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<mods:title id="D99E7766754915D6519A623715FDB9AE">Two new species of Cyrtodactylus (Squamata: Gekkonidae) from northern Laos including new finding and expanded diagnosis of C. bansocensis</mods:title>
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<mods:namePart id="E7AAF71A2A1B501A46B6DEDB87D93077">Luu, Vinh Quang</mods:namePart>
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<mods:namePart id="D623190550BA0381E2DEEB4BEAC62E87">Sitthivong, Saly</mods:namePart>
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<mods:namePart id="3178BE97727CDDBB1DB5A10BAD936F25">Le, Minh Duc</mods:namePart>
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<mods:affiliation id="D3B7051CDC28927ABC7FDF3335E3D83D">Faculty of Environmental Sciences, University of Science, Vietnam National University Hanoi, 334 Nguyen Trai Road, Hanoi, Vietnam. le. duc. minh @ hus. edu. vn; https: // orcid. org / 0000 - 0002 - 2953 - 2815. &amp; Central Institute for Natural Resources and Environmental Studies, Vietnam National University Hanoi, 19 Le Thanh Tong, Hanoi, Vietnam. &amp; Department of Herpetology, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam. nqt 2 @ yahoo. com; https: // orcid. org / 0000 - 0002 - 6601 - 0880</mods:affiliation>
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<mods:namePart id="64A2FB7E83C5ED3F9F392E228EAC01AC">Nguyen, Truong Quang</mods:namePart>
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Redescription of 
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<emphasis id="99CBF407BF27FFBC293C8848FD7DAD41" bold="true" box="[367,686,294,321]" italics="true" pageId="17" pageNumber="520">Cyrtodactylus bansocensis</emphasis>
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including the first record of a female
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<emphasis id="99CBF407BF27FFBC28C488FDFE92ADAD" box="[151,321,403,429]" italics="true" pageId="17" pageNumber="520">C. bansocensis</emphasis>
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was described based on two adult males from 
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(
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Luu 
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2016c
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on the new specimen collected in about 
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distance from the 
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locality, including for the first time a female individual, the specimen is described in the following including an expanded diagnosis of the species
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.
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<emphasis id="99CBF407BF27FFBC28948890FE6DAE18" bold="true" box="[199,446,510,536]" pageId="17" pageNumber="520">Specimen examined.</emphasis>
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adult female 
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.2016.4 (
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) collected by 
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in 
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from 
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, 
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(
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, 
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, at an elevation of 
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a.s.l.
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.
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<emphasis id="99CBF407BF27FFBC28948B28FE2FAE61" bold="true" box="[199,508,582,609]" pageId="17" pageNumber="520">Morphological characters</emphasis>
. Adult female, SVL 
<quantity id="6C4785F0BF27FFBC2AB08B29FC99AE61" box="[739,842,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="6.38" pageId="17" pageNumber="520" unit="mm" value="63.8">63.8 mm</quantity>
; tail length 
<quantity id="6C4785F0BF27FFBC2B9F8B29FBE0AE61" box="[972,1075,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="9.459999999999999" pageId="17" pageNumber="520" unit="mm" value="94.6">94.6 mm</quantity>
. The examined specimen differs from the original description of 
<bibRefCitation id="CF2E55E4BF27FFBC2A538B05FD1FAE86" author="Luu, V. Q. &amp; Nguyen, T. Q. &amp; Le, M. D. &amp; Bonkowski, M. &amp; Ziegler, T." box="[512,716,619,646]" pageId="17" pageNumber="520" pagination="87 - 102" refId="ref17490" refString="Luu, V. Q., Nguyen, T. Q., Le, M. D., Bonkowski, M. &amp; Ziegler, T. (2016 c) A new species of karst-dwelling bent-toed gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa, 4079 (1), 87 - 102. https: // doi. org / 10.11646 / zootaxa. 4079.1.6" type="journal article" year="2016">
Luu 
<emphasis id="99CBF407BF27FFBC2A668B05FDB5AE85" box="[565,614,619,645]" italics="true" pageId="17" pageNumber="520">et al</emphasis>
. (2016c)
</bibRefCitation>
by having more supralabials and infralabials (11 or 12 vs 8–10; 10 vs. 8, respectively), more dorsal tubercle rows (20 vs. 14 or 15) and more scales around midbody (102 vs. 86 or 87). For further morphological data see 
<tableCitation id="E63D1DAEBF27FFBC2A058BDDFD7FAECD" box="[598,684,691,717]" captionStart="TABLE 6" captionStartId="17.[151,239,764,788]" captionText="TABLE 6. Selected morphological characters and scalation of the female specimen of Cyrtodactylus bansocensis. For abbreviations see Material and Methods. Measurements in mm; * = character given right/left." pageId="17" pageNumber="520">Table 6</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="FFC0789DBF27FFBC28C48B92FBAEAF38" pageId="17" pageNumber="520">
<paragraph id="AB002815BF27FFBC28C48B92FBAEAF38" blockId="17.[151,1436,764,825]" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28C48B92FED8AF14" bold="true" box="[151,267,764,788]" pageId="17" pageNumber="520">TABLE 6.</emphasis>
Selected morphological characters and scalation of the female specimen of 
<taxonomicName id="6CBF5396BF27FFBC2C1A8B92FABBAF14" authorityName="Luu, Nguyen, Le, Bonkowski &amp; Ziegler" authorityYear="2016" box="[1097,1384,764,788]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="bansocensis">
<emphasis id="99CBF407BF27FFBC2C1A8B92FABBAF14" box="[1097,1384,764,788]" italics="true" pageId="17" pageNumber="520">Cyrtodactylus bansocensis</emphasis>
</taxonomicName>
. For abbreviations see Material and Methods. Measurements in mm; * = character given right/left.
</paragraph>
</caption>
<paragraph id="AB002815BF27FFBC299D8A27FB5BAA85" pageId="17" pageNumber="520">
<table id="D9BFDAB5BF27005228F58A27FAC2AA86" box="[166,1297,841,1670]" gridcols="4" gridrows="21" pageId="17" pageNumber="520">
<tr id="158F2A57BF27005228F58A27FAC2AF61" box="[166,1297,841,865]" gridrow="0" pageId="17" pageNumber="520" rowspan-0="1" rowspan-2="1">
<th id="565E432BBF270052299D8A27FDA4AF61" box="[462,631,841,865]" gridcol="1" gridrow="0" pageId="17" pageNumber="520">VNUF R.2016.4</th>
<th id="565E432BBF2700522C3B8A27FAC2AF61" box="[1128,1297,841,865]" gridcol="3" gridrow="0" pageId="17" pageNumber="520">VNUF R.2016.4</th>
</tr>
<tr id="158F2A57BF27005228F58A1EFAC2AF89" box="[166,1297,880,905]" gridrow="1" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58A1EFF25AF89" box="[166,246,880,905]" gridcol="0" gridrow="1" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58A1EFF1FAF88" bold="true" box="[166,204,880,904]" pageId="17" pageNumber="520">Sex</emphasis>
</th>
<td id="565E432BBF270052299D8A1EFDA4AF89" box="[462,631,880,905]" gridcol="1" gridrow="1" pageId="17" pageNumber="520">Female</td>
<td id="565E432BBF2700522B428A1EFCBEAF89" box="[785,877,880,905]" gridcol="2" gridrow="1" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428A1FFCE9AF89" bold="true" box="[785,826,881,905]" pageId="17" pageNumber="520">ML</emphasis>
</td>
<td id="565E432BBF2700522C3B8A1EFAC2AF89" box="[1128,1297,880,905]" gridcol="3" gridrow="1" pageId="17" pageNumber="520">2.7</td>
</tr>
<tr id="158F2A57BF27005228F58AF6FAC2AFB1" box="[166,1297,920,945]" gridrow="2" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58AF6FF25AFB1" box="[166,246,920,945]" gridcol="0" gridrow="2" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58AF6FF04AFB0" bold="true" box="[166,215,920,944]" pageId="17" pageNumber="520">SVL</emphasis>
</th>
<td id="565E432BBF270052299D8AF6FDA4AFB1" box="[462,631,920,945]" gridcol="1" gridrow="2" pageId="17" pageNumber="520">63.8</td>
<td id="565E432BBF2700522B428AF6FCBEAFB1" box="[785,877,920,945]" gridcol="2" gridrow="2" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428AF6FC9FAFB0" bold="true" box="[785,844,920,944]" pageId="17" pageNumber="520">SPL*</emphasis>
</td>
<td id="565E432BBF2700522C3B8AF6FAC2AFB1" box="[1128,1297,920,945]" gridcol="3" gridrow="2" pageId="17" pageNumber="520">11/12</td>
</tr>
<tr id="158F2A57BF27005228F58AAFFAC2AFDA" box="[166,1297,961,986]" gridrow="3" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58AAFFF25AFDA" box="[166,246,961,986]" gridcol="0" gridrow="3" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58AAFFF01AFD9" bold="true" box="[166,210,961,985]" pageId="17" pageNumber="520">TaL</emphasis>
</th>
<td id="565E432BBF270052299D8AAFFDA4AFDA" box="[462,631,961,986]" gridcol="1" gridrow="3" pageId="17" pageNumber="520">94.6</td>
<td id="565E432BBF2700522B428AAFFCBEAFDA" box="[785,877,961,986]" gridcol="2" gridrow="3" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428AAFFC9BAFD9" bold="true" box="[785,840,961,985]" pageId="17" pageNumber="520">IFL*</emphasis>
</td>
<td id="565E432BBF2700522C3B8AAFFAC2AFDA" box="[1128,1297,961,986]" gridcol="3" gridrow="3" pageId="17" pageNumber="520">10/10</td>
</tr>
<tr id="158F2A57BF27005228F58A87FAC2A802" box="[166,1297,1001,1026]" gridrow="4" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58A87FF25A802" box="[166,246,1001,1026]" gridcol="0" gridrow="4" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58A87FF1FA801" bold="true" box="[166,204,1001,1025]" pageId="17" pageNumber="520">AG</emphasis>
</th>
<td id="565E432BBF270052299D8A87FDA4A802" box="[462,631,1001,1026]" gridcol="1" gridrow="4" pageId="17" pageNumber="520">26.4</td>
<td id="565E432BBF2700522B428A87FCBEA802" box="[785,877,1001,1026]" gridcol="2" gridrow="4" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428A87FCE3A801" bold="true" box="[785,816,1001,1025]" pageId="17" pageNumber="520">N*</emphasis>
</td>
<td id="565E432BBF2700522C3B8A87FAC2A802" box="[1128,1297,1001,1026]" gridcol="3" gridrow="4" pageId="17" pageNumber="520">3/3</td>
</tr>
<tr id="158F2A57BF27005228F58D7CFAC2A82A" box="[166,1297,1042,1066]" gridrow="5" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58D7CFF25A82A" box="[166,246,1042,1066]" gridcol="0" gridrow="5" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58D7CFF18A82A" bold="true" box="[166,203,1042,1066]" pageId="17" pageNumber="520">HL</emphasis>
</th>
<td id="565E432BBF270052299D8D7CFDA4A82A" box="[462,631,1042,1066]" gridcol="1" gridrow="5" pageId="17" pageNumber="520">17.8</td>
<td id="565E432BBF2700522B428D7CFCBEA82A" box="[785,877,1042,1066]" gridcol="2" gridrow="5" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428D7CFCC8A82A" bold="true" box="[785,795,1042,1066]" pageId="17" pageNumber="520">I</emphasis>
</td>
<td id="565E432BBF2700522C3B8D7CFAC2A82A" box="[1128,1297,1042,1066]" gridcol="3" gridrow="5" pageId="17" pageNumber="520">0</td>
</tr>
<tr id="158F2A57BF27005228F58D57FAC2A852" box="[166,1297,1081,1106]" gridrow="6" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58D57FF25A852" box="[166,246,1081,1106]" gridcol="0" gridrow="6" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58D54FF00A852" bold="true" box="[166,211,1082,1106]" pageId="17" pageNumber="520">HW</emphasis>
</th>
<td id="565E432BBF270052299D8D57FDA4A852" box="[462,631,1081,1106]" gridcol="1" gridrow="6" pageId="17" pageNumber="520">13.1</td>
<td id="565E432BBF2700522B428D57FCBEA852" box="[785,877,1081,1106]" gridcol="2" gridrow="6" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428D57FCBEA851" bold="true" box="[785,877,1081,1105]" pageId="17" pageNumber="520">SC5SPL</emphasis>
</td>
<td id="565E432BBF2700522C3B8D57FAC2A852" box="[1128,1297,1081,1106]" gridcol="3" gridrow="6" pageId="17" pageNumber="520">35</td>
</tr>
<tr id="158F2A57BF27005228F58D0CFAC2A87A" box="[166,1297,1122,1146]" gridrow="7" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58D0CFF25A87A" box="[166,246,1122,1146]" gridcol="0" gridrow="7" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58D0CFF1EA87A" bold="true" box="[166,205,1122,1146]" pageId="17" pageNumber="520">HH</emphasis>
</th>
<td id="565E432BBF270052299D8D0CFDA4A87A" box="[462,631,1122,1146]" gridcol="1" gridrow="7" pageId="17" pageNumber="520">7.8</td>
<td id="565E432BBF2700522B428D0CFCBEA87A" box="[785,877,1122,1146]" gridcol="2" gridrow="7" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428D0CFCFCA87A" bold="true" box="[785,815,1122,1146]" pageId="17" pageNumber="520">IO</emphasis>
</td>
<td id="565E432BBF2700522C3B8D0CFAC2A87A" box="[1128,1297,1122,1146]" gridcol="3" gridrow="7" pageId="17" pageNumber="520">35</td>
</tr>
<tr id="158F2A57BF27005228F58DE4FAC2A8A3" box="[166,1297,1162,1187]" gridrow="8" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58DE4FF25A8A3" box="[166,246,1162,1187]" gridcol="0" gridrow="8" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58DE4FF16A8A2" bold="true" box="[166,197,1162,1186]" pageId="17" pageNumber="520">SE</emphasis>
</th>
<td id="565E432BBF270052299D8DE4FDA4A8A3" box="[462,631,1162,1187]" gridcol="1" gridrow="8" pageId="17" pageNumber="520">8.1</td>
<td id="565E432BBF2700522B428DE4FCBEA8A3" box="[785,877,1162,1187]" gridcol="2" gridrow="8" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428DE4FCEBA8A2" bold="true" box="[785,824,1162,1186]" pageId="17" pageNumber="520">PM</emphasis>
</td>
<td id="565E432BBF2700522C3B8DE4FAC2A8A3" box="[1128,1297,1162,1187]" gridcol="3" gridrow="8" pageId="17" pageNumber="520">2</td>
</tr>
<tr id="158F2A57BF27005228F58DDCFAC2A8CB" box="[166,1297,1202,1227]" gridrow="9" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58DDCFF25A8CB" box="[166,246,1202,1227]" gridcol="0" gridrow="9" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58DDDFF1BA8CB" bold="true" box="[166,200,1203,1227]" pageId="17" pageNumber="520">EE</emphasis>
</th>
<td id="565E432BBF270052299D8DDCFDA4A8CB" box="[462,631,1202,1227]" gridcol="1" gridrow="9" pageId="17" pageNumber="520">5.1</td>
<td id="565E432BBF2700522B428DDCFCBEA8CB" box="[785,877,1202,1227]" gridcol="2" gridrow="9" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428DDCFCE7A8CA" bold="true" box="[785,820,1202,1226]" pageId="17" pageNumber="520">GP</emphasis>
</td>
<td id="565E432BBF2700522C3B8DDCFAC2A8CB" box="[1128,1297,1202,1227]" gridcol="3" gridrow="9" pageId="17" pageNumber="520">7</td>
</tr>
<tr id="158F2A57BF27005228F58DB5FAC2A8F3" box="[166,1297,1243,1267]" gridrow="10" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58DB5FF25A8F3" box="[166,246,1243,1267]" gridcol="0" gridrow="10" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58DB5FF07A8F3" bold="true" box="[166,212,1243,1267]" pageId="17" pageNumber="520">IND</emphasis>
</th>
<td id="565E432BBF270052299D8DB5FDA4A8F3" box="[462,631,1243,1267]" gridcol="1" gridrow="10" pageId="17" pageNumber="520">2.1</td>
<td id="565E432BBF2700522B428DB5FCBEA8F3" box="[785,877,1243,1267]" gridcol="2" gridrow="10" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428DB5FC95A8F3" bold="true" box="[785,838,1243,1267]" pageId="17" pageNumber="520">DTR</emphasis>
</td>
<td id="565E432BBF2700522C3B8DB5FAC2A8F3" box="[1128,1297,1243,1267]" gridcol="3" gridrow="10" pageId="17" pageNumber="520">20</td>
</tr>
<tr id="158F2A57BF27005228F58C6DFAC2A91B" box="[166,1297,1283,1307]" gridrow="11" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58C6DFF25A91B" box="[166,246,1283,1307]" gridcol="0" gridrow="11" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58C6DFF05A91B" bold="true" box="[166,214,1283,1307]" pageId="17" pageNumber="520">IOD</emphasis>
</th>
<td id="565E432BBF270052299D8C6DFDA4A91B" box="[462,631,1283,1307]" gridcol="1" gridrow="11" pageId="17" pageNumber="520">6.0</td>
<td id="565E432BBF2700522B428C6DFCBEA91B" box="[785,877,1283,1307]" gridcol="2" gridrow="11" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428C6DFC85A91B" bold="true" box="[785,854,1283,1307]" pageId="17" pageNumber="520">GSDT</emphasis>
</td>
<td id="565E432BBF2700522C3B8C6DFAC2A91B" box="[1128,1297,1283,1307]" gridcol="3" gridrow="11" pageId="17" pageNumber="520">8–9</td>
</tr>
<tr id="158F2A57BF27005228F58C45FAC2A944" box="[166,1297,1323,1348]" gridrow="12" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58C45FF25A944" box="[166,246,1323,1348]" gridcol="0" gridrow="12" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58C45FF1FA943" bold="true" box="[166,204,1323,1347]" pageId="17" pageNumber="520">OD</emphasis>
</th>
<td id="565E432BBF270052299D8C45FDA4A944" box="[462,631,1323,1348]" gridcol="1" gridrow="12" pageId="17" pageNumber="520">5.0</td>
<td id="565E432BBF2700522B428C45FCBEA944" box="[785,877,1323,1348]" gridcol="2" gridrow="12" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428C45FC9AA943" bold="true" box="[785,841,1323,1347]" pageId="17" pageNumber="520">SMC</emphasis>
</td>
<td id="565E432BBF2700522C3B8C45FAC2A944" box="[1128,1297,1323,1348]" gridcol="3" gridrow="12" pageId="17" pageNumber="520">204</td>
</tr>
<tr id="158F2A57BF27005228F58C3DFAC2A96C" box="[166,1297,1363,1388]" gridrow="13" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58C3DFF25A96C" box="[166,246,1363,1388]" gridcol="0" gridrow="13" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58C3AFF1AA96C" bold="true" box="[166,201,1364,1388]" pageId="17" pageNumber="520">ED</emphasis>
</th>
<td id="565E432BBF270052299D8C3DFDA4A96C" box="[462,631,1363,1388]" gridcol="1" gridrow="13" pageId="17" pageNumber="520">1.8</td>
<td id="565E432BBF2700522B428C3DFCBEA96C" box="[785,877,1363,1388]" gridcol="2" gridrow="13" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428C3DFCE4A96B" bold="true" box="[785,823,1363,1387]" pageId="17" pageNumber="520">MS</emphasis>
</td>
<td id="565E432BBF2700522C3B8C3DFAC2A96C" box="[1128,1297,1363,1388]" gridcol="3" gridrow="13" pageId="17" pageNumber="520">102</td>
</tr>
<tr id="158F2A57BF27005228F58C12FAC2A994" box="[166,1297,1404,1428]" gridrow="14" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58C12FF25A994" box="[166,246,1404,1428]" gridcol="0" gridrow="14" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58C12FF25A994" bold="true" box="[166,246,1404,1428]" pageId="17" pageNumber="520">ForeaL</emphasis>
</th>
<td id="565E432BBF270052299D8C12FDA4A994" box="[462,631,1404,1428]" gridcol="1" gridrow="14" pageId="17" pageNumber="520">10.6</td>
<td id="565E432BBF2700522B428C12FCBEA994" box="[785,877,1404,1428]" gridcol="2" gridrow="14" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428C12FCF0A994" bold="true" box="[785,803,1404,1428]" pageId="17" pageNumber="520">V</emphasis>
</td>
<td id="565E432BBF2700522C3B8C12FAC2A994" box="[1128,1297,1404,1428]" gridcol="3" gridrow="14" pageId="17" pageNumber="520">35</td>
</tr>
<tr id="158F2A57BF27005228F58CCAFAC2A9BC" box="[166,1297,1444,1468]" gridrow="15" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58CCAFF25A9BC" box="[166,246,1444,1468]" gridcol="0" gridrow="15" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58CCAFF16A9BC" bold="true" box="[166,197,1444,1468]" pageId="17" pageNumber="520">SL</emphasis>
</th>
<td id="565E432BBF270052299D8CCAFDA4A9BC" box="[462,631,1444,1468]" gridcol="1" gridrow="15" pageId="17" pageNumber="520">13.7</td>
<td id="565E432BBF2700522B428CCAFCBEA9BC" box="[785,877,1444,1468]" gridcol="2" gridrow="15" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428CCAFC9EA9BC" bold="true" box="[785,845,1444,1468]" pageId="17" pageNumber="520">LD1*</emphasis>
</td>
<td id="565E432BBF2700522C3B8CCAFAC2A9BC" box="[1128,1297,1444,1468]" gridcol="3" gridrow="15" pageId="17" pageNumber="520">15/15</td>
</tr>
<tr id="158F2A57BF27005228F58CA2FAC2A9E5" box="[166,1297,1484,1509]" gridrow="16" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58CA2FF25A9E5" box="[166,246,1484,1509]" gridcol="0" gridrow="16" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58CA2FF0DA9E4" bold="true" box="[166,222,1484,1508]" pageId="17" pageNumber="520">LeF4</emphasis>
</th>
<td id="565E432BBF270052299D8CA2FDA4A9E5" box="[462,631,1484,1509]" gridcol="1" gridrow="16" pageId="17" pageNumber="520">6.0</td>
<td id="565E432BBF2700522B428CA2FCBEA9E5" box="[785,877,1484,1509]" gridcol="2" gridrow="16" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428CA2FC9DA9E4" bold="true" box="[785,846,1484,1508]" pageId="17" pageNumber="520">LD4*</emphasis>
</td>
<td id="565E432BBF2700522C3B8CA2FAC2A9E5" box="[1128,1297,1484,1509]" gridcol="3" gridrow="16" pageId="17" pageNumber="520">18/20</td>
</tr>
<tr id="158F2A57BF27005228F58C9AFAC2AA0D" box="[166,1297,1524,1549]" gridrow="17" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58C9AFF25AA0D" box="[166,246,1524,1549]" gridcol="0" gridrow="17" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58C9AFF0CAA0C" bold="true" box="[166,223,1524,1548]" pageId="17" pageNumber="520">LeT4</emphasis>
</th>
<td id="565E432BBF270052299D8C9AFDA4AA0D" box="[462,631,1524,1549]" gridcol="1" gridrow="17" pageId="17" pageNumber="520">7.9</td>
<td id="565E432BBF2700522B428C9AFCBEAA0D" box="[785,877,1524,1549]" gridcol="2" gridrow="17" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428C9AFC99AA0C" bold="true" box="[785,842,1524,1548]" pageId="17" pageNumber="520">LT1*</emphasis>
</td>
<td id="565E432BBF2700522C3B8C9AFAC2AA0D" box="[1128,1297,1524,1549]" gridcol="3" gridrow="17" pageId="17" pageNumber="520">16/15</td>
</tr>
<tr id="158F2A57BF27005228F58F72FAC2AA35" box="[166,1297,1564,1589]" gridrow="18" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58F72FF25AA35" box="[166,246,1564,1589]" gridcol="0" gridrow="18" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58F73FF02AA35" bold="true" box="[166,209,1565,1589]" pageId="17" pageNumber="520">RW</emphasis>
</th>
<td id="565E432BBF270052299D8F72FDA4AA35" box="[462,631,1564,1589]" gridcol="1" gridrow="18" pageId="17" pageNumber="520">2.7</td>
<td id="565E432BBF2700522B428F72FCBEAA35" box="[785,877,1564,1589]" gridcol="2" gridrow="18" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428F72FC9AAA34" bold="true" box="[785,841,1564,1588]" pageId="17" pageNumber="520">LT4*</emphasis>
</td>
<td id="565E432BBF2700522C3B8F72FAC2AA35" box="[1128,1297,1564,1589]" gridcol="3" gridrow="18" pageId="17" pageNumber="520">21/20</td>
</tr>
<tr id="158F2A57BF27005228F58F2BFAC2AA5D" box="[166,1297,1605,1629]" gridrow="19" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58F2BFF25AA5D" box="[166,246,1605,1629]" gridcol="0" gridrow="19" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58F2BFF1FAA5D" bold="true" box="[166,204,1605,1629]" pageId="17" pageNumber="520">RH</emphasis>
</th>
<td id="565E432BBF270052299D8F2BFDA4AA5D" box="[462,631,1605,1629]" gridcol="1" gridrow="19" pageId="17" pageNumber="520">1.6</td>
<td id="565E432BBF2700522B428F2BFCBEAA5D" box="[785,877,1605,1629]" gridcol="2" gridrow="19" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428F2BFC8DAA5D" bold="true" box="[785,862,1605,1629]" pageId="17" pageNumber="520">PP+FP</emphasis>
</td>
<td id="565E432BBF2700522C3B8F2BFAC2AA5D" box="[1128,1297,1605,1629]" gridcol="3" gridrow="19" pageId="17" pageNumber="520">16</td>
</tr>
<tr id="158F2A57BF27005228F58F03FAC2AA86" box="[166,1297,1645,1670]" gridrow="20" pageId="17" pageNumber="520">
<th id="565E432BBF27005228F58F03FF25AA86" box="[166,246,1645,1670]" gridcol="0" gridrow="20" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28F58F03FF04AA85" bold="true" box="[166,215,1645,1669]" pageId="17" pageNumber="520">MW</emphasis>
</th>
<td id="565E432BBF270052299D8F03FDA4AA86" box="[462,631,1645,1670]" gridcol="1" gridrow="20" pageId="17" pageNumber="520">3.2</td>
<td id="565E432BBF2700522B428F03FCBEAA86" box="[785,877,1645,1670]" gridcol="2" gridrow="20" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC2B428F03FC9EAA85" bold="true" box="[785,845,1645,1669]" pageId="17" pageNumber="520">PAT*</emphasis>
</td>
<td id="565E432BBF2700522C3B8F03FAC2AA86" box="[1128,1297,1645,1670]" gridcol="3" gridrow="20" pageId="17" pageNumber="520">4/5</td>
</tr>
</table>
</paragraph>
<subSubSection id="E3A57B9EBF27FFBC28948FDBFB9CABCC" pageId="17" pageNumber="520" type="diagnosis">
<paragraph id="AB002815BF27FFBC28948FDBFB9CABCC" blockId="17.[151,1436,1717,2032]" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28948FDBFE6DAACF" bold="true" box="[199,446,1717,1743]" pageId="17" pageNumber="520">Expanded diagnosis.</emphasis>
As the original description of 
<taxonomicName id="6CBF5396BF27FFBC2B4B8FDBFB90AACF" authorityName="Luu, Nguyen, Le, Bonkowski &amp; Ziegler" authorityYear="2016" box="[792,1091,1717,1743]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="bansocensis">
<emphasis id="99CBF407BF27FFBC2B4B8FDBFB90AACF" box="[792,1091,1717,1743]" italics="true" pageId="17" pageNumber="520">Cyrtodactylus bansocensis</emphasis>
</taxonomicName>
was based on two adult males (
<bibRefCitation id="CF2E55E4BF27FFBC28CC8FB7FE8AAAF4" author="Luu, V. Q. &amp; Nguyen, T. Q. &amp; Le, M. D. &amp; Bonkowski, M. &amp; Ziegler, T." box="[159,345,1753,1780]" pageId="17" pageNumber="520" pagination="87 - 102" refId="ref17490" refString="Luu, V. Q., Nguyen, T. Q., Le, M. D., Bonkowski, M. &amp; Ziegler, T. (2016 c) A new species of karst-dwelling bent-toed gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa, 4079 (1), 87 - 102. https: // doi. org / 10.11646 / zootaxa. 4079.1.6" type="journal article" year="2016">
Luu 
<emphasis id="99CBF407BF27FFBC28878FB4FEDEAAF3" box="[212,269,1753,1779]" italics="true" pageId="17" pageNumber="520">et al.</emphasis>
2016c
</bibRefCitation>
) we herein expand the diagnosis of this species as follows: medium size, SVL reaching 74.0 mm; dorsal pattern with four light transverse bands between limb insertions; supranasals in contact; 14–20 irregular dorsal tubercle rows at midbody; lateral skin fold present without tubercles; 34 or 35 ventral scale rows between ventrolateral folds; 34 precloacal and femoral pores in a continuous row in males (or interrupted by poreless scales); 16 precloacal and femoral pores in a continuous row in females (two outer femoral pores just pitted on each side), enlarged femoral and precloacal scales present; 4–7 postcloacal tubercles on each side; dorsal tubercles present at base of tail; subcaudal scales transversely enlarged (modified after 
<bibRefCitation id="CF2E55E4BF27FFBC2BD48EDCFB90ABCC" author="Luu, V. Q. &amp; Nguyen, T. Q. &amp; Le, M. D. &amp; Bonkowski, M. &amp; Ziegler, T." box="[903,1091,1969,1996]" pageId="17" pageNumber="520" pagination="87 - 102" refId="ref17490" refString="Luu, V. Q., Nguyen, T. Q., Le, M. D., Bonkowski, M. &amp; Ziegler, T. (2016 c) A new species of karst-dwelling bent-toed gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa, 4079 (1), 87 - 102. https: // doi. org / 10.11646 / zootaxa. 4079.1.6" type="journal article" year="2016">
Luu 
<emphasis id="99CBF407BF27FFBC2BE88EDCFC20ABCB" box="[955,1011,1969,1995]" italics="true" pageId="17" pageNumber="520">et al.</emphasis>
2016c
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E3A57B9EBF27FFBF28948EBBFF3EACD5" lastPageId="18" lastPageNumber="521" pageId="17" pageNumber="520" type="biology_ecology">
<paragraph id="AB002815BF27FFBF28948EBBFF3EACD5" blockId="17.[151,1436,1717,2032]" lastBlockId="18.[151,1437,150,285]" lastPageId="18" lastPageNumber="521" pageId="17" pageNumber="520">
<emphasis id="99CBF407BF27FFBC28948EBBFE18ABEF" bold="true" box="[199,459,2005,2031]" pageId="17" pageNumber="520">Natural history notes.</emphasis>
The female specimen was collected at night (20:20) on a limestone cliff, ca. 
<quantity id="6C4785F0BF27FFBC2D738EBBFAB3ABF0" box="[1312,1376,2005,2032]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="520" unit="m" value="1.5">1.5 m</quantity>
from the ground. The surrounding habitat was karst forest. The relative humidity was 52% and the air temperature was 25.5°C.
</paragraph>
</subSubSection>
<subSubSection id="E3A57B9EBF24FFBF289489B0FE75AD1D" pageId="18" pageNumber="521" type="distribution">
<paragraph id="AB002815BF24FFBF289489B0FE75AD1D" blockId="18.[151,1437,150,285]" pageId="18" pageNumber="521">
<emphasis id="99CBF407BF24FFBF289489B0FEB2ACF8" bold="true" box="[199,353,222,248]" pageId="18" pageNumber="521">Distribution.</emphasis>
The female specimen of 
<taxonomicName id="6CBF5396BF24FFBF2A2489B1FC72ACF9" authorityName="Luu, Nguyen, Le, Bonkowski &amp; Ziegler" authorityYear="2016" box="[631,929,223,249]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="521" phylum="Chordata" rank="species" species="bansocensis">
<emphasis id="99CBF407BF24FFBF2A2489B1FC72ACF9" box="[631,929,223,249]" italics="true" pageId="18" pageNumber="521">Cyrtodactylus bansocensis</emphasis>
</taxonomicName>
was found close to the 
<typeStatus id="740496B7BF24FFBF2CF089B1FB00ACF9" box="[1187,1235,223,249]" pageId="18" pageNumber="521">type</typeStatus>
locality in 
<collectingRegion id="697BE6F7BF24FFBF2D1D89B1FE8EAD1D" country="Laos" name="Khammouan" pageId="18" pageNumber="521">Khammouane Province</collectingRegion>
, 
<collectingCountry id="D3A86885BF24FFBF293B886DFE71AD1D" box="[360,418,259,285]" name="Laos" pageId="18" pageNumber="521">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>