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<document id="8705A875E0D8BC9A3A432CFF16749ABC" ID-DOI="10.11646/zootaxa.4087.1.1" ID-GBIF-Dataset="17164bda-a943-496a-b044-1958db25c495" ID-ISSN="1175-5326" ID-Zenodo-Dep="399511" ID-ZooBank="C510CF71-0039-478A-91ED-BFD65B6FE0BE" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457771752054" checkinUser="plazi" docAuthor="Elena P. Kameneva &amp; Valery A. Kroneyev" docDate="2016" docId="5101BA35FFE3FFBAFF1AE568D464F817" docLanguage="en" docName="15545-5067-1-PB.pdf.imf" docOrigin="Zootaxa 4087 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Physiphora rugosa Kameneva &amp; Kroneyev, 2016, sp. n." docType="treatment" docVersion="9" lastPageNumber="68" masterDocId="AD38C24DFFA2FFF9FF8DE104D70CFF88" masterDocTitle="Revision of the Genus Physiphora Fallén 1810 (Diptera: Ulidiidae: Ulidiinae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="66" updateTime="1698184504803" updateUser="plazi">
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<mods:titleInfo id="391172011EECED7BC70903444CC1FBD6">
<mods:title id="48ACE03460A66F8D18CA03A484BBBBD8">Revision of the Genus Physiphora Fallén 1810 (Diptera: Ulidiidae: Ulidiinae)</mods:title>
</mods:titleInfo>
<mods:name id="0D6F7EA0A2A7BEC175ED14AD577FB428" type="personal">
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<mods:namePart id="6281922D65899B6E00F677F13D661B30">Elena P. Kameneva</mods:namePart>
</mods:name>
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<mods:namePart id="629472CEC1F501206F573342F33D3AD1">Valery A. Kroneyev</mods:namePart>
</mods:name>
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<mods:title id="B5DB8F3FA355215D4CFF3E83402833C8">Zootaxa</mods:title>
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<mods:date id="2F09823B85C33C8005867317D28EB16A">2016</mods:date>
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<mods:number id="2AD1AD6ED49B91DA1122A1DBCC99A44B">4087</mods:number>
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<mods:identifier id="44A12663E7A1C79EBD1BBB7090A037EB" type="DOI">10.11646/zootaxa.4087.1.1</mods:identifier>
<mods:identifier id="42DA28193D6114ADF59E7DC4594638B1" type="GBIF-Dataset">17164bda-a943-496a-b044-1958db25c495</mods:identifier>
<mods:identifier id="96D5DFA20C5479D66CE0843EF6769269" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5A1F78F4810EEA1F11299AAC205A328E" type="Zenodo-Dep">399511</mods:identifier>
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<treatment id="5101BA35FFE3FFBAFF1AE568D464F817" ID-DOI="http://doi.org/10.5281/zenodo.6066621" ID-GBIF-Taxon="117897957" ID-Zenodo-Dep="6066621" LSID="urn:lsid:plazi:treatment:5101BA35FFE3FFBAFF1AE568D464F817" httpUri="http://treatment.plazi.org/id/5101BA35FFE3FFBAFF1AE568D464F817" lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66">
<subSubSection id="91B258A8FFE3FFB8FF1AE568D650FB2F" pageId="65" pageNumber="66" type="nomenclature">
<paragraph id="D9170B23FFE3FFB8FF1AE568D6C9FB0E" blockId="65.[151,453,1132,1191]" box="[151,453,1132,1158]" pageId="65" pageNumber="66">
<heading id="825FBC4FFFE3FFB8FF1AE568D6C9FB0E" bold="true" box="[151,453,1132,1158]" fontSize="11" level="1" pageId="65" pageNumber="66" reason="1">
<emphasis id="EBDCD731FFE3FFB8FF1AE568D6C9FB0E" bold="true" box="[151,453,1132,1158]" pageId="65" pageNumber="66">
<taxonomicName id="1EA870A0FFE3FFB8FF1AE568D670FB0E" ID-CoL="8DC6M" box="[151,380,1132,1158]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="rugosa" status="sp. nov.">
<emphasis id="EBDCD731FFE3FFB8FF1AE568D670FB0E" bold="true" box="[151,380,1132,1158]" italics="true" pageId="65" pageNumber="66">Physiphora rugosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFFE3FFB8FE0EE568D6C9FB0E" box="[387,453,1132,1158]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D9170B23FFE3FFB8FF1AE58AD650FB2F" blockId="65.[151,453,1132,1191]" box="[151,348,1166,1191]" pageId="65" pageNumber="66">
<figureCitation id="419317A6FFE3FFB8FF1AE58AD65BFB2F" box="[151,343,1166,1191]" captionStart-0="FIGURES 313 316" captionStart-1="FIGURES 317 322" captionStartId-0="66.[151,264,1164,1185]" captionStartId-1="67.[151,264,1362,1383]" captionTargetBox-0="[186,1379,346,1109]" captionTargetBox-1="[159,1426,368,1337]" captionTargetId-0="figure@66.[151,1436,301,1142]" captionTargetId-1="figure@67.[151,1436,358,1339]" captionTargetPageId-0="66" captionTargetPageId-1="67" captionText-0="FIGURES 313 316. Physiphora rugosa: 313, habitus, left; 314, head and thorax, dorsal; 315, head, anterior; 316, wing. Scale 1 mm." captionText-1="FIGURES 317 322. Physiphora rugosa: 317, ♂ phallus (a — fragment of preglans, enlarged); 318, glans, enlarged; 319, epandrium, posterior; 320, epandrium, anterior; 321, hypandrium, ventral; 322, ejaculatory apodeme." httpUri-0="https://zenodo.org/record/399614/files/figure.png" httpUri-1="https://zenodo.org/record/399615/files/figure.png" pageId="65" pageNumber="66">Figures 313322</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="91B258A8FFE3FFB8FF1AE5D2D31EFAD4" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="D9170B23FFE3FFB8FF1AE5D2D31EFAD4" blockId="65.[151,1436,1238,2022]" pageId="65" pageNumber="66">
<emphasis id="EBDCD731FFE3FFB8FF1AE5D2D7F3FB67" bold="true" box="[151,255,1238,1263]" pageId="65" pageNumber="66">Material</emphasis>
.
<emphasis id="EBDCD731FFE3FFB8FE96E5D3D656FB66" bold="true" box="[283,346,1239,1262]" pageId="65" pageNumber="66">
<typeStatus id="0613B581FFE3FFB8FE96E5D3D658FB66" box="[283,340,1239,1262]" pageId="65" pageNumber="66">Type</typeStatus>
.
</emphasis>
<typeStatus id="0613B581FFE3FFB8FEFCE5D2D6D5FB67" box="[369,473,1238,1263]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
♂:
<emphasis id="EBDCD731FFE3FFB8FDADE5D2D5CBFB67" bold="true" box="[544,711,1238,1263]" pageId="65" pageNumber="66">
<collectingCountry id="A1BF4BB3FFE3FFB8FDADE5D2D5CBFB67" box="[544,711,1238,1263]" name="South Africa" pageId="65" pageNumber="66">South Africa</collectingCountry>
</emphasis>
: Eastern Cape: “Grahamstown / E.C.P. /
<date id="AD162DE3FFE3FFB8FB79E5D2D23FFB67" box="[1268,1331,1238,1263]" pageId="65" pageNumber="66">11.iii.</date>
[19]55 / P. S. Greathead”, “
<typeStatus id="0613B581FFE3FFB8FEE4E5FFD6DAFA9C" box="[361,470,1275,1300]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
”, “
<typeStatus id="0613B581FFE3FFB8FE74E5FFD56FFA9C" box="[505,611,1275,1300]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
<taxonomicName id="1EA870A0FFE3FFB8FDE6E5FFD4AFFA9C" authority="Steyskal" authorityName="Steyskal" box="[619,931,1275,1300]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="rugosa">Physiphora rugosa Steyskal</taxonomicName>
64” [red label] (BMNH).
<typeStatus id="0613B581FFE3FFB8FB54E5F8D246FA9B" box="[1241,1354,1276,1299]" pageId="65" pageNumber="66" type="paratype">Paratypes</typeStatus>
:
<emphasis id="EBDCD731FFE3FFB8FADBE5FFD7EFFAB0" bold="true" pageId="65" pageNumber="66">
<collectingCountry id="A1BF4BB3FFE3FFB8FADBE5FFD7EFFAB0" name="South Africa" pageId="65" pageNumber="66">South Africa</collectingCountry>
</emphasis>
:
<specimenCount id="CFAEC0AAFFE3FFB8FF74E41BD613FAB0" box="[249,287,1311,1336]" pageId="65" pageNumber="66" type="generic">1♂</specimenCount>
: “Cape Province, Matjesfontein, 1621.
<date id="AD162DE3FFE3FFB8FD62E41BD433FAB0" box="[751,831,1311,1336]" pageId="65" pageNumber="66" value="1928-10">x.1928</date>
”, “S. Africa, R. E. Turner, 1928—491” (BMNH) (dissected); 1♀: “Resolution / Albany Distr /
<date id="AD162DE3FFE3FFB8FD19E447D41DFAD3" box="[660,785,1347,1371]" pageId="65" pageNumber="66" value="1928-03-23">23.III.1928</date>
/ A.Walton” (USNM).
</paragraph>
</subSubSection>
<subSubSection id="91B258A8FFE3FFB8FF4AE463D4D4F816" pageId="65" pageNumber="66" type="diagnosis">
<paragraph id="D9170B23FFE3FFB8FF4AE463D4D4F816" blockId="65.[151,1436,1238,2022]" pageId="65" pageNumber="66">
<emphasis id="EBDCD731FFE3FFB8FF4AE463D636FA08" bold="true" box="[199,314,1383,1408]" pageId="65" pageNumber="66">Diagnosis</emphasis>
.
<taxonomicName id="1EA870A0FFE3FFB8FEC4E46CD6BFFAF7" box="[329,435,1384,1407]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="rugosa" status="sp. nov.">
<emphasis id="EBDCD731FFE3FFB8FEC4E46CD65BFAF7" box="[329,343,1384,1407]" italics="true" pageId="65" pageNumber="66">P</emphasis>
.
<emphasis id="EBDCD731FFE3FFB8FEE8E46CD6BFFAF7" box="[357,435,1384,1407]" italics="true" pageId="65" pageNumber="66">rugosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFFE3FFB8FE36E46CD6FBFA08" box="[443,503,1383,1408]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
is similar to
<taxonomicName id="1EA870A0FFE3FFB8FD08E46CD5F0FA08" box="[645,764,1383,1408]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="EBDCD731FFE3FFB8FD08E46CD59FFAF7" box="[645,659,1384,1407]" italics="true" pageId="65" pageNumber="66">P</emphasis>
.
<emphasis id="EBDCD731FFE3FFB8FD2FE463D5F0FA08" box="[674,764,1383,1408]" italics="true" pageId="65" pageNumber="66">obscura</emphasis>
</taxonomicName>
and
<taxonomicName id="1EA870A0FFE3FFB8FCB9E46CD493FAF7" box="[820,927,1384,1407]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="tarsata">
<emphasis id="EBDCD731FFE3FFB8FCB9E46CD493FAF7" box="[820,927,1384,1407]" italics="true" pageId="65" pageNumber="66">P. tarsata</emphasis>
</taxonomicName>
in the combination of the fore basitarsomere white, mesonotum densely rugulose, with shining black postpronotal lobe and margins of transverse suture, differing from them by the frons strongly wrinkled in anterior half (smooth in
<taxonomicName id="1EA870A0FFE3FFB8FBB4E4B4D3BFFA40" box="[1081,1203,1455,1480]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="EBDCD731FFE3FFB8FBB4E4B4D34BFA4F" box="[1081,1095,1456,1479]" italics="true" pageId="65" pageNumber="66">P</emphasis>
.
<emphasis id="EBDCD731FFE3FFB8FBD4E4ABD3BFFA40" box="[1113,1203,1455,1480]" italics="true" pageId="65" pageNumber="66">obscura</emphasis>
</taxonomicName>
, deeply pitted in
<taxonomicName id="1EA870A0FFE3FFB8FA0AE4B4D7EAFA63" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="tarsata">
<emphasis id="EBDCD731FFE3FFB8FA0AE4B4D7EAFA63" italics="true" pageId="65" pageNumber="66">P. tarsata</emphasis>
</taxonomicName>
) and mesonotum subshining green to violet on scutum and blue dull blue to black on scutellum; it also differs from
<taxonomicName id="1EA870A0FFE3FFB8FEA5E4FCD698F987" box="[296,404,1528,1551]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="tarsata">
<emphasis id="EBDCD731FFE3FFB8FEA5E4FCD698F987" box="[296,404,1528,1551]" italics="true" pageId="65" pageNumber="66">P. tarsata</emphasis>
</taxonomicName>
by the facial carina with λ-shaped white microtrichose area (in
<taxonomicName id="1EA870A0FFE3FFB8FBF9E4FCD3EDF987" box="[1140,1249,1528,1551]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="tarsata">
<emphasis id="EBDCD731FFE3FFB8FBF9E4FCD38EF987" box="[1140,1154,1528,1551]" italics="true" pageId="65" pageNumber="66">P</emphasis>
.
<emphasis id="EBDCD731FFE3FFB8FB1FE4FCD3EDF987" box="[1170,1249,1528,1551]" italics="true" pageId="65" pageNumber="66">tarsata</emphasis>
</taxonomicName>
face with entire transverse microtrichose band). It is similar to
<taxonomicName id="1EA870A0FFE3FFB8FD2EE718D414F9BB" box="[675,792,1564,1587]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="spriggsi" status="sp. nov.">
<emphasis id="EBDCD731FFE3FFB8FD2EE718D414F9BB" box="[675,792,1564,1587]" italics="true" pageId="65" pageNumber="66">P. spriggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFFE3FFB8FCADE718D450F9BB" box="[800,860,1564,1587]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
in having wrinkled or deeply pitted frons and wide facial rigge, clearly differing from that species by creamy fore metatarsus and facial carna with λ-shaped mark (respectively, black and entirely devoided microtrichia in
<taxonomicName id="1EA870A0FFE3FFB8FCADE760D49AF9F3" box="[800,918,1636,1659]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="spriggsi" status="sp. nov.">
<emphasis id="EBDCD731FFE3FFB8FCADE760D49AF9F3" box="[800,918,1636,1659]" italics="true" pageId="65" pageNumber="66">P. spriggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFFE3FFB8FC10E760D4D7F9F4" box="[925,987,1635,1660]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
).
<taxonomicName id="1EA870A0FFE3FFB8FC7CE760D3E8F9F4" authority="Macquart" authorityName="Macquart" box="[1009,1252,1635,1660]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="tarsata">
<emphasis id="EBDCD731FFE3FFB8FC7CE760D350F9F3" box="[1009,1116,1636,1659]" italics="true" pageId="65" pageNumber="66">P. tarsata</emphasis>
(Macquart)
</taxonomicName>
, another species with wrinkled or pitted frons and white fore metatarsus, differs by face with entire transverse microtrichose band and mesonotum subshining greenish, and frons deeply and widely pitted in anterior half and wrinkled posteriorly (uniformly deep wrinkled over whole frons in
<taxonomicName id="1EA870A0FFE3FFB8FD21E7D4D41BF96F" box="[684,791,1744,1767]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="rugosa" status="sp. nov.">
<emphasis id="EBDCD731FFE3FFB8FD21E7D4D5B6F96F" box="[684,698,1744,1767]" italics="true" pageId="65" pageNumber="66">P</emphasis>
.
<emphasis id="EBDCD731FFE3FFB8FD44E7D4D41BF96F" box="[713,791,1744,1767]" italics="true" pageId="65" pageNumber="66">rugosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFFE3FFB8FCADE7D4D453F960" box="[800,863,1743,1768]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
).
<taxonomicName id="1EA870A0FFE3FFB8FCFBE7D4D4EEF96F" box="[886,994,1744,1767]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="rugosa" status="sp. nov.">
<emphasis id="EBDCD731FFE3FFB8FCFBE7D4D488F96F" box="[886,900,1744,1767]" italics="true" pageId="65" pageNumber="66">P</emphasis>
.
<emphasis id="EBDCD731FFE3FFB8FC19E7D4D4EEF96F" box="[916,994,1744,1767]" italics="true" pageId="65" pageNumber="66">rugosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFFE3FFB8FC66E7D4D324F96F" box="[1003,1064,1744,1767]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
is similar to
<taxonomicName id="1EA870A0FFE3FFB8FB4FE7D4D224F96F" box="[1218,1320,1744,1767]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="aperta">
<emphasis id="EBDCD731FFE3FFB8FB4FE7D4D224F96F" box="[1218,1320,1744,1767]" italics="true" pageId="65" pageNumber="66">P. aperta</emphasis>
</taxonomicName>
in having wing with brownish veins and brownish yellow pterostigma, differing from it by narrowly closed cell r4+5 and different coloration of head and body.
<taxonomicName id="1EA870A0FFE3FFB8FDC1E61ED5BBF8B9" box="[588,695,1818,1841]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="rugosa" status="sp. nov.">
<emphasis id="EBDCD731FFE3FFB8FDC1E61ED5BBF8B9" box="[588,695,1818,1841]" italics="true" pageId="65" pageNumber="66">P. rugosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFFE3FFB8FD32E61ED5F0F8B9" box="[703,764,1818,1841]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
differs also from
<taxonomicName id="1EA870A0FFE3FFB8FC44E61ED34CF8B9" box="[969,1088,1818,1841]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="spriggsi" status="sp. nov.">
<emphasis id="EBDCD731FFE3FFB8FC44E61ED34CF8B9" box="[969,1088,1818,1841]" italics="true" pageId="65" pageNumber="66">P. spriggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFFE3FFB8FBC4E61ED38BF8B9" box="[1097,1159,1818,1841]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
, which often has rugose frons and face, by narrow facial carina with microtrichose pattern (in
<taxonomicName id="1EA870A0FFE3FFB8FC28E63BD317F8DE" box="[933,1051,1855,1878]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="spriggsi" status="sp. nov.">
<emphasis id="EBDCD731FFE3FFB8FC28E63BD317F8DE" box="[933,1051,1855,1878]" italics="true" pageId="65" pageNumber="66">P. spriggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFFE3FFB8FBAEE63BD36CF8DE" box="[1059,1120,1855,1878]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
, as wide as antennal groove and devoided of microtrichia) and subshining green mesonotum (in
<taxonomicName id="1EA870A0FFE3FFB8FC07E666D4F3F8F1" box="[906,1023,1890,1913]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="spriggsi" status="sp. nov.">
<emphasis id="EBDCD731FFE3FFB8FC07E666D4F3F8F1" box="[906,1023,1890,1913]" italics="true" pageId="65" pageNumber="66">P. spriggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFFE3FFB8FB88E666D34EF8F1" box="[1029,1090,1890,1913]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
, shining golden green), as well as preglans with wide black spines (in
<taxonomicName id="1EA870A0FFE3FFB8FDC5E683D5B1F816" box="[584,701,1927,1950]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="spriggsi" status="sp. nov.">
<emphasis id="EBDCD731FFE3FFB8FDC5E683D5B1F816" box="[584,701,1927,1950]" italics="true" pageId="65" pageNumber="66">P. spriggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0EF6A4AFFE3FFB8FD49E683D40DF816" box="[708,769,1927,1950]" pageId="65" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
, bare and smooth).
</paragraph>
</subSubSection>
<subSubSection id="91B258A8FFE3FFBAFF4AE6ADD53EF887" lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66" type="description">
<paragraph id="D9170B23FFE3FFBBFF4AE6ADD690FF7F" blockId="65.[151,1436,1238,2022]" lastBlockId="66.[151,1436,151,248]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<emphasis id="EBDCD731FFE3FFB8FF4AE6ADD65DF84A" bold="true" box="[199,337,1961,1986]" pageId="65" pageNumber="66">Description</emphasis>
. Head (
<figureCitation id="419317A6FFE3FFB8FE21E6ADD543F84A" box="[428,591,1961,1986]" captionStart="FIGURES 313 316" captionStartId="66.[151,264,1164,1185]" captionTargetBox="[186,1379,346,1109]" captionTargetId="figure@66.[151,1436,301,1142]" captionTargetPageId="66" captionText="FIGURES 313 316. Physiphora rugosa: 313, habitus, left; 314, head and thorax, dorsal; 315, head, anterior; 316, wing. Scale 1 mm." httpUri="https://zenodo.org/record/399614/files/figure.png" pageId="65" pageNumber="66">Figs. 314315</figureCitation>
) black, with dark brown frons, face and gena. Frons 1.01.1 times as long as wide, dark brown, subshining with moderately small round parafrontal microtichose spot not reaching anterior margin (
<figureCitation id="419317A6FFE0FFBBFF7AE193D655FF38" box="[247,345,151,176]" captionStart="FIGURES 313 316" captionStartId="66.[151,264,1164,1185]" captionTargetBox="[186,1379,346,1109]" captionTargetId="figure@66.[151,1436,301,1142]" captionTargetPageId="66" captionText="FIGURES 313 316. Physiphora rugosa: 313, habitus, left; 314, head and thorax, dorsal; 315, head, anterior; 316, wing. Scale 1 mm." httpUri="https://zenodo.org/record/399614/files/figure.png" pageId="66" pageNumber="67">Fig. 315</figureCitation>
), deeply wrinkled or pitted, without expressed calluses, flat or slightly convex, finely yellowish setulose. Vertical plates and ocellar triangle shining black; 2 pairs of black orbital setae; ocellar setae lateroclinate, as long as orbital setae.
</paragraph>
<caption id="8DD75BABFFE0FFBBFF1AE588D7D5FB48" httpUri="https://zenodo.org/record/399614/files/figure.png" pageId="66" pageNumber="67" targetBox="[186,1379,346,1109]" targetPageId="66">
<paragraph id="D9170B23FFE0FFBBFF1AE588D7D5FB48" blockId="66.[151,1436,1163,1216]" pageId="66" pageNumber="67">
<emphasis id="EBDCD731FFE0FFBBFF1AE588D660FB29" bold="true" box="[151,364,1164,1185]" pageId="66" pageNumber="67">FIGURES 313316.</emphasis>
<taxonomicName id="1EA870A0FFE0FFBBFEFEE58FD538FB29" box="[371,564,1163,1185]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="EBDCD731FFE0FFBBFEFEE58FD538FB29" box="[371,564,1163,1185]" italics="true" pageId="66" pageNumber="67">Physiphora rugosa</emphasis>
</taxonomicName>
: 313, habitus, left; 314, head and thorax, dorsal; 315, head, anterior; 316, wing. Scale 1 mm.
</paragraph>
</caption>
<paragraph id="D9170B23FFE0FFBBFF4AE5EAD6BAF9CD" blockId="66.[151,1437,1260,1995]" pageId="66" pageNumber="67">
Face brown, facial carina very wide, with steep lateral margins and wrinkled surface, brown or brown with black pattern, with white λ-shaped microtrichose area widely separated from microtrichose antennal grooves; epistome wrinkled, shining black, lateral sides of face moderately high, black. Gena brown, half as high as eye; facial ridge and parafacial subshining brown, more or less wrinkled, each with narrow white microtrichose stripe; gena posteriorly without microtrichose mark separating it from partly brownish yellow postgena (
<figureCitation id="419317A6FFE0FFBBFAA5E47ED29CFA1B" box="[1320,1424,1402,1427]" captionStart="FIGURES 313 316" captionStartId="66.[151,264,1164,1185]" captionTargetBox="[186,1379,346,1109]" captionTargetId="figure@66.[151,1436,301,1142]" captionTargetPageId="66" captionText="FIGURES 313 316. Physiphora rugosa: 313, habitus, left; 314, head and thorax, dorsal; 315, head, anterior; 316, wing. Scale 1 mm." httpUri="https://zenodo.org/record/399614/files/figure.png" pageId="66" pageNumber="67">Fig. 313</figureCitation>
). Occiput black with partly brown postgenae. Medial vertical seta 0.4 times as long as frons width, 1.5 times as long as lateral vertical, 4 times as long as ocellar, orbital, and 2.5 times as long as postocellar setae. Antenna yellowish brown; flagellomere 1 laterally brown, rounded apically, 1.82 times as long as wide, greyish microtrichose; arista bare, yellowish brown in basal 1/6, remainder black. Clypeus black. Palp black, grey microtrichose and black setose. Mouthparts black.
</paragraph>
<paragraph id="D9170B23FFE0FFBBFF4AE74BD525F80C" blockId="66.[151,1437,1260,1995]" pageId="66" pageNumber="67">
Thorax (
<figureCitation id="419317A6FFE0FFBBFEA7E74BD682F9E0" box="[298,398,1615,1640]" captionStart="FIGURES 313 316" captionStartId="66.[151,264,1164,1185]" captionTargetBox="[186,1379,346,1109]" captionTargetId="figure@66.[151,1436,301,1142]" captionTargetPageId="66" captionText="FIGURES 313 316. Physiphora rugosa: 313, habitus, left; 314, head and thorax, dorsal; 315, head, anterior; 316, wing. Scale 1 mm." httpUri="https://zenodo.org/record/399614/files/figure.png" pageId="66" pageNumber="67">Fig. 314</figureCitation>
). Scutum black, roughly rugulose, subshining, with golden-green to green (in
<typeStatus id="0613B581FFE0FFBBFA9DE74BD27FF9E0" box="[1296,1395,1615,1640]" pageId="66" pageNumber="67" type="holotype">holotype</typeStatus>
) or cyan to deep violet sheen; antepronotum, postpronotal lobe, posterior surface of notopleural triangle, supra-alar and postalar parts of scutum, anterior half of anepisternum and katepisternum, and whole anepimeron shining black; posterodorsal parts of anepisternum and katepisternum rugulose; postscutellum black, gray microtrichose; posteroventral margin of scutellum without microtrichose area; anatergite, katatergite and lateroventral parts of mediotergite sparsely grey microtrichose; medial part of mediotergite finely shagreened, with green or golden reflection. Scutum with acrostichal, dorsocentral, and intra-alar rows of setulae; acrostichal seta indistinguishable or lacking; dorsocentral setae hair-like; other setae moderately long, black: one postprononal, 2 postsutural supraalar, one intra-alar and one postalar.
</paragraph>
<paragraph id="D9170B23FFE0FFBBFF4AE68AD418F842" blockId="66.[151,1437,1260,1995]" pageId="66" pageNumber="67">Scutellum densely rugulose, usually with deep blue reflection or pitchy black, apparently bare or with indistinguishable setulae; 2 pairs of black scutellar setae.</paragraph>
<paragraph id="D9170B23FFE1FFBAFF4AE193D7C6FE97" blockId="67.[151,1436,151,287]" pageId="67" pageNumber="68">
Wing. Pale yellowish or hyaline, with partly brownish veins; pterostigma brownish yellow, cell r4+5 narrowly closed; apical section of M slightly arcuate (
<figureCitation id="419317A6FFE1FFBAFD0BE1BAD5EBFF5F" box="[646,743,190,215]" captionStart="FIGURES 313 316" captionStartId="66.[151,264,1164,1185]" captionTargetBox="[186,1379,346,1109]" captionTargetId="figure@66.[151,1436,301,1142]" captionTargetPageId="66" captionText="FIGURES 313 316. Physiphora rugosa: 313, habitus, left; 314, head and thorax, dorsal; 315, head, anterior; 316, wing. Scale 1 mm." httpUri="https://zenodo.org/record/399614/files/figure.png" pageId="67" pageNumber="68">Fig. 316</figureCitation>
). Postero-apical extension of cell cup 1.41.5 times as long as vein A1+CuA2, and 3 times as long as transverse section of vein CuA2. Calypters with white fringe. Length: 3.43.6 mm.
</paragraph>
<caption id="8DD75BABFFE1FFBAFF1AE456D394FA0E" httpUri="https://zenodo.org/record/399615/files/figure.png" pageId="67" pageNumber="68" targetBox="[159,1426,368,1337]" targetPageId="67">
<paragraph id="D9170B23FFE1FFBAFF1AE456D394FA0E" blockId="67.[151,1435,1361,1414]" pageId="67" pageNumber="68">
<emphasis id="EBDCD731FFE1FFBAFF1AE456D67CFAEF" bold="true" box="[151,368,1362,1383]" pageId="67" pageNumber="68">FIGURES 317322.</emphasis>
<taxonomicName id="1EA870A0FFE1FFBAFEF7E455D532FAEF" box="[378,574,1361,1383]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="EBDCD731FFE1FFBAFEF7E455D532FAEF" box="[378,574,1361,1383]" italics="true" pageId="67" pageNumber="68">Physiphora rugosa</emphasis>
</taxonomicName>
: 317, ♂ phallus (a — fragment of preglans, enlarged); 318, glans, enlarged; 319, epandrium, posterior; 320, epandrium, anterior; 321, hypandrium, ventral; 322, ejaculatory apodeme.
</paragraph>
</caption>
<paragraph id="D9170B23FFE1FFBAFF4AE4B0D3BEF99C" blockId="67.[151,1437,1459,1951]" pageId="67" pageNumber="68">Legs. Black except fore tarsus with basitarsomere creamy white in basal 4/5; mid- and hind femora and tibiae entirely brown or black; mid and hind tarsi yellow except 2 or 3 last tarsomere dark brown; all setae black; fore femur not swollen, postero-ventrally with 56 almost non-thickened short setae in apical half.</paragraph>
<paragraph id="D9170B23FFE1FFBAFF4AE71AD4C2F9D4" blockId="67.[151,1437,1459,1951]" pageId="67" pageNumber="68">Abdomen. Both tergites and sternites shining black, laterally finely shagreened, with slight golden reflection; all setulae black; abdominal tergite 1 basally sparsely grey microtrichose.</paragraph>
<paragraph id="D9170B23FFE1FFBAFF4AE762D512F964" blockId="67.[151,1437,1459,1951]" pageId="67" pageNumber="68">
<collectingCountry id="A1BF4BB3FFE1FFBAFF4AE762D60DF9F7" box="[199,257,1638,1663]" name="Malta" pageId="67" pageNumber="68">Male</collectingCountry>
postabdomen brown to black; epandrium as on
<figureCitation id="419317A6FFE1FFBAFC97E762D476F9F7" box="[794,890,1638,1663]" captionStart="FIGURES 317 322" captionStartId="67.[151,264,1362,1383]" captionTargetBox="[159,1426,368,1337]" captionTargetId="figure@67.[151,1436,358,1339]" captionTargetPageId="67" captionText="FIGURES 317 322. Physiphora rugosa: 317, ♂ phallus (a — fragment of preglans, enlarged); 318, glans, enlarged; 319, epandrium, posterior; 320, epandrium, anterior; 321, hypandrium, ventral; 322, ejaculatory apodeme." httpUri="https://zenodo.org/record/399615/files/figure.png" pageId="67" pageNumber="68">Fig. 319</figureCitation>
, cerci with short triangular nipple-like structures (
<figureCitation id="419317A6FFE1FFBAFF12E78FD60CF92C" box="[159,256,1675,1700]" captionStart="FIGURES 317 322" captionStartId="67.[151,264,1362,1383]" captionTargetBox="[159,1426,368,1337]" captionTargetId="figure@67.[151,1436,358,1339]" captionTargetPageId="67" captionText="FIGURES 317 322. Physiphora rugosa: 317, ♂ phallus (a — fragment of preglans, enlarged); 318, glans, enlarged; 319, epandrium, posterior; 320, epandrium, anterior; 321, hypandrium, ventral; 322, ejaculatory apodeme." httpUri="https://zenodo.org/record/399615/files/figure.png" pageId="67" pageNumber="68">Fig. 320</figureCitation>
), phallus with stipe slightly longer than preglans and glans; preglans with rough black spines or swellings (
<figureCitation id="419317A6FFE1FFBAFF12E7AAD60EF94F" box="[159,258,1710,1735]" captionStart="FIGURES 317 322" captionStartId="67.[151,264,1362,1383]" captionTargetBox="[159,1426,368,1337]" captionTargetId="figure@67.[151,1436,358,1339]" captionTargetPageId="67" captionText="FIGURES 317 322. Physiphora rugosa: 317, ♂ phallus (a — fragment of preglans, enlarged); 318, glans, enlarged; 319, epandrium, posterior; 320, epandrium, anterior; 321, hypandrium, ventral; 322, ejaculatory apodeme." httpUri="https://zenodo.org/record/399615/files/figure.png" pageId="67" pageNumber="68">Fig. 317</figureCitation>
a); glans with 56 short lobes (
<figureCitation id="419317A6FFE1FFBAFD0EE7AAD5E6F94F" box="[643,746,1710,1735]" captionStart="FIGURES 317 322" captionStartId="67.[151,264,1362,1383]" captionTargetBox="[159,1426,368,1337]" captionTargetId="figure@67.[151,1436,358,1339]" captionTargetPageId="67" captionText="FIGURES 317 322. Physiphora rugosa: 317, ♂ phallus (a — fragment of preglans, enlarged); 318, glans, enlarged; 319, epandrium, posterior; 320, epandrium, anterior; 321, hypandrium, ventral; 322, ejaculatory apodeme." httpUri="https://zenodo.org/record/399615/files/figure.png" pageId="67" pageNumber="68">Fig. 318</figureCitation>
). Hypandrium (
<figureCitation id="419317A6FFE1FFBAFC2EE7AAD30BF94F" box="[931,1031,1710,1735]" captionStart="FIGURES 317 322" captionStartId="67.[151,264,1362,1383]" captionTargetBox="[159,1426,368,1337]" captionTargetId="figure@67.[151,1436,358,1339]" captionTargetPageId="67" captionText="FIGURES 317 322. Physiphora rugosa: 317, ♂ phallus (a — fragment of preglans, enlarged); 318, glans, enlarged; 319, epandrium, posterior; 320, epandrium, anterior; 321, hypandrium, ventral; 322, ejaculatory apodeme." httpUri="https://zenodo.org/record/399615/files/figure.png" pageId="67" pageNumber="68">Fig. 321</figureCitation>
) asymmetric, with well developed triangular vanes of phallapodeme).
</paragraph>
<paragraph id="D9170B23FFE1FFBAFF4AE7F2D53EF887" blockId="67.[151,1437,1459,1951]" box="[199,562,1782,1807]" pageId="67" pageNumber="68">Female terminalia not dissected.</paragraph>
</subSubSection>
<subSubSection id="91B258A8FFE1FFBAFF4AE61FD6F3F8BC" box="[199,511,1819,1844]" pageId="67" pageNumber="68" type="distribution">
<paragraph id="D9170B23FFE1FFBAFF4AE61FD6F3F8BC" blockId="67.[151,1437,1459,1951]" box="[199,511,1819,1844]" pageId="67" pageNumber="68">
<emphasis id="EBDCD731FFE1FFBAFF4AE61FD655F8BC" bold="true" box="[199,345,1819,1844]" pageId="67" pageNumber="68">Distribution</emphasis>
.
<collectingCountry id="A1BF4BB3FFE1FFBAFEEAE61FD6F6F8BC" box="[359,506,1819,1844]" name="South Africa" pageId="67" pageNumber="68">South Africa</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="91B258A8FFE1FFBAFF4AE63AD694F8DF" box="[199,408,1854,1879]" pageId="67" pageNumber="68" type="biology_ecology">
<paragraph id="D9170B23FFE1FFBAFF4AE63AD694F8DF" blockId="67.[151,1437,1459,1951]" box="[199,408,1854,1879]" pageId="67" pageNumber="68">
<emphasis id="EBDCD731FFE1FFBAFF4AE63AD62DF8DF" bold="true" box="[199,289,1854,1879]" pageId="67" pageNumber="68">Biology</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="91B258A8FFE1FFBAFF4AE660D464F817" pageId="67" pageNumber="68" type="etymology">
<paragraph id="D9170B23FFE1FFBAFF4AE660D464F817" blockId="67.[151,1437,1459,1951]" pageId="67" pageNumber="68">
<emphasis id="EBDCD731FFE1FFBAFF4AE660D64AF8F3" bold="true" box="[199,326,1892,1915]" pageId="67" pageNumber="68">Etymology</emphasis>
. Specific epithet
<taxonomicName id="1EA870A0FFE1FFBAFD84E660D55BF8F3" box="[521,599,1892,1915]" class="Insecta" family="Ulidiidae" genus="Physiphora" kingdom="Animalia" order="Diptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="EBDCD731FFE1FFBAFD84E660D55BF8F3" box="[521,599,1892,1915]" italics="true" pageId="67" pageNumber="68">rugosa</emphasis>
</taxonomicName>
means “wrinkled” in Latin and reflects the structure of frons and face. The unpublished name originally proposed by G. C. Steyskal is used.
</paragraph>
</subSubSection>
</treatment>
</document>