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<document id="1F0928808212FB03A64BC86FC41CA185" ID-CLB-Dataset="8764" ID-DOI="10.11646/zootaxa.4822.4.3" ID-GBIF-Dataset="67b665b5-74e4-4d65-b375-a07f33a7aa75" ID-ISSN="1175-5326" ID-Zenodo-Dep="4452362" ID-ZooBank="4C7F87B9-D1C1-4821-AB37-BBA3FA59F55C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609339354294" checkinUser="plazi" docAuthor="Schneider, Nicole, Luu, Vinh Quang, Sitthivong, Saly, Teynié, Alexandre, Le, Minh Duc, Nguyen, Truong Quang &amp; Ziegler, Thomas" docDate="2020" docId="23169903BF24FFBE28C48C5DFA4FAF8A" docLanguage="en" docName="zootaxa.4822.4.3.pdf" docOrigin="Zootaxa 4822 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus Gray 1827" docType="key" docVersion="8" lastPageNumber="522" masterDocId="DF2FE17BBF36FFAD2853896EFFD3AC00" masterDocTitle="Two new species of Cyrtodactylus (Squamata: Gekkonidae) from northern Laos including new finding and expanded diagnosis of C. bansocensis" masterLastPageNumber="530" masterPageNumber="503" pageNumber="521" updateTime="1732485091614" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CFCBC4E75EA08544B359BF29F3EB0435">Two new species of Cyrtodactylus (Squamata: Gekkonidae) from northern Laos including new finding and expanded diagnosis of C. bansocensis</mods:title>
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<mods:namePart id="6611985E5EB3B1802DBA3E19834CDD0C">Schneider, Nicole</mods:namePart>
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<mods:affiliation id="32CD94C10774A9EE1F3252EEC06B0175">AG Zoologischer Garten Köln, Riehler Strasse 173, D- 50735 Cologne, Germany. https: // orcid. org / 0000 - 0002 - 4797 - 609 X. &amp; Institute of Zoology, University of Cologne, Zülpicher Strasse 47 b, D- 50674 Cologne, Germany. schneider. nicole 87 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6588 - 229 X.</mods:affiliation>
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<mods:namePart id="95A0F5E0A2FDBF47A6A4187907FFA26D">Luu, Vinh Quang</mods:namePart>
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<mods:affiliation id="9B7F50257B918BB0DA21A0C0D881716A">Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam. qvinhfuv @ yahoo. com. au; https: // orcid. org / 0000 - 0002 - 0634 - 1338.</mods:affiliation>
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<mods:namePart id="74FBB7D21B039D120EB4CAFCF2C671EE">Sitthivong, Saly</mods:namePart>
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<mods:affiliation id="39F07A0A912F0A47CD1050416AC415A9">Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam. qvinhfuv @ yahoo. com. au; https: // orcid. org / 0000 - 0002 - 0634 - 1338. &amp; Faculty of Forestry, National University of Laos, Dong Dok Campus, Vientiane, Lao PDR. sitthivong _ frt @ yahoo. co. th; https: // orcid. org / 0000 - 0002 - 9993 - 2136</mods:affiliation>
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<mods:namePart id="F14D15C9FC36ACC3F59070690A649CB8">Teynié, Alexandre</mods:namePart>
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<mods:affiliation id="4C4E38467906DD727B6A2AA7EAFC4982">Société d’Histoire Naturelle Alcide d’Orbigny, 57, rue de Gergovie, 63170 Aubière, France. ateynie @ shnao. net; https: // orcid. org / 0000 - 0002 - 6588 - 229 X.</mods:affiliation>
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<mods:namePart id="FDE70DB52592D2B68ED2CB031A8C53CD">Le, Minh Duc</mods:namePart>
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<mods:affiliation id="F0291E930123270681D8DC7A3A0CFE2A">Faculty of Environmental Sciences, University of Science, Vietnam National University Hanoi, 334 Nguyen Trai Road, Hanoi, Vietnam. le. duc. minh @ hus. edu. vn; https: // orcid. org / 0000 - 0002 - 2953 - 2815. &amp; Central Institute for Natural Resources and Environmental Studies, Vietnam National University Hanoi, 19 Le Thanh Tong, Hanoi, Vietnam. &amp; Department of Herpetology, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam. nqt 2 @ yahoo. com; https: // orcid. org / 0000 - 0002 - 6601 - 0880</mods:affiliation>
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<mods:namePart id="9796B914B53DEB4DC6873D6A37641227">Nguyen, Truong Quang</mods:namePart>
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<mods:namePart id="4B9E5787949557B10E834E1630CEAB0E">Ziegler, Thomas</mods:namePart>
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<mods:affiliation id="9A7040ACE3E4666D99CD0448F20C1369">AG Zoologischer Garten Köln, Riehler Strasse 173, D- 50735 Cologne, Germany. https: // orcid. org / 0000 - 0002 - 4797 - 609 X. &amp; Institute of Zoology, University of Cologne, Zülpicher Strasse 47 b, D- 50674 Cologne, Germany. schneider. nicole 87 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6588 - 229 X.</mods:affiliation>
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<mods:title id="B6195CA594318DD7EEF61B96D7DB21A1">Zootaxa</mods:title>
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<mods:date id="F6BDAF299ED56CF52B5A89B056FEDB3D">2020</mods:date>
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<mods:number id="1F4894DE6283AC15380EAF2051CF3BE6">2020-08-07</mods:number>
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<mods:number id="863B9D9D2DFC0C4C9BBAAF344CBCB276">4822</mods:number>
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<mods:identifier id="2DC497E6F826263C36C09A689ADF50AB" type="DOI">10.11646/zootaxa.4822.4.3</mods:identifier>
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<treatment id="23169903BF24FFBE28C48C5DFA4FAF8A" ID-DOI="http://doi.org/10.5281/zenodo.4452366" ID-GBIF-Taxon="177010390" ID-Zenodo-Dep="4452366" LSID="urn:lsid:plazi:treatment:23169903BF24FFBE28C48C5DFA4FAF8A" httpUri="http://treatment.plazi.org/id/23169903BF24FFBE28C48C5DFA4FAF8A" lastPageId="19" lastPageNumber="522" pageId="18" pageNumber="521" scope_class="Squamata" scope_family="Gekkonidae">
<subSubSection id="E3A57B9EBF24FFBF28C48C5DFC5FA94D" box="[151,908,1330,1357]" pageId="18" pageNumber="521" type="nomenclature">
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<heading id="F0489F79BF24FFBF28C48C5DFC5FA94D" bold="true" box="[151,908,1330,1357]" fontSize="11" level="1" pageId="18" pageNumber="521" reason="1">
<emphasis id="99CBF407BF24FFBF28C48C5DFC5FA94D" bold="true" box="[151,908,1330,1357]" pageId="18" pageNumber="521">
Key to Laotian species of
<taxonomicName id="6CBF5396BF24FFBF29848C5CFDAEA94D" ID-CoL="CD3HY" ID-ENA="96732" authorityName="Gray" authorityYear="1827" box="[471,637,1330,1357]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="521" phylum="Chordata" rank="genus">
<emphasis id="99CBF407BF24FFBF29848C5CFDAEA94D" bold="true" box="[471,637,1330,1357]" italics="true" pageId="18" pageNumber="521">Cyrtodactylus</emphasis>
</taxonomicName>
(referring to
<tableCitation id="E63D1DAEBF24FFBF2B748C5DFC50A94D" box="[807,899,1331,1357]" captionStart="TABLE 2" captionStartId="6.[151,239,465,491]" captionTargetBox="[165,2018,614,1682]" captionText="TABLE 2. Morphological characters, distribution and habitat information of all Laotian Cyrtodactylus species. Abbreviations are as follows: SVL: snout–vent length (in mm); TaL: Tail length (in mm); SPL: Supralabials; IFL: Infralabials; SMC: along the midbody from mental shield to anterior edge of cloaca; MS: midbody scales; V: ventral scales; DTR: dorsal tubercle rows; EPS: enlarged precloacal scales; EFS: enlarged femoral scales; FP: femoral pores; PP: precloacal pores; LD4/LT4: subdigital lamellae under fourth finger/toe; *=regenerated tail;?=unknown character." httpUri="http://table.plazi.org/id/FFC0789DBF30FFAB28C488BFFD1AAE57" pageId="18" pageNumber="521" tableUuid="FFC0789DBF30FFAB28C488BFFD1AAE57">Table 2</tableCitation>
)
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<paragraph id="AB002815BF24FFBF28C48C16FA4FA98D" blockId="18.[151,1437,1400,2038]" box="[151,1436,1400,1422]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48C16FA4FA98D" box="[151,1436,1400,1422]" pageId="18" pageNumber="521">
1 Precloacal-femoral pores in males in separated series..........................................
<taxonomicName id="6CBF5396BF24FFBF2C8B8C16FA4FA98D" authorityName="Sitthivong, Luu, Ha, Nguyen, Le &amp; Ziegler" authorityYear="2019" box="[1240,1436,1400,1421]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="521" phylum="Chordata" rank="species" species="muangfuangensis">
<emphasis id="99CBF407BF24FFBF2C8B8C16FA4FA98D" box="[1240,1436,1400,1421]" italics="true" pageId="18" pageNumber="521">C. muangfuangensis</emphasis>
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</paragraph>
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<keyLead id="104E8B20BF24FFBF28C48CF8FA4FA9AA" box="[151,1436,1429,1451]" pageId="18" pageNumber="521">- Precloacal-femoral pores in males in contiguous series or precloacal-femoral pores absent in males.................... 2</keyLead>
</paragraph>
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<keyLead id="104E8B20BF24FFBF28C48CDDFA4FA9C8" box="[151,1436,1459,1481]" pageId="18" pageNumber="521">2 Enlarged subcaudals present............................................................................. 3</keyLead>
</paragraph>
<paragraph id="AB002815BF24FFBF28C48CBEFA4FA9E6" blockId="18.[151,1437,1400,2038]" box="[151,1436,1488,1510]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48CBEFA4FA9E6" box="[151,1436,1488,1510]" pageId="18" pageNumber="521">- Enlarged subcaudals absent............................................................................ 15</keyLead>
</paragraph>
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<keyStep id="104B30B0BF24FFBF28C48C83FA4FAA20" pageId="18" pageNumber="521">
<paragraph id="AB002815BF24FFBF28C48C83FA4FAA02" blockId="18.[151,1437,1400,2038]" box="[151,1436,1517,1539]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48C83FA4FAA02" box="[151,1436,1517,1539]" pageId="18" pageNumber="521">3 Precloacal-femoral pores present in males.................................................................. 4</keyLead>
</paragraph>
<paragraph id="AB002815BF24FFBF28C48F65FA4FAA20" blockId="18.[151,1437,1400,2038]" box="[151,1436,1547,1569]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48F65FA4FAA20" box="[151,1436,1547,1569]" pageId="18" pageNumber="521">- Precloacal-femoral pores absent or unknown in males....................................................... 19</keyLead>
</paragraph>
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<keyStep id="104B30B0BF24FFBF28C48F46FA4FAA5B" pageId="18" pageNumber="521">
<paragraph id="AB002815BF24FFBF28C48F46FA4FAA3E" blockId="18.[151,1437,1400,2038]" box="[151,1436,1576,1598]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48F46FA4FAA3E" box="[151,1436,1576,1598]" pageId="18" pageNumber="521">4 Dorsum blotched..................................................................................... 5</keyLead>
</paragraph>
<paragraph id="AB002815BF24FFBF28C48F28FA4FAA5B" blockId="18.[151,1437,1400,2038]" box="[151,1436,1605,1627]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48F28FA4FAA5B" box="[151,1436,1605,1627]" pageId="18" pageNumber="521">- Dorsum banded....................................................................................... 6</keyLead>
</paragraph>
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<paragraph id="AB002815BF24FFBF28C48F0CFA4FAA78" blockId="18.[151,1437,1400,2038]" box="[151,1436,1634,1657]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48F0CFA4FAA78" box="[151,1436,1634,1657]" pageId="18" pageNumber="521">
5 Precloacal pores absent in females, Supralabials 11–16................................................
<taxonomicName id="6CBF5396BF24FFBF2D6B8F0DFA4FAA78" authorityName="Ulber" authorityYear="1993" box="[1336,1436,1635,1656]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="521" phylum="Chordata" rank="species" species="jarujini">
<emphasis id="99CBF407BF24FFBF2D6B8F0DFA4FAA78" box="[1336,1436,1635,1656]" italics="true" pageId="18" pageNumber="521">C. jarujini</emphasis>
</taxonomicName>
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</paragraph>
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<keyLead id="104E8B20BF24FFBF28C48FEEFA4FAA95" box="[151,1436,1664,1686]" pageId="18" pageNumber="521">
- Precloacal pores in females 0–14, Supralabials 9–11................................................
<taxonomicName id="6CBF5396BF24FFBF2D4B8FEEFA4FAA95" authorityName="Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov &amp; Chulisov" authorityYear="2014" box="[1304,1436,1664,1685]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="521" phylum="Chordata" rank="species" species="multiporus">
<emphasis id="99CBF407BF24FFBF2D4B8FEEFA4FAA95" box="[1304,1436,1664,1685]" italics="true" pageId="18" pageNumber="521">C. multiporus</emphasis>
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<keyLead id="104E8B20BF24FFBF28C48FF3FA4FAAB2" box="[151,1436,1693,1715]" pageId="18" pageNumber="521">
6 Ventrals 27–29................................................................................
<taxonomicName id="6CBF5396BF24FFBF2D6E8FF0FA4FAAB2" authorityName="Luu, Calame, Nguyen, Le, Bonkowski &amp; Ziegler" authorityYear="2016" box="[1341,1436,1693,1715]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="521" phylum="Chordata" rank="species" species="rufford">
<emphasis id="99CBF407BF24FFBF2D6E8FF0FA4FAAB2" box="[1341,1436,1693,1715]" italics="true" pageId="18" pageNumber="521">C. rufford</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="AB002815BF24FFBF28C48FD5FA4FAAD0" blockId="18.[151,1437,1400,2038]" box="[151,1436,1723,1745]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48FD5FA4FAAD0" box="[151,1436,1723,1745]" pageId="18" pageNumber="521">- Ventrals 31 or more................................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF24FFBF28C48FB6FA4FAB0A" pageId="18" pageNumber="521">
<paragraph id="AB002815BF24FFBF28C48FB6FA4FAAED" blockId="18.[151,1437,1400,2038]" box="[151,1436,1752,1774]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48FB6FA4FAAED" box="[151,1436,1752,1774]" pageId="18" pageNumber="521">7 Precloacal-femoral pores in males 20–29................................................................... 8</keyLead>
</paragraph>
<paragraph id="AB002815BF24FFBF28C48F98FA4FAB0A" blockId="18.[151,1437,1400,2038]" box="[151,1436,1781,1803]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48F98FA4FAB0A" box="[151,1436,1781,1803]" pageId="18" pageNumber="521">- Precloacal-femoral pores in males 34 or more............................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF24FFBF28C48E7DFA4EAB45" pageId="18" pageNumber="521">
<paragraph id="AB002815BF24FFBF28C48E7DFA4FAB28" blockId="18.[151,1437,1400,2038]" box="[151,1436,1811,1833]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48E7DFA4FAB28" box="[151,1436,1811,1833]" pageId="18" pageNumber="521">
8 Precloacal pores (pitted scales) in females present................................................
<taxonomicName id="6CBF5396BF24FFBF2D5C8E7DFA4FAB28" authorityName="Luu, Bonkowski, Nguyen, Le, Schneider, Ngo &amp; Ziegler" authorityYear="2016" box="[1295,1436,1811,1832]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="521" phylum="Chordata" rank="species" species="sommerladi">
<emphasis id="99CBF407BF24FFBF2D5C8E7DFA4FAB28" box="[1295,1436,1811,1832]" italics="true" pageId="18" pageNumber="521">C. sommerladi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="AB002815BF24FFBF28C48E5EFA4EAB45" blockId="18.[151,1437,1400,2038]" box="[151,1437,1840,1862]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48E5EFA4EAB45" box="[151,1437,1840,1862]" pageId="18" pageNumber="521">
- Precloacal pores in females absent.............................................................
<taxonomicName id="6CBF5396BF24FFBF2D558E5EFA4EAB45" authorityName="Luu, Calame, Nguyen, Bonkowski &amp; Ziegler" authorityYear="2016" box="[1286,1437,1840,1861]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="521" phylum="Chordata" rank="species" species="soudthichaki">
<emphasis id="99CBF407BF24FFBF2D558E5EFA4EAB45" box="[1286,1437,1840,1861]" italics="true" pageId="18" pageNumber="521">C. soudthichaki</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF24FFBF28C48E23FA4FAB80" pageId="18" pageNumber="521">
<paragraph id="AB002815BF24FFBF28C48E23FA4FAB62" blockId="18.[151,1437,1400,2038]" box="[151,1436,1869,1891]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48E23FA4FAB62" box="[151,1436,1869,1891]" pageId="18" pageNumber="521">
9 Precloacal-femoral pores in males 34, Midbody scales 86–87.......................................
<taxonomicName id="6CBF5396BF24FFBF2D5F8E20FA4FAB62" authorityName="Luu, Nguyen, Le, Bonkowski &amp; Ziegler" authorityYear="2016" box="[1292,1436,1869,1891]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="521" phylum="Chordata" rank="species" species="bansocensis">
<emphasis id="99CBF407BF24FFBF2D5F8E20FA4FAB62" box="[1292,1436,1869,1891]" italics="true" pageId="18" pageNumber="521">C. bansocensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="AB002815BF24FFBF28C48E05FA4FAB80" blockId="18.[151,1437,1400,2038]" box="[151,1436,1898,1921]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48E05FA4FAB80" box="[151,1436,1898,1921]" pageId="18" pageNumber="521">- Precloacal-femoral pores in males 35 or more.............................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF24FFBF28C48EE6FA4FABBA" pageId="18" pageNumber="521">
<paragraph id="AB002815BF24FFBF28C48EE6FA4FAB9D" blockId="18.[151,1437,1400,2038]" box="[151,1436,1928,1950]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48EE6FA4FAB9D" box="[151,1436,1928,1950]" pageId="18" pageNumber="521">10 Precloacal-femoral pores in females 0–34................................................................. 11</keyLead>
</paragraph>
<paragraph id="AB002815BF24FFBF28C48EC8FA4FABBA" blockId="18.[151,1437,1400,2038]" box="[151,1436,1957,1979]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48EC8FA4FABBA" box="[151,1436,1957,1979]" pageId="18" pageNumber="521">
- Precloacal-femoral pores in females 38............................................................
<taxonomicName id="6CBF5396BF24FFBF2D678EC8FA4FABBA" authorityName="Luu, Bonkowski, Nguyen, Le, Schneider, Ngo &amp; Ziegler" authorityYear="2016" box="[1332,1436,1957,1979]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="521" phylum="Chordata" rank="species" species="calamei">
<emphasis id="99CBF407BF24FFBF2D678EC8FA4FABBA" box="[1332,1436,1957,1979]" italics="true" pageId="18" pageNumber="521">C. calamei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="AB002815BF24FFBF28C48EADFA4FABD8" blockId="18.[151,1437,1400,2038]" box="[151,1436,1986,2009]" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBF28C48EADFA4FABD8" box="[151,1436,1986,2009]" pageId="18" pageNumber="521">
<keyStep id="104B30B0BF24FFBF28C48EADFA4FABD8" box="[151,1436,1986,2009]" pageId="18" pageNumber="521">11 Midbody scales 155–172.............................................................................. 12</keyStep>
</keyLead>
</paragraph>
<paragraph id="AB002815BF24FFBE28C48E8EFA4FACAD" blockId="18.[151,1437,1400,2038]" lastBlockId="19.[151,1437,151,907]" lastPageId="19" lastPageNumber="522" pageId="18" pageNumber="521">
<keyLead id="104E8B20BF24FFBE28C48E8EFA4FACAD" lastPageId="19" lastPageNumber="522" pageId="18" pageNumber="521">
- Midbody scales 179 or more........................................................................... 13
<keyStep id="104B30B0BF25FFBE28C489F6FA4FACAD" box="[151,1436,151,173]" pageId="19" pageNumber="522">
12 Ventrals 31–32, Precloacal-femoral pores in males 44, in females 21......................................
<taxonomicName id="6CBF5396BF25FFBE2D6F89F6FA4FACAD" authorityName="Luu, Calame, Bonkowski, Nguyen &amp; Ziegler" authorityYear="2014" box="[1340,1436,152,173]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="jaegeri">
<emphasis id="99CBF407BF25FFBE2D6F89F6FA4FACAD" box="[1340,1436,152,173]" italics="true" pageId="19" pageNumber="522">C. jaegeri</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C489DBFA4FACCA" blockId="19.[151,1437,151,907]" box="[151,1436,181,203]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C489DBFA4FACCA" box="[151,1436,181,203]" pageId="19" pageNumber="522">
- Ventrals 32–38, Precloacal-femoral pores in males 40– 44, in females 0–17........................
<taxonomicName id="6CBF5396BF25FFBE2C8889DBFA4FACCA" authorityName="Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov &amp; Chulisov" authorityYear="2014" box="[1243,1436,181,202]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="khammouanensis">
<emphasis id="99CBF407BF25FFBE2C8889DBFA4FACCA" box="[1243,1436,181,202]" italics="true" pageId="19" pageNumber="522">C. khammouanensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<keyStep id="104B30B0BF25FFBE28C489BCFA4FAD05" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C489BCFA4FACE7" blockId="19.[151,1437,151,907]" box="[151,1436,210,232]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C489BCFA4FACE7" box="[151,1436,210,232]" pageId="19" pageNumber="522">
13 SVL ≤
<quantity id="6C4785F0BF25FFBE297489BCFEAAACE7" box="[295,377,210,231]" metricMagnitude="-2" metricUnit="m" metricValue="7.12" pageId="19" pageNumber="522" unit="mm" value="71.2">71.2mm</quantity>
, Dorsal tubercle rows 19–24.....................................................
<taxonomicName id="6CBF5396BF25FFBE2D5889BDFA4FACE7" authorityName="Ngo &amp; Pauwels" authorityYear="2010" box="[1291,1436,210,232]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="lomyenensis">
<emphasis id="99CBF407BF25FFBE2D5889BDFA4FACE7" box="[1291,1436,210,232]" italics="true" pageId="19" pageNumber="522">C. lomyenensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C4899EFA4FAD05" blockId="19.[151,1437,151,907]" box="[151,1436,239,261]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C4899EFA4FAD05" box="[151,1436,239,261]" pageId="19" pageNumber="522">
- SVL&gt;
<quantity id="6C4785F0BF25FFBE297B899EFEB4AD05" box="[296,359,240,261]" metricMagnitude="-2" metricUnit="m" metricValue="7.2" pageId="19" pageNumber="522" unit="mm" value="72.0">72mm</quantity>
, Dorsal tubercle rows 14–20................................................................ 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF25FFBE28C48863FA4FAD3F" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C48863FA4FAD22" blockId="19.[151,1437,151,907]" box="[151,1436,269,291]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48863FA4FAD22" box="[151,1436,269,291]" pageId="19" pageNumber="522">
14 Dorsal tubercle rows 16–20, Precloacal-femoral pores in females 24–34..................................
<taxonomicName id="6CBF5396BF25FFBE2D7A8863FA4FAD22" authorityName="Nazarov, Poyarkov, Orlov, N.S.Nguyen, Milto, Martynov, Konstantinov &amp; Chulisov" authorityYear="2014" box="[1321,1436,269,290]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="darevskii">
<emphasis id="99CBF407BF25FFBE2D7A8863FA4FAD22" box="[1321,1436,269,290]" italics="true" pageId="19" pageNumber="522">C. darevskii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C48845FA4FAD3F" blockId="19.[151,1437,151,907]" box="[151,1436,298,320]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48845FA4FAD3F" box="[151,1436,298,320]" pageId="19" pageNumber="522">
- Dorsal tubercle rows 14–19, Precloacal-femoral pores in females 0–28..............................
<taxonomicName id="6CBF5396BF25FFBE2CA38845FA4FAD3F" authorityName="Luu, Bonkowski, Nguyen, Le, Schneider, Ngo &amp; Ziegler" authorityYear="2016" box="[1264,1436,298,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="hinnamnoensis">
<emphasis id="99CBF407BF25FFBE2CA38845FA4FAD3F" box="[1264,1436,298,320]" italics="true" pageId="19" pageNumber="522">C. hinnamnoensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF25FFBE28C48829FA4FAD7A" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C48829FA4FAD5D" blockId="19.[151,1437,151,907]" box="[151,1436,327,349]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48829FA4FAD5D" box="[151,1436,327,349]" pageId="19" pageNumber="522">15 Precloacal pores in females present...................................................................... 16</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C4880BFA4FAD7A" blockId="19.[151,1437,151,907]" box="[151,1436,357,379]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C4880BFA4FAD7A" box="[151,1436,357,379]" pageId="19" pageNumber="522">- Precloacal pores in females absent....................................................................... 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF25FFBE28C488ECFA4FADB4" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C488ECFA4EAD97" blockId="19.[151,1437,151,907]" box="[151,1437,386,408]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C488ECFA4EAD97" box="[151,1437,386,408]" pageId="19" pageNumber="522">
16 Dorsum banded.............................................................................
<taxonomicName id="6CBF5396BF25FFBE2D4988EDFA4EAD97" box="[1306,1437,386,408]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="wayakonei">
<emphasis id="99CBF407BF25FFBE2D4988EDFA4EAD97" box="[1306,1437,386,408]" italics="true" pageId="19" pageNumber="522">C. wayakonei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C488CEFA4FADB4" blockId="19.[151,1437,151,907]" box="[151,1436,415,437]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C488CEFA4FADB4" box="[151,1436,415,437]" pageId="19" pageNumber="522">
- Dorsum blotched..................................................................
<taxonomicName id="6CBF5396BF25FFBE2CFD88CEFA4FADB4" authorityName="Rosler, Vu, Nguyen, Ngo &amp; Ziegler" authorityYear="2008" box="[1198,1436,415,437]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="pseudoquadrivirgatus">
<emphasis id="99CBF407BF25FFBE2CFD88CEFA4FADB4" box="[1198,1436,415,437]" italics="true" pageId="19" pageNumber="522">C. pseudoquadrivirgatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF25FFBE28C488D3FA4FADEF" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C488D3FA4EADD2" blockId="19.[151,1437,151,907]" box="[151,1437,445,467]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C488D3FA4EADD2" box="[151,1437,445,467]" pageId="19" pageNumber="522">
17 Dorsum banded.............................................................................
<taxonomicName id="6CBF5396BF25FFBE2D4988D3FA4EADD2" authorityName="Schneider, Nguyen, Le, Nophaseud, Bonkowski &amp; Ziegler" authorityYear="2014" box="[1306,1437,445,466]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="vilaphongi">
<emphasis id="99CBF407BF25FFBE2D4988D3FA4EADD2" box="[1306,1437,445,466]" italics="true" pageId="19" pageNumber="522">C. vilaphongi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C488B5FA4FADEF" blockId="19.[151,1437,151,907]" box="[151,1436,474,496]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C488B5FA4FADEF" box="[151,1436,474,496]" pageId="19" pageNumber="522">- Dorsum blotched.................................................................................... 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF25FFBE28C48896FA4EAE2B" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C48896FA4FAE0C" blockId="19.[151,1437,151,907]" box="[151,1436,503,525]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48896FA4FAE0C" box="[151,1436,503,525]" pageId="19" pageNumber="522">
18 Ventrals 30..................................................................................
<taxonomicName id="6CBF5396BF25FFBE2D798896FA4FAE0C" authorityName="David, Teynie &amp; Ohler" authorityYear="2004" box="[1322,1436,503,525]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="buchardi">
<emphasis id="99CBF407BF25FFBE2D798896FA4FAE0C" box="[1322,1436,503,525]" italics="true" pageId="19" pageNumber="522">C. buchardi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C48B7BFA4EAE2B" blockId="19.[151,1437,151,907]" box="[151,1437,533,555]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48B7BFA4EAE2B" box="[151,1437,533,555]" pageId="19" pageNumber="522">
- Ventrals 40–50................................................................................
<taxonomicName id="6CBF5396BF25FFBE2D698B7BFA4EAE2B" authorityName="Heidrich, Rosler, Vu, Bohme &amp; Ziegler" authorityYear="2007" box="[1338,1437,533,555]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="cryptus">
<emphasis id="99CBF407BF25FFBE2D698B7BFA4EAE2B" box="[1338,1437,533,555]" italics="true" pageId="19" pageNumber="522">C. cryptus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF25FFBE28C48B5CFA4FAE65" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C48B5CFA4FAE47" blockId="19.[151,1437,151,907]" box="[151,1436,562,584]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48B5CFA4FAE47" box="[151,1436,562,584]" pageId="19" pageNumber="522">19 Dorsum blotched.................................................................................... 20</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C48B3EFA4FAE65" blockId="19.[151,1437,151,907]" box="[151,1436,591,613]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48B3EFA4FAE65" box="[151,1436,591,613]" pageId="19" pageNumber="522">- Dorsum banded...................................................................................... 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF25FFBE28C48B03FA4EAE9F" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C48B03FA4FAE82" blockId="19.[151,1437,151,907]" box="[151,1436,621,643]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48B03FA4FAE82" box="[151,1436,621,643]" pageId="19" pageNumber="522">
20 Precloacal pores present in females................................................................
<taxonomicName id="6CBF5396BF25FFBE2D138B03FA4FAE82" authorityName="David, Nguyen, Schneider &amp; Ziegler" authorityYear="2011" box="[1344,1436,621,642]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="teyniei">
<emphasis id="99CBF407BF25FFBE2D138B03FA4FAE82" box="[1344,1436,621,642]" italics="true" pageId="19" pageNumber="522">C. teyniei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C48BE5FA4EAE9F" blockId="19.[151,1437,151,907]" box="[151,1437,650,672]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48BE5FA4EAE9F" box="[151,1437,650,672]" pageId="19" pageNumber="522">
- Precloacal pores absent in females................................................................
<taxonomicName id="6CBF5396BF25FFBE2D7D8BE5FA4EAE9F" authorityName="Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov &amp; Chulisov" authorityYear="2014" box="[1326,1437,650,672]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="spelaeus">
<emphasis id="99CBF407BF25FFBE2D7D8BE5FA4EAE9F" box="[1326,1437,650,672]" italics="true" pageId="19" pageNumber="522">C. spelaeus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF25FFBE28C48BC6FA4FAEDA" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C48BC6FA4FAEBD" blockId="19.[151,1437,151,907]" box="[151,1436,679,701]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48BC6FA4FAEBD" box="[151,1436,679,701]" pageId="19" pageNumber="522">21 Femoral pores present in males......................................................................... 22</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C48BABFA4FAEDA" blockId="19.[151,1437,151,907]" box="[151,1436,709,731]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48BABFA4FAEDA" box="[151,1436,709,731]" pageId="19" pageNumber="522">- Femoral pores absent in males.......................................................................... 24</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF25FFBE28C48B8CFA4FAF14" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C48B8CFA4FAEF7" blockId="19.[151,1437,151,907]" box="[151,1436,738,760]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48B8CFA4FAEF7" box="[151,1436,738,760]" pageId="19" pageNumber="522">22 Precloacal pores in males 7–10......................................................................... 23</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C48A6EFA4FAF14" blockId="19.[151,1437,151,907]" box="[151,1436,767,789]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48A6EFA4FAF14" box="[151,1436,767,789]" pageId="19" pageNumber="522">
- Precloacal pores in males 14.................................................................
<taxonomicName id="6CBF5396BF25FFBE2D568A6EFA4FAF14" authorityName="Ulber" authorityYear="1993" box="[1285,1436,767,789]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="99CBF407BF25FFBE2D568A6EFA4FAF14" box="[1285,1436,767,789]" italics="true" pageId="19" pageNumber="522">C. interdigitalis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF25FFBE28C48A73FA4EAF4F" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C48A73FA4FAF32" blockId="19.[151,1437,151,907]" box="[151,1436,797,819]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48A73FA4FAF32" box="[151,1436,797,819]" pageId="19" pageNumber="522">
23 Precloacal pores in females 7............................................................
<taxonomicName id="6CBF5396BF25FFBE2C808A73FA99AF32" authority="Schneider &amp; Luu &amp; Sitthivong &amp; Teynié &amp; Le &amp; Nguyen &amp; Ziegler, 2020" authorityName="Schneider &amp; Luu &amp; Sitthivong &amp; Teynié &amp; Le &amp; Nguyen &amp; Ziegler" authorityYear="2020" box="[1235,1354,797,818]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="ngoiensis" status="sp. nov.">
<emphasis id="99CBF407BF25FFBE2C808A73FA99AF32" box="[1235,1354,797,818]" italics="true" pageId="19" pageNumber="522">C. ngoiensis</emphasis>
</taxonomicName>
<emphasis id="99CBF407BF25FFBE2D038A73FA4FAF32" bold="true" box="[1360,1436,797,818]" pageId="19" pageNumber="522">
<taxonomicNameLabel id="82F8497CBF25FFBE2D038A73FA4FAF32" box="[1360,1436,797,818]" pageId="19" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C48A55FA4EAF4F" blockId="19.[151,1437,151,907]" box="[151,1437,826,848]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48A55FA4EAF4F" box="[151,1437,826,848]" pageId="19" pageNumber="522">
- Precloacal pores in females 10-12............................................................
<taxonomicName id="6CBF5396BF25FFBE2D518A55FA4EAF4F" authorityName="Nazarov, Pauwels, Konstantinov, Chulisov, Orlov and Poyarkov" authorityYear="2018" box="[1282,1437,826,848]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="thathomensis">
<emphasis id="99CBF407BF25FFBE2D518A55FA4EAF4F" box="[1282,1437,826,848]" italics="true" pageId="19" pageNumber="522">C. thathomensis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="104B30B0BF25FFBE28C48A36FA4FAF8A" pageId="19" pageNumber="522">
<paragraph id="AB002815BF25FFBE28C48A36FA4FAF6C" blockId="19.[151,1437,151,907]" box="[151,1436,855,877]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48A36FA4FAF6C" box="[151,1436,855,877]" pageId="19" pageNumber="522">
24 Precloacal pores in males 4, in females 4–6...........................................................
<taxonomicName id="6CBF5396BF25FFBE2D178A36FA4FAF6C" authorityName="Schneider, Nguyen, Schmitz, Kingsada, Auer &amp; Ziegler" authorityYear="2011" box="[1348,1436,855,877]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="pageli">
<emphasis id="99CBF407BF25FFBE2D178A36FA4FAF6C" box="[1348,1436,855,877]" italics="true" pageId="19" pageNumber="522">C. pageli</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="AB002815BF25FFBE28C48A1BFA4FAF8A" blockId="19.[151,1437,151,907]" box="[151,1436,885,907]" pageId="19" pageNumber="522">
<keyLead id="104E8B20BF25FFBE28C48A1BFA4FAF8A" box="[151,1436,885,907]" pageId="19" pageNumber="522">
- Precloacal pores in males 6.........................................................
<taxonomicName id="6CBF5396BF25FFBE2CCD8A1BFA99AF8A" authority="Schneider &amp; Luu &amp; Sitthivong &amp; Teynié &amp; Le &amp; Nguyen &amp; Ziegler, 2020" authorityName="Schneider &amp; Luu &amp; Sitthivong &amp; Teynié &amp; Le &amp; Nguyen &amp; Ziegler" authorityYear="2020" box="[1182,1354,885,906]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="522" phylum="Chordata" rank="species" species="houaphanensis" status="sp. nov.">
<emphasis id="99CBF407BF25FFBE2CCD8A1BFA99AF8A" box="[1182,1354,885,906]" italics="true" pageId="19" pageNumber="522">C. houaphanensis</emphasis>
</taxonomicName>
<emphasis id="99CBF407BF25FFBE2D038A1BFA4FAF8A" bold="true" box="[1360,1436,885,906]" pageId="19" pageNumber="522">
<taxonomicNameLabel id="82F8497CBF25FFBE2D038A1BFA4FAF8A" box="[1360,1436,885,906]" pageId="19" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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