237 lines
33 KiB
XML
237 lines
33 KiB
XML
<document id="272B4E5877EDF4CE860E10E79D36EAD3" ID-CLB-Dataset="25284" ID-DOI="10.5852/ejt.2019.550" ID-GBIF-Dataset="dc3253d4-edc8-481b-966e-791e40856534" ID-Zenodo-Dep="3476402" ID-ZooBank="F8707194-B55E-48CA-8FE0-4CD0D023C217" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570541136797" checkinUser="plazi" docAuthor="Johansson, Niklas & Cederberg, Björn" docDate="2019" docId="A270EE7EFC28FFBFF360ACF93719FE30" docLanguage="en" docName="ejt-550_johansson_cederberg.pdf.imf" docOrigin="European Journal of Taxonomy 550" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophion ocellaris Ulbricht 1926" docType="treatment" docVersion="10" lastPageNumber="85" masterDocId="5E499606FC7CFFEAF13CAD503450FFC8" masterDocTitle="Review of the Swedish species of Ophion (Hymenoptera: Ichneumonidae: Ophioninae), with the description of 18 new species and an illustrated key to Swedish species" masterLastPageNumber="136" masterPageNumber="1" pageNumber="85" updateTime="1698758453468" updateUser="ExternalLinkService">
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<mods:title id="5DF4E0B94743BF3CD9F51749385FA553">Review of the Swedish species of Ophion (Hymenoptera: Ichneumonidae: Ophioninae), with the description of 18 new species and an illustrated key to Swedish species</mods:title>
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<mods:namePart id="0D5729BBCC22856B0033D22B8977A89B">Johansson, Niklas</mods:namePart>
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<mods:namePart id="18D0E4C542C8E8DDE5D0A2AE26181601">Cederberg, Björn</mods:namePart>
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<mods:title id="135744F92D26DF5A24C24AB1B1EA7FB4">European Journal of Taxonomy</mods:title>
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<mods:part id="B9093B9A715BCFD4D65A0996E2350DB2">
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<mods:date id="6C5D8307F9B9B63153A396B5B05E921E">2019</mods:date>
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<mods:number id="708E3D5C334B85BF82FBF6C4312CA73F">2019-09-12</mods:number>
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<mods:number id="6F80ABBB83AB4D49B2AA9FF7C65DA702">550</mods:number>
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<mods:identifier id="1AE794339B23AAFEECCDE77D430B8569" type="DOI">10.5852/ejt.2019.550</mods:identifier>
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<treatment id="A270EE7EFC28FFBFF360ACF93719FE30" ID-DOI="http://doi.org/10.5281/zenodo.3477019" ID-GBIF-Taxon="159660608" ID-Zenodo-Dep="3477019" LSID="urn:lsid:plazi:treatment:A270EE7EFC28FFBFF360ACF93719FE30" httpUri="http://treatment.plazi.org/id/A270EE7EFC28FFBFF360ACF93719FE30" lastPageId="85" lastPageNumber="85" pageId="84" pageNumber="85">
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<subSubSection id="62C30CE3FC28FFBEF360ACF93788FE0C" box="[604,984,425,452]" pageId="84" pageNumber="85" type="nomenclature">
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<paragraph id="2A665F68FC28FFBEF360ACF93788FE0C" blockId="84.[604,984,425,487]" box="[604,984,425,452]" pageId="84" pageNumber="85">
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<heading id="712EE804FC28FFBEF360ACF93788FE0C" box="[604,984,425,452]" centered="true" fontSize="11" level="2" pageId="84" pageNumber="85" reason="2">
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<taxonomicName id="EDD924EBFC28FFBEF360ACF93788FE0C" ID-CoL="49YK5" authority="Ulbricht, 1926" authorityName="Ulbricht" authorityYear="1926" box="[604,984,425,452]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="ocellaris">
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<emphasis id="18AD837AFC28FFBEF360ACF93775FE0C" bold="true" box="[604,805,425,452]" italics="true" pageId="84" pageNumber="85">Ophion ocellaris</emphasis>
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<bibRefCitation id="4E482299FC28FFBEF210ACFA3788FE0C" author="Ulbricht" box="[812,984,426,452]" firstAuthor="Ulbricht" pageId="84" pageNumber="85" pagination="1 - 30" refId="ref66549" refString="Ulbricht 1926. Niederrheinische Ichneumoniden. 4 Nachtrag. Mitteilungen der Naturwissenschaftliches Museum Crefeld 1926: 1 - 30" type="journal article" year="1926">Ulbricht, 1926</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="62C30CE3FC28FFBEF385AC9D372AFE2F" box="[697,890,461,487]" pageId="84" pageNumber="85" type="description">
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<paragraph id="2A665F68FC28FFBEF385AC9D372AFE2F" blockId="84.[604,984,425,487]" box="[697,890,461,487]" pageId="84" pageNumber="85">
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<figureCitation id="B2E243EDFC28FFBEF385AC9D3747FE2F" box="[697,791,461,487]" captionStart="Fig" captionStartId="15.[189,232,1746,1772]" captionTargetBox="[371,1220,293,1682]" captionTargetId="figure@15.[340,1247,262,1708]" captionTargetPageId="15" captionText="Fig. 7. A–D. Occipital carina posterior view. A. Ophion confusus Johansson sp. nov., holotype, ♀ (NHRS-HEVA000008637). B. O. mocsaryi Brauns, 1889. C. O. ocellaris Ulbricht, 1926. D. O. areolaris Brauns, 1889. E–G. Hind trochanter and trochantellus dorsal view. E. O. slaviceki Kriechbaumer, 1892. F. O. obscuratus Fabricius, 1798. G. O. costatus Ratzeburg, 1848. H–I. Mandible ventral view. H. O. obscuratus. I. O. slaviceki. J–K. First tergite, aberration O. artemisiae Boie, 1855. J. Dorsal view. K. Lateral view. L–M. Head in posterior view showing generalized shape of occipital carina. L. O. obscuratus. M. O. variegatus Rudow, 1883. N–O. Central flagellomeres in males. N. O. obscuratus. O. O. variegatus." figureDoi="http://doi.org/10.5281/zenodo.3476416" httpUri="https://zenodo.org/record/3476416/files/figure.png" pageId="84" pageNumber="85">Figs 7C</figureCitation>
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,
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<figureCitation id="B2E243EDFC28FFBEF218AC9D3717FE2F" box="[804,839,461,487]" captionStart="Fig" captionStartId="16.[189,232,1605,1631]" captionTargetBox="[285,1312,292,1544]" captionTargetId="figure@16.[261,1325,266,1568]" captionTargetPageId="16" captionText="Fig. 8. A–C.Apical flagellomeres in female. A. Ophion parvulusKriechbaumer, 1879. B. O. paraparvulus Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008696). C. O. tenuicornis Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008743). D–F. Apical flagellomeres in female (pilosity removed). D. O. ellenae Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008655). E. O. arenarius Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008606). F. O. inclinans Johansson sp. nov., holotype, ♀ (NHRS-HEVA000008667). G–H. Apical hind tarsal segments dorsal view in female. G. O. parvulus. H. O. costatus Ratzeburg, 1848. I–J. Generalized punctation below antennal sockets in anterior view. I. O. confusus Johansson sp. nov., holotype, ♀ (NHRS-HEVA000008637). J. O. mocsaryi Brauns, 1889. K–L. Posterior segments of female metasoma lateral view. K. O. areolaris Brauns, 1889. L. O. ocellaris Ulbricht, 1926. M. Central flagellomeres in female of O. brevicornis Morley, 1915. N–O. Basal flagellomeres. N. O. confusus Johansson sp. nov., holotype, ♀ (NHRS-HEVA000008637). O. O. mocsaryi." figureDoi="http://doi.org/10.5281/zenodo.3476420" httpUri="https://zenodo.org/record/3476420/files/figure.png" pageId="84" pageNumber="85">8L</figureCitation>
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,
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<figureCitation id="B2E243EDFC28FFBEF26FAC9D372AFE2F" box="[851,890,461,487]" captionStart="Fig" captionStartId="18.[189,232,1739,1765]" captionTargetBox="[305,1281,298,1666]" captionTargetId="figure@18.[263,1325,265,1697]" captionTargetPageId="18" captionText="Fig. 10. Generalized carination of propodeum in females, posterodorsal view. A. Ophion luteus (Linnaeus, 1758)/O. scutellaris Thomson, 1888/O. obscuratus Fabricius, 1792. B. O. costatus Ratzeburg, 1848. C. O. brocki Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008633). D. O. confusus Johansson sp. nov., holotype, ♀ (NHRS-HEVA000008637). E. O. crassicornis Brock, 1982. F. O. artemisiae Boie, 1855. G. O. ellenae Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008655). H. O. perkinsi Brock, 1982. I. O. ocellaris Ulbricht, 1926. J. O. areolaris Brauns, 1889. K. O. dispar Brauns, 1895. L. O. tenuicornis Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008743). M. O. longigena Thomson, 1888. N. O. brevicornis Morley, 1915. O. O. forticornis Morley, 1915." figureDoi="http://doi.org/10.5281/zenodo.3476424" httpUri="https://zenodo.org/record/3476424/files/figure.png" pageId="84" pageNumber="85">
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<date id="5E6779A8FC28FFBEF26FAC9D372AFE2F" box="[851,890,461,487]" pageId="84" pageNumber="85" value="2010-01">10I</date>
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</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="62C30CE3FC28FFBEF181AF5E363CFD84" pageId="84" pageNumber="85" type="reference_group">
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||
<paragraph id="2A665F68FC28FFBEF181AF5E3639FDE1" blockId="84.[189,620,526,588]" box="[189,617,526,553]" pageId="84" pageNumber="85">
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<treatmentCitationGroup id="0AC97846FC28FFBEF181AF5E3639FDE1" box="[189,617,526,553]" pageId="84" pageNumber="85">
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<taxonomicName id="EDD924EBFC28FFBEF181AF5E3635FDE1" ID-CoL="49YK5" authority="Ulbricht, 1926: 17" authorityName="Ulbricht" authorityPageNumber="17" authorityYear="1926" box="[189,613,526,553]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="ocellaris">
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<emphasis id="18AD837AFC28FFBEF181AF5E35D3FDE0" box="[189,387,526,552]" italics="true" pageId="84" pageNumber="85">Ophion ocellaris</emphasis>
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<treatmentCitation id="AB787979FC28FFBEF0B6AF5E3635FDE1" author="Ulbricht" box="[394,613,526,553]" page="17" pageId="84" pageNumber="85" year="1926">
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<bibRefCitation id="4E482299FC28FFBEF0B6AF5E3635FDE1" author="Ulbricht" box="[394,613,526,553]" firstAuthor="Ulbricht" pageId="84" pageNumber="85" pagination="1 - 30" refId="ref66549" refString="Ulbricht 1926. Niederrheinische Ichneumoniden. 4 Nachtrag. Mitteilungen der Naturwissenschaftliches Museum Crefeld 1926: 1 - 30" type="journal article" year="1926">Ulbricht, 1926: 17</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="2A665F68FC28FFBEF181AF62363CFD84" blockId="84.[189,620,526,588]" box="[189,620,562,588]" pageId="84" pageNumber="85">
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<treatmentCitationGroup id="0AC97846FC28FFBEF181AF62363CFD84" box="[189,620,562,588]" pageId="84" pageNumber="85">
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<taxonomicName id="EDD924EBFC28FFBEF181AF623630FD84" ID-CoL="49YHJ" authority="Hellen, 1926: 14" authorityName="Hellen." authorityPageNumber="14" authorityYear="1926" box="[189,608,562,588]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="fuscicollis">
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<emphasis id="18AD837AFC28FFBEF181AF6235C3FD84" box="[189,403,562,588]" italics="true" pageId="84" pageNumber="85">Ophion fuscicollis</emphasis>
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<treatmentCitation id="AB787979FC28FFBEF0A7AF623630FD84" author="Hellen W." box="[411,608,562,588]" page="14" pageId="84" pageNumber="85" year="1926">
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<bibRefCitation id="4E482299FC28FFBEF0A7AF623630FD84" author="Hellen" box="[411,608,562,588]" firstAuthor="Hellen" pageId="84" pageNumber="85" pagination="1 - 27" refId="ref64600" refString="Hellen W. 1926. Beitrage zur Kenntnis der Ichneumoniden Finlands II. Subfam. Ophioninae und Anomaloninae. Acta Societatis pro Fauna et Flora Fennica 56 (6): 1 - 27." type="journal article" year="1926">Hellén, 1926: 14</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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||
?
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="62C30CE3FC28FFBEF181AF233134FD74" pageId="84" pageNumber="85" type="materials_examined">
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<paragraph id="2A665F68FC28FFBEF181AF2335FAFD45" blockId="84.[189,426,627,653]" box="[189,426,627,653]" pageId="84" pageNumber="85">
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<heading id="712EE804FC28FFBEF181AF2335FAFD45" bold="true" box="[189,426,627,653]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
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||
<emphasis id="18AD837AFC28FFBEF181AF2335FAFD45" bold="true" box="[189,426,627,653]" pageId="84" pageNumber="85">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2A665F68FC28FFBEF181AFF23134FD74" blockId="84.[189,1380,674,701]" box="[189,1380,674,701]" pageId="84" pageNumber="85">
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<materialsCitation id="9AB15535FC28FFBEF181AFF23134FD74" ID-GBIF-Occurrence="2425024605" box="[189,1380,674,701]" collectionCode="ZMH" country="Sweden" location="Sweden" pageId="84" pageNumber="85" specimenCount="91" specimenCount-female="91" typeStatus="holotype">
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<typeStatus id="F562E1CAFC28FFBEF181AFF2357DFD74" box="[189,301,674,700]" pageId="84" pageNumber="85" type="holotype">Holotype</typeStatus>
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, ♂, of
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<taxonomicName id="EDD924EBFC28FFBEF041AFF23604FD74" authorityName="Hellen." authorityYear="1926" box="[381,596,674,700]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="fuscicollis">
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<emphasis id="18AD837AFC28FFBEF041AFF23604FD74" box="[381,596,674,700]" italics="true" pageId="84" pageNumber="85">Ophion fuscicollis</emphasis>
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</taxonomicName>
|
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(
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<collectionCode id="4CC8C7ADFC28FFBEF35FAFF236F6FD74" box="[611,678,674,700]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="84" pageNumber="85" type="Museum">ZMH</collectionCode>
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||
);
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||
<specimenCount id="3CDF94E1FC28FFBEF383AFF23741FD74" box="[703,785,674,700]" pageId="84" pageNumber="85" type="female">83 ♀♀</specimenCount>
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, 11 ♂♂ (
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||
<collectingCountry id="52CE1FF8FC28FFBEF242AFF237B1FD75" box="[894,993,674,701]" name="Sweden" pageId="84" pageNumber="85">Sweden</collectingCountry>
|
||
);
|
||
<specimenCount id="3CDF94E1FC28FFBEF2C9AFF23071FD74" box="[1013,1057,674,700]" pageId="84" pageNumber="85" type="female">1 ♀</specimenCount>
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||
(
|
||
<collectingCountry id="52CE1FF8FC28FFBEF50DAFF230DEFD75" box="[1073,1166,674,701]" name="Croatia" pageId="84" pageNumber="85">Croatia</collectingCountry>
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||
);
|
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<specimenCount id="3CDF94E1FC28FFBEF59EAFF230B4FD74" box="[1186,1252,674,700]" pageId="84" pageNumber="85" type="female">7 ♀♀</specimenCount>
|
||
(
|
||
<collectingCountry id="52CE1FF8FC28FFBEF5CFAFF23105FD74" box="[1267,1365,674,700]" name="Norway" pageId="84" pageNumber="85">Norway</collectingCountry>
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||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62C30CE3FC28FFBEF181AFB336CBFA1D" pageId="84" pageNumber="85" type="diagnosis">
|
||
<paragraph id="2A665F68FC28FFBEF181AFB33566FD35" blockId="84.[189,310,739,765]" box="[189,310,739,765]" pageId="84" pageNumber="85">
|
||
<heading id="712EE804FC28FFBEF181AFB33566FD35" bold="true" box="[189,310,739,765]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
|
||
<emphasis id="18AD837AFC28FFBEF181AFB33566FD35" bold="true" box="[189,310,739,765]" pageId="84" pageNumber="85">Diagnosis</emphasis>
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||
</heading>
|
||
</paragraph>
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||
<paragraph id="2A665F68FC28FFBEF181AE4230EDFB44" blockId="84.[189,1399,785,1164]" pageId="84" pageNumber="85">
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||
Fore wing length
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||
<quantity id="ED21F28DFC28FFBEF0B9AE413654FCE4" box="[389,516,785,812]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="84" pageNumber="85" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
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. Antenna in both sexes with 51–55 flagellomeres. First flagellomere 3.5 times as long as wide. Head narrowed behind eyes. Occipital carina entirely absent (
|
||
<figureCitation id="B2E243EDFC28FFBEF50BAE6530C1FC87" box="[1079,1169,821,847]" captionStart="Fig" captionStartId="15.[189,232,1746,1772]" captionTargetBox="[371,1220,293,1682]" captionTargetId="figure@15.[340,1247,262,1708]" captionTargetPageId="15" captionText="Fig. 7. A–D. Occipital carina posterior view. A. Ophion confusus Johansson sp. nov., holotype, ♀ (NHRS-HEVA000008637). B. O. mocsaryi Brauns, 1889. C. O. ocellaris Ulbricht, 1926. D. O. areolaris Brauns, 1889. E–G. Hind trochanter and trochantellus dorsal view. E. O. slaviceki Kriechbaumer, 1892. F. O. obscuratus Fabricius, 1798. G. O. costatus Ratzeburg, 1848. H–I. Mandible ventral view. H. O. obscuratus. I. O. slaviceki. J–K. First tergite, aberration O. artemisiae Boie, 1855. J. Dorsal view. K. Lateral view. L–M. Head in posterior view showing generalized shape of occipital carina. L. O. obscuratus. M. O. variegatus Rudow, 1883. N–O. Central flagellomeres in males. N. O. obscuratus. O. O. variegatus." figureDoi="http://doi.org/10.5281/zenodo.3476416" httpUri="https://zenodo.org/record/3476416/files/figure.png" pageId="84" pageNumber="85">Fig. 7C</figureCitation>
|
||
). Malar space about 0.1 times as long as mandibular base in female and male. Mandibular gape right-angled, with internal angles. Wing membrane strongly yellowish. Mesopleuron polished and punctate. Interstices between punctures 1–2 times their diameter. Pleurosternal angles strongly obtuse, weakly defined and obviously anterior to sternal angles (as in
|
||
<figureCitation id="B2E243EDFC28FFBEF31FAE92362BFC14" box="[547,635,962,988]" captionStart="Fig" captionStartId="17.[189,232,1849,1875]" captionTargetBox="[189,1398,640,1797]" captionTargetId="figure@17.[189,1398,640,1799]" captionTargetPageId="17" captionText="Fig. 9. Epicnemial carina, antero-ventral view of female. A. Ophion splendens Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008708). B. O. mocsaryi Brauns, 1889. C. O. dispar Brauns, 1895. D. O. parvulus Kriechbaumer, 1879. E. O. slaviceki Kriechbaumer, 1892. F. O. angularis Johansson & Cederberg sp. nov., paratype, ♀ (MZLU 6366:3). G. O. crassicornis Brock, 1982. H. O. areolaris Brauns, 1889. I. O. kallanderi Johansson sp. nov., paratype, ♀ (MZLU 6371:5)." figureDoi="http://doi.org/10.5281/zenodo.3476422" httpUri="https://zenodo.org/record/3476422/files/figure.png" pageId="84" pageNumber="85">Fig. 9H</figureCitation>
|
||
). Scutellum without lateral carinae, rectangular with parallel sides (as in
|
||
<figureCitation id="B2E243EDFC28FFBEF038AEB5350FFBC8" box="[260,351,997,1024]" captionStart="Fig" captionStartId="14.[189,232,1841,1867]" captionTargetBox="[273,1272,270,1727]" captionTargetId="figure@14.[189,1398,270,1796]" captionTargetPageId="14" captionText="Fig. 6. A–D. Scutellum, dorsal view. A. Ophion luteus (Linnaeus, 1758). B. O. areolaris Brauns, 1889. C. O. inclinans Johansson sp. nov., holotype, ♀ (NHRS-HEVA000008667). D. O. dispar Brauns, 1895. E–H. Firsttergite, lateral view.E. O.inclinansJohanssonsp.nov.,holotype,♀ (NHRS-HEVA000008667). F. O. dispar. G. O. artemisiae Boie, 1855. H. O. luteus. I–J. Shape of male parameres, lateral view. I. Ophion sp. cf. pseudocostatus Meyer, 1935. J. O. artemisiae." figureDoi="http://doi.org/10.5281/zenodo.3476414" httpUri="https://zenodo.org/record/3476414/files/figure.png" pageId="84" pageNumber="85">Fig. 6B</figureCitation>
|
||
). Propodeum coriaceous with carina strongly reduced and only posterior transverse carina present and often interrupted or weak centrally. Propodeum centrally with transverse wrinkles (
|
||
<figureCitation id="B2E243EDFC28FFBEF435A9583139FBEA" box="[1289,1385,1032,1058]" captionStart="Fig" captionStartId="18.[189,232,1739,1765]" captionTargetBox="[305,1281,298,1666]" captionTargetId="figure@18.[263,1325,265,1697]" captionTargetPageId="18" captionText="Fig. 10. Generalized carination of propodeum in females, posterodorsal view. A. Ophion luteus (Linnaeus, 1758)/O. scutellaris Thomson, 1888/O. obscuratus Fabricius, 1792. B. O. costatus Ratzeburg, 1848. C. O. brocki Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008633). D. O. confusus Johansson sp. nov., holotype, ♀ (NHRS-HEVA000008637). E. O. crassicornis Brock, 1982. F. O. artemisiae Boie, 1855. G. O. ellenae Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008655). H. O. perkinsi Brock, 1982. I. O. ocellaris Ulbricht, 1926. J. O. areolaris Brauns, 1889. K. O. dispar Brauns, 1895. L. O. tenuicornis Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008743). M. O. longigena Thomson, 1888. N. O. brevicornis Morley, 1915. O. O. forticornis Morley, 1915." figureDoi="http://doi.org/10.5281/zenodo.3476424" httpUri="https://zenodo.org/record/3476424/files/figure.png" pageId="84" pageNumber="85">
|
||
Fig.
|
||
<date id="5E6779A8FC28FFBEF403A9583139FBEA" box="[1343,1385,1032,1058]" pageId="84" pageNumber="85" value="2010-01">10I</date>
|
||
</figureCitation>
|
||
). Female metasoma shorter than in
|
||
<emphasis id="18AD837AFC28FFBEF36CA97C36B6FB8D" box="[592,742,1067,1094]" italics="true" pageId="84" pageNumber="85">
|
||
<taxonomicName id="EDD924EBFC28FFBEF36CA97C36B3FB8D" authorityName="Brauns" authorityYear="1889" box="[592,739,1067,1094]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="areolaris">O. areolaris</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Tergite
|
||
<quantity id="ED21F28DFC28FFBEF275A97B37C5FB8D" box="[841,917,1067,1094]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="84" pageNumber="85" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4–5 in</quantity>
|
||
lateral view about 0.75 times as long as wide. Spiracles of tergite 2–7 more close to the base of the tergite (
|
||
<figureCitation id="B2E243EDFC28FFBEF289A91E305DFBA1" box="[949,1037,1102,1129]" captionStart="Fig" captionStartId="16.[189,232,1605,1631]" captionTargetBox="[285,1312,292,1544]" captionTargetId="figure@16.[261,1325,266,1568]" captionTargetPageId="16" captionText="Fig. 8. A–C.Apical flagellomeres in female. A. Ophion parvulusKriechbaumer, 1879. B. O. paraparvulus Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008696). C. O. tenuicornis Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008743). D–F. Apical flagellomeres in female (pilosity removed). D. O. ellenae Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008655). E. O. arenarius Johansson sp. nov., paratype, ♀ (NHRS-HEVA000008606). F. O. inclinans Johansson sp. nov., holotype, ♀ (NHRS-HEVA000008667). G–H. Apical hind tarsal segments dorsal view in female. G. O. parvulus. H. O. costatus Ratzeburg, 1848. I–J. Generalized punctation below antennal sockets in anterior view. I. O. confusus Johansson sp. nov., holotype, ♀ (NHRS-HEVA000008637). J. O. mocsaryi Brauns, 1889. K–L. Posterior segments of female metasoma lateral view. K. O. areolaris Brauns, 1889. L. O. ocellaris Ulbricht, 1926. M. Central flagellomeres in female of O. brevicornis Morley, 1915. N–O. Basal flagellomeres. N. O. confusus Johansson sp. nov., holotype, ♀ (NHRS-HEVA000008637). O. O. mocsaryi." figureDoi="http://doi.org/10.5281/zenodo.3476420" httpUri="https://zenodo.org/record/3476420/files/figure.png" pageId="84" pageNumber="85">Fig. 8L</figureCitation>
|
||
). Male parameres longer, about 1.5 times as long as wide and more rounded dorsally in lateral view than in
|
||
<taxonomicName id="EDD924EBFC28FFBEF516A92230E6FB44" authorityName="Brauns" authorityYear="1889" box="[1066,1206,1138,1164]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="areolaris">
|
||
<emphasis id="18AD837AFC28FFBEF516A92230E6FB44" box="[1066,1206,1138,1164]" italics="true" pageId="84" pageNumber="85">O. areolaris</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2A665F68FC28FFBEF181A9E33544FB05" blockId="84.[189,276,1203,1229]" box="[189,276,1203,1229]" pageId="84" pageNumber="85">
|
||
<heading id="712EE804FC28FFBEF181A9E33544FB05" bold="true" box="[189,276,1203,1229]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
|
||
<emphasis id="18AD837AFC28FFBEF181A9E33544FB05" bold="true" box="[189,276,1203,1229]" pageId="84" pageNumber="85">Colour</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2A665F68FC28FFBEF181A9B2372BFAD7" blockId="84.[189,1399,1250,1311]" pageId="84" pageNumber="85">Body testaceous, sometimes with fuscous areas on meso- and metasoma. Mandibular teeth black. Stemmaticum black or dark brown, ovipositor sheath black.</paragraph>
|
||
<paragraph id="2A665F68FC28FFBEF181A8163536FAA9" blockId="84.[189,358,1350,1377]" box="[189,358,1350,1377]" pageId="84" pageNumber="85">
|
||
<heading id="712EE804FC28FFBEF181A8163536FAA9" bold="true" box="[189,358,1350,1377]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
|
||
<emphasis id="18AD837AFC28FFBEF181A8163536FAA9" bold="true" box="[189,358,1350,1377]" pageId="84" pageNumber="85">DNA barcode</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2A665F68FC28FFBEF181A82536CBFA1D" blockId="84.[189,1399,1397,1493]" pageId="84" pageNumber="85">
|
||
The DNA barcode sequences of four Swedish specimens of
|
||
<taxonomicName id="EDD924EBFC28FFBEF245A825306EFA47" authorityName="Ulbricht" authorityYear="1926" box="[889,1086,1397,1423]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="ocellaris">
|
||
<emphasis id="18AD837AFC28FFBEF245A825306EFA47" box="[889,1086,1397,1423]" italics="true" pageId="84" pageNumber="85">Ophion ocellaris</emphasis>
|
||
</taxonomicName>
|
||
are available at the BOLD systems database (www.boldsystems.org, BIN. BOLD: ADG0270/ADG3721. Specimen codes: STI- NJBC: Clade 1; 02, Clade 2; 01, 03–04).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62C30CE3FC28FFBEF181A8AD314DF944" pageId="84" pageNumber="85" type="biology_ecology">
|
||
<paragraph id="2A665F68FC28FFBEF181A8AD3570F9DF" blockId="84.[189,288,1533,1559]" box="[189,288,1533,1559]" pageId="84" pageNumber="85">
|
||
<heading id="712EE804FC28FFBEF181A8AD3570F9DF" bold="true" box="[189,288,1533,1559]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
|
||
<emphasis id="18AD837AFC28FFBEF181A8AD3570F9DF" bold="true" box="[189,288,1533,1559]" pageId="84" pageNumber="85">Ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2A665F68FC28FFBEF181AB7B314DF944" blockId="84.[189,1399,1579,1676]" pageId="84" pageNumber="85">
|
||
This species occurs in late summer and autumn, late June–early September and the hosts seem to be various species of
|
||
<taxonomicName id="EDD924EBFC28FFBEF0A0AB1E3673F9A0" authorityName="Boisduval" authorityYear="1828" box="[412,547,1614,1640]" class="Insecta" family="Drepanidae" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="family">Drepanidae</taxonomicName>
|
||
(subfamily
|
||
<taxonomicName id="EDD924EBFC28FFBEF388AB1E376DF9A1" authorityName="Smith" authorityYear="1893" box="[692,829,1614,1641]" class="Insecta" family="Drepanidae" kingdom="Animalia" order="Lepidoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="subFamily" subFamily="Thyatirinae">Thyatirinae</taxonomicName>
|
||
) according to
|
||
<bibRefCitation id="4E482299FC28FFBEF2D6AB1E30DDF9A1" author="Brock" box="[1002,1165,1614,1641]" firstAuthor="Brock" pageId="84" pageNumber="85" pagination="57 - 97" refId="ref63572" refString="Brock J. P. 1982. A systematic study of the genus Ophion in Britain (Hymenoptera, Ichneumonidae). Tijdschrift voor Entomologie 125: 57 - 97." type="journal article" year="1982">Brock (1982)</bibRefCitation>
|
||
. It has mainly been collected in boreal woodland habitats, but has also been attracted to light in semi-open habitats.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62C30CE3FC28FFBEF181ABE3358FF8D7" pageId="84" pageNumber="85" type="distribution">
|
||
<paragraph id="2A665F68FC28FFBEF181ABE3358DF905" blockId="84.[189,477,1715,1741]" box="[189,477,1715,1741]" pageId="84" pageNumber="85">
|
||
<heading id="712EE804FC28FFBEF181ABE3358DF905" bold="true" box="[189,477,1715,1741]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
|
||
<emphasis id="18AD837AFC28FFBEF181ABE3358DF905" bold="true" box="[189,477,1715,1741]" pageId="84" pageNumber="85">
|
||
Distribution in
|
||
<collectingCountry id="52CE1FF8FC28FFBEF041ABE3358DF905" box="[381,477,1715,1741]" name="Sweden" pageId="84" pageNumber="85">Sweden</collectingCountry>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2A665F68FC28FFBEF181ABB2358FF8D7" blockId="84.[189,1398,1762,1823]" pageId="84" pageNumber="85">This species is quite common and occurs over most of the country, but is apparently missing in high alpine areas in the north.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62C30CE3FC28FFBFF181AA173719FE30" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="discussion">
|
||
<paragraph id="2A665F68FC28FFBEF181AA17357FF8A9" blockId="84.[189,303,1863,1889]" box="[189,303,1863,1889]" pageId="84" pageNumber="85">
|
||
<heading id="712EE804FC28FFBEF181AA17357FF8A9" bold="true" box="[189,303,1863,1889]" fontSize="11" level="3" pageId="84" pageNumber="85" reason="3">
|
||
<emphasis id="18AD837AFC28FFBEF181AA17357FF8A9" bold="true" box="[189,303,1863,1889]" pageId="84" pageNumber="85">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2A665F68FC28FFBFF181AA253719FE30" blockId="84.[189,1399,1909,2006]" lastBlockId="85.[189,1399,267,505]" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
|
||
The
|
||
<typeStatus id="F562E1CAFC28FFBEF1C8AA25350BF847" box="[244,347,1909,1935]" pageId="84" pageNumber="85" type="holotype">holotype</typeStatus>
|
||
male of
|
||
<taxonomicName id="EDD924EBFC28FFBEF0F6AA2536F3F847" authorityName="Hellen." authorityYear="1926" box="[458,675,1909,1935]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="fuscicollis">
|
||
<emphasis id="18AD837AFC28FFBEF0F6AA2536F3F847" box="[458,675,1909,1935]" italics="true" pageId="84" pageNumber="85">Ophion fuscicollis</emphasis>
|
||
</taxonomicName>
|
||
was studied and although access to the type of
|
||
<taxonomicName id="EDD924EBFC28FFBEF5D5AA253127F847" authorityName="Ulbricht" authorityYear="1926" box="[1257,1399,1909,1935]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="ocellaris">
|
||
<emphasis id="18AD837AFC28FFBEF5D5AA253127F847" box="[1257,1399,1909,1935]" italics="true" pageId="84" pageNumber="85">O. ocellaris</emphasis>
|
||
</taxonomicName>
|
||
was denied, the opinion presented here is that the species are synonymous. However, the actual date of publication for the description of either of the species is uncertain and the question of which name has priority over the other remains. In this study the more commonly used name
|
||
<taxonomicName id="EDD924EBFC29FFBFF570AC5C3088FEED" authorityName="Ulbricht" authorityYear="1926" box="[1100,1240,267,294]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="ocellaris">
|
||
<emphasis id="18AD837AFC29FFBFF570AC5C3088FEED" box="[1100,1240,267,294]" italics="true" pageId="85" pageNumber="86">O. ocellaris</emphasis>
|
||
</taxonomicName>
|
||
is used. Also of interest, is that one of the four barcoded specimens forms a separate BIN with a series of specimens from
|
||
<collectingCountry id="52CE1FF8FC29FFBFF1C0AC0235C9FEA4" box="[252,409,338,364]" name="United Kingdom" pageId="85" pageNumber="86">Great Britain</collectingCountry>
|
||
. The existence of two distinct clades could indicate the existence of a cryptic species. Despite studying a rather large amount of material, no reliable morphological differences have been detected. The discovery of a presumably cryptic species could of course also mean that the proposed synonymy between
|
||
<taxonomicName id="EDD924EBFC29FFBFF091ACEC366AFE1D" authorityName="Ulbricht" authorityYear="1926" box="[429,570,443,470]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="ocellaris">
|
||
<emphasis id="18AD837AFC29FFBFF091ACEC366AFE1D" box="[429,570,443,470]" italics="true" pageId="85" pageNumber="86">O. ocellaris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EDD924EBFC29FFBFF34BACEC3745FE1D" authorityName="Hellen." authorityYear="1926" box="[631,789,443,470]" class="Insecta" family="Ichneumonidae" genus="Ophion" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="fuscicollis">
|
||
<emphasis id="18AD837AFC29FFBFF34BACEC3745FE1D" box="[631,789,443,470]" italics="true" pageId="85" pageNumber="86">O. fuscicollis</emphasis>
|
||
</taxonomicName>
|
||
is more complicated. Therefore this study refrains from a formal publication of the synonymy at this time.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |