261 lines
24 KiB
XML
261 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.818.30477" ID-GBIF-Dataset="560ded1c-9e7b-4ae1-b777-318025a8cee0" ID-PMC="PMC6354008" ID-Pensoft-Pub="1313-2970-818-89" ID-PubMed="30723380" ID-ZBK="85650B90B4DD4FE08C16FD34BA805C07" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-818-89" ModsDocOrigin="ZooKeys 818" ModsDocTitle="The extraordinary genus Myja is not a tergipedid, but related to the Facelinidae s. str. with the addition of two new species from Japan (Mollusca, Nudibranchia)" checkinTime="1548263243176" checkinUser="pensoft" docAuthor="Martynov, Alexander, Mehrotra, Rahul, Chavanich, Suchana, Nakano, Rie, Kashio, Sho, Lundin, Kennet, Picton, Bernard & Korshunova, Tatiana" docDate="2019" docId="A24C709C72EB4D05360AF6635A80FC83" docLanguage="en" docName="ZooKeys 818: 89-116" docOrigin="ZooKeys 818" docSource="http://dx.doi.org/10.3897/zookeys.818.30477" docTitle="Myja longicornis Bergh 1896" docType="treatment" docVersion="4" lastPageNumber="89" masterDocId="9F71FFA00A781D2DF927657CFFBBFA46" masterDocTitle="The extraordinary genus Myja is not a tergipedid, but related to the Facelinidae s. str. with the addition of two new species from Japan (Mollusca, Nudibranchia)" masterLastPageNumber="116" masterPageNumber="89" pageNumber="89" updateTime="1668166777296" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The extraordinary genus Myja is not a tergipedid, but related to the Facelinidae s. str. with the addition of two new species from Japan (Mollusca, Nudibranchia)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Martynov, Alexander</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mehrotra, Rahul</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chavanich, Suchana</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nakano, Rie</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kashio, Sho</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lundin, Kennet</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Picton, Bernard</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Korshunova, Tatiana</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>818</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>89</mods:start>
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<mods:end>116</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.818.30477</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.818.30477</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-818-89</mods:identifier>
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<mods:identifier type="ZBK">85650B90B4DD4FE08C16FD34BA805C07</mods:identifier>
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<mods:identifier type="ZooBank">85650B90B4DD4FE08C16FD34BA805C07</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="154125891" LSID="urn:lsid:plazi:treatment:A24C709C72EB4D05360AF6635A80FC83" httpUri="http://treatment.plazi.org/id/A24C709C72EB4D05360AF6635A80FC83" lastPageNumber="89" pageId="0" pageNumber="89">
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<subSubSection pageId="0" pageNumber="89" type="nomenclature">
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<paragraph pageId="0" pageNumber="89">
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<taxonomicName authority="Bergh, 1896" authorityName="Bergh" authorityYear="1896" class="Gastropoda" family="Tergipedidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja longicornis" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="longicornis">Myja cf. longicornis Bergh, 1896</taxonomicName>
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Figs 1, 4
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="materials_examined">
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<paragraph pageId="0" pageNumber="89">Material.</paragraph>
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<paragraph pageId="0" pageNumber="89">1 specimen, ZMMU Op-667, 6 mm long (fixed), Thailand, Koh Samaesan, 21 June 2018, depth 8 -16 m, soft sediment habitats, hydroids, collectors Rahul Mehrotra, Suchana Chavanich. 2 specimens, ZMMU Op-668, ca. 3 and 2 mm (fixed) same locality and collectors.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="locality">
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<paragraph pageId="0" pageNumber="89">Locality.</paragraph>
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<paragraph pageId="0" pageNumber="89">Thailand, Chonburi, Koh Samaesan.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="diagnosis">
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<paragraph pageId="0" pageNumber="89">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="89">Up to eight ceratal rows, ground colour translucent greyish, ceratal cores light to dark greyish, ceratal tops dull reddish, apices with white spot, anterior cerata with prominent reddish basal spot (distributed over the whole surface in some cerata), white gonad spherules moderately dense, sparse white spots in the first half of the dorsal part, cerata moderately widened at top without smaller separate cupola-shaped tip, central tooth with sharp to pitted top and numerous lateral denticles, up to 23 small denticles, irregular in size, no distinct furrows and ridges on the teeth surfaces, no accessory penial gland, penis unarmed.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="description">
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<paragraph pageId="0" pageNumber="89">Description.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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Body very elongate, up to 6 mm in preserved length (up to 10 mm alive) (Fig. 1A, B). Rhinophores similar in size to oral tentacles, smooth. Dorsal papillae cylindrical, forming eight ceratal rows along dorsal edges. Apices of papillae moderately to slightly widened, without cupola-shaped appendage (Fig. 1C). Notal edge absent. Anal opening acleioproctic on right side before first posterior ceratal rows. Reproductive openings lateral, below first anterior and second posterior rows of cerata. Ground colour translucent greyish. Oral tentacles and rhinophores with scattered opaque white dots. Digestive gland in the cerata (ceratal cores) light to dark greyish, digestive gland in upper part of cerata with reddish internal spot, apices with white spot. Anterior cerata with prominent reddish basal spot in some cerata distributed over its whole surface. Central branches of digestive gland visible through dorsal part of body greyish. Numerous small, moderately dense white gonads appear as white spherules that shine through dorsal surface. Jaws broadly triangular with prominent anterior wings, masticatory borders smooth (Fig. 1D). Radula uniserial, very small compared to pharynx internal volume (Fig. 1E, indicated by an arrow). Radular formula 13
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<normalizedToken originalValue="×">x</normalizedToken>
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0.1.0. Central tooth narrowly triangular, with sharp to pitted top (Fig. 1F, G, H) and up to 23 denticles that are small and irregular in size, without distinct dorsal denticle furrows, only sometimes with fine rib-like structures (Fig. 1H).
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</paragraph>
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<paragraph pageId="0" pageNumber="89">Reproductive system diaulic (Fig. 4A). Ampulla moderate in size (Fig. 4A, am). Vas deferens short, without distinct prostatic portion (Fig. 4A, vd), penial sheath widened (Fig. 4A, psh), penis unarmed, with elevations (Fig. 4A, p). Single proximal receptaculum seminis very large, oval (Fig. 4A, rsp).</paragraph>
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<caption pageId="0" pageNumber="89">
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<paragraph pageId="0" pageNumber="89">
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Figure 1. Comparison of
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja longicornis" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="longicornis">Myja longicornis</taxonomicName>
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Bergh, 1896 with other aeolidacean taxa that have been proposed to have relationships with it (
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<taxonomicName class="Gastropoda" family="Calmidae" genus="Calma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calma" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">Calma</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Tergipedidae" genus="Tergipes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tergipes" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">Tergipes</taxonomicName>
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) and which are covered by present analysis [type species of the genus
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Facelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Facelina" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">Facelina</taxonomicName>
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,
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<taxonomicName lsidName="F. auriculata" pageId="0" pageNumber="89" rank="species" species="auriculata">F. auriculata</taxonomicName>
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(
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<normalizedToken originalValue="Müller">Mueller</normalizedToken>
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, 1776)].
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<normalizedToken originalValue="A–H">A-H</normalizedToken>
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja longicornis" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="longicornis">Myja cf. longicornis</taxonomicName>
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from Thailand, living animal ca. 10 mm in length A dorsal view of hydroids in situ B lateral view of hydroids in situ (left), egg mass on the hydroid (right) C details of cerata D smooth masticatory processes of jaws (indicated by arrows), SEM E pharynx, dissected dorsally to show very narrow radula (indicated by an arrow), LM F whole radula, SEM G anteriormost part of radula to show sacoglossan-like small knife-shaped teeth, SEM H teeth from the middle part of radula, SEM; I, J
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja longicornis" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="longicornis">Myja longicornis</taxonomicName>
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Bergh, 1896 external view and radula (anterior part), reproduced from the first description by
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<bibRefCitation author="Bergh, R" journalOrPublisher="Revue Suisse de Zoologie et Annales de Musee d'Histoire Naturelle de Geneve" pageId="0" pageNumber="89" pagination="385 - 394" title="Eolidiens d'Amboine. Voyage de MM. M. Bedot et C. Pictet dans l'Archipel Malais." volume="4" year="1896">Bergh (1896)</bibRefCitation>
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;
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<normalizedToken originalValue="K–O">K-O</normalizedToken>
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Facelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Facelina auriculata" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="auriculata">Facelina auriculata</taxonomicName>
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jaws and radula of a specimen from UK, collected together with neotype K masticatory process (well-defined denticles indicated by arrow), SEM L radula (arrow) on odontophore, to show that anteriormost teeth are not reduced, LM M anterior part of radula to show that teeth are not reduced N anteriormost tooth of radula O two anterior teeth of radula P radula (middle part) of
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<taxonomicName class="Gastropoda" family="Calmidae" genus="Calma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calma glaucoides" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="glaucoides">Calma glaucoides</taxonomicName>
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(Alder & Hancock, 1854) from Norway Q radula (middle part)
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<taxonomicName class="Gastropoda" family="Tergipedidae" genus="Tergipes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tergipes tergipes" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="tergipes">Tergipes tergipes</taxonomicName>
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(
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<normalizedToken originalValue="Forsskål">Forsskal</normalizedToken>
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in Niebuhr, 1775). Scale bars: 20
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(D, N, O, Q); 50
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(F, K); 10
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(G, P); 5
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(H); 100
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(M). Photographs of living specimens by Chanon Ngernthongdee and Siwat Worachananant, SEM images by AV Martynov. Figures I and J are reproduced from
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<bibRefCitation author="Bergh, R" journalOrPublisher="Revue Suisse de Zoologie et Annales de Musee d'Histoire Naturelle de Geneve" pageId="0" pageNumber="89" pagination="385 - 394" title="Eolidiens d'Amboine. Voyage de MM. M. Bedot et C. Pictet dans l'Archipel Malais." volume="4" year="1896">Bergh (1896)</bibRefCitation>
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, the publication not currently under copyright.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="biology_ecology">
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<paragraph pageId="0" pageNumber="89">Biology.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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Subtidal, highly cryptic on
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<taxonomicName class="Hydrozoa" family="Pennariidae" genus="Pennaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pennaria disticha" order="Anthoathecata" pageId="0" pageNumber="89" phylum="Cnidaria" rank="species" species="disticha">Pennaria disticha</taxonomicName>
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hydroids in soft sediment habitats beyond the coral reef or on the same hydroids at the reef edge (Fig. 1A, B). Egg mass is a long narrow ribbon, white, laid directly onto host hydroids (Fig. 1B).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="distribution">
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<paragraph pageId="0" pageNumber="89">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="89">Presently found only at Koh Samaesan, Thailand, but expected to be found in neighbouring regions of the Indo-West Pacific.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="remarks">
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<paragraph pageId="0" pageNumber="89">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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Thai specimens show closeness to the type species of the genus
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja longicornis" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="longicornis">Myja longicornis</taxonomicName>
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from Ambon (Indonesia) in such features as the apically widened cerata, only a single pair of anterior cerata, acleioproctic anus, winged jaws, and small uniserial radula. Therefore, studying these specimens allows us to reveal the phylogenetic and taxonomic position of the genus
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">Myja</taxonomicName>
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via both morphological and molecular means. However, while
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. cf. longicornis</taxonomicName>
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from Thailand is similar to the type species of the genus
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">Myja</taxonomicName>
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,
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. longicornis</taxonomicName>
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from the type locality in Ambon as described in the original description by
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<bibRefCitation author="Bergh, R" journalOrPublisher="Revue Suisse de Zoologie et Annales de Musee d'Histoire Naturelle de Geneve" pageId="0" pageNumber="89" pagination="385 - 394" title="Eolidiens d'Amboine. Voyage de MM. M. Bedot et C. Pictet dans l'Archipel Malais." volume="4" year="1896">Bergh (1896)</bibRefCitation>
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, there are differences in several external and internal characters which do not allow us to identify the Thai material as
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. longicornis</taxonomicName>
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and
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<normalizedToken originalValue="Bergh’s">Bergh's</normalizedToken>
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figures are reproduced here (Fig. 1I, H). We therefore record here the specimens from Thailand as
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. cf. longicornis</taxonomicName>
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. The distinguishing features of
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. cf. longicornis</taxonomicName>
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are predominantly greyish without the green digestive gland branches both in the body and in the cerata, as was clearly indicated for
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. longicornis</taxonomicName>
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in the original description (Bergh, 1896: 389, 390). It has a reddish and not brown-chocolate basal spot at anterior pair of cerata, and similar reddish (and not brown) pigment at ceratal apices. Furthermore, the radula of
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. longicornis</taxonomicName>
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as depicted in
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<bibRefCitation author="Bergh, R" journalOrPublisher="Revue Suisse de Zoologie et Annales de Musee d'Histoire Naturelle de Geneve" pageId="0" pageNumber="89" pagination="385 - 394" title="Eolidiens d'Amboine. Voyage de MM. M. Bedot et C. Pictet dans l'Archipel Malais." volume="4" year="1896">Bergh (1896</bibRefCitation>
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; reproduced here Fig. 1J) has more distinct lateral denticles, which are lower in number (10), compared to
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. cf. longicornis</taxonomicName>
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(at least 23) (see Fig. 1H). We suspect that there is hidden species diversity in the genus
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">Myja</taxonomicName>
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of the Indo-West Pacific. Specimens collected in 2016 reveal the presence of at least two more species of the genus
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">Myja</taxonomicName>
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, which differ from
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. cf. longicornis</taxonomicName>
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based on morphological and molecular data and from
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. longicornis</taxonomicName>
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according to the morphological data, are described as new to science,
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja karin" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="karin">Myja karin</taxonomicName>
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sp. n. (see Fig. 2) and
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<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja hyotan" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="hyotan">Myja hyotan</taxonomicName>
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sp. n. (see Fig. 3). Minimum uncorrected p-distances of the COI marker which separate
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. cf. longicornis</taxonomicName>
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from
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<taxonomicName lsidName="M. karin" pageId="0" pageNumber="89" rank="species" species="karin">M. karin</taxonomicName>
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sp. n. are 11.9%. Minimum uncorrected p-distances of the 16S marker which separate
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<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. cf. longicornis</taxonomicName>
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from
|
||
<taxonomicName lsidName="M. karin" pageId="0" pageNumber="89" rank="species" species="karin">M. karin</taxonomicName>
|
||
sp. n. are 3.71% and from
|
||
<taxonomicName lsidName="M. hyotan" pageId="0" pageNumber="89" rank="species" species="hyotan">M. hyotan</taxonomicName>
|
||
sp. n. are 2.55%. Minimum uncorrected p-distances of the H3 marker which separate
|
||
<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. cf. longicornis</taxonomicName>
|
||
from
|
||
<taxonomicName lsidName="M. karin" pageId="0" pageNumber="89" rank="species" species="karin">M. karin</taxonomicName>
|
||
sp. n. are 4.28% and from
|
||
<taxonomicName lsidName="M. hyotan" pageId="0" pageNumber="89" rank="species" species="hyotan">M. hyotan</taxonomicName>
|
||
sp. n. are 3.36%, whereas p-distances between the two specimens of
|
||
<taxonomicName lsidName="M. longicornis" pageId="0" pageNumber="89" rank="species" species="longicornis">M. cf. longicornis</taxonomicName>
|
||
for COI, 16S, and H3 markers are 0.2%, 0%, and 0% respectively.
|
||
</paragraph>
|
||
<caption pageId="0" pageNumber="89">
|
||
<paragraph pageId="0" pageNumber="89">
|
||
Figure 2.
|
||
<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja karin" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="karin">Myja karin</taxonomicName>
|
||
sp. n.
|
||
<normalizedToken originalValue="A–D">A-D</normalizedToken>
|
||
holotype A dorsal view B ventral view C lateral view D animal with egg mass E details of cerata F lateral view on hydroids in situ G, H veligers;
|
||
<normalizedToken originalValue="I–N">I-N</normalizedToken>
|
||
paratype I jaws J smooth masticatory processes of jaws (inidicated by arrows), SEM K radula on odontophore, to show narrow teeth and reduced anteriormost teeth (arrow), LM L whole radula, SEM M teeth from the middle part of radula N anterior teeth. Scale bars: 100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(I); 50
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(J, L); 10
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(M); 5
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(N). Photographs of living specimens by TA Korshunova and AV Martynov, SEM images by AV Martynov.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="0" pageNumber="89">
|
||
<paragraph pageId="0" pageNumber="89">
|
||
Figure 3.
|
||
<taxonomicName class="Gastropoda" family="Facelinidae" genus="Myja" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myja hyotan" order="Nudibranchia" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="hyotan">Myja hyotan</taxonomicName>
|
||
sp. n., holotype. A dorsal view B ventral view C lateral view D details of cerata E dorsal view on hydroids in situ F jaw G smooth masticatory processes of jaws (indicated by arrows), SEM H radula on odontophore, to show reduced anteriormost teeth (arrow), LM I anterior teeth with strongly reduced anteriormost teeth, SEM J teeth from the middle part of radula K posterior part of radula to show smooth teeth. Scale bars: 100
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(F); 50
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(G, I); 10
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(J, K). Photographs of living specimens by TA Korshunova and AV Martynov, SEM images by AV Martynov.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |