragraph id="6F41874FFFBE4D60C2C0F99F2963F9B7" blockId="7.[151,1436,1573,1632]" pageId="7" pageNumber="27">
<emphasis id="5D8A5B5DFFBE4D60C2C0F99F2B6BF9EA" bold="true" box="[151,358,1573,1597]" pageId="7" pageNumber="27">FIGURES 3033.</emphasis>
<taxonomicName id="A8FEFCCCFFBE4D60C339F99C288AF9EA" box="[366,647,1574,1597]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="27" phylum="Arthropoda" rank="species" species="aspinosus" status="sp. nov.">
<emphasis id="5D8A5B5DFFBE4D60C339F99C288AF9EA" box="[366,647,1574,1597]" italics="true" pageId="7" pageNumber="27">Anoplodesmus aspinosus</emphasis>
</taxonomicName>
<emphasis id="5D8A5B5DFFBE4D60C0D8F99F28FFF9EA" bold="true" box="[655,754,1573,1597]" pageId="7" pageNumber="27">
<taxonomicNameLabel id="46B9E626FFBE4D60C0D8F99F28E1F9EA" box="[655,748,1573,1597]" pageId="7" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
right gonopod of male paratype from FuShan (dorsomedial, medial, ventral and lateral views, respectively). Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="6F41874FFFBE4D60C293F9312AC8F8C2" blockId="7.[151,1437,1675,2037]" pageId="7" pageNumber="27">
Sterna like in
<taxonomicName id="A8FEFCCCFFBE4D60C33DF9362BE0F972" box="[362,493,1676,1701]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="27" phylum="Arthropoda" rank="species" species="spiniger" status="sp. nov.">
<emphasis id="5D8A5B5DFFBE4D60C33DF9362BE0F972" box="[362,493,1676,1701]" italics="true" pageId="7" pageNumber="27">A. spiniger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46B9E626FFBE4D60C3A3F931285EF972" box="[500,595,1675,1701]" pageId="7" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
, but cones a little smaller (
<figureCitation id="F7C59BCAFFBE4D60C1C4F93129E1F972" box="[915,1004,1675,1701]" captionStart="FIGURES 23 29" captionStartId="6.[151,272,1853,1877]" captionTargetBox="[163,1434,190,1833]" captionTargetId="figure@6.[155,1440,183,1840]" captionTargetPageId="6" captionText="FIGURES 23 29. Anoplodesmus aspinosus sp. nov., paratypes from GuanU. 23, habitus of male and female (dorsal view); 24, anterior part of male body (dorsolateral view); 25 &amp; 26, middle part of male body (dorsal and lateral views, respectively); 27, caudal part of male body (lateral view); 28, telson (dorsal view); 29, sternal region of male midbody segments. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/194030/files/figure.png" pageId="7" pageNumber="27">Fig. 29</figureCitation>
); sternal lobe between male coxae 4 subtrapeziform (
<figureCitation id="F7C59BCAFFBE4D60C337F90B2BB6F91C" box="[352,443,1713,1739]" captionStart="FIGURES 17 22" captionStartId="5.[151,272,1435,1459]" captionTargetBox="[157,1431,302,1405]" captionTargetId="figure@5.[151,1436,279,1426]" captionTargetPageId="5" captionText="FIGURES 17 22. Anoplodesmus aspinosus sp. nov., male paratype from GuanU. 17, segment 10 (lateral view); 18, sternal lobe between coxae 4 (subventral view); 19 22, left gonopod (medial, ventral, lateral and dorsal views, respectively). Scale bar = 1.0 mm for figures 17 &amp; 18; 0.5 mm for figures 19 22." httpUri="https://zenodo.org/record/194029/files/figure.png" pageId="7" pageNumber="27">Fig. 18</figureCitation>
). Male legs with prefemora evidently incrassate dorsally and very densely setose ventrally, nearly all male tibiae and tarsi except several posteriormost ones with clear brushes (
<figureCitation id="F7C59BCAFFBE4D60C6A0F96C2F5FF927" box="[1271,1362,1750,1776]" captionStart="FIGURES 17 22" captionStartId="5.[151,272,1435,1459]" captionTargetBox="[157,1431,302,1405]" captionTargetId="figure@5.[151,1436,279,1426]" captionTargetPageId="5" captionText="FIGURES 17 22. Anoplodesmus aspinosus sp. nov., male paratype from GuanU. 17, segment 10 (lateral view); 18, sternal lobe between coxae 4 (subventral view); 19 22, left gonopod (medial, ventral, lateral and dorsal views, respectively). Scale bar = 1.0 mm for figures 17 &amp; 18; 0.5 mm for figures 19 22." httpUri="https://zenodo.org/record/194029/files/figure.png" pageId="7" pageNumber="27">Figs 17</figureCitation>
,
<figureCitation id="F7C59BCAFFBE4D60C737F96C2AB5F8C2" captionStart="FIGURES 23 29" captionStartId="6.[151,272,1853,1877]" captionTargetBox="[163,1434,190,1833]" captionTargetId="figure@6.[155,1440,183,1840]" captionTargetPageId="6" captionText="FIGURES 23 29. Anoplodesmus aspinosus sp. nov., paratypes from GuanU. 23, habitus of male and female (dorsal view); 24, anterior part of male body (dorsolateral view); 25 &amp; 26, middle part of male body (dorsal and lateral views, respectively); 27, caudal part of male body (lateral view); 28, telson (dorsal view); 29, sternal region of male midbody segments. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/194030/files/figure.png" pageId="7" pageNumber="27">26 &amp; 27</figureCitation>
).
</paragraph>
<paragraph id="6F41874FFFBE4D60C293F89B2F42F822" blockId="7.[151,1437,1675,2037]" pageId="7" pageNumber="27">
Gonopods (
<figureCitation id="F7C59BCAFFBE4D60C318F89B2BD7F8EC" box="[335,474,1824,1851]" captionStart="FIGURES 17 22" captionStartId="5.[151,272,1435,1459]" captionTargetBox="[157,1431,302,1405]" captionTargetId="figure@5.[151,1436,279,1426]" captionTargetPageId="5" captionText="FIGURES 17 22. Anoplodesmus aspinosus sp. nov., male paratype from GuanU. 17, segment 10 (lateral view); 18, sternal lobe between coxae 4 (subventral view); 19 22, left gonopod (medial, ventral, lateral and dorsal views, respectively). Scale bar = 1.0 mm for figures 17 &amp; 18; 0.5 mm for figures 19 22." httpUri="https://zenodo.org/record/194029/files/figure.png" pageId="7" pageNumber="27">
Figs 19
<emphasis id="5D8A5B5DFFBE4D60C3FDF89A2BB4F8ED" bold="true" box="[426,441,1824,1850]" pageId="7" pageNumber="27"></emphasis>
22
</figureCitation>
,
<figureCitation id="F7C59BCAFFBE4D60C3B1F89B283FF8EC" box="[486,562,1825,1851]" captionStart="FIGURES 30 33" captionStartId="7.[151,274,1573,1597]" captionTargetBox="[160,1436,186,1559]" captionTargetId="figure@7.[155,1440,181,1566]" captionTargetPageId="7" captionText="FIGURES 30 33. Anoplodesmus aspinosus sp. nov., right gonopod of male paratype from FuShan (dorsomedial, medial, ventral and lateral views, respectively). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/194031/files/figure.png" pageId="7" pageNumber="27">3033</figureCitation>
) very complex. Coxite elongate but relatively stout, subcylindrical, with only two setae distoventrally. Telopodite falcate; femorite evidently expanded both near base (
<emphasis id="5D8A5B5DFFBE4D60C6A4F8FC2F07F8B7" bold="true" box="[1267,1290,1862,1888]" pageId="7" pageNumber="27">lo</emphasis>
) and distad, about as long as solenophore, with a clear oblique impression on medial face and a similar longitudinal impression on lateral face; solenophore (
<emphasis id="5D8A5B5DFFBE4D60C022F82A28ACF87D" bold="true" box="[629,673,1936,1962]" pageId="7" pageNumber="27">sph</emphasis>
) distinctly set off from femorite by a lateral sulcus, twisted, with both a lamina medialis and, especially, a lamina lateralis well-developed and fringed, ending up in a large, complex, membranous structure, sheathing much but far from all of an extremely long solenomere (
<emphasis id="5D8A5B5DFFBE4D60C77CF8612F33F822" bold="true" box="[1323,1342,2011,2037]" pageId="7" pageNumber="27">sl</emphasis>
).
</paragraph>
<paragraph id="6F41874FFFB14D6FC293FF2D2E24FF65" blockId="8.[151,1436,151,290]" box="[196,1065,151,178]" pageId="8" pageNumber="28">
<emphasis id="5D8A5B5DFFB14D6FC293FF2D2B1AFF66" bold="true" box="[196,279,151,177]" pageId="8" pageNumber="28">Name:</emphasis>
To emphasize the absence of a distofemoral spine on the gonopod.
</paragraph>
</subSubSection>
<subSubSection id="27E4D4C4FFB14D6FC2C0FF072BF3F8D7" pageId="8" pageNumber="28" type="discussion">
<paragraph id="6F41874FFFB14D6FC2C0FF0729B2FEF5" blockId="8.[151,1436,151,290]" pageId="8" pageNumber="28">
<emphasis id="5D8A5B5DFFB14D6FC2C0FF072B19FF00" bold="true" box="[151,276,189,215]" pageId="8" pageNumber="28">Remarks:</emphasis>
<taxonomicName id="A8FEFCCCFFB14D6FC34AFF072BB8FF01" box="[285,437,189,214]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="28" phylum="Arthropoda" rank="species" species="aspinosus" status="sp. nov.">
<emphasis id="5D8A5B5DFFB14D6FC34AFF072BB8FF01" box="[285,437,189,214]" italics="true" pageId="8" pageNumber="28">A. aspinosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46B9E626FFB14D6FC3EAFF072817FF00" box="[445,538,189,215]" pageId="8" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
seems to populate only Taiwans northern and central parts, likely being a mid-montane species ranging between 500 and
<quantity id="A8062AAAFFB14D6FC080FF58293FFF2B" box="[727,818,226,252]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="28" unit="m" value="2000.0">2000 m</quantity>
a.s.l., as opposed to
<taxonomicName id="A8FEFCCCFFB14D6FC667FF582EB9FF2C" box="[1072,1204,226,251]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="28" phylum="Arthropoda" rank="species" species="spiniger" status="sp. nov.">
<emphasis id="5D8A5B5DFFB14D6FC667FF582EB9FF2C" box="[1072,1204,226,251]" italics="true" pageId="8" pageNumber="28">A. spiniger</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46B9E626FFB14D6FC6EAFF582F16FF2B" box="[1213,1307,226,252]" pageId="8" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
which has only been found in lowlands in the southern part of the island (Map).
</paragraph>
<paragraph id="6F41874FFFB14D6FC2C0F9722BF3F8D7" blockId="8.[151,1437,1736,1792]" pageId="8" pageNumber="28">
<emphasis id="5D8A5B5DFFB14D6FC2C0F9722AD3F937" bold="true" box="[151,222,1736,1760]" pageId="8" pageNumber="28">MAP.</emphasis>
Distribution of
<taxonomicName id="A8FEFCCCFFB14D6FC3F4F972285DF908" box="[419,592,1736,1759]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="5D8A5B5DFFB14D6FC3F4F972285DF908" box="[419,592,1736,1759]" italics="true" pageId="8" pageNumber="28">Anoplodesmus</emphasis>
</taxonomicName>
species in
<collectingCountry id="17E9C7DFFFB14D6FC08EF9722939F908" box="[729,820,1736,1759]" name="Taiwan" pageId="8" pageNumber="28">Taiwan</collectingCountry>
. Triangle,
<taxonomicName id="A8FEFCCCFFB14D6FC1E0F9722EC2F908" box="[951,1231,1736,1759]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="28" phylum="Arthropoda" rank="species" species="spiniger" status="sp. nov.">
<emphasis id="5D8A5B5DFFB14D6FC1E0F9722EC2F908" box="[951,1231,1736,1759]" italics="true" pageId="8" pageNumber="28">Anoplodesmus spiniger</emphasis>
</taxonomicName>
<emphasis id="5D8A5B5DFFB14D6FC68FF9722F34F937" bold="true" box="[1240,1337,1736,1760]" pageId="8" pageNumber="28">
<taxonomicNameLabel id="46B9E626FFB14D6FC68FF9722F34F937" box="[1240,1337,1736,1760]" pageId="8" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Square,
<taxonomicName id="A8FEFCCCFFB14D6FC2C0F9522BAFF928" box="[151,418,1768,1791]" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="28" phylum="Arthropoda" rank="species" species="aspinosus" status="sp. nov.">
<emphasis id="5D8A5B5DFFB14D6FC2C0F9522BAFF928" box="[151,418,1768,1791]" italics="true" pageId="8" pageNumber="28">Anoplodesmus aspinosus</emphasis>
</taxonomicName>
<emphasis id="5D8A5B5DFFB14D6FC3FEF9522BF3F8D7" bold="true" box="[425,510,1768,1792]" pageId="8" pageNumber="28">
<taxonomicNameLabel id="46B9E626FFB14D6FC3FEF9522BF3F8D7" box="[425,510,1768,1792]" pageId="8" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>